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<document ID-DOI="http://dx.doi.org/10.3897/BDJ.4.e7720" ID-PMC="PMC4910500" ID-Pensoft-Pub="1314-2828--7720" ID-PubMed="27346952" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1314-2828--e7720" ModsDocOrigin="Biodiversity Data Journal " ModsDocTitle="Taxonomic revision of Rochefortia Sw. (Ehretiaceae, Boraginales)" checkinTime="1553126155575" checkinUser="pensoft" docAuthor="Irimia, Ramona-Elena &amp; Gottschling, Marc" docDate="2016" docId="7E465052BDD2FE17D4BDF4F1E4E150B1" docLanguage="en" docName="BiodivDatJour 4: e7720" docOrigin="Biodiversity Data Journal 4" docSource="http://dx.doi.org/10.3897/BDJ.4.e7720" docTitle="Rochefortia spinosa Urb." docType="treatment" docVersion="4" lastPageNumber="7720" masterDocId="FFB11146FFEDFFDFFF9CC8652A49F76B" masterDocTitle="Taxonomic revision of Rochefortia Sw. (Ehretiaceae, Boraginales)" masterLastPageNumber="7720" masterPageNumber="7720" pageNumber="7720" updateTime="1668126848301" updateUser="ExternalLinkService">
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<mods:title>Taxonomic revision of Rochefortia Sw. (Ehretiaceae, Boraginales)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Irimia, Ramona-Elena</mods:namePart>
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<mods:namePart>Gottschling, Marc</mods:namePart>
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<mods:title>Biodiversity Data Journal</mods:title>
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<mods:date>2016</mods:date>
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<mods:number>4</mods:number>
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<mods:start>7720</mods:start>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/BDJ.4.e7720</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2828--7720</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:7E465052BDD2FE17D4BDF4F1E4E150B1" httpUri="http://treatment.plazi.org/id/7E465052BDD2FE17D4BDF4F1E4E150B1" lastPageNumber="7720" pageId="0" pageNumber="7720">
<subSubSection pageId="0" pageNumber="7720" type="nomenclature">
<paragraph pageId="0" pageNumber="7720">
<taxonomicName authority="(Jacq.) Urb." class="Magnoliopsida" family="Ehretiaceae" genus="Rochefortia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rochefortia spinosa" order="Boraginales" pageId="0" pageNumber="7720" phylum="Tracheophyta" rank="species" species="spinosa">Rochefortia spinosa (Jacq.) Urb.</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="7720" type="reference_group">
<paragraph pageId="0" pageNumber="7720">
<taxonomicName class="Magnoliopsida" family="Ehretiaceae" genus="Rochefortia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rochefortia spinosa" order="Boraginales" pageId="0" pageNumber="7720" phylum="Tracheophyta" rank="species" species="spinosa">Rochefortia spinosa</taxonomicName>
<taxonomicName class="Magnoliopsida" family="Ehretiaceae" genus="Rochefortia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rochefortia spinosa" order="Boraginales" pageId="0" pageNumber="7720" phylum="Tracheophyta" rank="species" species="spinosa">Rochefortia spinosa</taxonomicName>
Feddes Repert. 13: 472. 1915
<taxonomicName class="Magnoliopsida" family="Ehretiaceae" genus="Ehretia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ehretia spinosa" order="Boraginales" pageId="0" pageNumber="7720" phylum="Tracheophyta" rank="species" species="spinosa">Ehretia spinosa</taxonomicName>
Enum. syst. pl.: 14. 1760Select. stirp. amer. hist.: 46, pl. CLXXX 18. 1763
<taxonomicName class="Magnoliopsida" family="Ehretiaceae" genus="Lutrostylis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Lutrostylis spinosa" order="Boraginales" pageId="0" pageNumber="7720" phylum="Tracheophyta" rank="species" species="spinosa">Lutrostylis spinosa</taxonomicName>
Gen. Hist. 4: 391. 1838pl. CLXXX 18 in Jacq., Select. stirp. amer. hist. 1763
</paragraph>
<paragraph pageId="0" pageNumber="7720">
<taxonomicName class="Magnoliopsida" family="Ehretiaceae" genus="Rochefortia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rochefortia spinosa" order="Boraginales" pageId="0" pageNumber="7720" phylum="Tracheophyta" rank="species" species="spinosa">Rochefortia spinosa</taxonomicName>
<taxonomicName class="Magnoliopsida" family="Ehretiaceae" genus="Ehretia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ehretia fasciculata" order="Boraginales" pageId="0" pageNumber="7720" phylum="Tracheophyta" rank="species" species="fasciculata">Ehretia fasciculata</taxonomicName>
Nov. gen. sp. 3: 66. 1818
<taxonomicName class="Magnoliopsida" family="Ehretiaceae" genus="Crematomia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Crematomia fasciculata" order="Boraginales" pageId="0" pageNumber="7720" phylum="Tracheophyta" rank="species" species="fasciculata">Crematomia fasciculata</taxonomicName>
Ann. Mag. Nat. Hist., ser. 4 3: 312-313. 1869
<taxonomicName class="Magnoliopsida" family="Ehretiaceae" genus="Morelosia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Morelosia fasciculata" order="Boraginales" pageId="0" pageNumber="7720" phylum="Tracheophyta" rank="species" species="fasciculata">Morelosia fasciculata</taxonomicName>
Revis. gen. Pl. 2: 439. 1891
<taxonomicName class="Magnoliopsida" family="Ehretiaceae" genus="Bourreria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bourreria fasciculata" order="Boraginales" pageId="0" pageNumber="7720" phylum="Tracheophyta" rank="species" species="fasciculata">Bourreria fasciculata</taxonomicName>
Engl. &amp; Prantl, Nat. Pflanzenfam. 4 (3a): 87. 1893A.J.A. Bonpland &amp; F.W.H.A. von Humboldt 92P-670679
</paragraph>
<paragraph pageId="0" pageNumber="7720">
<taxonomicName class="Magnoliopsida" family="Ehretiaceae" genus="Rochefortia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rochefortia spinosa" order="Boraginales" pageId="0" pageNumber="7720" phylum="Tracheophyta" rank="species" species="spinosa">Rochefortia spinosa</taxonomicName>
<taxonomicName class="Magnoliopsida" family="Ehretiaceae" genus="Rochefortia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rochefortia jacquinii" order="Boraginales" pageId="0" pageNumber="7720" phylum="Tracheophyta" rank="species" species="jacquinii">Rochefortia jacquinii</taxonomicName>
Abh.
<normalizedToken originalValue="Königl">Koenigl</normalizedToken>
. Ges. Wiss.
<normalizedToken originalValue="Göttingen">Goettingen</normalizedToken>
7: 253. 1857J. Goudot s.n.GOET-395K
</paragraph>
<paragraph pageId="0" pageNumber="7720">
<taxonomicName class="Magnoliopsida" family="Ehretiaceae" genus="Rochefortia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rochefortia spinosa" order="Boraginales" pageId="0" pageNumber="7720" phylum="Tracheophyta" rank="species" species="spinosa">Rochefortia spinosa</taxonomicName>
<taxonomicName class="Magnoliopsida" family="Ehretiaceae" genus="Rochefortia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rochefortia fasciculata" order="Boraginales" pageId="0" pageNumber="7720" phylum="Tracheophyta" rank="species" species="fasciculata">Rochefortia fasciculata</taxonomicName>
Engl. &amp; Prantl, Nat. Pflanzenfam. 4 (3a): 89. 1893J.J. Triana 3744COL-112389US-110770
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="7720" type="description">
<paragraph pageId="0" pageNumber="7720">Description</paragraph>
<paragraph pageId="0" pageNumber="7720">Trees up to 10.0 m tall or smaller shrubs 2.0-5.0 m tall, crown arching, galls absent; indument glabrescent, young twigs variously pilose through glabrescent; bark grey light to grey brown, longitudinally fissured, wood light; thorns 0.80-1.0 cm long, robust, somewhat acute, sometimes slightly curved at tips, simple, scattered, alternate, glabrous. Leaves fasciculate, occasionally stipulate (2 deciduous stipules, as stated by Lasser 758: US!), dark green, dark pale, sometimes red when older; petiole 0.2-1.5 cm long, slender, hirsute, rarely glabrous; blade 0.9-10.8 cm long, 0.3-4.5 cm wide, obovate, widely obovate, rarely elliptic, coriaceous, primary veins prominent, secondary veins 5-6, tertiary veins reticulate; base cuneate or sometimes rounded, rarely oblique; apex retuse, rounded, obcordate, emarginate, acute, occasionally mucronulate; adaxial surface glabrous or with scattered trichomes (especially on the midrib and at tips), rugose, with cystotliths, abaxial surface glabrous or with stiff trichomes on the midrib and tips. Inflorescence terminal or axillary, secondarily branched, branches slender, glabrescent, pedicels 0.30-0.40 long. Calyx 0.18-0.25 cm long, coriaceous, hirsute outside, glabrous inside, lobes 0.15-0.18 cm long, 0.10-0.20 cm wide, divided from the base, triangulate, apex slightly acute, margin strigose. Corolla 0.35-0.40 cm long, white, yellowish, orange, rarely green (Ferrari 706: F!), tube funnel-shaped, 0.18-0.20 cm long, lobes 0.20-0.25 cm long, ovate, glabrous on both sides, slightly ciliate at tips. Anthers of male flower 0.15-0.30 cm long, filaments 0.20-0.23 cm long, adnate to the corolla tube for 0.10-0.12 cm, pollen present; anthers of female flower 0.08-0.10 cm long, filaments 0.06-0.10 cm long, adnate to the corolla tube for 0.02-0.03 cm, pollen absent. Ovary of male flower subglobose, stylodia 2, branches 0.07-0.10 cm long, glabrous, with strigose, grey trichomes at tips; ovary of female flower globose, 0.12-0.14 cm long, stylodia 2, 0.18-0.22 cm long, slightly unequal, 2 well developed stigmas, cotyliform. Fruit 0.50-0.80 cm tall, 0.60-0.80 cm wide, globose, green when immature, yellowish reddish through red and dark purple during maturation; style accrescent, not persistent; pyrene 0.40-0.60 cm tall, 0.30-0.40 cm wide, 0.15-0.30 cm deep, with various ridges on the abaxial surface.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="7720" type="distribution">
<paragraph pageId="0" pageNumber="7720">Distribution</paragraph>
<paragraph pageId="0" pageNumber="7720">Northern South America (symbol &quot;&quot; in Fig. 3) in deciduous costal dry forests, in medium and high moist forests, on dry steep cliffs and limestone soils at altitudes between 0-1300 m.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="7720" type="ecology">
<paragraph pageId="0" pageNumber="7720">Ecology</paragraph>
<paragraph pageId="0" pageNumber="7720">
Flowering Jan,
<normalizedToken originalValue="MarMay">Mar-May</normalizedToken>
, Jul, Sep, Nov; fruiting
<normalizedToken originalValue="FebDec">Feb-Dec</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="7720" type="taxon discussion">
<paragraph pageId="0" pageNumber="7720">Taxon discussion</paragraph>
<paragraph pageId="0" pageNumber="7720">
<taxonomicName class="Magnoliopsida" family="Ehretiaceae" genus="Rochefortia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rochefortia spinosa" order="Boraginales" pageId="0" pageNumber="7720" phylum="Tracheophyta" rank="species" species="spinosa">Rochefortia spinosa</taxonomicName>
is among the species with the largest leaf size after
<taxonomicName lsidName="R. lundellii." pageId="0" pageNumber="7720" rank="species" species="lundellii.">R. lundellii.</taxonomicName>
Specimens across different geographical areas are relatively homogenous displaying a low variability of morphological traits (e.g., few collections have leave lengths shorter than 5.5 cm). The species is not sympatric with any other species but morphologically, it resembles
<taxonomicName lsidName="R. lundellii" pageId="0" pageNumber="7720" rank="species" species="lundellii">R. lundellii</taxonomicName>
. It can be distinguished from the latter by the presence of 2 stylodia and generally by the less extensive developed thorns on stems. In the past,
<taxonomicName lsidName="R. spinosa" pageId="0" pageNumber="7720" rank="species" species="spinosa">R. spinosa</taxonomicName>
was one of the most confusing
<taxonomicName class="Magnoliopsida" family="Ehretiaceae" genus="Rochefortia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rochefortia" order="Boraginales" pageId="0" pageNumber="7720" phylum="Tracheophyta" rank="genus">Rochefortia</taxonomicName>
species and was believed being distributed across multiple islands of the Caribbean. However, molecular data (
<bibRefCitation author="Irimia, Ramona Elena" journalOrPublisher="Plant Systematics and Evolution" pageId="0" pageNumber="7720" pagination="1509 - 1516" title="Strong biogeographic signal in the phylogenetic relationships of Rochefortia Sw. (Ehretiaceae, Boraginales)" volume="301" year="2015">Irimia et al. 2015</bibRefCitation>
) indicate that the species is restricted to northern South America (primarily Colombia and Venezuela, but also Peru though with few collections available).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="7720" type="notes">
<paragraph pageId="0" pageNumber="7720">Notes</paragraph>
<paragraph pageId="0" pageNumber="7720">
Representative specimens examined. - COLOMBIA.
<normalizedToken originalValue="Bolívar">Bolivar</normalizedToken>
: Cartagena, Tierra Bomba,
<geoCoordinate direction="north" orientation="latitude" precision="925" value="10.333333">10°20'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-75.53333">75°32'W</geoCoordinate>
, 4 Sep 1989 (fr), Cuadros 4332 (MO!); San
<normalizedToken originalValue="Juán">Juan</normalizedToken>
de Nepomuceno, Santuario Nacional de los Colorados, 70 km SW of Cartagena,
<geoCoordinate direction="north" orientation="latitude" precision="925" value="9.966666">9°58'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-75.166664">75°10'W</geoCoordinate>
, 13 Jan 1988 (fl), Gentry et al. 60688 (MO!); Cundinamarca: bw Viota and Girardot, elev 320-560 m,
<geoCoordinate direction="north" orientation="latitude" precision="925" value="4.4333334">4°26'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-74.583336">74°35'W</geoCoordinate>
, Aug 1964 (fr), Saravia Toledo 4652 (COL!); Huila: La Bodega, Cordillera Oriental, elev 650 m, 3°16'N, 74°54'W, 21 Nov 1944 (fr), Little 8954 (COL! US!); Carretera a San Antonio, El Silencio Quebrada San Roque, elev
<normalizedToken originalValue="±">+/-</normalizedToken>
900 m,
<geoCoordinate direction="north" orientation="latitude" precision="925" value="3.35">3°21'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-74.78333">74°47'W</geoCoordinate>
, 28 Sep 1990 (fr), Llanos H &amp; Camacho 1824 (COL!); Villavieja, Desierto de la Tatacoa,
<geoCoordinate direction="north" orientation="latitude" precision="925" value="3.2166667">3°13'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-75.166664">75°10'W</geoCoordinate>
[retroactively inferred], 23 Apr 2003 (♂ fl), Figueroa et al. 246 (COL! COL!); Magdalena: Valle del
<normalizedToken originalValue="Río">Rio</normalizedToken>
Cesare (parte occidental), SW of Los Venados, elev 60 m, 10°N, 73°42'W [retroactively inferred], Feb 1961 (fr), Dugand 5590 (COL! US! US!). - PERU. San Martin: Valley of
<normalizedToken originalValue="Río">Rio</normalizedToken>
Huallaga, 29 km S of Tarapoto, near El Abra, 6°40'S, 76°20'W [herbarium label], 6 Feb 1984 (fr), Gentry &amp; Smith 45027 (MO!); Juan Guerra, elev 720 m, 6°37'S, 76°27'W [retroactively inferred], Dec 1929 (fr), Williams 6874 (F!). - VENEZUELA.
<normalizedToken originalValue="Anzoátegui">Anzoategui</normalizedToken>
: Alrededores de
<normalizedToken originalValue="Píritu">Piritu</normalizedToken>
,
<geoCoordinate direction="north" orientation="latitude" precision="925" value="10.216666">10°13'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-65.51667">65°31'W</geoCoordinate>
[retroactively inferred], 18 Aug 1965 (fr),
<taxonomicName lsidName="" pageId="0" pageNumber="7720" rank="tribe" tribe="Agostini">Agostini</taxonomicName>
526 (US!); Aragua: Zamora, 4 km N of Villa de Cura,
<geoCoordinate direction="north" orientation="latitude" precision="925" value="10.55">10°33'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-67.46667">67°28'W</geoCoordinate>
[retroactively inferred], 23 Mar 1969 (♀ fl, fr), Ferrari &amp; Bunting 713 (F!);
<normalizedToken originalValue="Bolívar">Bolivar</normalizedToken>
: Kavanayen, elev 1300 m,
<geoCoordinate direction="north" orientation="latitude" precision="925" value="5.6">5°36'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-61.5">61°30'W</geoCoordinate>
[retroactively inferred], 27 May 1946 (♂ fl) Lasser 758 (US!); Carabobo: Valencia, elev 400 m,
<geoCoordinate direction="north" orientation="latitude" precision="925" value="10.75">10°45'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-68.35">68°21'W</geoCoordinate>
[retroactively inferred], 14 Sep 1959 (fr), Williams 12198 (F!); Portuguesa: Guanare, junction bw rivers Tucupido and Guanare,
<geoCoordinate direction="north" orientation="latitude" precision="925" value="9.533334">9°32'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-69.816666">69°49'W</geoCoordinate>
[retroactively inferred], 15 Apr 1982 (fr), Utrera &amp; Stergios 152 (MO!), Utrera &amp; Stergios 148 (MO!); Miranda, bordeando Laguna Grande,
<geoCoordinate direction="north" orientation="latitude" precision="925" value="10.55">10°33'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-66.066666">66°04'W</geoCoordinate>
[herbarium label], 27 May 1981 (fl), Berry 3736 (MO!); Monagas: Dos Caminos, via Maturin,
<geoCoordinate direction="north" orientation="latitude" precision="925" value="9.566667">9°34'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-64.4">63°84'W</geoCoordinate>
[retroactively inferred], Jul 1969 (♀ fl, fr), Aristeguieta et al. 7162 (F! US!), Aristeguieta et al. 7163 (F! US!); Sucre: Guarapoturo, via Chacopata,
<geoCoordinate direction="north" orientation="latitude" precision="925" value="10.6">10°36'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-63.65">63°39'W</geoCoordinate>
[retroactively inferred], 13 Sep 1984 (fl),
<normalizedToken originalValue="Cumaná">Cumana</normalizedToken>
Campos 2458 (MO!); Peninsula de Araya, ca 4 km west of Caimancito, elev 10 m,
<geoCoordinate direction="north" orientation="latitude" precision="925" value="10.616667">10°37'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-63.95">63°57'W</geoCoordinate>
, 20 May 1981 (fr), Liesner &amp;
<normalizedToken originalValue="González">Gonzalez</normalizedToken>
12160 (MO!); Trujillo: Valera, Loma de Moron,
<geoCoordinate direction="north" orientation="latitude" precision="925" value="9.316667">9°19'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-70.566666">70°34'W</geoCoordinate>
[retroactively inferred], 18 Nov 1922 (♂ fl), Pittier 10718 (US!); Vargas, Parroquia Catia la Mar, Escuela Naval,
<geoCoordinate direction="north" orientation="latitude" precision="925" value="10.6">10°36'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-67.03333">67°02'W</geoCoordinate>
, 20 Oct 1988 (fr),
<normalizedToken originalValue="Ramírez">Ramirez</normalizedToken>
2576 (MO!); Yaracuy: Pena, Cambural, bw Yaritagua and Barquisimeto, elev 350-400 m,
<geoCoordinate direction="north" orientation="latitude" precision="925" value="10.116667">10°07'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-69.183334">69°11'W</geoCoordinate>
, 8 Nov 2006 (♀ fl), Meier et al. 14018 (M!).
</paragraph>
<subSection pageId="0" pageNumber="7720" type="common names">
<paragraph pageId="0" pageNumber="7720">Common names</paragraph>
<paragraph pageId="0" pageNumber="7720">
<normalizedToken originalValue="“macarao”">&quot;macarao&quot;</normalizedToken>
, &quot;pega paloma&quot; and
<normalizedToken originalValue="“tachuelo”">&quot;tachuelo&quot;</normalizedToken>
in Venezuela and &quot;cruceto macho&quot; and &quot;tatacun o revienta puerco&quot; in Colombia.
</paragraph>
</subSection>
<subSection pageId="0" pageNumber="7720" type="economic uses">
<paragraph pageId="0" pageNumber="7720">Economic uses</paragraph>
<paragraph pageId="0" pageNumber="7720">
In Venezuela, the species is occasionally cultivated and forms hedgerows together with
<taxonomicName genus="Yagales" lsidName="Yagales guamacho" pageId="0" pageNumber="7720" rank="species" species="guamacho">guamacho</taxonomicName>
(probably
<taxonomicName class="Magnoliopsida" family="Cactaceae" genus="Pereskia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pereskia guamacho" order="Caryophyllales" pageId="0" pageNumber="7720" phylum="Tracheophyta" rank="species" species="guamacho">Pereskia guamacho</taxonomicName>
F.A.C.Weber:
<taxonomicName family="Cactaceae" lsidName="Yagales" pageId="0" pageNumber="7720" rank="family">Cactaceae</taxonomicName>
, as noted on Ferrari &amp; Bunting 713: F!).
</paragraph>
</subSection>
</subSubSection>
</treatment>
</document>