304 lines
33 KiB
XML
304 lines
33 KiB
XML
<document id="4B1D305503FF8F9931D55C5EAD0AFD5F" ID-DOI="10.11646/zootaxa.3856.2.2" ID-GBIF-Dataset="24c01b66-c0d2-458b-a7fe-0e3de779a8e8" ID-ISSN="1175-5326" ID-Zenodo-Dep="250932" ID-ZooBank="ABC0E7D7-2E22-4605-A97B-0F1E632F3930" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460889728808" checkinUser="plazi" docAuthor="Bota, Cornelio Andrés" docDate="2014" docId="4E56376DFFA4FF9646CB63222DBAFE7A" docLanguage="en" docName="zt03856p210.pdf" docOrigin="Zootaxa 3856 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhionaeschna caligo Bota, 2014, sp. nov." docType="treatment" docVersion="8" lastPageNumber="200" masterDocId="B26F4F15FFA3FF9E465C607B2E0DFFB6" masterDocTitle="A brief look at the Odonata from the Páramo ecosystems in Colombia, with the descriptions of Oxyallagma colombianum sp. nov. and Rhionaeschna caligo sp. nov. (Odonata: Coenagrionidae, Aeshnidae, Libellulidae)" masterLastPageNumber="210" masterPageNumber="192" pageNumber="199" updateTime="1698359463704" updateUser="plazi">
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<mods:title id="5B53EC4AC7FE1D6D6A8DB76E990C2D79">A brief look at the Odonata from the Páramo ecosystems in Colombia, with the descriptions of Oxyallagma colombianum sp. nov. and Rhionaeschna caligo sp. nov. (Odonata: Coenagrionidae, Aeshnidae, Libellulidae)</mods:title>
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<mods:name id="41F3A070252669D1920B0C9EB21FDA54" type="personal">
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<mods:roleTerm id="FF2E250FF9A42CC87CD75BBA63860561">Author</mods:roleTerm>
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<mods:namePart id="971EF04B448B2E61ABDA8E26529E3274">Bota, Cornelio Andrés</mods:namePart>
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<mods:typeOfResource id="5E7E24015835E5AF9A55B46988649268">text</mods:typeOfResource>
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<mods:title id="BF7229BFECD6B085634E5F1432EF3FD5">Zootaxa</mods:title>
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<mods:part id="EB427CD749DE760EF2E7EFBD9C46B6EE">
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<mods:date id="0AF07E5F5C57E0B547F96EF015581029">2014</mods:date>
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<mods:detail id="32EFB10179E8FE76E5991F730DC223DA" type="volume">
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<mods:number id="4BCE3E735EBDD8C382F000A6F05B4AD1">3856</mods:number>
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<mods:detail id="C2B691A9C7B9728D4D685B0C5D9D1B5D" type="issue">
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<mods:number id="9F56304C4954382C8F6606091455060D">2</mods:number>
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<mods:classification id="1262683C8A2A484899DCF511F9B0E93B">journal article</mods:classification>
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<mods:identifier id="A5BC918A4371E14A9885749F7DC56E1C" type="DOI">10.11646/zootaxa.3856.2.2</mods:identifier>
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<treatment id="4E56376DFFA4FF9646CB63222DBAFE7A" ID-DOI="http://doi.org/10.5281/zenodo.6141385" ID-GBIF-Taxon="119586698" ID-Zenodo-Dep="6141385" LSID="urn:lsid:plazi:treatment:4E56376DFFA4FF9646CB63222DBAFE7A" httpUri="http://treatment.plazi.org/id/4E56376DFFA4FF9646CB63222DBAFE7A" lastPageId="8" lastPageNumber="200" pageId="7" pageNumber="199">
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<subSubSection id="8EE5D5F0FFA4FF9946CB63222A48FC23" pageId="7" pageNumber="199" type="nomenclature">
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<paragraph id="C640867BFFA4FF9946CB63222FF6FCC5" blockId="7.[151,1093,857,917]" box="[151,507,857,883]" pageId="7" pageNumber="199">
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<heading id="9D083117FFA4FF9946CB63222FF6FCC5" bold="true" box="[151,507,857,883]" fontSize="11" level="1" pageId="7" pageNumber="199" reason="1">
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<emphasis id="F48B5A69FFA4FF9946CB63222FF6FCC5" bold="true" box="[151,507,857,883]" pageId="7" pageNumber="199">
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<taxonomicName id="01FFFDF8FFA4FF9946CB63222F98FCC5" ID-CoL="9X49R" box="[151,405,857,883]" class="Insecta" family="Aeshnidae" genus="Rhionaeschna" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="199" phylum="Arthropoda" rank="species" species="caligo" status="sp. nov.">
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<emphasis id="F48B5A69FFA4FF9946CB63222F98FCC5" bold="true" box="[151,405,857,883]" italics="true" pageId="7" pageNumber="199">Rhionaeschna caligo</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="EFB8E712FFA4FF9947C163222FF6FCC5" box="[413,507,857,883]" pageId="7" pageNumber="199" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="C640867BFFA4FF9946CB63072A48FC23" blockId="7.[151,1093,857,917]" box="[151,1093,892,917]" pageId="7" pageNumber="199">
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<figureCitation id="5EC49AFEFFA4FF9946CB63072EEDFC22" box="[151,224,892,917]" captionStart="FIGURE 1" captionStartId="9.[151,250,1956,1978]" captionTargetBox="[151,1435,195,1933]" captionTargetId="figure@9.[151,1435,195,1934]" captionTargetPageId="9" captionText="FIGURE 1. Rhionaeschna caligo: (a) Male paratype (Páramo El Morro); (c) Male paratype (Páramo Sabanas). (b) Rhionaeschna marchali: Male (Tolima, Murillo)." httpUri="https://zenodo.org/record/250933/files/figure.png" pageId="7" pageNumber="199">Figs. 1</figureCitation>
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a (Habitus ♂), 1c (Frontal ♂), 2b
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<emphasis id="F48B5A69FFA4FF99440963062C6EFC22" box="[597,611,893,916]" italics="true" pageId="7" pageNumber="199">–</emphasis>
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e (Diagnostic traits), Table 1 (coordinates).
