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<mods:title id="CAEAC472AF8FBCC5858A888AD5393276">The millipede genus Epanerchodus Attems, 1901 in continental China, with descriptions of seven new cavernicolous species (Diplopoda, Polydesmida, Polydesmidae)</mods:title>
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<mods:namePart id="3E25B773F37492DF4482A3047FCF2311">Liu, Weixin</mods:namePart>
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<treatment id="9005EB48422D8E4048DAF94604C4F993" ID-DOI="http://doi.org/10.5281/zenodo.5973784" ID-GBIF-Taxon="148692465" ID-Zenodo-Dep="5973784" LSID="urn:lsid:plazi:treatment:9005EB48422D8E4048DAF94604C4F993" httpUri="http://treatment.plazi.org/id/9005EB48422D8E4048DAF94604C4F993" lastPageId="22" lastPageNumber="75" pageId="20" pageNumber="73">
<subSubSection id="50B609D5422D8E4248DAF9460735F975" pageId="20" pageNumber="73" type="nomenclature">
<paragraph id="18135A5E422D8E4248DAF94604D2F92A" blockId="20.[151,763,1701,1761]" box="[151,763,1701,1727]" pageId="20" pageNumber="73">
<heading id="435BED32422D8E4248DAF94604D2F92A" bold="true" box="[151,763,1701,1727]" fontSize="11" level="1" pageId="20" pageNumber="73" reason="1">
<taxonomicName id="DFAC21DD422D8E4248DAF94604D2F92A" ID-CoL="6FN5L" authority="Golovatch &amp; Geoffroy, 2014" authorityName="Golovatch &amp; Geoffroy" authorityYear="2014" box="[151,763,1701,1727]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="20" pageNumber="73" phylum="Arthropoda" rank="species" species="lipsae">
<emphasis id="2AD8864C422D8E4248DAF94604D2F92A" bold="true" box="[151,763,1701,1727]" pageId="20" pageNumber="73">
<emphasis id="2AD8864C422D8E4248DAF94607BDF92A" bold="true" box="[151,404,1701,1727]" italics="true" pageId="20" pageNumber="73">Epanerchodus lipsae</emphasis>
<bibRefCitation id="7C3D27AF422D8E4249D6F94604D2F92A" box="[411,763,1701,1727]" pageId="20" pageNumber="73" refId="ref12900" refString="Golovatch, S. I. &amp; Geoffroy, J. - J. (2014) On some new or poorly-known species of the millipede family Polydesmidae from southern China (Diplopoda: Polydesmida). Russian Entomological Journal, 23 (2), 91 - 105." type="journal article">Golovatch &amp; Geoffroy, 2014</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="18135A5E422D8E4248DAF92B0735F975" blockId="20.[151,763,1701,1761]" box="[151,284,1736,1761]" pageId="20" pageNumber="73">
<figureCitation id="809746DB422D8E4248DAF92B0731F975" box="[151,280,1736,1761]" captionStart-0="FIGURE 18" captionStart-1="FIGURE 19" captionStartId-0="21.[151,250,1967,1989]" captionStartId-1="22.[151,250,1185,1207]" captionTargetBox-0="[313,1273,245,1908]" captionTargetBox-1="[163,1416,219,1152]" captionTargetId-0="figure@21.[303,1283,245,1908]" captionTargetId-1="figure@22.[151,1436,193,1164]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionText-0="FIGURE 18. Epanerchodus lipsae Golovatch and Geoffroy 2014, male from Wolong Dong Cave. A: habitus, ventral view. B: habitus, dorsal view. Scale bar: 0.5 mm." captionText-1="FIGURE 19. Epanerchodus lipsae Golovatch and Geoffroy 2014, male from Wolong Dong Cave. A: right gonopod, lateral view. B: right gonopod, mesal view. Abbreviations: a = spiniform process; cl = clivus; en = endomere; ex = exomere; p = process." httpUri-0="https://zenodo.org/record/1458372/files/figure.png" httpUri-1="https://zenodo.org/record/1458374/files/figure.png" pageId="20" pageNumber="73">Figs 1819</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="50B609D5422D8E4248DAF8F3058AF8B3" box="[151,931,1808,1830]" pageId="20" pageNumber="73" type="reference_group">
<paragraph id="18135A5E422D8E4248DAF8F3058AF8B3" blockId="20.[151,931,1808,1830]" box="[151,931,1808,1830]" pageId="20" pageNumber="73">
<treatmentCitationGroup id="38BC7D70422D8E4248DAF8F3058AF8B3" box="[151,931,1808,1830]" pageId="20" pageNumber="73">
<taxonomicName id="DFAC21DD422D8E4248DAF8F304F2F8B3" ID-CoL="6FN5L" authority="Golovatch and Geoffroy, 2014: 102" authorityName="Golovatch and Geoffroy" authorityPageNumber="102" authorityYear="2014" box="[151,731,1808,1830]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="20" pageNumber="73" phylum="Arthropoda" rank="species" species="lipsae">
<emphasis id="2AD8864C422D8E4248DAF8F30743F8B3" box="[151,362,1808,1830]" italics="true" pageId="20" pageNumber="73">Epanerchodus lipsae</emphasis>
<treatmentCitation id="990D7C4F422D8E42493CF8F304F2F8B3" author="" box="[369,731,1808,1830]" page="102" pageId="20" pageNumber="73">
<bibRefCitation id="7C3D27AF422D8E42493CF8F30480F8B3" box="[369,681,1808,1830]" pageId="20" pageNumber="73" refId="ref12900" refString="Golovatch, S. I. &amp; Geoffroy, J. - J. (2014) On some new or poorly-known species of the millipede family Polydesmidae from southern China (Diplopoda: Polydesmida). Russian Entomological Journal, 23 (2), 91 - 105." type="journal article">Golovatch and Geoffroy, 2014</bibRefCitation>
: 102
</treatmentCitation>
</taxonomicName>
, figs 1720, map 3.
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</paragraph>
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<subSubSection id="50B609D5422D8E4048DAF8B104C4F993" lastPageId="22" lastPageNumber="75" pageId="20" pageNumber="73" type="materials_examined">
<paragraph id="18135A5E422D8E4248DAF8B1031AF84D" blockId="20.[151,1436,1874,2008]" pageId="20" pageNumber="73">
<materialsCitation id="A8C45003422D8E4248DAF8B10506F81A" ID-GBIF-Occurrence="1917355113" collectingDate="2014-07-15" collectionCode="SCAU" country="China" county="Beichuan County" location="Xunlongshan Mountain" municipality="Mianyang City" pageId="20" pageNumber="73" specimenCount="14" specimenCount-female="8" specimenCount-male="6" stateProvince="Sichuan Province">
<emphasis id="2AD8864C422D8E4248DAF8B1075FF8FE" bold="true" box="[151,374,1874,1899]" pageId="20" pageNumber="73">Material examined</emphasis>
:
<specimenCount id="0EAA91D7422D8E4249CBF8B107F7F8FE" box="[390,478,1874,1899]" pageId="20" pageNumber="73" type="male">6 males</specimenCount>
,
<specimenCount id="0EAA91D7422D8E4249A6F8B0047FF8FE" box="[491,598,1874,1899]" pageId="20" pageNumber="73" type="female">8 females</specimenCount>
(
<collectionCode id="7EBDC29B422D8E424A2BF8B0049BF8FE" box="[614,690,1875,1899]" pageId="20" pageNumber="73">SCAU</collectionCode>
) from
<collectingCountry id="60BB1ACE422D8E424B4CF8B1056CF8FE" box="[769,837,1874,1899]" name="China" pageId="20" pageNumber="73">China</collectingCountry>
,
<collectingRegion id="DA6894BC422D8E424B1CF8B1023FF8FE" box="[849,1046,1874,1899]" country="China" name="Sichuan" pageId="20" pageNumber="73">Sichuan Province</collectingRegion>
,
<collectingMunicipality id="F877C024422D8E424C6FF8B102E3F8FE" box="[1058,1226,1874,1899]" pageId="20" pageNumber="73">Mianyang City</collectingMunicipality>
,
<collectingCounty id="F17222D2422D8E424C9AF8B103B1F8FE" box="[1239,1432,1874,1899]" pageId="20" pageNumber="73">Beichuan County</collectingCounty>
,
<location id="1D730C85422D8E4248DAF895078BF81A" LSID="urn:lsid:plazi:treatment:9005EB48422D8E4048DAF94604C4F993:1D730C85422D8E4248DAF895078BF81A" box="[151,418,1910,1935]" country="China" county="Beichuan County" municipality="Mianyang City" name="Xunlongshan Mountain" pageId="20" pageNumber="73" stateProvince="Sichuan Province">Xunlongshan Mountain</location>
,
<location id="1D730C85422D8E4249FFF89504BEF81A" LSID="urn:lsid:plazi:treatment:9005EB48422D8E4048DAF94604C4F993:1D730C85422D8E4249FFF89504BEF81A" box="[434,663,1910,1936]" country="China" county="Beichuan County" municipality="Mianyang City" name="Wolong Dong Cave" pageId="20" pageNumber="73" stateProvince="Sichuan Province">Wolong Dong Cave</location>
,
<collectingDate id="7C568576422D8E424AEAF8950506F81A" box="[679,815,1910,1935]" pageId="20" pageNumber="73" value="2014-07-15">
<date id="6C127C9E422D8E424AEAF8950506F81A" box="[679,815,1910,1935]" pageId="20" pageNumber="73" value="2014-07-15">15.VII.2014</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="A8C45003422D8E424B0DF894040EF826" ID-GBIF-Occurrence="1917355146" collectingDate="2014-07-16" collectionCode="SCAU" country="China" location="Yuanwang Dong Cave" pageId="20" pageNumber="73" specimenCount="34" specimenCount-female="26" specimenCount-male="8" stateProvince="Sichuan Province">
<specimenCount id="0EAA91D7422D8E424B0DF89405B2F81A" box="[832,923,1910,1935]" pageId="20" pageNumber="73" type="male">8 males</specimenCount>
,
<specimenCount id="0EAA91D7422D8E424BE1F8950201F81A" box="[940,1064,1910,1935]" pageId="20" pageNumber="73" type="female">26 females</specimenCount>
(
<collectionCode id="7EBDC29B422D8E424C76F89402AEF81A" box="[1083,1159,1911,1935]" pageId="20" pageNumber="73">SCAU</collectionCode>
) from the same county,
<location id="1D730C85422D8E4248DAF87807BBF826" LSID="urn:lsid:plazi:treatment:9005EB48422D8E4048DAF94604C4F993:1D730C85422D8E4248DAF87807BBF826" box="[151,402,1947,1971]" country="China" name="Yuanwang Dong Cave" pageId="20" pageNumber="73" stateProvince="Sichuan Province">Yuanwang Dong Cave</location>
,
<collectingDate id="7C568576422D8E4249D3F879040EF826" box="[414,551,1946,1971]" pageId="20" pageNumber="73" value="2014-07-16">
<date id="6C127C9E422D8E4249D3F879040EF826" box="[414,551,1946,1971]" pageId="20" pageNumber="73" value="2014-07-16">16.VII.2014</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="A8C45003422D8E424A7EF878031AF84D" ID-GBIF-Occurrence="1917355126" collectingDate="2014-07-19" collectionCode="SCAU" country="China" county="Jiangyou City" location="Shuimen Dong Cave" municipality="Hanzeng Town" pageId="20" pageNumber="73" specimenCount="29" specimenCount-female="19" specimenCount-male="10" stateProvince="Sichuan Province">
<specimenCount id="0EAA91D7422D8E424A7EF87804B0F826" box="[563,665,1946,1971]" pageId="20" pageNumber="73" type="male">10 males</specimenCount>
,
<specimenCount id="0EAA91D7422D8E424AE8F8780535F826" box="[677,796,1946,1971]" pageId="20" pageNumber="73" type="female">19 females</specimenCount>
(
<collectionCode id="7EBDC29B422D8E424B61F8780550F826" box="[812,889,1947,1971]" pageId="20" pageNumber="73">SCAU</collectionCode>
) from
<collectingRegion id="DA6894BC422D8E424B88F87902A0F826" box="[965,1161,1946,1971]" country="China" name="Sichuan" pageId="20" pageNumber="73">Sichuan Province</collectingRegion>
,
<collectingCounty id="F17222D2422D8E424CD9F8790306F826" box="[1172,1327,1946,1971]" pageId="20" pageNumber="73">Jiangyou City</collectingCounty>
,
<collectingMunicipality id="F877C024422D8E424D77F87806F1F842" pageId="20" pageNumber="73">Hanzeng Town</collectingMunicipality>
,
<location id="1D730C85422D8E4248AEF85D07E3F842" LSID="urn:lsid:plazi:treatment:9005EB48422D8E4048DAF94604C4F993:1D730C85422D8E4248AEF85D07E3F842" box="[227,458,1982,2008]" country="China" county="Jiangyou City" municipality="Hanzeng Town" name="Shuimen Dong Cave" pageId="20" pageNumber="73" stateProvince="Sichuan Province">Shuimen Dong Cave</location>
,
<collectingDate id="7C568576422D8E424998F85C0474F842" box="[469,605,1983,2007]" pageId="20" pageNumber="73" value="2014-07-19">
<date id="6C127C9E422D8E424998F85C0474F842" box="[469,605,1983,2007]" pageId="20" pageNumber="73" value="2014-07-19">19.VII.2014</date>
</collectingDate>
, colls:
<location id="1D730C85422D8E424AE4F85D050EF842" LSID="urn:lsid:plazi:treatment:9005EB48422D8E4048DAF94604C4F993:1D730C85422D8E424AE4F85D050EF842" box="[681,807,1982,2007]" country="China" county="Jiangyou City" municipality="Hanzeng Town" name="Weixin Liu" pageId="20" pageNumber="73" stateProvince="Sichuan Province">Weixin Liu</location>
,
<location id="1D730C85422D8E424B7FF85D0592F842" LSID="urn:lsid:plazi:treatment:9005EB48422D8E4048DAF94604C4F993:1D730C85422D8E424B7FF85D0592F842" box="[818,955,1982,2007]" country="China" county="Jiangyou City" municipality="Hanzeng Town" name="Haomin Yin" pageId="20" pageNumber="73" stateProvince="Sichuan Province">Haomin Yin</location>
,
<location id="1D730C85422D8E424B8BF85D024DF84D" LSID="urn:lsid:plazi:treatment:9005EB48422D8E4048DAF94604C4F993:1D730C85422D8E424B8BF85D024DF84D" box="[966,1124,1982,2008]" country="China" county="Jiangyou City" municipality="Hanzeng Town" name="Sunbin Huang" pageId="20" pageNumber="73" stateProvince="Sichuan Province">Sunbin Huang</location>
and Xinhui Wang.
</materialsCitation>
</paragraph>
<caption id="4CD30AD6422C8E4348DAF84C0405F871" httpUri="https://zenodo.org/record/1458372/files/figure.png" pageId="21" pageNumber="74" startId="21.[151,250,1967,1989]" targetBox="[313,1273,245,1908]" targetPageId="21">
<paragraph id="18135A5E422C8E4348DAF84C0405F871" blockId="21.[151,1436,1967,2020]" pageId="21" pageNumber="74">
<emphasis id="2AD8864C422C8E4348DAF84C0709F850" bold="true" box="[151,288,1967,1989]" pageId="21" pageNumber="74">FIGURE 18.</emphasis>
<taxonomicName id="DFAC21DD422C8E43496BF84C051DF850" authority="Golovatch and Geoffroy 2014" authorityName="Golovatch and Geoffroy" authorityYear="2014" box="[294,820,1967,1989]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="21" pageNumber="74" phylum="Arthropoda" rank="species" species="lipsae">
<emphasis id="2AD8864C422C8E43496BF84C07D3F850" box="[294,506,1967,1989]" italics="true" pageId="21" pageNumber="74">Epanerchodus lipsae</emphasis>
Golovatch and Geoffroy 2014
</taxonomicName>
, male from Wolong Dong Cave.
<emphasis id="2AD8864C422C8E434CC9F84C02BFF850" bold="true" box="[1156,1174,1967,1989]" pageId="21" pageNumber="74">A</emphasis>
: habitus, ventral view.
<emphasis id="2AD8864C422C8E434DC9F84C03BCF850" bold="true" box="[1412,1429,1967,1989]" pageId="21" pageNumber="74">B</emphasis>
: habitus, dorsal view. Scale bar: 0.5 mm.
</paragraph>
</caption>
<caption id="4CD30AD6422F8E4048DAFB4206C3FB61" httpUri="https://zenodo.org/record/1458374/files/figure.png" pageId="22" pageNumber="75" startId="22.[151,250,1185,1207]" targetBox="[163,1416,219,1152]" targetPageId="22">
<paragraph id="18135A5E422F8E4048DAFB4206C3FB61" blockId="22.[151,1435,1185,1268]" pageId="22" pageNumber="75">
<emphasis id="2AD8864C422F8E4048DAFB420708FB22" bold="true" box="[151,289,1185,1207]" pageId="22" pageNumber="75">FIGURE 19.</emphasis>
<taxonomicName id="DFAC21DD422F8E404967FB420569FB22" authority="Golovatch and Geoffroy 2014" authorityName="Golovatch and Geoffroy" authorityYear="2014" box="[298,832,1185,1207]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="22" pageNumber="75" phylum="Arthropoda" rank="species" species="lipsae">
<emphasis id="2AD8864C422F8E404967FB4207D6FB22" box="[298,511,1185,1207]" italics="true" pageId="22" pageNumber="75">Epanerchodus lipsae</emphasis>
Golovatch and Geoffroy 2014
</taxonomicName>
, male from Wolong Dong Cave.
<emphasis id="2AD8864C422F8E404CD1FB420287FB22" bold="true" box="[1180,1198,1185,1207]" pageId="22" pageNumber="75">A</emphasis>
: right gonopod, lateral view.
<emphasis id="2AD8864C422F8E40489BFB5C06CEFB40" bold="true" box="[214,231,1215,1237]" pageId="22" pageNumber="75">B</emphasis>
: right gonopod, mesal view. Abbreviations: a = spiniform process; cl = clivus; en = endomere; ex = exomere; p = process.
</paragraph>
</caption>
<paragraph id="18135A5E422F8E4048DAFAD904C4F993" blockId="22.[151,1437,1338,1543]" pageId="22" pageNumber="75">
<emphasis id="2AD8864C422F8E4048DAFAD906F3FAC6" bold="true" box="[151,218,1338,1363]" pageId="22" pageNumber="75">Notes</emphasis>
: This species is diagnosed by the gonopod showing a peg-shaped exomere (
<emphasis id="2AD8864C422F8E404C31FAD902BFFAC6" bold="true" box="[1148,1174,1338,1363]" pageId="22" pageNumber="75">ex</emphasis>
) and a ribbon-shaped endomere (
<emphasis id="2AD8864C422F8E404958FABD0718FAE2" bold="true" box="[277,305,1374,1399]" pageId="22" pageNumber="75">en</emphasis>
). The endomere has an axe-shaped mesal process (
<emphasis id="2AD8864C422F8E404B3DFABD0556FAE2" bold="true" box="[880,895,1374,1399]" pageId="22" pageNumber="75">p</emphasis>
) and a long, flagelli- or spiniform process (
<emphasis id="2AD8864C422F8E404D24FABD035EFAE2" bold="true" box="[1385,1399,1374,1399]" pageId="22" pageNumber="75">a</emphasis>
) at its base (
<figureCitation id="809746DB422F8E4048B7FA610764FA0E" box="[250,333,1410,1435]" captionStart="FIGURE 19" captionStartId="22.[151,250,1185,1207]" captionTargetBox="[163,1416,219,1152]" captionTargetId="figure@22.[151,1436,193,1164]" captionTargetPageId="22" captionText="FIGURE 19. Epanerchodus lipsae Golovatch and Geoffroy 2014, male from Wolong Dong Cave. A: right gonopod, lateral view. B: right gonopod, mesal view. Abbreviations: a = spiniform process; cl = clivus; en = endomere; ex = exomere; p = process." httpUri="https://zenodo.org/record/1458374/files/figure.png" pageId="22" pageNumber="75">Fig. 19</figureCitation>
).
<taxonomicName id="DFAC21DD422F8E40492FFA610464FA0E" authorityName="Golovatch and Geoffroy" authorityYear="2014" box="[354,589,1410,1435]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="22" pageNumber="75" phylum="Arthropoda" rank="species" species="lipsae">
<emphasis id="2AD8864C422F8E40492FFA610464FA0E" box="[354,589,1410,1435]" italics="true" pageId="22" pageNumber="75">Epanerchodus lipsae</emphasis>
</taxonomicName>
has been described from a few caves in Jiangyou and Beichuan counties in northeastern
<collectingRegion id="DA6894BC422F8E404967FA4507DAFA2A" box="[298,499,1446,1471]" country="China" name="Sichuan" pageId="22" pageNumber="75">Sichuan Province</collectingRegion>
. The material examined here comes from several additional caves and agrees with the original description in morphological characters (
<bibRefCitation id="7C3D27AF422F8E404ABEFA290214FA76" box="[755,1085,1482,1507]" pageId="22" pageNumber="75" refId="ref12900" refString="Golovatch, S. I. &amp; Geoffroy, J. - J. (2014) On some new or poorly-known species of the millipede family Polydesmidae from southern China (Diplopoda: Polydesmida). Russian Entomological Journal, 23 (2), 91 - 105." type="journal article">Golovatch &amp; Geoffroy 2014</bibRefCitation>
); only the body length is more variable in these specimens:
<emphasis id="2AD8864C422F8E404994FA0C07DDF993" box="[473,500,1519,1542]" italics="true" pageId="22" pageNumber="75">ca</emphasis>
1628
<emphasis id="2AD8864C422F8E404A0AFA0C044FF993" box="[583,614,1519,1542]" italics="true" pageId="22" pageNumber="75">vs.</emphasis>
<quantity id="DF54F7BB422F8E404A21FA0D04C1F993" box="[620,744,1518,1543]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" metricValueMax="2.6" metricValueMin="2.2" pageId="22" pageNumber="75" unit="mm" value="24.0" valueMax="26.0" valueMin="22.0">2226 mm</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>