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<document id="E61AC5ADB725FB1AB3B37606DFC311AB" ID-CLB-Dataset="29036" ID-DOI="10.11646/zootaxa.4459.1.2" ID-GBIF-Dataset="acdf8f3d-d29f-463e-a431-191275b69a80" ID-ISSN="1175-5326" ID-Zenodo-Dep="1458336" ID-ZooBank="67E956AB-04B1-4EF7-8CC0-E152F95D0563" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539287820689" checkinUser="plazi" docAuthor="Liu, Weixin &amp; Golovatch, Sergei" docDate="2018" docId="9005EB4842358E5848DAFAE80203FA09" docLanguage="en" docName="zootaxa.4459.1.2.pdf" docOrigin="Zootaxa 4459 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Epanerchodus parvus Liu &amp; Golovatch 2018, new species" docType="treatment" docVersion="7" lastPageNumber="67" masterDocId="6C3C933042398E56484DFFE30629FF95" masterDocTitle="The millipede genus Epanerchodus Attems, 1901 in continental China, with descriptions of seven new cavernicolous species (Diplopoda, Polydesmida, Polydesmidae)" masterLastPageNumber="84" masterPageNumber="53" pageNumber="65" updateTime="1698577720608" updateUser="ExternalLinkService">
<mods:mods id="006729870B51147882F49717D79103C7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="651067A7AC8D9F52E07ED6ADF5CB48DB">
<mods:title id="8F0477394BFC44376C7FB1D544368186">The millipede genus Epanerchodus Attems, 1901 in continental China, with descriptions of seven new cavernicolous species (Diplopoda, Polydesmida, Polydesmidae)</mods:title>
</mods:titleInfo>
<mods:name id="72ACDE4EDD5D96973C89DB1D4DC1DCB0" type="personal">
<mods:role id="37AFC44786E5CC967B11E3877DB12B98">
<mods:roleTerm id="80103DC7CEA9AE7243B1BC7CF204A96F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0545FE9FF730C58E25746330D4678FDA">Liu, Weixin</mods:namePart>
</mods:name>
<mods:name id="FF3A08413A4CE7AED3B5D44B9E4DA9EE" type="personal">
<mods:role id="9C635DB8B65497015563BCC9E611F76C">
<mods:roleTerm id="CD4B402B6F49540071A33B6174351A67">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C949F95FBAA5EA4130316BEE6910B987">Golovatch, Sergei</mods:namePart>
</mods:name>
<mods:typeOfResource id="FD44C3D8C9BBF5BA02F664922C9B3996">text</mods:typeOfResource>
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<mods:titleInfo id="083713854F612A7B4F6FE3EFA2D8C10D">
<mods:title id="30996E581B438AA34A066DA0B0388E1E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A7E5E2C2BDB669EBE096D3B3D6C66DDB">
<mods:date id="9DFAE6B66A8190DE005C25C55F18FBCC">2018</mods:date>
<mods:detail id="E0DC163D96A6617313FE87D4C29E326F" type="pubDate">
<mods:number id="8C808D96F16DD4CEB9E96201E7B8D472">2018-08-14</mods:number>
</mods:detail>
<mods:detail id="00FB0EF3F8CDB5FC7531812EA1ABEAC0" type="volume">
<mods:number id="7D2FC7C86C84D55E2FD31864715ADC26">4459</mods:number>
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<mods:detail id="977963C9504FF4B7B454D16184C9ABB9" type="issue">
<mods:number id="D0F6FDFE02FC78113C03E0B6DCD9BB31">1</mods:number>
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<mods:classification id="F887A3E7223571627A3C6E0EC0EEBA6D">journal article</mods:classification>
<mods:identifier id="21CFC889216AB67A3F279D0279068D76" type="CLB-Dataset">29036</mods:identifier>
<mods:identifier id="589DE7532C3EF60C8C86260140859ED4" type="DOI">10.11646/zootaxa.4459.1.2</mods:identifier>
<mods:identifier id="61B801EF68746EDF978EA5F19D3BBC18" type="GBIF-Dataset">acdf8f3d-d29f-463e-a431-191275b69a80</mods:identifier>
<mods:identifier id="A54060C151C9F8D491C052C752E8A636" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="CB9AFF10989F23484B77EA3079AED0C3" type="Zenodo-Dep">1458336</mods:identifier>
<mods:identifier id="E669F4D57F93D3F98EBEA8AA40FBE82A" type="ZooBank">67E956AB-04B1-4EF7-8CC0-E152F95D0563</mods:identifier>
</mods:mods>
<treatment id="9005EB4842358E5848DAFAE80203FA09" ID-DOI="http://doi.org/10.5281/zenodo.5973776" ID-GBIF-Taxon="148692460" ID-Zenodo-Dep="5973776" LSID="urn:lsid:plazi:treatment:9005EB4842358E5848DAFAE80203FA09" httpUri="http://treatment.plazi.org/id/9005EB4842358E5848DAFAE80203FA09" lastPageId="14" lastPageNumber="67" pageId="12" pageNumber="65">
<subSubSection id="50B609D542358E5A48DAFAE80733FAD3" pageId="12" pageNumber="65" type="nomenclature">
<paragraph id="18135A5E42358E5A48DAFAE80417FAB0" blockId="12.[151,574,1291,1350]" box="[151,574,1291,1317]" pageId="12" pageNumber="65">
<heading id="435BED3242358E5A48DAFAE80417FAB0" bold="true" box="[151,574,1291,1317]" fontSize="11" level="1" pageId="12" pageNumber="65" reason="1">
<emphasis id="2AD8864C42358E5A48DAFAE80417FAB0" bold="true" box="[151,574,1291,1317]" pageId="12" pageNumber="65">
<taxonomicName id="DFAC21DD42358E5A48DAFAE807B6FAB0" ID-CoL="6FN58" authority="Liu &amp; Golovatch, 2018" authorityName="Liu &amp; Golovatch" authorityYear="2018" box="[151,415,1291,1317]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="65" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
<emphasis id="2AD8864C42358E5A48DAFAE807B6FAB0" bold="true" box="[151,415,1291,1317]" italics="true" pageId="12" pageNumber="65">Epanerchodus parvus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="31EB3B3742358E5A49E3FAE80417FAB0" box="[430,574,1291,1317]" pageId="12" pageNumber="65" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="18135A5E42358E5A48DAFACE0733FAD3" blockId="12.[151,574,1291,1350]" box="[151,282,1325,1350]" pageId="12" pageNumber="65">
<figureCitation id="809746DB42358E5A48DAFACE073FFAD3" box="[151,278,1325,1350]" captionStart-0="FIGURE 10" captionStart-1="FIGURE 11" captionStartId-0="13.[151,250,1391,1413]" captionStartId-1="14.[151,250,1216,1238]" captionTargetBox-0="[190,1395,191,1378]" captionTargetBox-1="[172,1407,361,1167]" captionTargetId-0="figure@13.[188,1398,188,1383]" captionTargetId-1="figure@14.[151,1436,337,1195]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="FIGURE 10. Epanerchodus parvus n. sp., male paratype. A: anterior part of body, dorsal view. B: anterior part of body, ventral view. C: midbody rings, dorsal view. D: posterior part of body, dorsal view. E: midbody rings, ventral view. F: posterior part of body, ventral view. G: habitus, dorsal view. Scale bar: A–F, 0.1 mm; G, 0.2 mm." captionText-1="FIGURE 11. Epanerchodus parvus n. sp., male paratype. A: left gonopod, mesal view. B: left gonopod, lateral view. Abbreviations: cl = clivus; en = endomere; p = process." httpUri-0="https://zenodo.org/record/1458356/files/figure.png" httpUri-1="https://zenodo.org/record/1458358/files/figure.png" pageId="12" pageNumber="65">Figs 10–11</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="50B609D542358E5A48DAFA930757FA60" pageId="12" pageNumber="65" type="materials_examined">
<paragraph id="18135A5E42358E5A48DAFA930757FA60" blockId="12.[151,1437,1392,2029]" pageId="12" pageNumber="65">
<materialsCitation id="A8C4500342358E5A48DAFA930505FA44" ID-GBIF-Occurrence="1917355119" collectingDate="2014-06-29" collectionCode="SCAU" country="China" county="Cenggong County" location="Pingzhuang Town" municipality="Qiandongnan Autonomous Prefecture" pageId="12" pageNumber="65" specimenCount="1" stateProvince="Guizhou Province" typeStatus="holotype">
<emphasis id="2AD8864C42358E5A48DAFA930753FA1C" bold="true" box="[151,378,1392,1417]" pageId="12" pageNumber="65">Material examined</emphasis>
:
<typeStatus id="C717E4FC42358E5A49C7FA9307DBFA1C" box="[394,498,1392,1417]" pageId="12" pageNumber="65" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="7EBDC29B42358E5A4A0CFA9204A7FA1C" box="[577,654,1393,1417]" pageId="12" pageNumber="65">SCAU</collectionCode>
eGZ7-1) from
<collectingCountry id="60BB1ACE42358E5A4B73FA9305ADFA1C" box="[830,900,1392,1417]" name="China" pageId="12" pageNumber="65">China</collectingCountry>
,
<collectingRegion id="DA6894BC42358E5A4BDDFA930249FA1C" box="[912,1120,1392,1417]" country="China" name="Guizhou" pageId="12" pageNumber="65">Guizhou Province</collectingRegion>
,
<collectingMunicipality id="F877C02442358E5A4C21FA930724FA38" pageId="12" pageNumber="65">Qiandongnan Autonomous Prefecture</collectingMunicipality>
,
<collectingCounty id="F17222D242358E5A4954FA7607C1FA38" box="[281,488,1429,1453]" pageId="12" pageNumber="65">Cenggong County</collectingCounty>
,
<location id="1D730C8542358E5A49B9FA7704EBFA39" LSID="urn:lsid:plazi:treatment:9005EB4842358E5848DAFAE80203FA09:1D730C8542358E5A49B9FA7704EBFA39" box="[500,706,1428,1453]" country="China" county="Cenggong County" municipality="Qiandongnan Autonomous Prefecture" name="Pingzhuang Town" pageId="12" pageNumber="65" stateProvince="Guizhou Province">Pingzhuang Town</location>
,
<location id="1D730C8542358E5A4A82FA770201FA38" LSID="urn:lsid:plazi:treatment:9005EB4842358E5848DAFAE80203FA09:1D730C8542358E5A4A82FA770201FA38" box="[719,1064,1428,1453]" country="China" county="Cenggong County" municipality="Qiandongnan Autonomous Prefecture" name="Wanfuchangcheng Dong Cave" pageId="12" pageNumber="65" stateProvince="Guizhou Province">Wanfuchangcheng Dong Cave</location>
,
<collectingDate id="7C56857642358E5A4C78FA76029EFA38" box="[1077,1207,1429,1453]" pageId="12" pageNumber="65" value="2014-06-29">
<date id="6C127C9E42358E5A4C78FA76029EFA38" box="[1077,1207,1429,1453]" pageId="12" pageNumber="65" value="2014-06-29">29.VI.2014</date>
</collectingDate>
, colls:
<location id="1D730C8542358E5A4D45FA7703B1FA38" LSID="urn:lsid:plazi:treatment:9005EB4842358E5848DAFAE80203FA09:1D730C8542358E5A4D45FA7703B1FA38" box="[1288,1432,1428,1453]" country="China" county="Cenggong County" municipality="Qiandongnan Autonomous Prefecture" name="Mingyi Tian" pageId="12" pageNumber="65" stateProvince="Guizhou Province">Mingyi Tian</location>
,
<location id="1D730C8542358E5A48DAFA5B0731FA44" LSID="urn:lsid:plazi:treatment:9005EB4842358E5848DAFAE80203FA09:1D730C8542358E5A48DAFA5B0731FA44" box="[151,280,1464,1489]" country="China" county="Cenggong County" municipality="Qiandongnan Autonomous Prefecture" name="Weixin Liu" pageId="12" pageNumber="65" stateProvince="Guizhou Province">Weixin Liu</location>
,
<location id="1D730C8542358E5A496EFA5B0798FA44" LSID="urn:lsid:plazi:treatment:9005EB4842358E5848DAFAE80203FA09:1D730C8542358E5A496EFA5B0798FA44" box="[291,433,1464,1489]" country="China" county="Cenggong County" municipality="Qiandongnan Autonomous Prefecture" name="Haomin Yin" pageId="12" pageNumber="65" stateProvince="Guizhou Province">Haomin Yin</location>
,
<location id="1D730C8542358E5A49F6FA5B0474FA44" LSID="urn:lsid:plazi:treatment:9005EB4842358E5848DAFAE80203FA09:1D730C8542358E5A49F6FA5B0474FA44" box="[443,605,1464,1489]" country="China" county="Cenggong County" municipality="Qiandongnan Autonomous Prefecture" name="Sunbin Huang" pageId="12" pageNumber="65" stateProvince="Guizhou Province">Sunbin Huang</location>
and
<location id="1D730C8542358E5A4ADEFA5B0505FA44" LSID="urn:lsid:plazi:treatment:9005EB4842358E5848DAFAE80203FA09:1D730C8542358E5A4ADEFA5B0505FA44" box="[659,812,1464,1489]" country="China" county="Cenggong County" municipality="Qiandongnan Autonomous Prefecture" name="Xinhui Wang" pageId="12" pageNumber="65" stateProvince="Guizhou Province">Xinhui Wang</location>
</materialsCitation>
.
<materialsCitation id="A8C4500342358E5A4B7AFA5A0753FA60" ID-GBIF-Occurrence="1917355135" collectingDate="2014-06-29" collectionCode="SCAU" country="China" county="Cenggong County" location="Pingzhuang Town" municipality="Qiandongnan Autonomous Prefecture" pageId="12" pageNumber="65" specimenCount="28" specimenCount-female="20" specimenCount-male="8" stateProvince="Guizhou Province" typeStatus="paratype">
<typeStatus id="C717E4FC42358E5A4B7AFA5A0580FA44" box="[823,937,1465,1489]" pageId="12" pageNumber="65" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="0EAA91D742358E5A4BFEFA5A0225FA44" box="[947,1036,1464,1489]" pageId="12" pageNumber="65" type="male">8 males</specimenCount>
,
<specimenCount id="0EAA91D742358E5A4C55FA5A02BBFA44" box="[1048,1170,1464,1489]" pageId="12" pageNumber="65" type="female">20 females</specimenCount>
(
<collectionCode id="7EBDC29B42358E5A4CECFA5A02C4FA44" box="[1185,1261,1465,1489]" pageId="12" pageNumber="65">SCAU</collectionCode>
eGZ7-2), same data as the
<typeStatus id="C717E4FC42358E5A4959FA3F0753FA60" box="[276,378,1500,1525]" pageId="12" pageNumber="65" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="50B609D542358E5A488AF9E30530F8AC" pageId="12" pageNumber="65" type="diagnosis">
<paragraph id="18135A5E42358E5A488AF9E3049FF910" blockId="12.[151,1437,1392,2029]" pageId="12" pageNumber="65">
<emphasis id="2AD8864C42358E5A488AF9E30713F98C" bold="true" box="[199,314,1536,1561]" pageId="12" pageNumber="65">Diagnosis</emphasis>
: Adult males of
<taxonomicName id="DFAC21DD42358E5A4A43F9E104A9F98C" authority="Liu &amp; Golovatch, 2018" authorityName="Liu &amp; Golovatch" authorityYear="2018" box="[526,640,1538,1561]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="65" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
<emphasis id="2AD8864C42358E5A4A43F9E104A9F98C" box="[526,640,1538,1561]" italics="true" pageId="12" pageNumber="65">E. parvus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31EB3B3742358E5A4AC0F9E304FBF98C" box="[653,722,1536,1561]" pageId="12" pageNumber="65" rank="species">
<emphasis id="2AD8864C42358E5A4AC0F9E304FBF98C" bold="true" box="[653,722,1536,1561]" pageId="12" pageNumber="65">n. sp.</emphasis>
</taxonomicNameLabel>
are distinct from other
<taxonomicName id="DFAC21DD42358E5A4BB5F9E302B0F98C" baseAuthorityName="Golovatch" baseAuthorityYear="1991" box="[1016,1177,1536,1561]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="2AD8864C42358E5A4BB5F9E302B0F98C" box="[1016,1177,1536,1561]" italics="true" pageId="12" pageNumber="65">Epanerchodus</emphasis>
</taxonomicName>
species based on the following combination of characters: (1) body small (
<emphasis id="2AD8864C42358E5A4ABEF9C50527F9A8" box="[755,782,1574,1597]" italics="true" pageId="12" pageNumber="65">ca</emphasis>
<quantity id="DF54F7BB42358E5A4B58F9C705BAF9A8" box="[789,915,1572,1597]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="1.6" metricValueMin="1.3" pageId="12" pageNumber="65" unit="mm" value="14.5" valueMax="16.0" valueMin="13.0">13–16 mm</quantity>
); (2) collum broader than head (
<figureCitation id="809746DB42358E5A4CB3F9C7034EF9A8" box="[1278,1383,1572,1597]" captionStart="FIGURE 10" captionStartId="13.[151,250,1391,1413]" captionTargetBox="[190,1395,191,1378]" captionTargetId="figure@13.[188,1398,188,1383]" captionTargetPageId="13" captionText="FIGURE 10. Epanerchodus parvus n. sp., male paratype. A: anterior part of body, dorsal view. B: anterior part of body, ventral view. C: midbody rings, dorsal view. D: posterior part of body, dorsal view. E: midbody rings, ventral view. F: posterior part of body, ventral view. G: habitus, dorsal view. Scale bar: A–F, 0.1 mm; G, 0.2 mm." httpUri="https://zenodo.org/record/1458356/files/figure.png" pageId="12" pageNumber="65">Fig. 10A</figureCitation>
); (3) gonopodal clivus broadly triangular; (4) endomere at base with a small, digitiform, lateral process (
<emphasis id="2AD8864C42358E5A4CBDF9AB02D6F9F4" bold="true" box="[1264,1279,1608,1633]" pageId="12" pageNumber="65">p</emphasis>
), while its tip beak-shaped and denticulate near apex (
<figureCitation id="809746DB42358E5A4A1AF98F0480F910" box="[599,681,1644,1669]" captionStart="FIGURE 11" captionStartId="14.[151,250,1216,1238]" captionTargetBox="[172,1407,361,1167]" captionTargetId="figure@14.[151,1436,337,1195]" captionTargetPageId="14" captionText="FIGURE 11. Epanerchodus parvus n. sp., male paratype. A: left gonopod, mesal view. B: left gonopod, lateral view. Abbreviations: cl = clivus; en = endomere; p = process." httpUri="https://zenodo.org/record/1458358/files/figure.png" pageId="12" pageNumber="65">Fig. 11</figureCitation>
).
</paragraph>
<paragraph id="18135A5E42358E5A488AF9730530F8AC" blockId="12.[151,1437,1392,2029]" pageId="12" pageNumber="65">
The former character is similar to that observed in
<taxonomicName id="DFAC21DD42358E5A4B59F97105B4F93C" authority="Liu &amp; Golovatch, 2018" authorityName="Liu &amp; Golovatch" authorityYear="2018" box="[788,925,1680,1705]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="65" phylum="Arthropoda" rank="species" species="gladiatus" status="sp. nov.">
<emphasis id="2AD8864C42358E5A4B59F97105B4F93C" box="[788,925,1680,1705]" italics="true" pageId="12" pageNumber="65">E. gladiatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31EB3B3742358E5A4BEAF97305C1F93C" box="[935,1000,1680,1705]" pageId="12" pageNumber="65" rank="species">
<emphasis id="2AD8864C42358E5A4BEAF97305C1F93C" bold="true" box="[935,1000,1680,1705]" pageId="12" pageNumber="65">n. sp.</emphasis>
</taxonomicNameLabel>
,
<taxonomicName id="DFAC21DD42358E5A4BB4F971024EF93C" authority="Liu &amp; Golovatch, 2018" authorityName="Liu &amp; Golovatch" authorityYear="2018" box="[1017,1127,1680,1705]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="65" phylum="Arthropoda" rank="species" species="chutou" status="sp. nov.">
<emphasis id="2AD8864C42358E5A4BB4F971024EF93C" box="[1017,1127,1680,1705]" italics="true" pageId="12" pageNumber="65">E. chutou</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31EB3B3742358E5A4C3DF9730298F93C" box="[1136,1201,1680,1705]" pageId="12" pageNumber="65" rank="species">
<emphasis id="2AD8864C42358E5A4C3DF9730298F93C" bold="true" box="[1136,1201,1680,1705]" pageId="12" pageNumber="65">n. sp.</emphasis>
</taxonomicNameLabel>
(both from caves in
<collectingRegion id="DA6894BC42358E5A48DAF95706D3F958" box="[151,250,1716,1741]" country="China" name="Guizhou" pageId="12" pageNumber="65">Guizhou</collectingRegion>
,
<collectingCountry id="60BB1ACE42358E5A4947F9570766F958" box="[266,335,1716,1741]" name="China" pageId="12" pageNumber="65">China</collectingCountry>
) and
<taxonomicName id="DFAC21DD42358E5A49D9F95507D7F958" box="[404,510,1717,1741]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="65" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="2AD8864C42358E5A49D9F95507D7F958" box="[404,510,1717,1741]" italics="true" pageId="12" pageNumber="65">E. fuscus</emphasis>
</taxonomicName>
(from
<collectingRegion id="DA6894BC42358E5A4A19F9560484F959" box="[596,685,1717,1740]" country="China" name="Yunnan" pageId="12" pageNumber="65">Yunnan</collectingRegion>
,
<collectingCountry id="60BB1ACE42358E5A4AF1F957052DF958" box="[700,772,1716,1741]" name="China" pageId="12" pageNumber="65">China</collectingCountry>
). However,
<taxonomicName id="DFAC21DD42358E5A4BDFF9550228F958" authority="Liu &amp; Golovatch, 2018" authorityName="Liu &amp; Golovatch" authorityYear="2018" box="[914,1025,1718,1741]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="65" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
<emphasis id="2AD8864C42358E5A4BDFF9550228F958" box="[914,1025,1718,1741]" italics="true" pageId="12" pageNumber="65">E. parvus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31EB3B3742358E5A4C40F9570279F958" box="[1037,1104,1716,1741]" pageId="12" pageNumber="65" rank="species">
<emphasis id="2AD8864C42358E5A4C40F9570279F958" bold="true" box="[1037,1104,1716,1741]" pageId="12" pageNumber="65">n. sp.</emphasis>
</taxonomicNameLabel>
is distinguished not only in gonopodal structural details, but also by the narrower body: widths of pro- and metazonae 0.8–1.0 and
<quantity id="DF54F7BB42358E5A4D5BF93B03B4F965" box="[1302,1437,1752,1777]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="12" pageNumber="65" unit="mm" value="1.4" valueMax="1.5" valueMin="1.3">1.3–1.5 mm</quantity>
<emphasis id="2AD8864C42358E5A48DAF91D069CF880" box="[151,181,1790,1813]" italics="true" pageId="12" pageNumber="65">vs.</emphasis>
clearly broader in
<taxonomicName id="DFAC21DD42358E5A49C7F91D0438F880" authority="Liu &amp; Golovatch, 2018" authorityName="Liu &amp; Golovatch" authorityYear="2018" box="[394,529,1788,1813]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="65" phylum="Arthropoda" rank="species" species="gladiatus" status="sp. nov.">
<emphasis id="2AD8864C42358E5A49C7F91D0438F880" box="[394,529,1788,1813]" italics="true" pageId="12" pageNumber="65">E. gladiatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31EB3B3742358E5A4A55F91F047EF880" box="[536,599,1788,1813]" pageId="12" pageNumber="65" rank="species">
<emphasis id="2AD8864C42358E5A4A55F91F047EF880" bold="true" box="[536,599,1788,1813]" pageId="12" pageNumber="65">n. sp.</emphasis>
</taxonomicNameLabel>
,
<taxonomicName id="DFAC21DD42358E5A4A2BF91D04F8F880" authority="Liu &amp; Golovatch, 2018" authorityName="Liu &amp; Golovatch" authorityYear="2018" box="[614,721,1788,1813]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="65" phylum="Arthropoda" rank="species" species="chutou" status="sp. nov.">
<emphasis id="2AD8864C42358E5A4A2BF91D04F8F880" box="[614,721,1788,1813]" italics="true" pageId="12" pageNumber="65">E. chutou</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31EB3B3742358E5A4A94F91F0531F880" box="[729,792,1788,1813]" pageId="12" pageNumber="65" rank="species">
<emphasis id="2AD8864C42358E5A4A94F91F0531F880" bold="true" box="[729,792,1788,1813]" pageId="12" pageNumber="65">n. sp.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="DFAC21DD42358E5A4B02F91D059DF880" box="[847,948,1789,1813]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="65" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="2AD8864C42358E5A4B02F91D059DF880" box="[847,948,1789,1813]" italics="true" pageId="12" pageNumber="65">E. fuscus</emphasis>
</taxonomicName>
: widths of pro- and metazonae 1.2–1.3 and 2.5–3.0, 1–1.2 and 2.0, and 2.3 and 4.0 mm, respectively.
</paragraph>
</subSubSection>
<subSubSection id="50B609D542358E58488AF8A70722FAED" lastPageId="14" lastPageNumber="67" pageId="12" pageNumber="65" type="description">
<paragraph id="18135A5E42358E58488AF8A70241FE88" blockId="12.[151,1437,1392,2029]" lastBlockId="14.[151,1436,151,285]" lastPageId="14" lastPageNumber="67" pageId="12" pageNumber="65">
<emphasis id="2AD8864C42358E5A488AF8A70778F8C8" bold="true" box="[199,337,1860,1885]" pageId="12" pageNumber="65">Description</emphasis>
: Based on
<typeStatus id="C717E4FC42358E5A4998F8A6042CF8C8" box="[469,517,1861,1885]" pageId="12" pageNumber="65">type</typeStatus>
specimens. Lengths of body
<emphasis id="2AD8864C42358E5A4B15F8A5055AF8C8" box="[856,883,1862,1885]" italics="true" pageId="12" pageNumber="65">ca</emphasis>
<quantity id="DF54F7BB42358E5A4B31F8A70208F8C9" box="[892,1057,1860,1885]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="1.55" metricValueMin="1.35" pageId="12" pageNumber="65" unit="mm" value="14.5" valueMax="15.5" valueMin="13.5">13.5–15.5 mm</quantity>
(males) or
<quantity id="DF54F7BB42358E5A4CEAF8A7030BF8C9" box="[1191,1314,1860,1885]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="12" pageNumber="65" unit="mm" value="16.0" valueMax="17.0" valueMin="15.0">15–17 mm</quantity>
(females), widths of pro- and metazonae 0.8–1.0 and
<quantity id="DF54F7BB42358E5A4A3FF88B04D0F815" box="[626,761,1896,1921]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="12" pageNumber="65" unit="mm" value="1.4" valueMax="1.5" valueMin="1.3">1.3–1.5 mm</quantity>
(males) or 1.01–1.2 and
<quantity id="DF54F7BB42358E5A4C42F88A02BFF815" box="[1039,1174,1897,1921]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="12" pageNumber="65" unit="mm" value="1.75" valueMax="1.8" valueMin="1.7">1.7–1.8 mm</quantity>
(females).
<emphasis id="2AD8864C42358E5A4D5EF88B03BDF814" bold="true" box="[1299,1428,1896,1921]" pageId="12" pageNumber="65">Coloration</emphasis>
: in alcohol generally pallid. Mouthparts and gonopodal telopodites light yellowish.
<emphasis id="2AD8864C42358E5A4C1BF86F02BDF830" bold="true" box="[1110,1172,1932,1957]" pageId="12" pageNumber="65">Body</emphasis>
: Adults with 20 rings. Width: collum &lt;head = rings 2–3 &lt;4 &lt;5–15, thereafter body gradually tapering towards telson.
<emphasis id="2AD8864C42358E5A4CBBF853031CF85C" bold="true" box="[1270,1333,1968,1993]" pageId="12" pageNumber="65">Head</emphasis>
: clypeus densely pilose, vertex sparsely setose, epicranial suture conspicuous (
<figureCitation id="809746DB42358E5A4BECF837022EF878" box="[929,1031,2004,2029]" captionStart="FIGURE 10" captionStartId="13.[151,250,1391,1413]" captionTargetBox="[190,1395,191,1378]" captionTargetId="figure@13.[188,1398,188,1383]" captionTargetPageId="13" captionText="FIGURE 10. Epanerchodus parvus n. sp., male paratype. A: anterior part of body, dorsal view. B: anterior part of body, ventral view. C: midbody rings, dorsal view. D: posterior part of body, dorsal view. E: midbody rings, ventral view. F: posterior part of body, ventral view. G: habitus, dorsal view. Scale bar: A–F, 0.1 mm; G, 0.2 mm." httpUri="https://zenodo.org/record/1458356/files/figure.png" pageId="12" pageNumber="65">Fig. 10B</figureCitation>
). Antennae long, reaching behind posterior margin of ring 3 when extended posteriorly, slightly clavate (
<figureCitation id="809746DB42348E5B4BE1FA0C0248F99D" box="[940,1121,1519,1544]" captionStart="FIGURE 10" captionStartId="13.[151,250,1391,1413]" captionTargetBox="[190,1395,191,1378]" captionTargetId="figure@13.[188,1398,188,1383]" captionTargetPageId="13" captionText="FIGURE 10. Epanerchodus parvus n. sp., male paratype. A: anterior part of body, dorsal view. B: anterior part of body, ventral view. C: midbody rings, dorsal view. D: posterior part of body, dorsal view. E: midbody rings, ventral view. F: posterior part of body, ventral view. G: habitus, dorsal view. Scale bar: A–F, 0.1 mm; G, 0.2 mm." httpUri="https://zenodo.org/record/1458356/files/figure.png" pageId="13" pageNumber="66">Figs. 10A, B, G</figureCitation>
).
<emphasis id="2AD8864C42348E5B4C39FA0C032DF99D" bold="true" box="[1140,1284,1519,1544]" pageId="13" pageNumber="66">Exoskeleton</emphasis>
: Collum fanshaped, with a lateral incision on each side (
<figureCitation id="809746DB42348E5B4AEBF9F0053BF9B9" box="[678,786,1555,1580]" captionStart="FIGURE 10" captionStartId="13.[151,250,1391,1413]" captionTargetBox="[190,1395,191,1378]" captionTargetId="figure@13.[188,1398,188,1383]" captionTargetPageId="13" captionText="FIGURE 10. Epanerchodus parvus n. sp., male paratype. A: anterior part of body, dorsal view. B: anterior part of body, ventral view. C: midbody rings, dorsal view. D: posterior part of body, dorsal view. E: midbody rings, ventral view. F: posterior part of body, ventral view. G: habitus, dorsal view. Scale bar: A–F, 0.1 mm; G, 0.2 mm." httpUri="https://zenodo.org/record/1458356/files/figure.png" pageId="13" pageNumber="66">Fig. 10A</figureCitation>
). Paraterga narrow (
<figureCitation id="809746DB42348E5B4C4DF9F0027EF9B9" box="[1024,1111,1555,1580]" captionStart="FIGURE 10" captionStartId="13.[151,250,1391,1413]" captionTargetBox="[190,1395,191,1378]" captionTargetId="figure@13.[188,1398,188,1383]" captionTargetPageId="13" captionText="FIGURE 10. Epanerchodus parvus n. sp., male paratype. A: anterior part of body, dorsal view. B: anterior part of body, ventral view. C: midbody rings, dorsal view. D: posterior part of body, dorsal view. E: midbody rings, ventral view. F: posterior part of body, ventral view. G: habitus, dorsal view. Scale bar: A–F, 0.1 mm; G, 0.2 mm." httpUri="https://zenodo.org/record/1458356/files/figure.png" pageId="13" pageNumber="66">Fig. 10</figureCitation>
), midbody paraterga extend metatergite to
<emphasis id="2AD8864C42348E5B4973F9DB0770F9DA" box="[318,345,1592,1615]" italics="true" pageId="13" pageNumber="66">ca</emphasis>
1.5x width of prozonite. Paraterga on collum and rings 2–4 clearly upturned dorsally above a faintly convex dorsum, other paraterga flat. From ring 10 on, caudolateral corners of paraterga increasingly acuteangled, sharper and pointed on rings 15–19 (
<figureCitation id="809746DB42348E5B4AD9F99C04D7F90D" box="[660,766,1663,1688]" captionStart="FIGURE 10" captionStartId="13.[151,250,1391,1413]" captionTargetBox="[190,1395,191,1378]" captionTargetId="figure@13.[188,1398,188,1383]" captionTargetPageId="13" captionText="FIGURE 10. Epanerchodus parvus n. sp., male paratype. A: anterior part of body, dorsal view. B: anterior part of body, ventral view. C: midbody rings, dorsal view. D: posterior part of body, dorsal view. E: midbody rings, ventral view. F: posterior part of body, ventral view. G: habitus, dorsal view. Scale bar: A–F, 0.1 mm; G, 0.2 mm." httpUri="https://zenodo.org/record/1458356/files/figure.png" pageId="13" pageNumber="66">Fig. 10D</figureCitation>
). Integument shining, translucent, prozonae very delicately alveolate. Limbus regularly denticulate. Constriction between pro- and metazonae narrow, shallow and smooth. Metatergal sculpture faint, with three irregular transverse rows of setigerous, polygonal bosses. Sulcus between front and middle rows of setae a little deeper than that between middle and caudal rows. Tergal setae invisible. Three or four faint setigerous incisions at lateral margins of poreless and pore-bearing rings, respectively. Pore formula normal: 5, 7, 9, 10, 12, 13, 15–19, ozopores evident, dorsal, clearly set off from lateral margin and located between last and penultimate marginal incisions. Epiproct short, conical, pre-apical lateral papillae small. Hypoproct subtrapeziform, with two setigerous papillae. Pleurosternal carinae small, present only on ring 2. Sterna sparsely setose, cross-shaped impressions shallow. Legs long and slender (
<figureCitation id="809746DB42348E5B4B93F87C0218F82D" box="[990,1073,1951,1976]" captionStart="FIGURE 10" captionStartId="13.[151,250,1391,1413]" captionTargetBox="[190,1395,191,1378]" captionTargetId="figure@13.[188,1398,188,1383]" captionTargetPageId="13" captionText="FIGURE 10. Epanerchodus parvus n. sp., male paratype. A: anterior part of body, dorsal view. B: anterior part of body, ventral view. C: midbody rings, dorsal view. D: posterior part of body, dorsal view. E: midbody rings, ventral view. F: posterior part of body, ventral view. G: habitus, dorsal view. Scale bar: A–F, 0.1 mm; G, 0.2 mm." httpUri="https://zenodo.org/record/1458356/files/figure.png" pageId="13" pageNumber="66">Fig. 10</figureCitation>
), about 2.0–2.2 times as long as body ring height in both sexes, without sphaerotrichomes or sternal cones, prefemora not bulging laterally.
<emphasis id="2AD8864C42378E5848DAFF740739FF25" bold="true" box="[151,272,151,176]" pageId="14" pageNumber="67">Gonopods</emphasis>
: slender (
<figureCitation id="809746DB42378E5849C9FF7407F0FF25" box="[388,473,151,176]" captionStart="FIGURE 11" captionStartId="14.[151,250,1216,1238]" captionTargetBox="[172,1407,361,1167]" captionTargetId="figure@14.[151,1436,337,1195]" captionTargetPageId="14" captionText="FIGURE 11. Epanerchodus parvus n. sp., male paratype. A: left gonopod, mesal view. B: left gonopod, lateral view. Abbreviations: cl = clivus; en = endomere; p = process." httpUri="https://zenodo.org/record/1458358/files/figure.png" pageId="14" pageNumber="67">Fig. 11</figureCitation>
). Telopodite mostly suberect. Prefemur densely setose and nearly half the length of telopodite. Clivus (
<emphasis id="2AD8864C42378E584923FF5F07ABFF40" bold="true" box="[366,386,188,213]" pageId="14" pageNumber="67">cl</emphasis>
) broadly triangular. Endomere (
<emphasis id="2AD8864C42378E584AA0FF5F0520FF40" bold="true" box="[749,777,188,213]" pageId="14" pageNumber="67">en</emphasis>
) tip beak-shaped, denticulate near apex. Endomere at base with a small, digitiform, lateral process (
<emphasis id="2AD8864C42378E584A2EFF3C045BFF6D" bold="true" box="[611,626,223,248]" pageId="14" pageNumber="67">p</emphasis>
). Seminal groove starting mesally, then recurving laterad to run into an accessory seminal chamber, the latter opening on a hairy pulvillus. An exomere absent.
</paragraph>
<caption id="4CD30AD642348E5B48DAFA8C05EBFA57" httpUri="https://zenodo.org/record/1458356/files/figure.png" pageId="13" pageNumber="66" startId="13.[151,250,1391,1413]" targetBox="[190,1395,191,1378]" targetPageId="13">
<paragraph id="18135A5E42348E5B48DAFA8C05EBFA57" blockId="13.[151,1436,1391,1474]" pageId="13" pageNumber="66">
<emphasis id="2AD8864C42348E5B48DAFA8C0736FA10" bold="true" box="[151,287,1391,1413]" pageId="13" pageNumber="66">FIGURE 10.</emphasis>
<taxonomicName id="DFAC21DD42348E5B4969FA8C0429FA10" authority="Liu &amp; Golovatch, 2018" authorityName="Liu &amp; Golovatch" authorityYear="2018" box="[292,512,1391,1413]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="13" pageNumber="66" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
<emphasis id="2AD8864C42348E5B4969FA8C0429FA10" box="[292,512,1391,1413]" italics="true" pageId="13" pageNumber="66">Epanerchodus parvus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31EB3B3742348E5B4A4BFA8C0416FA10" box="[518,575,1391,1413]" pageId="13" pageNumber="66" rank="species">
<emphasis id="2AD8864C42348E5B4A4BFA8C0416FA10" bold="true" box="[518,575,1391,1413]" pageId="13" pageNumber="66">n. sp.</emphasis>
</taxonomicNameLabel>
, male paratype.
<emphasis id="2AD8864C42348E5B4AA8FA8C04DEFA10" bold="true" box="[741,759,1391,1413]" pageId="13" pageNumber="66">A</emphasis>
: anterior part of body, dorsal view.
<emphasis id="2AD8864C42348E5B4C13FA8C0246FA10" bold="true" box="[1118,1135,1391,1413]" pageId="13" pageNumber="66">B</emphasis>
: anterior part of body, ventral view.
<emphasis id="2AD8864C42348E5B489EFA6D06CCFA31" bold="true" box="[211,229,1422,1444]" pageId="13" pageNumber="66">C</emphasis>
: midbody rings, dorsal view.
<emphasis id="2AD8864C42348E5B4A5CFA6D040AFA31" bold="true" box="[529,547,1422,1444]" pageId="13" pageNumber="66">D</emphasis>
: posterior part of body, dorsal view.
<emphasis id="2AD8864C42348E5B4BDAFA6D0581FA31" bold="true" box="[919,936,1422,1444]" pageId="13" pageNumber="66">E</emphasis>
: midbody rings, ventral view.
<emphasis id="2AD8864C42348E5B4C91FA6D02C2FA31" bold="true" box="[1244,1259,1422,1444]" pageId="13" pageNumber="66">F</emphasis>
: posterior part of body, ventral view.
<emphasis id="2AD8864C42348E5B4910FA4F0758FA57" bold="true" box="[349,369,1452,1474]" pageId="13" pageNumber="66">G</emphasis>
: habitus, dorsal view. Scale bar: A–F, 0.1 mm; G, 0.2 mm.
</paragraph>
</caption>
<caption id="4CD30AD642378E5848DAFB2304E0FB60" httpUri="https://zenodo.org/record/1458358/files/figure.png" pageId="14" pageNumber="67" startId="14.[151,250,1216,1238]" targetBox="[172,1407,361,1167]" targetPageId="14">
<paragraph id="18135A5E42378E5848DAFB2304E0FB60" blockId="14.[151,1437,1216,1269]" pageId="14" pageNumber="67">
<emphasis id="2AD8864C42378E5848DAFB23070DFB43" bold="true" box="[151,292,1216,1238]" pageId="14" pageNumber="67">FIGURE 11.</emphasis>
<taxonomicName id="DFAC21DD42378E58497DFB230438FB42" authority="Liu &amp; Golovatch, 2018" authorityName="Liu &amp; Golovatch" authorityYear="2018" box="[304,529,1216,1239]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="67" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
<emphasis id="2AD8864C42378E58497DFB230438FB42" box="[304,529,1216,1239]" italics="true" pageId="14" pageNumber="67">Epanerchodus parvus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31EB3B3742378E584A53FB230474FB43" box="[542,605,1216,1238]" pageId="14" pageNumber="67" rank="species">
<emphasis id="2AD8864C42378E584A53FB230474FB43" bold="true" box="[542,605,1216,1238]" pageId="14" pageNumber="67">n. sp.</emphasis>
</taxonomicNameLabel>
, male paratype.
<emphasis id="2AD8864C42378E584B5BFB230501FB43" bold="true" box="[790,808,1216,1238]" pageId="14" pageNumber="67">A</emphasis>
: left gonopod, mesal view.
<emphasis id="2AD8864C42378E584C13FB230246FB43" bold="true" box="[1118,1135,1216,1238]" pageId="14" pageNumber="67">B</emphasis>
: left gonopod, lateral view. Abbreviations: cl = clivus; en = endomere; p = process.
</paragraph>
</caption>
<paragraph id="18135A5E42378E58488AFAD80722FAED" blockId="14.[151,1436,1339,1436]" pageId="14" pageNumber="67">
<emphasis id="2AD8864C42378E58488AFAD806D6FAC1" bold="true" box="[199,255,1339,1364]" pageId="14" pageNumber="67">Note</emphasis>
: Because of the long slender antennae and legs, and a depigmented cuticle, this species is most likely a troglobite.
</paragraph>
</subSubSection>
<subSubSection id="50B609D542378E58488AFA600203FA09" box="[199,1066,1411,1436]" pageId="14" pageNumber="67" type="etymology">
<paragraph id="18135A5E42378E58488AFA600203FA09" blockId="14.[151,1436,1339,1436]" box="[199,1066,1411,1436]" pageId="14" pageNumber="67">
<emphasis id="2AD8864C42378E58488AFA600761FA09" bold="true" box="[199,328,1411,1436]" pageId="14" pageNumber="67">Etymology</emphasis>
: To emphasize the small size of this species; masculine adjective.
</paragraph>
</subSubSection>
</treatment>
</document>
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