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</paragraph>
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</subSubSection>
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<subSubSection id="8EE5D5F0FFA4FF9946CB63BF2C8AFBB6" pageId="7" pageNumber="199" type="etymology">
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<paragraph id="C640867BFFA4FF9946CB63BF2C8AFBB6" blockId="7.[151,1437,964,2033]" pageId="7" pageNumber="199">
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<emphasis id="F48B5A69FFA4FF9946CB63BF2F13FC6B" bold="true" box="[151,286,964,989]" pageId="7" pageNumber="199">Etymology.</emphasis>
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From Latin
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<taxonomicName id="01FFFDF8FFA4FF9947F763BF2FFCFC6A" box="[427,497,964,988]" class="Insecta" family="Aeshnidae" genus="Rhionaeschna" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="199" phylum="Arthropoda" rank="species" species="caligo">
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<emphasis id="F48B5A69FFA4FF9947F763BF2FFCFC6A" box="[427,497,964,988]" italics="true" pageId="7" pageNumber="199">caligo</emphasis>
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</taxonomicName>
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meaning fog, to be treated as a noun in apposition. The name refers to this species’ habit of flying even on cold and foggy days.
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</paragraph>
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</subSubSection>
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<subSubSection id="8EE5D5F0FFA4FF99469B64772C96FBDA" pageId="7" pageNumber="199" type="materials_examined">
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<paragraph id="C640867BFFA4FF99469B64772C96FBDA" blockId="7.[151,1437,964,2033]" pageId="7" pageNumber="199">
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<emphasis id="F48B5A69FFA4FF99469B64772F63FB93" bold="true" box="[199,366,1036,1061]" pageId="7" pageNumber="199">
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<typeStatus id="194438D9FFA4FF99469B64772F0FFB93" box="[199,258,1036,1061]" pageId="7" pageNumber="199">Type</typeStatus>
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material
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</emphasis>
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(
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<specimenCount id="D0F94DF2FFA4FF99472364772FA4FB92" box="[383,425,1036,1060]" pageId="7" pageNumber="199" type="generic">8 ♂</specimenCount>
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)
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<collectingCountry id="BEE8C6EBFFA4FF9947E664772C23FB93" box="[442,558,1036,1061]" name="Colombia" pageId="7" pageNumber="199">
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<emphasis id="F48B5A69FFA4FF9947E664772C23FB93" bold="true" box="[442,558,1036,1061]" pageId="7" pageNumber="199">Colombia</emphasis>
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</collectingCountry>
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: Antioquia:
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<taxonomicName id="01FFFDF8FFA4FF9944E364772D19FB92" box="[703,788,1036,1060]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="210" phylum="Arthropoda" rank="species" species="paramo">Páramo</taxonomicName>
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El Morro:
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<specimenCount id="D0F94DF2FFA4FF9945CF64772DB2FB92" box="[915,959,1036,1060]" pageId="7" pageNumber="199" type="generic">2 ♂</specimenCount>
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,
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<date id="B241A0BBFFA4FF99459064772A7DFB93" box="[972,1136,1036,1061]" pageId="7" pageNumber="199" value="2012-06-15" valueMax="2012-06-17" valueMin="2012-06-15">
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15
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<emphasis id="F48B5A69FFA4FF9945B464762DFBFB92" box="[1000,1014,1037,1060]" italics="true" pageId="7" pageNumber="199">–</emphasis>
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17.vi.2012
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</date>
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, Leg: C. Bota, C. Moreno, J. Del Río, J. Caly & M. Moreno.
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<taxonomicName id="01FFFDF8FFA4FF99444C644B2C6BFBFE" box="[528,614,1072,1096]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="210" phylum="Arthropoda" rank="species" species="paramo">Páramo</taxonomicName>
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Sabanas:
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<specimenCount id="D0F94DF2FFA4FF99448A644B2D0FFBF1" box="[726,770,1072,1096]" pageId="7" pageNumber="199" type="generic">2 ♂</specimenCount>
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:
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<date id="B241A0BBFFA4FF994551644B2D7CFBFE" box="[781,881,1072,1096]" pageId="7" pageNumber="199" value="2011-10-03">3.x.2011</date>
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, Leg: C. Bota & J. D. Castaño.
