262 lines
32 KiB
XML
262 lines
32 KiB
XML
<document id="4AD46AFE630DF72F4466D895B39BFD87" ID-CLB-Dataset="29036" ID-DOI="10.11646/zootaxa.4459.1.2" ID-GBIF-Dataset="acdf8f3d-d29f-463e-a431-191275b69a80" ID-ISSN="1175-5326" ID-Zenodo-Dep="1458336" ID-ZooBank="67E956AB-04B1-4EF7-8CC0-E152F95D0563" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539287820689" checkinUser="plazi" docAuthor="Liu, Weixin & Golovatch, Sergei" docDate="2018" docId="9005EB4842378E4748DAFA070242FE77" docLanguage="en" docName="zootaxa.4459.1.2.pdf" docOrigin="Zootaxa 4459 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Epanerchodus gladiatus Liu & Golovatch 2018, new species" docType="treatment" docVersion="7" lastPageNumber="70" masterDocId="6C3C933042398E56484DFFE30629FF95" masterDocTitle="The millipede genus Epanerchodus Attems, 1901 in continental China, with descriptions of seven new cavernicolous species (Diplopoda, Polydesmida, Polydesmidae)" masterLastPageNumber="84" masterPageNumber="53" pageNumber="67" updateTime="1698577720608" updateUser="ExternalLinkService">
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<mods:mods id="AB568A4FEB2A9545EE1C6D3150D1A9B0" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="2B9CDA83D91940B9111CC71269FC440C">
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<mods:title id="15DDE703D7A5000A62F12597B301DA59">The millipede genus Epanerchodus Attems, 1901 in continental China, with descriptions of seven new cavernicolous species (Diplopoda, Polydesmida, Polydesmidae)</mods:title>
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<mods:name id="D926D89373D861F302022A4073C7DA5B" type="personal">
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<mods:roleTerm id="DE68B12929E4A1345E571F80D1AEFF79">Author</mods:roleTerm>
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<mods:namePart id="187C2BF3FFD89D2E6EA1C0E909046914">Liu, Weixin</mods:namePart>
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<mods:name id="F6A359BBE7B03B4AF0731BCC01760DA9" type="personal">
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<mods:roleTerm id="0F609FEB4596DCD3E848310AFC8E85D2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B60D082B079DE3D2C751AB53657090EF">Golovatch, Sergei</mods:namePart>
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<mods:typeOfResource id="0462BD2F0AF0651DA31DA29740EFF8A3">text</mods:typeOfResource>
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<mods:titleInfo id="7A8B504EEB405B850D74D9E1215EC67C">
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<mods:title id="CC49100941AF5E36EADB4CC5203B91CC">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="7EB6FDCB2A79C05E942A0C39D9F6EF20">
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<mods:date id="228E7B9056B025005948D40FD7670C1D">2018</mods:date>
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<mods:detail id="CD83382D1551225BD9C822B4260B6A27" type="pubDate">
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<mods:number id="405C74B10F1016DD860FCEA4CB5805CB">2018-08-14</mods:number>
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<mods:detail id="E918F1307AAC2178A50DB5455466C132" type="volume">
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<mods:number id="EE01DC8159B526BBB017B6E8D78BE035">4459</mods:number>
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</mods:detail>
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<mods:detail id="61DA34D99318C874BDA329EC16C7E4E7" type="issue">
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<mods:number id="DC8D216C6DC779C042F76D49B30C6397">1</mods:number>
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<mods:classification id="4AEE320E6C94B67E8426B287934A11BF">journal article</mods:classification>
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<mods:identifier id="7222F38810E429E28EE70A64EC06B5DB" type="CLB-Dataset">29036</mods:identifier>
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<mods:identifier id="6590DA53E9A62486F038DE241648DA9F" type="DOI">10.11646/zootaxa.4459.1.2</mods:identifier>
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<mods:identifier id="5A5698FDEB1FCDF868E1DC720A282C4B" type="GBIF-Dataset">acdf8f3d-d29f-463e-a431-191275b69a80</mods:identifier>
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<mods:identifier id="ACFF425603AF23938CFD1927020C8B00" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
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<treatment id="9005EB4842378E4748DAFA070242FE77" ID-DOI="http://doi.org/10.5281/zenodo.5973778" ID-GBIF-Taxon="148692459" ID-Zenodo-Dep="5973778" LSID="urn:lsid:plazi:treatment:9005EB4842378E4748DAFA070242FE77" httpUri="http://treatment.plazi.org/id/9005EB4842378E4748DAFA070242FE77" lastPageId="17" lastPageNumber="70" pageId="14" pageNumber="67">
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<subSubSection id="50B609D542378E5848DAFA070735F9B5" pageId="14" pageNumber="67" type="nomenclature">
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<paragraph id="18135A5E42378E5848DAFA070472FA6B" blockId="14.[151,603,1508,1568]" box="[151,603,1508,1534]" pageId="14" pageNumber="67">
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<heading id="435BED3242378E5848DAFA070472FA6B" bold="true" box="[151,603,1508,1534]" fontSize="11" level="1" pageId="14" pageNumber="67" reason="1">
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<emphasis id="2AD8864C42378E5848DAFA070472FA6B" bold="true" box="[151,603,1508,1534]" pageId="14" pageNumber="67">
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<taxonomicName id="DFAC21DD42378E5848DAFA070795FA6B" ID-CoL="6FN62" authority="Liu & Golovatch, 2018" authorityName="Liu & Golovatch" authorityYear="2018" box="[151,444,1508,1534]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="67" phylum="Arthropoda" rank="species" species="gladiatus" status="sp. nov.">
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<emphasis id="2AD8864C42378E5848DAFA070795FA6B" bold="true" box="[151,444,1508,1534]" italics="true" pageId="14" pageNumber="67">Epanerchodus gladiatus</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="31EB3B3742378E584986FA070472FA6B" box="[459,603,1508,1534]" pageId="14" pageNumber="67" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="18135A5E42378E5848DAF9E40735F9B5" blockId="14.[151,603,1508,1568]" box="[151,284,1543,1568]" pageId="14" pageNumber="67">
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<figureCitation id="809746DB42378E5848DAF9E40731F9B5" box="[151,280,1543,1568]" captionStart-0="FIGURE 12" captionStart-1="FIGURE 13" captionStartId-0="15.[151,250,1814,1836]" captionStartId-1="16.[151,250,1380,1402]" captionTargetBox-0="[298,1297,193,1802]" captionTargetBox-1="[177,1412,206,1340]" captionTargetId-0="figure@15.[292,1298,193,1802]" captionTargetId-1="figure@16.[151,1436,193,1359]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionText-0="FIGURE 12. Epanerchodus gladiatus n. sp., male paratype. A: habitus, dorsal view. B: habitus, ventral view. Scale bar: 0.2 mm." captionText-1="FIGURE 13. Epanerchodus gladiatus n. sp., male paratype. A: left gonopod, mesal view. B: left gonopod, lateral view. Abbreviations: a = outgrowth; cl = clivus; en = endomere; p = process." httpUri-0="https://zenodo.org/record/1458360/files/figure.png" httpUri-1="https://zenodo.org/record/1458362/files/figure.png" pageId="14" pageNumber="67">Figs 12–13</figureCitation>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="50B609D542378E5848DAF9AC0520F925" pageId="14" pageNumber="67" type="materials_examined">
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||
<paragraph id="18135A5E42378E5848DAF9AC0520F925" blockId="14.[151,1436,1615,2000]" pageId="14" pageNumber="67">
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<materialsCitation id="A8C4500342378E5848DAF9AC02EBF919" ID-GBIF-Occurrence="1917355156" collectingDate="2015-07-29" collectionCode="SCAU" country="China" county="Wuchuan County" location="Gaodong Village" municipality="Huangdu Town" pageId="14" pageNumber="67" specimenCount="1" stateProvince="Guizhou Province" typeStatus="holotype">
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<emphasis id="2AD8864C42378E5848DAF9AC0751F9FD" bold="true" box="[151,376,1615,1640]" pageId="14" pageNumber="67">Material examined</emphasis>
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:
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<typeStatus id="C717E4FC42378E5849CAF9AC07C6F9FD" box="[391,495,1615,1640]" pageId="14" pageNumber="67" type="holotype">Holotype</typeStatus>
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male (
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<collectionCode id="7EBDC29B42378E584A77F9B304AFF9FD" box="[570,646,1616,1640]" pageId="14" pageNumber="67">SCAU</collectionCode>
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eGZ13-1) from
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<collectingCountry id="60BB1ACE42378E584B73F9AC05ADF9FD" box="[830,900,1615,1640]" name="China" pageId="14" pageNumber="67">China</collectingCountry>
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,
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<collectingRegion id="DA6894BC42378E584BC2F9AC0274F9FD" box="[911,1117,1615,1640]" country="China" name="Guizhou" pageId="14" pageNumber="67">Guizhou Province</collectingRegion>
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,
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<collectingCounty id="F17222D242378E584C2AF9AC0302F9FD" box="[1127,1323,1615,1640]" pageId="14" pageNumber="67">Wuchuan County</collectingCounty>
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,
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<collectingMunicipality id="F877C02442378E584D78F9AC06F0F91E" pageId="14" pageNumber="67">Huangdu Town</collectingMunicipality>
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,
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<location id="1D730C8542378E5848A9F990078EF919" LSID="urn:lsid:plazi:treatment:9005EB4842378E4748DAFA070242FE77:1D730C8542378E5848A9F990078EF919" box="[228,423,1651,1676]" country="China" county="Wuchuan County" municipality="Huangdu Town" name="Gaodong Village" pageId="14" pageNumber="67" stateProvince="Guizhou Province">Gaodong Village</location>
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,
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<location id="1D730C8542378E5849FFF99004ADF919" LSID="urn:lsid:plazi:treatment:9005EB4842378E4748DAFA070242FE77:1D730C8542378E5849FFF99004ADF919" box="[434,644,1651,1676]" country="China" county="Wuchuan County" municipality="Huangdu Town" name="Yinshi Dong Cave" pageId="14" pageNumber="67" stateProvince="Guizhou Province">Yinshi Dong Cave</location>
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,
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<collectingDate id="7C56857642378E584ADDF9900535F919" box="[656,796,1651,1676]" pageId="14" pageNumber="67" value="2015-07-29">
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<date id="6C127C9E42378E584ADDF9900535F919" box="[656,796,1651,1676]" pageId="14" pageNumber="67" value="2015-07-29">29.VII.2015</date>
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</collectingDate>
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, colls:
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<location id="1D730C8542378E584B21F99005D1F919" LSID="urn:lsid:plazi:treatment:9005EB4842378E4748DAFA070242FE77:1D730C8542378E584B21F99005D1F919" box="[876,1016,1651,1676]" country="China" county="Wuchuan County" municipality="Huangdu Town" name="Mingyi Tian" pageId="14" pageNumber="67" stateProvince="Guizhou Province">Mingyi Tian</location>
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and
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<location id="1D730C8542378E584C7DF99002EBF919" LSID="urn:lsid:plazi:treatment:9005EB4842378E4748DAFA070242FE77:1D730C8542378E584C7DF99002EBF919" box="[1072,1218,1651,1676]" country="China" county="Wuchuan County" municipality="Huangdu Town" name="Jingli Cheng" pageId="14" pageNumber="67" stateProvince="Guizhou Province">Jingli Cheng</location>
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</materialsCitation>
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.
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<materialsCitation id="A8C4500342378E584C80F997052CF925" ID-GBIF-Occurrence="1917355117" collectingDate="2015-07-29" collectionCode="SCAU" country="China" county="Wuchuan County" location="Gaodong Village" municipality="Huangdu Town" pageId="14" pageNumber="67" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Guizhou Province" typeStatus="paratype">
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<typeStatus id="C717E4FC42378E584C80F9970316F919" box="[1229,1343,1652,1676]" pageId="14" pageNumber="67" type="paratype">Paratypes</typeStatus>
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||
:
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<specimenCount id="0EAA91D742378E584D04F99703B1F919" box="[1353,1432,1651,1676]" pageId="14" pageNumber="67" type="male">1 male</specimenCount>
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||
,
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||
<specimenCount id="0EAA91D742378E5848DAF97B072AF925" box="[151,259,1687,1712]" pageId="14" pageNumber="67" type="female">2 females</specimenCount>
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||
(
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||
<collectionCode id="7EBDC29B42378E58495CF97B0774F925" box="[273,349,1688,1712]" pageId="14" pageNumber="67">SCAU</collectionCode>
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eGZ13-1), same data as the
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<typeStatus id="C717E4FC42378E584AD2F974052CF925" box="[671,773,1687,1712]" pageId="14" pageNumber="67" type="holotype">holotype</typeStatus>
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</materialsCitation>
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||
.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="50B609D542378E58488AF958053AF845" pageId="14" pageNumber="67" type="diagnosis">
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<paragraph id="18135A5E42378E58488AF9580711F8D5" blockId="14.[151,1436,1615,2000]" pageId="14" pageNumber="67">
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<emphasis id="2AD8864C42378E58488AF9580713F941" bold="true" box="[199,314,1723,1748]" pageId="14" pageNumber="67">Diagnosis</emphasis>
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: Adult males of
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<taxonomicName id="DFAC21DD42378E584A4AF95F04BAF941" authority="Liu & Golovatch, 2018" authorityName="Liu & Golovatch" authorityYear="2018" box="[519,659,1723,1748]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="67" phylum="Arthropoda" rank="species" species="gladiatus" status="sp. nov.">
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<emphasis id="2AD8864C42378E584A4AF95F04BAF941" box="[519,659,1723,1748]" italics="true" pageId="14" pageNumber="67">E. gladiatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="31EB3B3742378E584AD3F95804CBF941" box="[670,738,1723,1748]" pageId="14" pageNumber="67" rank="species">
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<emphasis id="2AD8864C42378E584AD3F95804CBF941" bold="true" box="[670,738,1723,1748]" pageId="14" pageNumber="67">n. sp.</emphasis>
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</taxonomicNameLabel>
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are distinct from other
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<taxonomicName id="DFAC21DD42378E584BB2F9580289F941" baseAuthorityName="Golovatch" baseAuthorityYear="1991" box="[1023,1184,1723,1748]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="67" phylum="Arthropoda" rank="genus">
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<emphasis id="2AD8864C42378E584BB2F9580289F941" box="[1023,1184,1723,1748]" italics="true" pageId="14" pageNumber="67">Epanerchodus</emphasis>
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</taxonomicName>
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species based on the following combination of characters: (1) body small (
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<emphasis id="2AD8864C42378E584AB1F903053FF962" box="[764,790,1760,1783]" italics="true" pageId="14" pageNumber="67">ca</emphasis>
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||
<quantity id="DF54F7BB42378E584B6DF93C05B3F962" box="[800,922,1759,1784]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="14" pageNumber="67" unit="mm" value="16.0" valueMax="17.0" valueMin="15.0">15–17 mm</quantity>
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||
long); (2) gonopod suberect and slender; (3) endomere sword-shaped, with a small, rounded, ventral bump (
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||
<emphasis id="2AD8864C42378E584B17F8E00541F889" bold="true" box="[858,872,1795,1820]" pageId="14" pageNumber="67">a</emphasis>
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||
) and a prominent, digitiform, mesal process (
|
||
<emphasis id="2AD8864C42378E584D25F8E0035EF889" bold="true" box="[1384,1399,1795,1820]" pageId="14" pageNumber="67">p</emphasis>
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||
) at base (
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||
<figureCitation id="809746DB42378E584895F8C40702F8D5" box="[216,299,1831,1856]" captionStart="FIGURE 13" captionStartId="16.[151,250,1380,1402]" captionTargetBox="[177,1412,206,1340]" captionTargetId="figure@16.[151,1436,193,1359]" captionTargetPageId="16" captionText="FIGURE 13. Epanerchodus gladiatus n. sp., male paratype. A: left gonopod, mesal view. B: left gonopod, lateral view. Abbreviations: a = outgrowth; cl = clivus; en = endomere; p = process." httpUri="https://zenodo.org/record/1458362/files/figure.png" pageId="14" pageNumber="67">Fig. 13</figureCitation>
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||
).
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||
</paragraph>
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||
<paragraph id="18135A5E42378E58488AF8A8053AF845" blockId="14.[151,1436,1615,2000]" pageId="14" pageNumber="67">
|
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This new species is similar to
|
||
<taxonomicName id="DFAC21DD42378E584A68F8AF04BBF8F1" authority="Liu & Golovatch, 2018" authorityName="Liu & Golovatch" authorityYear="2018" box="[549,658,1867,1892]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="67" phylum="Arthropoda" rank="species" species="chutou" status="sp. nov.">
|
||
<emphasis id="2AD8864C42378E584A68F8AF04BBF8F1" box="[549,658,1867,1892]" italics="true" pageId="14" pageNumber="67">E. chutou</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="31EB3B3742378E584AD1F8A804F4F8F1" box="[668,733,1867,1892]" pageId="14" pageNumber="67" rank="species">
|
||
<emphasis id="2AD8864C42378E584AD1F8A804F4F8F1" bold="true" box="[668,733,1867,1892]" pageId="14" pageNumber="67">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(from a cave in
|
||
<collectingRegion id="DA6894BC42378E584BEDF8A8022AF8F1" box="[928,1027,1867,1892]" country="China" name="Guizhou" pageId="14" pageNumber="67">Guizhou</collectingRegion>
|
||
,
|
||
<collectingCountry id="60BB1ACE42378E584C5DF8A8027CF8F1" box="[1040,1109,1867,1892]" name="China" pageId="14" pageNumber="67">China</collectingCountry>
|
||
) in the endomere showing a basal outgrowth and a missing exomere. However,
|
||
<taxonomicName id="DFAC21DD42378E584AA2F8930553F81D" authority="Liu & Golovatch, 2018" authorityName="Liu & Golovatch" authorityYear="2018" box="[751,890,1903,1928]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="67" phylum="Arthropoda" rank="species" species="gladiatus" status="sp. nov.">
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||
<emphasis id="2AD8864C42378E584AA2F8930553F81D" box="[751,890,1903,1928]" italics="true" pageId="14" pageNumber="67">E. gladiatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="31EB3B3742378E584BC8F88C05E1F81D" box="[901,968,1903,1928]" pageId="14" pageNumber="67" rank="species">
|
||
<emphasis id="2AD8864C42378E584BC8F88C05E1F81D" bold="true" box="[901,968,1903,1928]" pageId="14" pageNumber="67">n. sp.</emphasis>
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||
</taxonomicNameLabel>
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||
is distinguished by the endomere being sword-shaped, with a small, rounded, ventral bump (
|
||
<emphasis id="2AD8864C42378E584B59F870050BF839" bold="true" box="[788,802,1939,1964]" pageId="14" pageNumber="67">a</emphasis>
|
||
) (
|
||
<figureCitation id="809746DB42378E584B0FF87005B0F839" box="[834,921,1939,1964]" captionStart="FIGURE 13" captionStartId="16.[151,250,1380,1402]" captionTargetBox="[177,1412,206,1340]" captionTargetId="figure@16.[151,1436,193,1359]" captionTargetPageId="16" captionText="FIGURE 13. Epanerchodus gladiatus n. sp., male paratype. A: left gonopod, mesal view. B: left gonopod, lateral view. Abbreviations: a = outgrowth; cl = clivus; en = endomere; p = process." httpUri="https://zenodo.org/record/1458362/files/figure.png" pageId="14" pageNumber="67">Fig. 13</figureCitation>
|
||
)
|
||
<emphasis id="2AD8864C42378E584BE2F87705E7F83E" box="[943,974,1940,1963]" italics="true" pageId="14" pageNumber="67">vs.</emphasis>
|
||
endomere hoe-shaped, with a slender, spiniform, basal process (
|
||
<emphasis id="2AD8864C42378E5849FBF85407EDF845" bold="true" box="[438,452,1975,2000]" pageId="14" pageNumber="67">a</emphasis>
|
||
) in
|
||
<taxonomicName id="DFAC21DD42378E5849BFF85B0477F845" authority="Liu & Golovatch, 2018" authorityName="Liu & Golovatch" authorityYear="2018" box="[498,606,1975,2000]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="67" phylum="Arthropoda" rank="species" species="chutou" status="sp. nov.">
|
||
<emphasis id="2AD8864C42378E5849BFF85B0477F845" box="[498,606,1975,2000]" italics="true" pageId="14" pageNumber="67">E. chutou</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="31EB3B3742378E584A29F854048AF845" box="[612,675,1975,2000]" pageId="14" pageNumber="67" rank="species">
|
||
<emphasis id="2AD8864C42378E584A29F854048AF845" bold="true" box="[612,675,1975,2000]" pageId="14" pageNumber="67">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="809746DB42378E584AFFF854052FF845" box="[690,774,1975,2000]" captionStart="FIGURE 15" captionStartId="18.[151,250,1462,1484]" captionTargetBox="[173,1421,710,1439]" captionTargetId="figure@18.[170,1422,688,1452]" captionTargetPageId="18" captionText="FIGURE 15. Epanerchodus chutou n. sp., male paratype. A: right gonopod, lateral view. B: right gonopod, mesal view. Abbreviations: a = spiniform process; cl = clivus; en = endomere; p = process." httpUri="https://zenodo.org/record/1458366/files/figure.png" pageId="14" pageNumber="67">Fig. 15</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="4CD30AD642368E5948DAF8F506EFF8DE" httpUri="https://zenodo.org/record/1458360/files/figure.png" pageId="15" pageNumber="68" startId="15.[151,250,1814,1836]" targetBox="[298,1297,193,1802]" targetPageId="15">
|
||
<paragraph id="18135A5E42368E5948DAF8F506EFF8DE" blockId="15.[151,1436,1814,1867]" pageId="15" pageNumber="68">
|
||
<emphasis id="2AD8864C42368E5948DAF8F50708F8B9" bold="true" box="[151,289,1814,1836]" pageId="15" pageNumber="68">FIGURE 12.</emphasis>
|
||
<taxonomicName id="DFAC21DD42368E594964F8F50409F8B9" authority="Liu & Golovatch, 2018" authorityName="Liu & Golovatch" authorityYear="2018" box="[297,544,1814,1836]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="68" phylum="Arthropoda" rank="species" species="gladiatus" status="sp. nov.">
|
||
<emphasis id="2AD8864C42368E594964F8F50409F8B9" box="[297,544,1814,1836]" italics="true" pageId="15" pageNumber="68">Epanerchodus gladiatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="31EB3B3742368E594A65F8F5044AF8B9" box="[552,611,1814,1836]" pageId="15" pageNumber="68" rank="species">
|
||
<emphasis id="2AD8864C42368E594A65F8F5044AF8B9" bold="true" box="[552,611,1814,1836]" pageId="15" pageNumber="68">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, male paratype.
|
||
<emphasis id="2AD8864C42368E594B42F8F50508F8B9" bold="true" box="[783,801,1814,1836]" pageId="15" pageNumber="68">A</emphasis>
|
||
: habitus, dorsal view.
|
||
<emphasis id="2AD8864C42368E594C46F8F50235F8B9" bold="true" box="[1035,1052,1814,1836]" pageId="15" pageNumber="68">B</emphasis>
|
||
: habitus, ventral view. Scale bar: 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="4CD30AD642298E4648DAFA87054CFA0C" httpUri="https://zenodo.org/record/1458362/files/figure.png" pageId="16" pageNumber="69" startId="16.[151,250,1380,1402]" targetBox="[177,1412,206,1340]" targetPageId="16">
|
||
<paragraph id="18135A5E42298E4648DAFA87054CFA0C" blockId="16.[151,1437,1380,1433]" pageId="16" pageNumber="69">
|
||
<emphasis id="2AD8864C42298E4648DAFA87070DFAEF" bold="true" box="[151,292,1380,1402]" pageId="16" pageNumber="69">FIGURE 13.</emphasis>
|
||
<taxonomicName id="DFAC21DD42298E464963FA870401FAEF" authority="Liu & Golovatch, 2018" authorityName="Liu & Golovatch" authorityYear="2018" box="[302,552,1380,1402]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="16" pageNumber="69" phylum="Arthropoda" rank="species" species="gladiatus" status="sp. nov.">
|
||
<emphasis id="2AD8864C42298E464963FA870401FAEF" box="[302,552,1380,1402]" italics="true" pageId="16" pageNumber="69">Epanerchodus gladiatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="31EB3B3742298E464A7EFA870459FAEF" box="[563,624,1380,1402]" pageId="16" pageNumber="69" rank="species">
|
||
<emphasis id="2AD8864C42298E464A7EFA870459FAEF" bold="true" box="[563,624,1380,1402]" pageId="16" pageNumber="69">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, male paratype.
|
||
<emphasis id="2AD8864C42298E464B69FA87051FFAEF" bold="true" box="[804,822,1380,1402]" pageId="16" pageNumber="69">A</emphasis>
|
||
: left gonopod, mesal view.
|
||
<emphasis id="2AD8864C42298E464C29FA87025CFAEF" bold="true" box="[1124,1141,1380,1402]" pageId="16" pageNumber="69">B</emphasis>
|
||
: left gonopod, lateral view. Abbreviations: a = outgrowth; cl = clivus; en = endomere; p = process.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="50B609D542298E4748D2FA230725FE2B" lastPageId="17" lastPageNumber="70" pageId="16" pageNumber="69" type="description">
|
||
<paragraph id="18135A5E42298E4748D2FA230529FEED" blockId="16.[151,1437,1472,2031]" lastBlockId="17.[151,1437,151,482]" lastPageId="17" lastPageNumber="70" pageId="16" pageNumber="69">
|
||
<emphasis id="2AD8864C42298E4648D2FA230700FA4C" bold="true" box="[159,297,1472,1497]" pageId="16" pageNumber="69">Description</emphasis>
|
||
: Based on
|
||
<typeStatus id="C717E4FC42298E4649E7FA2207F3FA4C" box="[426,474,1473,1497]" pageId="16" pageNumber="69">type</typeStatus>
|
||
specimens. Lengths of both sexes
|
||
<emphasis id="2AD8864C42298E464B2BFA2205A9FA4D" box="[870,896,1473,1496]" italics="true" pageId="16" pageNumber="69">ca</emphasis>
|
||
<quantity id="DF54F7BB42298E464BC4FA23022CFA4D" box="[905,1029,1472,1497]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="16" pageNumber="69" unit="mm" value="16.0" valueMax="17.0" valueMin="15.0">15–17 mm</quantity>
|
||
, widths of pro- and metazonae
|
||
<quantity id="DF54F7BB42298E464D26FA2206D8FA6C" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="16" pageNumber="69" unit="mm" value="1.25" valueMax="1.3" valueMin="1.2">1.2– 1.3 mm</quantity>
|
||
and 2.5–3.0 mm.
|
||
<emphasis id="2AD8864C42298E464986FA020465FA6F" bold="true" box="[459,588,1505,1530]" pageId="16" pageNumber="69">Coloration</emphasis>
|
||
: in alcohol generally pallid. Mouthparts and gonopodal telopodites light yellowish.
|
||
<emphasis id="2AD8864C42298E46495FF9E00779F989" bold="true" box="[274,336,1539,1564]" pageId="16" pageNumber="69">Body</emphasis>
|
||
: Adults with 20 rings. Width: head << collum <ring 3 <4 <2 <5–15, thereafter body gradually tapering towards telson.
|
||
<emphasis id="2AD8864C42298E4649E0F9C707C4F9A8" bold="true" box="[429,493,1572,1597]" pageId="16" pageNumber="69">Head</emphasis>
|
||
: clypeus densely pilose, vertex sparsely setose, epicranial suture conspicuous (
|
||
<figureCitation id="809746DB42298E464D22F9C706E0F9CB" captionStart="FIGURE 12" captionStartId="15.[151,250,1814,1836]" captionTargetBox="[298,1297,193,1802]" captionTargetId="figure@15.[292,1298,193,1802]" captionTargetPageId="15" captionText="FIGURE 12. Epanerchodus gladiatus n. sp., male paratype. A: habitus, dorsal view. B: habitus, ventral view. Scale bar: 0.2 mm." httpUri="https://zenodo.org/record/1458360/files/figure.png" pageId="16" pageNumber="69">Fig. 12B</figureCitation>
|
||
). Antennae long, reaching behind posterior margin of ring 3 when extended posteriorly, slightly clavate (
|
||
<figureCitation id="809746DB42298E464D22F9A6069FF9EA" captionStart="FIGURE 12" captionStartId="15.[151,250,1814,1836]" captionTargetBox="[298,1297,193,1802]" captionTargetId="figure@15.[292,1298,193,1802]" captionTargetPageId="15" captionText="FIGURE 12. Epanerchodus gladiatus n. sp., male paratype. A: habitus, dorsal view. B: habitus, ventral view. Scale bar: 0.2 mm." httpUri="https://zenodo.org/record/1458360/files/figure.png" pageId="16" pageNumber="69">Fig. 12</figureCitation>
|
||
).
|
||
<emphasis id="2AD8864C42298E464881F9840774F915" bold="true" box="[204,349,1639,1664]" pageId="16" pageNumber="69">Exoskeleton</emphasis>
|
||
: Collum fan-shaped, with a lateral incision on each side. Paraterga evident (
|
||
<figureCitation id="809746DB42298E464C80F9840308F9EA" box="[1229,1313,1639,1664]" captionStart="FIGURE 12" captionStartId="15.[151,250,1814,1836]" captionTargetBox="[298,1297,193,1802]" captionTargetId="figure@15.[292,1298,193,1802]" captionTargetPageId="15" captionText="FIGURE 12. Epanerchodus gladiatus n. sp., male paratype. A: habitus, dorsal view. B: habitus, ventral view. Scale bar: 0.2 mm." httpUri="https://zenodo.org/record/1458360/files/figure.png" pageId="16" pageNumber="69">Fig. 12</figureCitation>
|
||
), midbody paraterga extend metatergite to
|
||
<emphasis id="2AD8864C42298E464A49F96A0436F935" box="[516,543,1673,1696]" italics="true" pageId="16" pageNumber="69">ca</emphasis>
|
||
2.1x width of prozonite. Paraterga on collum and rings 2–7 clearly upturned dorsally above a faintly convex dorsum, other paraterga flat (
|
||
<figureCitation id="809746DB42298E464BDEF94905C6F957" box="[915,1007,1706,1731]" captionStart="FIGURE 12" captionStartId="15.[151,250,1814,1836]" captionTargetBox="[298,1297,193,1802]" captionTargetId="figure@15.[292,1298,193,1802]" captionTargetPageId="15" captionText="FIGURE 12. Epanerchodus gladiatus n. sp., male paratype. A: habitus, dorsal view. B: habitus, ventral view. Scale bar: 0.2 mm." httpUri="https://zenodo.org/record/1458360/files/figure.png" pageId="16" pageNumber="69">Fig. 12</figureCitation>
|
||
). Caudolateral corners of paraterga increasingly acute-angled, especially strongly projecting posteriorly past tergal margin on rings 17–19 (
|
||
<figureCitation id="809746DB42298E464D6AF92803A6F971" box="[1319,1423,1739,1764]" captionStart="FIGURE 12" captionStartId="15.[151,250,1814,1836]" captionTargetBox="[298,1297,193,1802]" captionTargetId="figure@15.[292,1298,193,1802]" captionTargetPageId="15" captionText="FIGURE 12. Epanerchodus gladiatus n. sp., male paratype. A: habitus, dorsal view. B: habitus, ventral view. Scale bar: 0.2 mm." httpUri="https://zenodo.org/record/1458360/files/figure.png" pageId="16" pageNumber="69">Fig. 12A</figureCitation>
|
||
). Integument shining, translucent, prozonae very delicately alveolate. Limbus regularly denticulate. Constriction between pro- and metazonae narrow, shallow and smooth. Metatergal sculpture faint, with three irregular transverse rows of setigerous, polygonal bosses. Sulcus between front and middle rows of setae a little deeper than that between middle and caudal rows. Tergal setae very short, present sometimes only on collum, ring 2 and ring 19.Three or four faint setigerous incisions at lateral margins of poreless and pore-bearing rings, respectively. Pore formula normal: 5, 7, 9, 10, 12, 13, 15–19, ozopores evident, dorsal, clearly set off from lateral margin and located between last and penultimate marginal incisions. Epiproct short, conical, pre-apical lateral papillae small. Hypoproct subtrapeziform, with two setigerous papillae. Pleurosternal carinae small, present only on ring 2. Sterna sparsely setose, cross-shaped impressions shallow. Legs long and slender, about 2.2–2.5 times as long as body ring height in both sexes, without sphaerotrichomes or sternal cones, prefemora not bulging laterally.
|
||
<emphasis id="2AD8864C42288E474D56FF5A03BDFF47" bold="true" box="[1307,1428,185,210]" pageId="17" pageNumber="70">Gonopods</emphasis>
|
||
: slender (
|
||
<figureCitation id="809746DB42288E4748B4FF390765FF66" box="[249,332,218,243]" captionStart="FIGURE 13" captionStartId="16.[151,250,1380,1402]" captionTargetBox="[177,1412,206,1340]" captionTargetId="figure@16.[151,1436,193,1359]" captionTargetPageId="16" captionText="FIGURE 13. Epanerchodus gladiatus n. sp., male paratype. A: left gonopod, mesal view. B: left gonopod, lateral view. Abbreviations: a = outgrowth; cl = clivus; en = endomere; p = process." httpUri="https://zenodo.org/record/1458362/files/figure.png" pageId="17" pageNumber="70">Fig. 13</figureCitation>
|
||
). Telopodite linear, erect. Prefemur densely setose, about half as long as telopodite. Clivus (
|
||
<emphasis id="2AD8864C42288E474D24FF390354FF66" bold="true" box="[1385,1405,218,243]" pageId="17" pageNumber="70">cl</emphasis>
|
||
) a small bump. Endomere (
|
||
<emphasis id="2AD8864C42288E4749F9FF1F07E6FE80" bold="true" box="[436,463,252,277]" pageId="17" pageNumber="70">en</emphasis>
|
||
) slender, sword-shaped, broadened at midway and distally, but constricted between. Endomere apex subacuminate, with a small, rounded, ventral bump (
|
||
<emphasis id="2AD8864C42288E474BD1FEFE0583FEA3" bold="true" box="[924,938,285,310]" pageId="17" pageNumber="70">a</emphasis>
|
||
) and a prominent, digitiform, mesal process (
|
||
<emphasis id="2AD8864C42288E4748EDFEDD0686FEC2" bold="true" box="[160,175,318,343]" pageId="17" pageNumber="70">p</emphasis>
|
||
) at base. Seminal groove starting mesally, then recurving laterad to run into an accessory seminal chamber, the latter opening on a hairy pulvillus. An exomere absent.
|
||
</paragraph>
|
||
<paragraph id="18135A5E42288E47488AFE620725FE2B" blockId="17.[151,1437,151,482]" pageId="17" pageNumber="70">
|
||
<emphasis id="2AD8864C42288E47488AFE6206D6FE0F" bold="true" box="[199,255,385,410]" pageId="17" pageNumber="70">Note</emphasis>
|
||
: Due to the long and slender antennae and legs, and a depigmented cuticle, this species is most likely a troglobite.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="50B609D542288E47488AFE2A0242FE77" box="[199,1131,457,482]" pageId="17" pageNumber="70" type="etymology">
|
||
<paragraph id="18135A5E42288E47488AFE2A0242FE77" blockId="17.[151,1437,151,482]" box="[199,1131,457,482]" pageId="17" pageNumber="70">
|
||
<emphasis id="2AD8864C42288E47488AFE2A0761FE77" bold="true" box="[199,328,457,482]" pageId="17" pageNumber="70">Etymology</emphasis>
|
||
: To emphasize the endomere being sword-shaped; masculine adjective.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |