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<document id="E2EF56B98DEA5B0D3DBCF033D648A958" ID-DOI="10.25674/415" ID-ISSN="2509-9523" ID-Zenodo-Dep="13194032" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722631177589" checkinUser="felipe" docAuthor="Seifert, Bernhard" docDate="2024" docId="577887E6FF86FFD268CF5F2FFEBF366E" docLanguage="en" docName="SoilOrg.96.2.113-144.pdf" docOrigin="Soil Organisms (London, England) 96 (2)" docSource="http://dx.doi.org/10.25674/415" docStyle="DocumentStyle:EA2ECAC87EC21C07AB805D5B9B4601F2.2:SoilOrg.2018-.journal_article" docStyleId="EA2ECAC87EC21C07AB805D5B9B4601F2" docStyleName="SoilOrg.2018-.journal_article" docStyleVersion="2" docTitle="Cardiocondyla yoruba Rigato 2002" docType="treatment" docVersion="2" lastPageNumber="137" masterDocId="AB41FF9EFF91FFCA681B5A5FFFA8347E" masterDocTitle="A taxonomic revision of the ants of the Cardiocondyla wroughtonii group (Hymenoptera: Formicidae) with a checklist of the Cardiocondyla species of the world" masterLastPageNumber="144" masterPageNumber="113" pageNumber="136" updateTime="1722679537643" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="666FCB48115439D9B45004828D09B072">A taxonomic revision of the ants of the Cardiocondyla wroughtonii group (Hymenoptera: Formicidae) with a checklist of the Cardiocondyla species of the world</mods:title>
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<treatment id="577887E6FF86FFD268CF5F2FFEBF366E" LSID="urn:lsid:plazi:treatment:577887E6FF86FFD268CF5F2FFEBF366E" httpUri="http://treatment.plazi.org/id/577887E6FF86FFD268CF5F2FFEBF366E" lastPageId="24" lastPageNumber="137" pageId="23" pageNumber="136">
<subSubSection id="97CB657BFF86FFDD68CF5F2FFE1831D7" pageId="23" pageNumber="136" type="nomenclature">
<paragraph id="DF6E36F0FF86FFDD68CF5F2FFDF531F6" blockId="23.[212,605,1392,1416]" box="[212,605,1392,1416]" pageId="23" pageNumber="136">
<heading id="8426819CFF86FFDD68CF5F2FFDF531F6" box="[212,605,1392,1416]" fontSize="10" level="2" pageId="23" pageNumber="136" reason="4">
<taxonomicName id="18D14D73FF86FFDD68CF5F2FFDF531F6" authority="Rigato 2002" authorityName="Rigato" authorityYear="2002" box="[212,605,1392,1416]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="136" phylum="Arthropoda" rank="species" species="yoruba">
<emphasis id="EDA5EAE2FF86FFDD68CF5F2FFDF531F6" bold="true" box="[212,605,1392,1416]" pageId="23" pageNumber="136">
<emphasis id="EDA5EAE2FF86FFDD68CF5F2FFE6431F6" bold="true" box="[212,460,1392,1416]" italics="true" pageId="23" pageNumber="136">Cardiocondyla yoruba</emphasis>
<bibRefCitation id="BB404B01FF86FFDD69C85F2FFDF531F6" author="Rigato, F." box="[467,605,1392,1416]" pageId="23" pageNumber="136" pagination="167 - 173" refId="ref19389" refString="Rigato, F. (2002): Three new Afrotropical Cardiocondyla Emery, with a revised key to the workers (Hymenoptera Formicidae). - Bollettino della Societa Entomologica Italiana 134: 167 - 173." type="journal article" year="2002">Rigato 2002</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="DF6E36F0FF86FFDD68CF5FCEFE1831D7" blockId="23.[212,432,1425,1449]" box="[212,432,1425,1449]" pageId="23" pageNumber="136">
<heading id="8426819CFF86FFDD68CF5FCEFE1831D7" bold="true" box="[212,432,1425,1449]" fontSize="10" level="2" pageId="23" pageNumber="136" reason="3">
<emphasis id="EDA5EAE2FF86FFDD68CF5FCEFE1831D7" bold="true" box="[212,432,1425,1449]" pageId="23" pageNumber="136">
[
<typeStatus id="006A8852FF86FFDD68C05FCEFEA731D7" box="[219,271,1425,1449]" pageId="23" pageNumber="136">type</typeStatus>
investigation]
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="97CB657BFF86FFDD68CF5FEDFE1A330B" pageId="23" pageNumber="136" type="description">
<paragraph id="DF6E36F0FF86FFDD68CF5FEDFEB032F1" blockId="23.[189,802,1458,1976]" pageId="23" pageNumber="136">
<materialsCitation id="6FB93CADFF86FFDD68CF5FEDFEBC32F1" collectionCode="IITA, T, BMNH" country="Nigeria" county="Ibadan" location="This" municipality="Investigated" pageId="23" pageNumber="136" specimenCode="CASENT0901752" specimenCount="3" specimenCount-worker="3" stateProvince="Niger" typeStatus="holotype">
<location id="DA0E602BFF86FFDD68CF5FEDFEAD31B4" LSID="urn:lsid:plazi:treatment:577887E6FF86FFD268CF5F2FFEBF366E:DA0E602BFF86FFDD68CF5FEDFEAD31B4" box="[212,261,1458,1482]" country="Nigeria" county="Ibadan" municipality="Investigated" name="This" pageId="23" pageNumber="136" stateProvince="Niger">This</location>
taxon has been described from
<collectingRegion id="1D15F812FF86FFDD6A4D5FEDFD3D31B4" box="[598,661,1458,1482]" country="Nigeria" name="Niger" pageId="23" pageNumber="136">Niger</collectingRegion>
.
<collectingMunicipality id="3F0AAC8AFF86FFDD6A855FEDFC8A31B4" box="[670,802,1458,1482]" pageId="23" pageNumber="136">Investigated</collectingMunicipality>
were
<specimenCount id="C9D7FD79FF86FFDD68E05F8CFE543195" box="[251,508,1491,1515]" count="3" pageId="23" pageNumber="136" type="worker" typeStatus="paratype">three paratype workers</specimenCount>
on the same pin with the site equal to the
<typeStatus id="006A8852FF86FFDD69725FABFE6F3272" box="[361,455,1524,1548]" pageId="23" pageNumber="136" type="holotype">holotype</typeStatus>
(see
<specimenCode id="8F779E8BFF86FFDD69E65FABFD6B3272" box="[509,707,1524,1548]" collectionCode="CASENT" pageId="23" pageNumber="136">CASENT0901752</specimenCode>
in www. antweb.org), labelled
<collectingCountry id="A7C67660FF86FFDD69D85C4AFD9D3253" box="[451,565,1557,1581]" name="Nigeria" pageId="23" pageNumber="136">NIGERIA</collectingCountry>
<collectingCounty id="360F4E7CFF86FFDD6A5C5C4AFD3B3253" box="[583,659,1557,1581]" pageId="23" pageNumber="136">Ibadan</collectingCounty>
,
<collectionCode id="B9C0AE35FF86FFDD6AB15C4AFD4D3253" box="[682,741,1557,1581]" country="Nigeria" httpUri="http://grbio.org/cool/8h49-s777" name="International Institute of Tropical Agriculture" pageId="23" pageNumber="136">IITA</collectionCode>
.xi.87,
<collectionCode id="B9C0AE35FF86FFDD68A65C69FF663230" box="[189,206,1590,1614]" name="Tavera, Department of Geology and Geophysics" pageId="23" pageNumber="136">T</collectionCode>
.Noyes (handwritten),
<taxonomicName id="18D14D73FF86FFDD69C45C69FD763230" authorityName="Seifert" authorityYear="2024" box="[479,734,1590,1614]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="136" phylum="Arthropoda" rank="species" species="yoruba" status="sp. nov.">Cardiocondyla yoruba</taxonomicName>
<taxonomicNameLabel id="F6965799FF86FFDD6AF45C69FC893233" box="[751,801,1590,1613]" pageId="23" pageNumber="136" rank="species">n.sp.</taxonomicNameLabel>
<typeStatus id="006A8852FF86FFDD68A65C08FEEA3211" box="[189,322,1623,1647]" pageId="23" pageNumber="136">PARATYPI</typeStatus>
, det. F.
<bibRefCitation id="BB404B01FF86FFDD69895C09FDB73211" author="Rigato, F." box="[402,543,1622,1647]" pageId="23" pageNumber="136" pagination="167 - 173" refId="ref19389" refString="Rigato, F. (2002): Three new Afrotropical Cardiocondyla Emery, with a revised key to the workers (Hymenoptera Formicidae). - Bollettino della Societa Entomologica Italiana 134: 167 - 173." type="journal article" year="2002">Rigato 2002</bibRefCitation>
(handwritten),
<collectionCode id="B9C0AE35FF86FFDD6ACB5C08FC893210" box="[720,801,1623,1646]" pageId="23" pageNumber="136">BMNH</collectionCode>
London
</materialsCitation>
.
</paragraph>
<paragraph id="DF6E36F0FF86FFDD68CF5CC7FEAA336D" blockId="23.[189,802,1458,1976]" pageId="23" pageNumber="136">
<emphasis id="EDA5EAE2FF86FFDD68CF5CC7FE6632CE" bold="true" box="[212,462,1688,1712]" pageId="23" pageNumber="136">Allmaterialexamined</emphasis>
.
<materialsCitation id="6FB93CADFF86FFDD69CC5CC7FF56336D" country="Ivory Coast" location="Bondoukou" pageId="23" pageNumber="136" specimenCount="11" specimenCount-worker="11">
Morphometricallyinvestigated were two samples with
<specimenCount id="C9D7FD79FF86FFDD69DE5CE5FDCD32AF" box="[453,613,1721,1745]" count="7" pageId="23" pageNumber="136" type="worker">seven workers</specimenCount>
: the
<typeStatus id="006A8852FF86FFDD6A875CE5FD6432AC" box="[668,716,1722,1746]" pageId="23" pageNumber="136">type</typeStatus>
sample plus a sample of
<specimenCount id="C9D7FD79FF86FFDD699A5C85FDA6328C" box="[385,526,1754,1778]" count="4" pageId="23" pageNumber="136" type="worker">four workers</specimenCount>
from
<location id="DA0E602BFF86FFDD6A435C85FD7D328C" LSID="urn:lsid:plazi:treatment:577887E6FF86FFD268CF5F2FFEBF366E:DA0E602BFF86FFDD6A435C85FD7D328C" box="[600,725,1754,1778]" country="Ivory Coast" name="Bondoukou" pageId="23" pageNumber="136">Bondoukou</location>
/
<collectingCountry id="A7C67660FF86FFDD6AFC5C85FF56336D" name="Ivory Coast" pageId="23" pageNumber="136">Ivory Coast</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="DF6E36F0FF86FFDD68CF5D44FE1A330B" blockId="23.[189,802,1458,1976]" pageId="23" pageNumber="136">
<emphasis id="EDA5EAE2FF86FFDD68CF5D44FE01334A" bold="true" box="[212,425,1819,1844]" pageId="23" pageNumber="136">Geographic range</emphasis>
. Known so far only from West Africa:
<collectingCountry id="A7C67660FF86FFDD69155D62FEFD332B" box="[270,341,1853,1877]" name="Ghana" pageId="23" pageNumber="136">Ghana</collectingCountry>
(by image evaluation of CASENT0901752),
<collectingCountry id="A7C67660FF86FFDD68A65D01FE923308" box="[189,314,1886,1910]" name="Ivory Coast" pageId="23" pageNumber="136">Ivory Coast</collectingCountry>
and
<collectingCountry id="A7C67660FF86FFDD69745D02FE06330B" box="[367,430,1885,1909]" name="Niger" pageId="23" pageNumber="136">Niger</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="97CB657BFF86FFD268CF5D21FDC83535" lastPageId="24" lastPageNumber="137" pageId="23" pageNumber="136" type="diagnosis">
<paragraph id="DF6E36F0FF86FFDD68CF5D21FC8A33C9" blockId="23.[189,802,1458,1976]" pageId="23" pageNumber="136">
<emphasis id="EDA5EAE2FF86FFDD68CF5D21FEEB33E8" bold="true" box="[212,323,1918,1942]" pageId="23" pageNumber="136">Diagnosis</emphasis>
: --Worker (
<tableCitation id="9253034BFF86FFDD69D75D21FDB033E8" box="[460,536,1918,1942]" captionStart="Table 2" captionStartId="26.[150,203,1476,1497]" captionText="Table 2. Workers of the C. wroughtonii group with mean FL/FR&gt; 1.040 and originating from Africa and theArabian Peninsula. Sequence of data: arithmetic mean ± standard deviation [minimum, maximum]." pageId="23" pageNumber="136">Tab. 2</tableCitation>
, Figs 610, key). Very small, CS 380 µm. Head very long, CL/CW 1.247.
</paragraph>
<paragraph id="DF6E36F0FF86FFDD6B235AAEFB143210" blockId="23.[824,1437,241,1976]" pageId="23" pageNumber="136">Median third of anterior clypeal margin straight, median third of occipital margin slightly concave. Postocular distance large, PoOc/CL 0.459. Frons relatively narrow (FRS/CS 0.245); frontal carinae slightly diverging frontal of FRS level (FL/FR 1.063), caudal of FRS level parallel or slightly converging. Eye rather small, EYE/CS 0.240 and with scattered microsetae of 67 µm length. Scape very short, SL/CS 0.724. Metanotal depression shallow (MGr/CS 1.27 %). Propodeal spines short (SP/CS 0.120) and almost triangular in lateral view, their axis in profile deviating from longitudinal mesosomal axis by 25°, in dorsal view not diverging and their bases moderately wide (SPBA/CS 0.275). Petiole rather narrow and moderately high (PeW/CS 0.274, PeH/CS 0.334); in profile with a concave anterior face and semicircular dorsum; in dorsal view with an almost globular node, which is only slightly longer than wide. Postpetiole narrow and low (PpW/CS 0.427, PpH/CS 0.288); postpetiolar sternite anterolaterally with a rounded lobiform protrusion on each side which clearly elevates above the level of median surface of sternite; the surface of the sternite thus appearing deeply concave in frontal view. Postpetiole in dorsal view with a straight to slightly concave anterior margin and convex sides. Lateral parts of clypeus each with 23 rugulae, central part slightly carinulate-reticulate. Frontal laminae with weak microsculpture, consisting of an irregular mixture of microrugulate, foveolate and microreticulate elements. Whole vertex foveolate, mediad of the eyes with densely-arranged, flat-bottomed foveolae of 1416 µm diameter. The largest foveolae show a well-demarcated but flat central tubercule of 68 µm diameter around the hair bases (Fig. 9). Whole surface of mesosoma foveolate-microreticulate; petiole strongly and postpetiolar tergite less strongly microreticulate. First gaster tergite with moderately long and dense pubescence (PLG/CS 6.89 %, sqPDG 3.86) and a more strongly developed microreticum, with the hair bases placed in microfoveolae of 46 µm diameter (Fig. 10). The microreticulum is an optical illusion in perpendicular view on the surface that is produced by the margins of roof-tile-like surface structures. Color: whole dorsum of gaster blackish brown and all remaining body parts yellowish or yellowish-brown.</paragraph>
<paragraph id="DF6E36F0FF86FFD26B545C28FDC83535" blockId="23.[824,1437,241,1976]" lastBlockId="24.[151,764,241,529]" lastPageId="24" lastPageNumber="137" pageId="23" pageNumber="136">
<emphasis id="EDA5EAE2FF86FFDD6B545C28FBFC32F1" bold="true" box="[847,1108,1655,1679]" pageId="23" pageNumber="136">Taxonomic comments.</emphasis>
<taxonomicName id="18D14D73FF86FFDD6C7A5C27FB7B32F1" authorityName="Seifert" authorityYear="2024" box="[1121,1235,1655,1679]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="136" phylum="Arthropoda" rank="species" species="yoruba">
<emphasis id="EDA5EAE2FF86FFDD6C7A5C27FB7B32F1" box="[1121,1235,1655,1679]" italics="true" pageId="23" pageNumber="136">C. yoruba</emphasis>
</taxonomicName>
is inseparable in a PCA from
<taxonomicName id="18D14D73FF86FFDD6BD05CC6FB9232CE" authorityName="Seifert" authorityYear="2024" box="[971,1082,1689,1712]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="136" phylum="Arthropoda" rank="species" species="yemeni">
<emphasis id="EDA5EAE2FF86FFDD6BD05CC6FB9232CE" box="[971,1082,1689,1712]" italics="true" pageId="23" pageNumber="136">C. yemeni</emphasis>
</taxonomicName>
considering all 16 NUMOBAT characters: the
<typeStatus id="006A8852FF86FFDD6BFA5CE5FBB932AC" box="[993,1041,1722,1746]" pageId="23" pageNumber="136">type</typeStatus>
samples of
<taxonomicName id="18D14D73FF86FFDD6C825CE5FB4D32AF" authorityName="Seifert" authorityYear="2024" box="[1177,1253,1722,1745]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="136" phylum="Arthropoda" rank="species" species="yemeni">
<emphasis id="EDA5EAE2FF86FFDD6C825CE5FB4D32AF" box="[1177,1253,1722,1745]" italics="true" pageId="23" pageNumber="136">yemeni</emphasis>
</taxonomicName>
and
<taxonomicName id="18D14D73FF86FFDD6D045CE6FAC432AF" authorityName="Seifert" authorityYear="2024" box="[1311,1388,1721,1745]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="136" phylum="Arthropoda" rank="species" species="yoruba">
<emphasis id="EDA5EAE2FF86FFDD6D045CE6FAC432AF" box="[1311,1388,1721,1745]" italics="true" pageId="23" pageNumber="136">yoruba</emphasis>
</taxonomicName>
and the non-type sample of
<taxonomicName id="18D14D73FF86FFDD6C535C85FB3D328C" authorityName="Seifert" authorityYear="2024" box="[1096,1173,1754,1778]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="136" phylum="Arthropoda" rank="species" species="yoruba">
<emphasis id="EDA5EAE2FF86FFDD6C535C85FB3D328C" box="[1096,1173,1754,1778]" italics="true" pageId="23" pageNumber="136">yoruba</emphasis>
</taxonomicName>
from
<collectingCountry id="A7C67660FF86FFDD6CFB5C85FAF5328F" box="[1248,1373,1754,1778]" name="Ivory Coast" pageId="23" pageNumber="136">Ivory coast</collectingCountry>
form a coherent cluster in a PCA (
<figureCitation id="47EA2A75FF86FFDD6C615CA4FB62336D" box="[1146,1226,1787,1811]" captionStart="Figure 54" captionStartId="24.[151,215,1886,1908]" captionTargetBox="[151,1399,1160,1864]" captionTargetId="figure-477@24.[169,1205,1180,1850]" captionTargetPageId="24" captionText="Figure 54. Principal component analysis of worker individuals considering all 16 morphometric characters. Triangles: Cardiocondyla weserka (grey: type of weserka, black: type of neferka, white: no type). Grey discs: syntypes of Cardiocondyla yemeni (Black rhombs: types of Cardiocondyla yoruba. White rhombs: C. yoruba sample from Comoe / Ivory coast). Black squares: Holotype and eight paratypes of Cardiocondyla heinzei n.sp. from Comoe / Ivory coast." figureDoi="http://doi.org/10.5281/zenodo.13194052" httpUri="https://zenodo.org/record/13194052/files/figure.png" pageId="23" pageNumber="136">Fig. 54</figureCitation>
). However, I stood back from synonymization of
<taxonomicName id="18D14D73FF86FFDD6C605D43FB60334A" authorityName="Seifert" authorityYear="2024" box="[1147,1224,1820,1844]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="136" phylum="Arthropoda" rank="species" species="yoruba">
<emphasis id="EDA5EAE2FF86FFDD6C605D43FB60334A" box="[1147,1224,1820,1844]" italics="true" pageId="23" pageNumber="136">yoruba</emphasis>
</taxonomicName>
because of the very remote allopatric ranges, the blackish brown gaster not observed so far in the
<taxonomicName id="18D14D73FF86FFDD6C2A5D01FBD5330B" authorityName="Seifert" authorityYear="2024" box="[1073,1149,1886,1909]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="136" phylum="Arthropoda" rank="species" species="yemeni">
<emphasis id="EDA5EAE2FF86FFDD6C2A5D01FBD5330B" box="[1073,1149,1886,1909]" italics="true" pageId="23" pageNumber="136">yemeni</emphasis>
</taxonomicName>
samples from
<collectingCountry id="A7C67660FF86FFDD6D395D01FAC5330B" box="[1314,1389,1886,1909]" name="Yemen" pageId="23" pageNumber="136">Yemen</collectingCountry>
and
<collectingCountry id="A7C67660FF86FFDD6B235D21FCD233E8" box="[824,890,1918,1942]" name="Oman" pageId="23" pageNumber="136">Oman</collectingCountry>
and because of the stronger microsculpture of
<taxonomicName id="18D14D73FF86FFDD6B235DC0FC2D33C9" authorityName="Seifert" authorityYear="2024" box="[824,901,1951,1975]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="136" phylum="Arthropoda" rank="species" species="yoruba">
<emphasis id="EDA5EAE2FF86FFDD6B235DC0FC2D33C9" box="[824,901,1951,1975]" italics="true" pageId="23" pageNumber="136">yoruba</emphasis>
</taxonomicName>
in particular on first gaster tergite. We have to wait if
<taxonomicName id="18D14D73FF89FFD268F85AAEFE983577" authorityName="Seifert" authorityYear="2024" box="[227,304,241,265]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="137" phylum="Arthropoda" rank="species" species="yoruba">
<emphasis id="EDA5EAE2FF89FFD268F85AAEFE983577" box="[227,304,241,265]" italics="true" pageId="24" pageNumber="137">yoruba</emphasis>
</taxonomicName>
might be more convingly demonstrated as a separate cryptic species when plenty of samples were analyzed morphometrically or genetically.
</paragraph>
</subSubSection>
<subSubSection id="97CB657BFF89FFD268B55B0BFEBF366E" pageId="24" pageNumber="137" type="biology_ecology">
<paragraph id="DF6E36F0FF89FFD268B55B0BFEBF366E" blockId="24.[151,764,241,529]" pageId="24" pageNumber="137">
<emphasis id="EDA5EAE2FF89FFD268B55B0BFEA33512" bold="true" box="[174,267,340,364]" pageId="24" pageNumber="137">Biology.</emphasis>
The species is seems to follow other
<taxonomicName id="18D14D73FF89FFD2688C5B2AFE3D35F3" box="[151,405,373,397]" pageId="24" pageNumber="137">
<emphasis id="EDA5EAE2FF89FFD2688C5B2AFE9135F3" box="[151,313,373,397]" italics="true" pageId="24" pageNumber="137">Cardiocondyla</emphasis>
species
</taxonomicName>
by living in anthropogenously disturbed areas. As habitats were reported a grassy patch adjacent to a paved road in the urban center of Accra /
<collectingCountry id="A7C67660FF89FFD2688C5B87FF76358E" box="[151,222,472,496]" name="Ghana" pageId="24" pageNumber="137">Ghana</collectingCountry>
and a path in the suburban area of Bondoukou /
<collectingCountry id="A7C67660FF89FFD2688C5BA6FEBB366E" box="[151,275,505,529]" name="Ivory Coast" pageId="24" pageNumber="137">Ivory coast</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>