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<taxonomicName id="01FFFDF8FFA4FF994292644B2B29FBFE" box="[1230,1316,1072,1096]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="210" phylum="Arthropoda" rank="species" species="paramo">Páramo</taxonomicName>
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<collectingCountry id="BEE8C6EBFFA4FF99437664542B94FBFE" box="[1322,1433,1071,1096]" name="Republic of the Congo" pageId="7" pageNumber="199">El Congo</collectingCountry>
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:
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<specimenCount id="D0F94DF2FFA4FF9946CB642F2ECEFBDA" box="[151,195,1108,1132]" pageId="7" pageNumber="199" type="generic">4 ♂</specimenCount>
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,
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<date id="B241A0BBFFA4FF994693642F2F7CFBDB" box="[207,369,1108,1133]" pageId="7" pageNumber="199" value="2011-09-10" valueMax="2011-09-14" valueMin="2011-09-10">
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10
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<emphasis id="F48B5A69FFA4FF9946B7642E2EF4FBDA" box="[235,249,1109,1132]" italics="true" pageId="7" pageNumber="199">–</emphasis>
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14.ix.2011
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</date>
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, Leg: L. Ríos & J. Zapata.
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</paragraph>
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</subSubSection>
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<subSubSection id="8EE5D5F0FFA4FF99469B640C2DD9F8AE" pageId="7" pageNumber="199" type="description">
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<paragraph id="C640867BFFA4FF99469B640C2A35F966" blockId="7.[151,1437,964,2033]" pageId="7" pageNumber="199">
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<emphasis id="F48B5A69FFA4FF99469B640C2C2EFB26" bold="true" box="[199,547,1143,1168]" pageId="7" pageNumber="199">
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Description.
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<typeStatus id="194438D9FFA4FF994703640C2FDDFB26" box="[351,464,1143,1168]" pageId="7" pageNumber="199" type="holotype">Holotype</typeStatus>
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:
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<emphasis id="F48B5A69FFA4FF994780640C2C17FB26" bold="true" box="[476,538,1143,1168]" italics="true" pageId="7" pageNumber="199">Head</emphasis>
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:
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</emphasis>
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Labium brown. Labrum light blue, with lateral margins yellow, and ventral and dorsal margin dark brown. Clypeus light blue, with lateral margins yellow. Frons yellow, light blue spots lateral to yellow area surrounding T-spot stem; T-spot stem narrower than vertex, approximately parallel sided, transverse arms about as wide as stem; vertex black with green spot at anterior margin on each side; brown stripe on frontoclypeal groove not widened to fronto
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<emphasis id="F48B5A69FFA4FF99442565722C8FFA96" box="[633,642,1289,1312]" italics="true" pageId="7" pageNumber="199">-</emphasis>
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ocular groove (
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<figureCitation id="5EC49AFEFFA4FF994570657C2D66FA96" box="[812,875,1287,1312]" captionStart="FIGURE 1" captionStartId="9.[151,250,1956,1978]" captionTargetBox="[151,1435,195,1933]" captionTargetId="figure@9.[151,1435,195,1934]" captionTargetPageId="9" captionText="FIGURE 1. Rhionaeschna caligo: (a) Male paratype (Páramo El Morro); (c) Male paratype (Páramo Sabanas). (b) Rhionaeschna marchali: Male (Tolima, Murillo)." httpUri="https://zenodo.org/record/250933/files/figure.png" pageId="7" pageNumber="199">Fig. 1</figureCitation>
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b).
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<emphasis id="F48B5A69FFA4FF9945CF657C2DE4FA96" bold="true" box="[915,1001,1287,1312]" italics="true" pageId="7" pageNumber="199">Thorax</emphasis>
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: Brown with pale pterothoracic stripes yellowish green, with indentations as in
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<figureCitation id="5EC49AFEFFA4FF99440065572C96FAF2" box="[604,667,1324,1349]" captionStart="FIGURE 1" captionStartId="9.[151,250,1956,1978]" captionTargetBox="[151,1435,195,1933]" captionTargetId="figure@9.[151,1435,195,1934]" captionTargetPageId="9" captionText="FIGURE 1. Rhionaeschna caligo: (a) Male paratype (Páramo El Morro); (c) Male paratype (Páramo Sabanas). (b) Rhionaeschna marchali: Male (Tolima, Murillo)." httpUri="https://zenodo.org/record/250933/files/figure.png" pageId="7" pageNumber="199">Fig. 1</figureCitation>
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a; legs black except for trochanters, coxae, and basal 75% of femora reddish brown, and basal 50% of inner surface of femur I yellowish green. Wings smoky, darker at base and along costa; membranule dark except basal 50% pale; Pt whitish on ventral side and brown on dorsal side. Ax in FW: 19 right, 20 left; HW: 13 right, 12 left; Px in FW: 12 right, 13 left; HW: 15; HW with two rows of cells between
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<collectingCountry id="BEE8C6EBFFA4FF99433065E32B9EFA06" box="[1388,1427,1432,1456]" name="Poland" pageId="7" pageNumber="199">RP</collectingCountry>
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1 and
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<collectingCountry id="BEE8C6EBFFA4FF99469A65C72EE4FA62" box="[198,233,1468,1492]" name="Poland" pageId="7" pageNumber="199">RP</collectingCountry>
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2 beginning under pterostigma.
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<emphasis id="F48B5A69FFA4FF99441265C72CB1FA63" bold="true" box="[590,700,1468,1493]" italics="true" pageId="7" pageNumber="199">Abdomen</emphasis>
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: brown with sterna of S9
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<emphasis id="F48B5A69FFA4FF99458665C62DE5FA62" box="[986,1000,1469,1492]" italics="true" pageId="7" pageNumber="199">–</emphasis>
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10 posterior to genital opercula brown, and yellowish green and light blue spots as in
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<figureCitation id="5EC49AFEFFA4FF9944FD65A42CECFA4E" box="[673,737,1503,1528]" captionStart="FIGURE 1" captionStartId="9.[151,250,1956,1978]" captionTargetBox="[151,1435,195,1933]" captionTargetId="figure@9.[151,1435,195,1934]" captionTargetPageId="9" captionText="FIGURE 1. Rhionaeschna caligo: (a) Male paratype (Páramo El Morro); (c) Male paratype (Páramo Sabanas). (b) Rhionaeschna marchali: Male (Tolima, Murillo)." httpUri="https://zenodo.org/record/250933/files/figure.png" pageId="7" pageNumber="199">Fig. 1</figureCitation>
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a. Outer margin of ventral terga VII linear. Ventral tubercle of S1 rounded in lateral view, genital lobe lower than 30% of its length; anterior lamina spine as long as about two times its basal width (
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<figureCitation id="5EC49AFEFFA4FF9947D7665C2FC3F9F6" box="[395,462,1575,1600]" captionStart="FIGURE 2" captionStartId="10.[151,250,1953,1975]" captionTargetBox="[157,1433,199,1931]" captionTargetId="figure@10.[151,1436,193,1932]" captionTargetPageId="10" captionText="FIGURE 2. (a) Rhionaeschna peralta female (habitus). Rhionaeschna caligo: (b) Hamuli, ventroposterior view; (c) Caudal appendages lateral view; (d) Vesica spermalis ventral view; (e) left spine of anterior lamina. (f) Rhionaeschna joannisi male (habitus). Illustrations by Natalia Uribe Macias." httpUri="https://zenodo.org/record/250934/files/figure.png" pageId="7" pageNumber="199">Fig. 2</figureCitation>
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f); ventral portion of hamuli shorter than hamular fold (
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<figureCitation id="5EC49AFEFFA4FF994265665C2A71F9F6" box="[1081,1148,1575,1600]" captionStart="FIGURE 2" captionStartId="10.[151,250,1953,1975]" captionTargetBox="[157,1433,199,1931]" captionTargetId="figure@10.[151,1436,193,1932]" captionTargetPageId="10" captionText="FIGURE 2. (a) Rhionaeschna peralta female (habitus). Rhionaeschna caligo: (b) Hamuli, ventroposterior view; (c) Caudal appendages lateral view; (d) Vesica spermalis ventral view; (e) left spine of anterior lamina. (f) Rhionaeschna joannisi male (habitus). Illustrations by Natalia Uribe Macias." httpUri="https://zenodo.org/record/250934/files/figure.png" pageId="7" pageNumber="199">Fig. 2</figureCitation>
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b), tip of hamular anterior process pointed (
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<figureCitation id="5EC49AFEFFA4FF99470B66372F91F9D3" box="[343,412,1612,1637]" captionStart="FIGURE 2" captionStartId="10.[151,250,1953,1975]" captionTargetBox="[157,1433,199,1931]" captionTargetId="figure@10.[151,1436,193,1932]" captionTargetPageId="10" captionText="FIGURE 2. (a) Rhionaeschna peralta female (habitus). Rhionaeschna caligo: (b) Hamuli, ventroposterior view; (c) Caudal appendages lateral view; (d) Vesica spermalis ventral view; (e) left spine of anterior lamina. (f) Rhionaeschna joannisi male (habitus). Illustrations by Natalia Uribe Macias." httpUri="https://zenodo.org/record/250934/files/figure.png" pageId="7" pageNumber="199">Fig. 2</figureCitation>
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b); two teeth on left auricle, two large and a third smaller and more external tooth on right auricle; genital lobe subtriangular, about as high as S1 tubercle. Vesica spermalis distal segment with basal folds in lateral and dorsal lobes as depicted in
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<figureCitation id="5EC49AFEFFA4FF99441166EF2C9CF91B" box="[589,657,1684,1709]" captionStart="FIGURE 2" captionStartId="10.[151,250,1953,1975]" captionTargetBox="[157,1433,199,1931]" captionTargetId="figure@10.[151,1436,193,1932]" captionTargetPageId="10" captionText="FIGURE 2. (a) Rhionaeschna peralta female (habitus). Rhionaeschna caligo: (b) Hamuli, ventroposterior view; (c) Caudal appendages lateral view; (d) Vesica spermalis ventral view; (e) left spine of anterior lamina. (f) Rhionaeschna joannisi male (habitus). Illustrations by Natalia Uribe Macias." httpUri="https://zenodo.org/record/250934/files/figure.png" pageId="7" pageNumber="199">Fig. 2</figureCitation>
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d. Dorso
|
||
<emphasis id="F48B5A69FFA4FF9944AE66EE2CF6F91A" box="[754,763,1685,1708]" italics="true" pageId="7" pageNumber="199">-</emphasis>
|
||
distal crest of male cercus smoothly convex, tip blunt (
|
||
<figureCitation id="5EC49AFEFFA4FF99433366EF2EAAF966" captionStart="FIGURE 2" captionStartId="10.[151,250,1953,1975]" captionTargetBox="[157,1433,199,1931]" captionTargetId="figure@10.[151,1436,193,1932]" captionTargetPageId="10" captionText="FIGURE 2. (a) Rhionaeschna peralta female (habitus). Rhionaeschna caligo: (b) Hamuli, ventroposterior view; (c) Caudal appendages lateral view; (d) Vesica spermalis ventral view; (e) left spine of anterior lamina. (f) Rhionaeschna joannisi male (habitus). Illustrations by Natalia Uribe Macias." httpUri="https://zenodo.org/record/250934/files/figure.png" pageId="7" pageNumber="199">Fig. 2</figureCitation>
|
||
c). Total length
|
||
<quantity id="01072B9EFFA4FF99470A66CC2FA5F966" box="[342,424,1719,1744]" metricMagnitude="-2" metricUnit="m" metricValue="6.2" pageId="7" pageNumber="199" unit="mm" value="62.0">62 mm</quantity>
|
||
. Abdomen length
|
||
<quantity id="01072B9EFFA4FF99442966CC2CCAF966" box="[629,711,1719,1744]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="7" pageNumber="199" unit="mm" value="45.0">45 mm</quantity>
|
||
. FW
|
||
<quantity id="01072B9EFFA4FF99455E66CC2D59F966" box="[770,852,1719,1744]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="7" pageNumber="199" unit="mm" value="45.0">45 mm</quantity>
|
||
. HW length
|
||
<quantity id="01072B9EFFA4FF9945BE66C32A39F966" box="[994,1076,1720,1744]" metricMagnitude="-2" metricUnit="m" metricValue="4.3" pageId="7" pageNumber="199" unit="mm" value="43.0">43 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="C640867BFFA4FF99469B66A72DD9F8AE" blockId="7.[151,1437,964,2033]" pageId="7" pageNumber="199">
|
||
<emphasis id="F48B5A69FFA4FF99469B66A72FD9F943" bold="true" box="[199,468,1756,1781]" pageId="7" pageNumber="199">
|
||
Variation in
|
||
<typeStatus id="194438D9FFA4FF99470266A72FD9F943" box="[350,468,1756,1781]" pageId="7" pageNumber="199" type="paratype">paratypes</typeStatus>
|
||
</emphasis>
|
||
: Ax in FW 18
|
||
<emphasis id="F48B5A69FFA4FF99442666A62C85F942" box="[634,648,1757,1780]" italics="true" pageId="7" pageNumber="199">–</emphasis>
|
||
22, in Hw 11
|
||
<emphasis id="F48B5A69FFA4FF99454066A62D27F942" box="[796,810,1757,1780]" italics="true" pageId="7" pageNumber="199">–</emphasis>
|
||
13. Px in FW 11
|
||
<emphasis id="F48B5A69FFA4FF9945B966A62DFEF942" box="[997,1011,1757,1780]" italics="true" pageId="7" pageNumber="199">–</emphasis>
|
||
15, in HW 13
|
||
<emphasis id="F48B5A69FFA4FF9942D266A62A91F942" box="[1166,1180,1757,1780]" italics="true" pageId="7" pageNumber="199">–</emphasis>
|
||
17. Total length
|
||
<quantity id="01072B9EFFA4FF99430966A72ECBF8AE" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="6.5" metricValueMin="6.2" pageId="7" pageNumber="199" unit="mm" value="63.5" valueMax="65.0" valueMin="62.0">
|
||
62
|
||
<emphasis id="F48B5A69FFA4FF99432E66A62B8DF942" box="[1394,1408,1757,1780]" italics="true" pageId="7" pageNumber="199">–</emphasis>
|
||
65 mm
|
||
</quantity>
|
||
. Abdomen length
|
||
<quantity id="01072B9EFFA4FF9947CF66842C02F8AE" box="[403,527,1791,1816]" metricMagnitude="-2" metricUnit="m" metricValue="4.6" metricValueMax="4.7" metricValueMin="4.5" pageId="7" pageNumber="199" unit="mm" value="46.0" valueMax="47.0" valueMin="45.0">
|
||
45
|
||
<emphasis id="F48B5A69FFA4FF9947F3677A2FB0F8AE" box="[431,445,1793,1816]" italics="true" pageId="7" pageNumber="199">–</emphasis>
|
||
47 mm
|
||
</quantity>
|
||
. FW
|
||
<quantity id="01072B9EFFA4FF994416677B2CCBF8AE" box="[586,710,1791,1816]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" metricValueMax="4.6" metricValueMin="4.4" pageId="7" pageNumber="199" unit="mm" value="45.0" valueMax="46.0" valueMin="44.0">
|
||
44
|
||
<emphasis id="F48B5A69FFA4FF99443A677A2C79F8AE" box="[614,628,1793,1816]" italics="true" pageId="7" pageNumber="199">–</emphasis>
|
||
46 mm
|
||
</quantity>
|
||
. HW length
|
||
<quantity id="01072B9EFFA4FF994508677B2DC2F8AE" box="[852,975,1792,1816]" metricMagnitude="-2" metricUnit="m" metricValue="4.25" metricValueMax="4.4" metricValueMin="4.1" pageId="7" pageNumber="199" unit="mm" value="42.5" valueMax="44.0" valueMin="41.0">
|
||
41
|
||
<emphasis id="F48B5A69FFA4FF994533677A2D70F8AE" box="[879,893,1793,1816]" italics="true" pageId="7" pageNumber="199">–</emphasis>
|
||
44 mm
|
||
</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8EE5D5F0FFA4FF99469B675F2D44F847" pageId="7" pageNumber="199" type="diagnosis">
|
||
<paragraph id="C640867BFFA4FF99469B675F2D44F847" blockId="7.[151,1437,964,2033]" pageId="7" pageNumber="199">
|
||
<emphasis id="F48B5A69FFA4FF99469B675F2F4CF88B" bold="true" box="[199,321,1828,1853]" pageId="7" pageNumber="199">Diagnosis.</emphasis>
|
||
This species fits within the
|
||
<taxonomicName id="01FFFDF8FFA4FF994425675F2D8DF88A" box="[633,896,1828,1852]" class="Insecta" family="Aeshnidae" genus="Rhionaeschna" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="199" phylum="Arthropoda" rank="species" species="punctata">
|
||
<emphasis id="F48B5A69FFA4FF994425675F2D8DF88A" box="[633,896,1828,1852]" italics="true" pageId="7" pageNumber="199">Rhionaeschna punctata</emphasis>
|
||
</taxonomicName>
|
||
group proposed by von
|
||
<bibRefCitation id="A26EFB8AFFA4FF9942D3675F2B95F88A" author="Ellenrieder" box="[1167,1432,1828,1853]" pageId="7" pageNumber="199" refString="von Ellenrieder, N. (2003) A synopsis of the neotropical species of " Aeshna " Fabricius: Genus Rhionaeschna Forster. Tijdschrift voor Entomologie, 146, 67 - 207. http: // dx. doi. org / 10.1163 / 22119434 - 900000120" type="journal article" year="2003">Ellenrieder (2003: 107)</bibRefCitation>
|
||
, and within this group it resembles
|
||
<taxonomicName id="01FFFDF8FFA4FF99444467322CB7F8D6" box="[536,698,1864,1888]" class="Insecta" family="Aeshnidae" genus="Rhionaeschna" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="199" phylum="Arthropoda" rank="species" species="demarmelsi">
|
||
<emphasis id="F48B5A69FFA4FF99444467322CB7F8D6" box="[536,698,1864,1888]" italics="true" pageId="7" pageNumber="199">R. demarmelsi</emphasis>
|
||
</taxonomicName>
|
||
because of the blunt tips of cercus (
|
||
<figureCitation id="5EC49AFEFFA4FF994217673C2A83F8D6" box="[1099,1166,1863,1888]" captionStart="FIGURE 2" captionStartId="10.[151,250,1953,1975]" captionTargetBox="[157,1433,199,1931]" captionTargetId="figure@10.[151,1436,193,1932]" captionTargetPageId="10" captionText="FIGURE 2. (a) Rhionaeschna peralta female (habitus). Rhionaeschna caligo: (b) Hamuli, ventroposterior view; (c) Caudal appendages lateral view; (d) Vesica spermalis ventral view; (e) left spine of anterior lamina. (f) Rhionaeschna joannisi male (habitus). Illustrations by Natalia Uribe Macias." httpUri="https://zenodo.org/record/250934/files/figure.png" pageId="7" pageNumber="199">Fig. 2</figureCitation>
|
||
c) and ventral portion of hamuli shorter than hamular fold (
|
||
<figureCitation id="5EC49AFEFFA4FF99447C67172C68F833" box="[544,613,1900,1925]" captionStart="FIGURE 2" captionStartId="10.[151,250,1953,1975]" captionTargetBox="[157,1433,199,1931]" captionTargetId="figure@10.[151,1436,193,1932]" captionTargetPageId="10" captionText="FIGURE 2. (a) Rhionaeschna peralta female (habitus). Rhionaeschna caligo: (b) Hamuli, ventroposterior view; (c) Caudal appendages lateral view; (d) Vesica spermalis ventral view; (e) left spine of anterior lamina. (f) Rhionaeschna joannisi male (habitus). Illustrations by Natalia Uribe Macias." httpUri="https://zenodo.org/record/250934/files/figure.png" pageId="7" pageNumber="199">Fig. 2</figureCitation>
|
||
b), but it clearly differs from
|
||
<taxonomicName id="01FFFDF8FFA4FF9945EB67162A56F832" box="[951,1115,1900,1924]" class="Insecta" family="Aeshnidae" genus="Rhionaeschna" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="199" phylum="Arthropoda" rank="species" species="demarmelsi">
|
||
<emphasis id="F48B5A69FFA4FF9945EB67162A56F832" box="[951,1115,1900,1924]" italics="true" pageId="7" pageNumber="199">R. demarmelsi</emphasis>
|
||
</taxonomicName>
|
||
by its anterior lamina spine being about twice as long as its basal width (
|
||
<figureCitation id="5EC49AFEFFA4FF9944F867F42CE4F81E" box="[676,745,1935,1960]" captionStart="FIGURE 2" captionStartId="10.[151,250,1953,1975]" captionTargetBox="[157,1433,199,1931]" captionTargetId="figure@10.[151,1436,193,1932]" captionTargetPageId="10" captionText="FIGURE 2. (a) Rhionaeschna peralta female (habitus). Rhionaeschna caligo: (b) Hamuli, ventroposterior view; (c) Caudal appendages lateral view; (d) Vesica spermalis ventral view; (e) left spine of anterior lamina. (f) Rhionaeschna joannisi male (habitus). Illustrations by Natalia Uribe Macias." httpUri="https://zenodo.org/record/250934/files/figure.png" pageId="7" pageNumber="199">Fig. 2</figureCitation>
|
||
e) rather than shorter than 1.5 times its basal width as in
|
||
<taxonomicName id="01FFFDF8FFA4FF9943D867EA2F17F87A" class="Insecta" family="Aeshnidae" genus="Rhionaeschna" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="199" phylum="Arthropoda" rank="species" species="demarmelsi">
|
||
<emphasis id="F48B5A69FFA4FF9943D867EA2F17F87A" italics="true" pageId="7" pageNumber="199">R. demarmelsi</emphasis>
|
||
</taxonomicName>
|
||
. Couplet 4 of Key M
|
||
<emphasis id="F48B5A69FFA4FF99444D67CE2C12F87A" box="[529,543,1973,1996]" italics="true" pageId="7" pageNumber="199">–</emphasis>
|
||
4 for males of the
|
||
<taxonomicName id="01FFFDF8FFA4FF9944AF67CF2DF0F87A" box="[755,1021,1972,1996]" class="Insecta" family="Aeshnidae" genus="Rhionaeschna" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="199" phylum="Arthropoda" rank="species" species="punctata">
|
||
<emphasis id="F48B5A69FFA4FF9944AF67CF2DF0F87A" box="[755,1021,1972,1996]" italics="true" pageId="7" pageNumber="199">Rhionaeschna punctata</emphasis>
|
||
</taxonomicName>
|
||
group in von
|
||
<bibRefCitation id="A26EFB8AFFA4FF9942FC67CF2B91F87B" author="Ellenrieder" box="[1184,1436,1972,1997]" pageId="7" pageNumber="199" refString="von Ellenrieder, N. (2003) A synopsis of the neotropical species of " Aeshna " Fabricius: Genus Rhionaeschna Forster. Tijdschrift voor Entomologie, 146, 67 - 207. http: // dx. doi. org / 10.1163 / 22119434 - 900000120" type="journal article" year="2003">Ellenrieder (2003: 76)</bibRefCitation>
|
||
can be modified as follows to accommodate this new species:
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8EE5D5F0FFABFF9646CB60EC2FF2FE33" pageId="8" pageNumber="200" type="key">
|
||
<key id="B26ED14AFFABFF9646CB60EC2B91FE89" pageId="8" pageNumber="200">
|
||
<keyStep id="7D0B9EDEFFABFF9646CB60EC2B96FEB2" pageId="8" pageNumber="200">
|
||
<paragraph id="C640867BFFABFF9646CB60EC2B96FF7C" blockId="8.[151,1436,151,319]" pageId="8" pageNumber="200">
|
||
<keyLead id="7D0E254EFFABFF9646CB60EC2B96FF7C" pageId="8" pageNumber="200">
|
||
4. Dorso
|
||
<emphasis id="F48B5A69FFABFF96474760E32F2AFF1B" box="[283,295,152,173]" italics="true" pageId="8" pageNumber="200">–</emphasis>
|
||
distal crest of cercus smoothly convex in lateral view (fig. 393b); ventral portion of hamuli (A) shorter than hamular fold (B) (fig. 311a)................................................................................... 5
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="C640867BFFABFF9646CB60A92B96FEB2" blockId="8.[151,1436,151,319]" pageId="8" pageNumber="200">
|
||
<keyLead id="7D0E254EFFABFF9646CB60A92B96FEB2" pageId="8" pageNumber="200">
|
||
– Dorso
|
||
<emphasis id="F48B5A69FFABFF96474760A92F2AFF51" box="[283,295,210,231]" italics="true" pageId="8" pageNumber="200">–</emphasis>
|
||
distal crest of cercus angulate in lateral view (figs. 390
|
||
<emphasis id="F48B5A69FFABFF96456660A92D4BFF51" box="[826,838,210,231]" italics="true" pageId="8" pageNumber="200">–</emphasis>
|
||
392b); ventral portion of hamuli (A) longer than hamular fold (B) (figs. 310, 312
|
||
<emphasis id="F48B5A69FFABFF9647CF608B2F92FEB3" box="[403,415,240,261]" italics="true" pageId="8" pageNumber="200">–</emphasis>
|
||
313)............................................................................... 6
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="7D0B9EDEFFABFF9646CB61762B91FE89" pageId="8" pageNumber="200">
|
||
<paragraph id="C640867BFFABFF9646CB61762B91FE94" blockId="8.[151,1436,151,319]" box="[151,1436,268,290]" pageId="8" pageNumber="200">
|
||
<keyLead id="7D0E254EFFABFF9646CB61762B91FE94" box="[151,1436,268,290]" pageId="8" pageNumber="200">
|
||
5. Anterior lamina spine as long as 30% of its basal width (fig. 353
|
||
<emphasis id="F48B5A69FFABFF96451761762D5AFE94" box="[843,855,269,290]" italics="true" pageId="8" pageNumber="200">–</emphasis>
|
||
354); Táchira State in
|
||
<collectingCountry id="BEE8C6EBFFABFF96427161762A9DFE94" box="[1069,1168,269,290]" name="Venezuela" pageId="8" pageNumber="200">Venezuela</collectingCountry>
|
||
(fig. 458)...
|
||
<taxonomicName id="01FFFDF8FFABFF96437061772B91FE94" ID-CoL="6X5FT" box="[1324,1436,268,290]" class="Insecta" family="Aeshnidae" genus="Rhionaeschna" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="200" phylum="Arthropoda" rank="species" species="demarmelsi">
|
||
<emphasis id="F48B5A69FFABFF96437061772B91FE94" box="[1324,1436,268,290]" italics="true" pageId="8" pageNumber="200">demarmelsi</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="C640867BFFABFF9646CB61512B91FE89" blockId="8.[151,1436,151,319]" box="[151,1436,297,319]" pageId="8" pageNumber="200">
|
||
<keyLead id="7D0E254EFFABFF9646CB61512B91FE89" box="[151,1436,297,319]" pageId="8" pageNumber="200">
|
||
– Anterior lamina spine as long as 2 times its basal width; Belmira
|
||
<taxonomicName id="01FFFDF8FFABFF96450E61512D96FE89" ID-CoL="53P8W" box="[850,923,298,319]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="210" phylum="Arthropoda" rank="species" species="paramo">Páramo</taxonomicName>
|
||
Complex in Antioquia Dep.,
|
||
<collectingCountry id="BEE8C6EBFFABFF9642E461512B1AFE89" box="[1208,1303,298,319]" name="Colombia" pageId="8" pageNumber="200">Colombia</collectingCountry>
|
||
....
|
||
<taxonomicName id="01FFFDF8FFABFF96433C61522B91FE89" ID-CoL="9X49R" box="[1376,1436,297,319]" class="Insecta" family="Aeshnidae" genus="Rhionaeschna" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="200" phylum="Arthropoda" rank="species" species="caligo">
|
||
<emphasis id="F48B5A69FFABFF96433C61522B91FE89" box="[1376,1436,297,319]" italics="true" pageId="8" pageNumber="200">caligo</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
<paragraph id="C640867BFFABFF96469B61172FF2FE33" blockId="8.[199,951,364,461]" box="[199,511,364,389]" pageId="8" pageNumber="200">Female and larva unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8EE5D5F0FFABFF96469B61F42DBAFE7A" pageId="8" pageNumber="200" type="distribution">
|
||
<paragraph id="C640867BFFABFF96469B61F42DBAFE7A" blockId="8.[199,951,364,461]" pageId="8" pageNumber="200">
|
||
<emphasis id="F48B5A69FFABFF96469B61F42F6DFE1E" bold="true" box="[199,352,399,424]" pageId="8" pageNumber="200">Distribution.</emphasis>
|
||
Belmira
|
||
<taxonomicName id="01FFFDF8FFABFF96479461EB2C10FE1E" box="[456,541,400,424]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="210" phylum="Arthropoda" rank="species" species="paramo">Páramo</taxonomicName>
|
||
complex, Antioquia,
|
||
<collectingCountry id="BEE8C6EBFFABFF96454D61F42D8EFE1E" box="[785,899,399,424]" name="Colombia" pageId="8" pageNumber="200">Colombia</collectingCountry>
|
||
.
|
||
<emphasis id="F48B5A69FFABFF96469B61CF2F2AFE7B" bold="true" box="[199,295,436,461]" pageId="8" pageNumber="200">Biology.</emphasis>
|
||
Male behavior is similar to that described for
|
||
<taxonomicName id="01FFFDF8FFABFF96457161CE2DBDFE7A" box="[813,944,436,460]" class="Insecta" family="Aeshnidae" genus="Rhionaeschna" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="200" phylum="Arthropoda" rank="species" species="marchali">
|
||
<emphasis id="F48B5A69FFABFF96457161CE2DBDFE7A" box="[813,944,436,460]" italics="true" pageId="8" pageNumber="200">R. marchali</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |