treatments-xml/data/2C/5F/93/2C5F9313693FD61598FAFF0DFE2AFEF7.xml

<document id="3906818947C591D0859B59A78741E0D7" ID="10.11646/zootaxa.4184.2.7" ID-DOI="10.11646/zootaxa.4184.2.7" ID-GBIF-Dataset="d45cdf93-6140-417b-a1d5-5acee38de05b" ID-ISSN="1175-5326" ID-Zenodo-Dep="164786" ID-ZooBank="24D9746D-2FE9-4CBC-A124-AB0CB3E5C602" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1478246841168" checkinUser="plazi" docAuthor="Nakahara, Shinichi, Vega, German &amp; Willmott, Keith R." docDate="2016" docId="2C5F9313693FD61598FAFF0DFE2AFEF7" docLanguage="en" docName="zootaxa.4184.2.7.pdf" docOrigin="Zootaxa 4184 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Euptychia favonius Nakahara, Vega &amp; Willmott, new species" docType="treatment" docVersion="9" lastPageNumber="365" masterDocId="D066EB6B693DD612986DFF9AFFEDFF92" masterDocTitle="Description of a new species of Euptychia Hübner, 1818 (Lepidoptera: Nymphalidae: Satyrinae) from the western Andes" masterLastPageNumber="366" masterPageNumber="358" pageNumber="360" updateTime="1698392110101" updateUser="plazi">
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<mods:title id="5E6EE2BDD4CC86427EB94CD23949A5D5">Description of a new species of Euptychia Hübner, 1818 (Lepidoptera: Nymphalidae: Satyrinae) from the western Andes</mods:title>
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<subSubSection id="ECEC718E693FD61098FAFF0DFEE7FF41" pageId="2" pageNumber="360" type="nomenclature">
<paragraph id="A4492205693FD61098FAFF0DFC6AFF20" blockId="2.[151,903,151,211]" box="[151,903,151,178]" pageId="2" pageNumber="360">
<heading id="FF019569693FD61098FAFF0DFC6AFF20" bold="true" box="[151,903,151,178]" fontSize="11" level="1" pageId="2" pageNumber="360" reason="1">
<emphasis id="9682FE17693FD61098FAFF0DFC6AFF20" bold="true" box="[151,903,151,178]" pageId="2" pageNumber="360">
<taxonomicName id="63F65986693FD61098FAFF0DFD06FF20" authority="Nakahara, Vega &amp; Willmott" authorityName="Nakahara, Vega &amp; Willmott" box="[151,747,151,178]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="360" phylum="Arthropoda" rank="species" species="favonius" status="sp. nov.">
<emphasis id="9682FE17693FD61098FAFF0DFE6FFF23" bold="true" box="[151,386,151,177]" italics="true" pageId="2" pageNumber="360">Euptychia favonius</emphasis>
Nakahara, Vega &amp; Willmott
</taxonomicName>
, 
<taxonomicNameLabel id="8DB1436C693FD6109A95FF02FC6AFF20" box="[760,903,152,178]" pageId="2" pageNumber="360" rank="species">new species</taxonomicNameLabel>
</emphasis>
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<paragraph id="A4492205693FD61098FAFF20FEE7FF41" blockId="2.[151,903,151,211]" box="[151,266,186,211]" pageId="2" pageNumber="360">
(
<figureCitation id="3CCD3E80693FD61098F2FF20FEECFF41" box="[159,257,186,211]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,250,1439,1461]" captionStartId-1="5.[151,250,1929,1951]" captionStartId-2="6.[151,250,1977,1999]" captionTargetBox-0="[175,1404,477,1408]" captionTargetBox-1="[223,1366,206,1875]" captionTargetBox-2="[163,1418,212,1938]" captionTargetId-0="figure@3.[151,1435,459,1415]" captionTargetId-1="figure@5.[212,1374,193,1888]" captionTargetId-2="figure@6.[151,1436,193,1956]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURE 1. Male and female specimens of E. favonius sp. nov. and E. meta, left dorsal and right ventral surfaces: A) E. favonius, HOLOTYPE male, western Ecuador; B) E. favonius, PARATYPE female, western Ecuador; C) E. meta, male, eastern Ecuador; D) E. meta, LECTOTYPE female, southeastern Peru." captionText-1="FIGURE 2. Genitalia of E. favonius: A) male genitalia, in lateral view; B) aedeagus, in lateral view; C) juxta, in posterior view; D) female genitalia, in posterior view; E) lamella antevaginalis, in ventral view; F) ductus bursae and corpus bursae, in dorsal view; G) signa (SN- 15 - 131; SN- 15 - 126) Scale bar = 1 mm." captionText-2="FIGURE 3. Distribution map for E. favonius." httpUri-0="https://zenodo.org/record/164787/files/figure.png" httpUri-1="https://zenodo.org/record/259867/files/figure.png" httpUri-2="https://zenodo.org/record/164789/files/figure.png" pageId="2" pageNumber="360">Figs 1–3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="ECEC718E693FD61098FAFE98FD34FE8A" box="[151,729,258,280]" pageId="2" pageNumber="360" type="description">
<paragraph id="A4492205693FD61098FAFE98FD34FE8A" blockId="2.[151,729,258,280]" box="[151,729,258,280]" pageId="2" pageNumber="360">
‘Undescribed 
<taxonomicName id="63F65986693FD6109948FE98FE66FE8A" authority="in Fratello" authorityName="in Fratello" authorityYear="2007" box="[293,395,258,280]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="360" phylum="Arthropoda" rank="genus">
<emphasis id="9682FE17693FD6109948FE98FE66FE8A" box="[293,395,258,280]" italics="true" pageId="2" pageNumber="360">Euptychia</emphasis>
</taxonomicName>
’ in 
<bibRefCitation id="C0675FF4693FD61099D8FE98FDADFE8A" author="Fratello" box="[437,576,258,280]" pageId="2" pageNumber="360" refString="Fratello, S. (2007) Eurytides and other Costa Rican Osa Peninsula surprises (Lepidoptera: Papilionoidea). Tropical Lepidoptera, 17 (2), 1 - 5." type="journal article" year="2007">Fratello 2007</bibRefCitation>
: figs. 3–6 (p.2)
</paragraph>
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<subSubSection id="ECEC718E693FD61098FAFEDFFD27FD35" pageId="2" pageNumber="360" type="diagnosis">
<paragraph id="A4492205693FD61098FAFEDFFD27FD35" blockId="2.[151,1437,325,2037]" pageId="2" pageNumber="360">
<emphasis id="9682FE17693FD61098FAFEDFFEFCFECC" bold="true" box="[151,273,325,350]" pageId="2" pageNumber="360">Diagnosis.</emphasis>
<taxonomicName id="63F65986693FD6109975FEDFFE1DFECF" box="[280,496,325,349]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="360" phylum="Arthropoda" rank="species" species="favonius">
<emphasis id="9682FE17693FD6109975FEDFFE1DFECF" box="[280,496,325,349]" italics="true" pageId="2" pageNumber="360">Euptychia favonius</emphasis>
</taxonomicName>
can be distinguished from 
<taxonomicName id="63F65986693FD6109B4FFEDFFC3DFECF" box="[802,976,325,349]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="360" phylum="Arthropoda" rank="species" species="meta">
<emphasis id="9682FE17693FD6109B4FFEDFFC3DFECF" box="[802,976,325,349]" italics="true" pageId="2" pageNumber="360">Euptychia meta</emphasis>
</taxonomicName>
by the following characters: 1) marginal lines of ventral forewing and ventral hindwing usually more undulating in 
<taxonomicName id="63F65986693FD6109BB6FEF0FBB4FE13" box="[987,1113,361,385]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="360" phylum="Arthropoda" rank="species" species="favonius">
<emphasis id="9682FE17693FD6109BB6FEF0FBB4FE13" box="[987,1113,361,385]" italics="true" pageId="2" pageNumber="360">E. favonius</emphasis>
</taxonomicName>
(less undulating in 
<taxonomicName id="63F65986693FD6109D58FEF0FA67FE13" box="[1333,1418,362,385]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="360" phylum="Arthropoda" rank="species" species="meta">
<emphasis id="9682FE17693FD6109D58FEF0FA67FE13" box="[1333,1418,362,385]" italics="true" pageId="2" pageNumber="360">E. meta</emphasis>
</taxonomicName>
); 2) outer ring of ventral concentric submarginal ocelli ringed with pale yellow and wider in 
<taxonomicName id="63F65986693FD6109CF2FE14FAF3FE37" box="[1183,1310,397,421]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="360" phylum="Arthropoda" rank="species" species="favonius">
<emphasis id="9682FE17693FD6109CF2FE14FAF3FE37" box="[1183,1310,397,421]" italics="true" pageId="2" pageNumber="360">E. favonius</emphasis>
</taxonomicName>
(ring paler yellowish white and narrower in 
<taxonomicName id="63F65986693FD6109A75FE28FD9CFE5B" box="[536,625,434,457]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="360" phylum="Arthropoda" rank="species" species="meta">
<emphasis id="9682FE17693FD6109A75FE28FD9CFE5B" box="[536,625,434,457]" italics="true" pageId="2" pageNumber="360">E. meta</emphasis>
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); 3) ventral hindwing discal and postdiscal bands relatively straighter (somewhat undulating in 
<taxonomicName id="63F65986693FD61099DFFE4CFDEAFE7F" box="[434,519,470,493]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="360" phylum="Arthropoda" rank="species" species="meta">
<emphasis id="9682FE17693FD61099DFFE4CFDEAFE7F" box="[434,519,470,493]" italics="true" pageId="2" pageNumber="360">E. meta</emphasis>
</taxonomicName>
); 4) ventral hindwing submarginal band narrower and more undulating in cells M2 and M 
<quantity id="630E8FE0693FD610988CFD91FEE9FD87" box="[225,260,508,537]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="2" pageNumber="360" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="63F65986693FD6109960FE67FE60FD86" box="[269,397,508,532]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="360" phylum="Arthropoda" rank="species" species="favonius">
<emphasis id="9682FE17693FD6109960FE67FE60FD86" box="[269,397,508,532]" italics="true" pageId="2" pageNumber="360">E. favonius</emphasis>
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(wider and less undulating in 
<taxonomicName id="63F65986693FD6109A81FE67FCAEFD86" box="[748,835,509,532]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="360" phylum="Arthropoda" rank="species" species="meta">
<emphasis id="9682FE17693FD6109A81FE67FCAEFD86" box="[748,835,509,532]" italics="true" pageId="2" pageNumber="360">E. meta</emphasis>
</taxonomicName>
); 5) lamella antevaginalis more elongated in ventral view in 
<taxonomicName id="63F65986693FD610989CFDB9FE9DFDA9" box="[241,368,547,571]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="360" phylum="Arthropoda" rank="species" species="favonius">
<emphasis id="9682FE17693FD610989CFDB9FE9DFDA9" box="[241,368,547,571]" italics="true" pageId="2" pageNumber="360">E. favonius</emphasis>
</taxonomicName>
compared to 
<taxonomicName id="63F65986693FD6109A64FDB9FDB2FDA8" box="[521,607,547,570]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="360" phylum="Arthropoda" rank="species" species="meta">
<emphasis id="9682FE17693FD6109A64FDB9FDB2FDA8" box="[521,607,547,570]" italics="true" pageId="2" pageNumber="360">E. meta</emphasis>
</taxonomicName>
; 6) spines of signa somewhat more developed in 
<taxonomicName id="63F65986693FD6109CE6FDB9FAE7FDA9" box="[1163,1290,547,571]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="360" phylum="Arthropoda" rank="species" species="favonius">
<emphasis id="9682FE17693FD6109CE6FDB9FAE7FDA9" box="[1163,1290,547,571]" italics="true" pageId="2" pageNumber="360">E. favonius</emphasis>
</taxonomicName>
compared to 
<taxonomicName id="63F65986693FD61098FAFDD2FF1DFDCD" box="[151,240,584,607]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="360" phylum="Arthropoda" rank="species" species="meta">
<emphasis id="9682FE17693FD61098FAFDD2FF1DFDCD" box="[151,240,584,607]" italics="true" pageId="2" pageNumber="360">E. meta</emphasis>
</taxonomicName>
. In addition, two other possibly consistent differences are the paler, more grayish ground color of the ventral wings and the broader and straighter dark ventral hindwing bands in 
<taxonomicName id="63F65986693FD6109C71FDF6FB73FD11" box="[1052,1182,619,643]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="360" phylum="Arthropoda" rank="species" species="favonius">
<emphasis id="9682FE17693FD6109C71FDF6FB73FD11" box="[1052,1182,619,643]" italics="true" pageId="2" pageNumber="360">E. favonius</emphasis>
</taxonomicName>
, but more material is needed to assess the reliability of these characters.
</paragraph>
</subSubSection>
<subSubSection id="ECEC718E693FD61698AAFD29FC07FB55" lastPageId="4" lastPageNumber="362" pageId="2" pageNumber="360" type="description">
<paragraph id="A4492205693FD61098AAFD29FC67FD5E" blockId="2.[151,1437,325,2037]" box="[199,906,691,716]" pageId="2" pageNumber="360">
<emphasis id="9682FE17693FD61098AAFD29FE51FD5E" bold="true" box="[199,444,691,716]" pageId="2" pageNumber="360">Description. MALE:</emphasis>
forewing length: 
<quantity id="630E8FE0693FD6109AE9FD29FD07FD59" box="[644,746,691,716]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" pageId="2" pageNumber="360" unit="mm" value="16.5">16.5 mm</quantity>
– 
<quantity id="630E8FE0693FD6109A94FD29FCA8FD59" box="[761,837,691,715]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="2" pageNumber="360" unit="mm" value="18.0">18 mm</quantity>
(n=5)
</paragraph>
<paragraph id="A4492205693FD61098AAFD4CFB7CFC36" blockId="2.[151,1437,325,2037]" pageId="2" pageNumber="360">
<emphasis id="9682FE17693FD61098AAFD4CFEFDFD7D" bold="true" box="[199,272,726,751]" pageId="2" pageNumber="360">Head:</emphasis>
Eyes with sparse brown hairs, with white scales at base; first segment of labial palpus brown, second segment length almost twice as great as eye depth, covered with short white hair-like scales and white scales laterally, as well as black scales along distal one-third of dorsal surface, ventrally adorned with black hair-like scales similar in length to segment width, third segment slightly shorter than one-fifth of second segment and covered with black scales laterally and ventrally, with creamy-white scales laterally; antennae approximately twofifths forewing length, with ca. 35 segments (n=2), distal 12–13 segments composing club.
</paragraph>
<paragraph id="A4492205693FD61098AAFC34FC8CFC7E" blockId="2.[151,1437,325,2037]" pageId="2" pageNumber="360">
<emphasis id="9682FE17693FD61098AAFC34FEC4FC55" bold="true" box="[199,297,942,967]" pageId="2" pageNumber="360">Thorax:</emphasis>
Dorsally and ventrally covered with dark brown scales and with additional long, sparse multi-colored hair-like scales dorsally; ventrally scattered with grayish scales.
</paragraph>
<paragraph id="A4492205693FD61098AAFC6CFC33FBC5" blockId="2.[151,1437,325,2037]" pageId="2" pageNumber="360">
<emphasis id="9682FE17693FD61098AAFC6CFEE5FB9D" bold="true" box="[199,264,1014,1039]" pageId="2" pageNumber="360">Legs:</emphasis>
Foreleg brown, tibia and femur almost same length, tarsus slightly shorter; midleg and hindleg with femur dorsally brown, ventrally pale buff, tibia and tarsus pale buff, terminal segments of tarsus brown, tarsus and tibia adorned with spines ventrally, tibial spurs absent at distal end of tibia.
</paragraph>
<paragraph id="A4492205693FD61098AAFBF9FC28FBEE" blockId="2.[151,1437,325,2037]" box="[199,965,1123,1148]" pageId="2" pageNumber="360">
<emphasis id="9682FE17693FD61098AAFBF9FEA8FBEE" bold="true" box="[199,325,1123,1148]" pageId="2" pageNumber="360">Abdomen:</emphasis>
Eighth tergite fully developed, similar to seventh tergite.
</paragraph>
<paragraph id="A4492205693FD61098AAFB1CFE77FABD" blockId="2.[151,1437,325,2037]" pageId="2" pageNumber="360">
<emphasis id="9682FE17693FD61098AAFB1CFE93FB0D" bold="true" box="[199,382,1158,1183]" pageId="2" pageNumber="360">Wing venation:</emphasis>
Most of forewing subcostal vein swollen; base of cubitus not swollen; forewing recurrent vein present in discal cell; hindwing humeral vein not developed. 
<emphasis id="9682FE17693FD6109B3FFB31FC32FB56" bold="true" box="[850,991,1195,1220]" pageId="2" pageNumber="360">Wing shape</emphasis>
(
<figureCitation id="3CCD3E80693FD6109B9CFB31FBDEFB51" box="[1009,1075,1195,1220]" captionStart="FIGURE 1" captionStartId="3.[151,250,1439,1461]" captionTargetBox="[175,1404,477,1408]" captionTargetId="figure@3.[151,1435,459,1415]" captionTargetPageId="3" captionText="FIGURE 1. Male and female specimens of E. favonius sp. nov. and E. meta, left dorsal and right ventral surfaces: A) E. favonius, HOLOTYPE male, western Ecuador; B) E. favonius, PARATYPE female, western Ecuador; C) E. meta, male, eastern Ecuador; D) E. meta, LECTOTYPE female, southeastern Peru." httpUri="https://zenodo.org/record/164787/files/figure.png" pageId="2" pageNumber="360">Fig. 1</figureCitation>
): Forewing subtriangular, apex rounded, costal margin convex, outer margin almost straight, inner margin straight; hindwing slightly elongate, outer margin slightly curved, inner margin slightly lobed near base and curved inward near tornus, anal lobe convex, slightly round.
</paragraph>
<paragraph id="A4492205693FD61098AAFAA1FBC2FAE5" blockId="2.[151,1437,325,2037]" pageId="2" pageNumber="360">
<emphasis id="9682FE17693FD61098AAFAA1FE6AFAC6" bold="true" box="[199,391,1339,1364]" pageId="2" pageNumber="360">Dorsal forewing</emphasis>
(
<figureCitation id="3CCD3E80693FD61099F5FAA1FE37FAC1" box="[408,474,1339,1364]" captionStart="FIGURE 1" captionStartId="3.[151,250,1439,1461]" captionTargetBox="[175,1404,477,1408]" captionTargetId="figure@3.[151,1435,459,1415]" captionTargetPageId="3" captionText="FIGURE 1. Male and female specimens of E. favonius sp. nov. and E. meta, left dorsal and right ventral surfaces: A) E. favonius, HOLOTYPE male, western Ecuador; B) E. favonius, PARATYPE female, western Ecuador; C) E. meta, male, eastern Ecuador; D) E. meta, LECTOTYPE female, southeastern Peru." httpUri="https://zenodo.org/record/164787/files/figure.png" pageId="2" pageNumber="360">Fig. 1</figureCitation>
): Light brown ground colour, darker towards apex, semi-translucent and thus subtly revealing ventral dark bands and ocelli, hair-like scales located at base of cell Cu2.
</paragraph>
<paragraph id="A4492205693FD61098AAFA1FFB21FA53" blockId="2.[151,1437,325,2037]" pageId="2" pageNumber="360">
<emphasis id="9682FE17693FD61098AAFA1FFE7FFA0C" bold="true" box="[199,402,1413,1438]" pageId="2" pageNumber="360">Dorsal hindwing</emphasis>
(
<figureCitation id="3CCD3E80693FD61099CBFA1FFE06FA0C" box="[422,491,1413,1438]" captionStart="FIGURE 1" captionStartId="3.[151,250,1439,1461]" captionTargetBox="[175,1404,477,1408]" captionTargetId="figure@3.[151,1435,459,1415]" captionTargetPageId="3" captionText="FIGURE 1. Male and female specimens of E. favonius sp. nov. and E. meta, left dorsal and right ventral surfaces: A) E. favonius, HOLOTYPE male, western Ecuador; B) E. favonius, PARATYPE female, western Ecuador; C) E. meta, male, eastern Ecuador; D) E. meta, LECTOTYPE female, southeastern Peru." httpUri="https://zenodo.org/record/164787/files/figure.png" pageId="2" pageNumber="360">Fig. 1</figureCitation>
): Ground colour similar to forewing, apex slightly darker, semitranslucent, thus subtly revealing ventral dark bands and ocelli, hair-like scales located in discal cell and cell Cu2.
</paragraph>
<paragraph id="A4492205693FD61098AAFA4AFD9FF81B" blockId="2.[151,1437,325,2037]" pageId="2" pageNumber="360">
<emphasis id="9682FE17693FD61098AAFA4AFE7DFA7B" bold="true" box="[199,400,1488,1513]" pageId="2" pageNumber="360">Ventral forewing</emphasis>
(
<figureCitation id="3CCD3E80693FD61099CDFA4AFE0DFA7B" box="[416,480,1488,1513]" captionStart="FIGURE 1" captionStartId="3.[151,250,1439,1461]" captionTargetBox="[175,1404,477,1408]" captionTargetId="figure@3.[151,1435,459,1415]" captionTargetPageId="3" captionText="FIGURE 1. Male and female specimens of E. favonius sp. nov. and E. meta, left dorsal and right ventral surfaces: A) E. favonius, HOLOTYPE male, western Ecuador; B) E. favonius, PARATYPE female, western Ecuador; C) E. meta, male, eastern Ecuador; D) E. meta, LECTOTYPE female, southeastern Peru." httpUri="https://zenodo.org/record/164787/files/figure.png" pageId="2" pageNumber="360">Fig. 1</figureCitation>
): Ground colour greyish; reddish-brown band extends posterior to swollen subcostal vein from radius to wing base; reddish-brown, straight discal band extends from radial vein, crossing discal cell in a slightly inward diagonal direction, narrower below cubital vein, and fading away before touching vein 2A; reddish-brown scales present along discocellular veins m1-m2 and m2-m3; reddish-brown postdiscal band extending from radial vein towards inner margin until reaching vein 2A, crossing wing in slightly outward direction, slightly narrowing towards posterior end; broad, faint, indistinct dark shading covering most of cells R5, M1, M2 and M3; concolorous sinuate submarginal band extending from apex towards tornus, broadening towards cell Cu1, slightly narrowing after cell Cu1; undulating marginal band, darker than previous four bands, extending from apex towards tornus; fringe brownish; ocellus in cell M1, extending across veins M1 and M2, black with one white pupil in center, tiny satellite ocellus present in cell M2, surrounded by two concentric rings, inner ring yellow, outer ring creamyyellow, wider than inner ring, small ocellus in cell M3 surrounded by two concentric rings, inner ring yellow, outer ring creamy-yellow, wider than inner ring.
</paragraph>
<paragraph id="A4492205693FD61198AAF80EFD9FFE06" blockId="2.[151,1437,325,2037]" lastBlockId="3.[151,1436,151,404]" lastPageId="3" lastPageNumber="361" pageId="2" pageNumber="360">
<emphasis id="9682FE17693FD61098AAF80EFE74F83F" bold="true" box="[199,409,1940,1965]" pageId="2" pageNumber="360">Ventral hindwing</emphasis>
(
<figureCitation id="3CCD3E80693FD61099C7F80EFE00F83E" box="[426,493,1940,1965]" captionStart="FIGURE 1" captionStartId="3.[151,250,1439,1461]" captionTargetBox="[175,1404,477,1408]" captionTargetId="figure@3.[151,1435,459,1415]" captionTargetPageId="3" captionText="FIGURE 1. Male and female specimens of E. favonius sp. nov. and E. meta, left dorsal and right ventral surfaces: A) E. favonius, HOLOTYPE male, western Ecuador; B) E. favonius, PARATYPE female, western Ecuador; C) E. meta, male, eastern Ecuador; D) E. meta, LECTOTYPE female, southeastern Peru." httpUri="https://zenodo.org/record/164787/files/figure.png" pageId="2" pageNumber="360">Fig. 1</figureCitation>
): Ground colour similar to forewing; regular reddish-brown band near wing base; discal band broader than forewing discal band, tapering posteriorly; postdiscal band parallel to discal band, concolorous, similar in width, anterior end extending distally along costa, posterior end bent basally along inner margin; broad, faint, indistinct dark shading covering around submarginal ocelli; submarginal band extending from apex towards tornus, anterior end occasionally fused with postdiscal band in cell Rs, undulating in cells M2 and M3, somewhat ‘M-shaped’, posterior end broadening and occasionally fused to postdiscal band in cell 2A; marginal line, darker, undulate, much thinner than submarginal band; fringe brownish; submarginal ocelli in cells M1, M2, M3 and Cu1 identical in appearance to forewing ocellus, ocelli in cells M2 and M3 smaller compared to those in M1 and Cu1, those ocelli in M1 and M2, M3 and Cu1 are each surrounded by two concentric rings, inner ring yellow, outer ring creamy-yellow, wider than inner ring.
</paragraph>
<caption id="F089728D693ED61198FAFA05FCFBFA60" httpUri="https://zenodo.org/record/164787/files/figure.png" pageId="3" pageNumber="361" targetBox="[175,1404,477,1408]" targetPageId="3">
<paragraph id="A4492205693ED61198FAFA05FCFBFA60" blockId="3.[151,1437,1439,1523]" pageId="3" pageNumber="361">
<emphasis id="9682FE17693ED61198FAFA05FEFBFA26" bold="true" box="[151,278,1439,1461]" pageId="3" pageNumber="361">FIGURE 1.</emphasis>
Male and female specimens of 
<taxonomicName id="63F65986693ED6119A06FA3BFD0DFA27" box="[619,736,1439,1462]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="361" phylum="Arthropoda" rank="species" species="favonius" status="sp. nov.">
<emphasis id="9682FE17693ED6119A06FA3BFD0DFA27" box="[619,736,1439,1462]" italics="true" pageId="3" pageNumber="361">E. favonius</emphasis>
</taxonomicName>
<emphasis id="9682FE17693ED6119A84FA3AFCD0FA27" bold="true" box="[745,829,1440,1461]" pageId="3" pageNumber="361">
<taxonomicNameLabel id="8DB1436C693ED6119A84FA3AFCD0FA27" box="[745,829,1440,1461]" pageId="3" pageNumber="361" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="63F65986693ED6119B19FA3BFC28FA24" box="[884,965,1441,1462]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="361" phylum="Arthropoda" rank="species" species="meta">
<emphasis id="9682FE17693ED6119B19FA3BFC28FA24" box="[884,965,1441,1462]" italics="true" pageId="3" pageNumber="361">E. meta</emphasis>
</taxonomicName>
, left dorsal and right ventral surfaces: A) 
<taxonomicName id="63F65986693ED6119DEBFA3BFF00FA46" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="361" phylum="Arthropoda" rank="species" species="favonius">
<emphasis id="9682FE17693ED6119DEBFA3BFF00FA46" italics="true" pageId="3" pageNumber="361">E. favonius</emphasis>
</taxonomicName>
, HOLOTYPE male, western Ecuador; B) 
<taxonomicName id="63F65986693ED6119AFBFA25FCE5FA46" box="[662,776,1470,1492]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="361" phylum="Arthropoda" rank="species" species="favonius">
<emphasis id="9682FE17693ED6119AFBFA25FCE5FA46" box="[662,776,1470,1492]" italics="true" pageId="3" pageNumber="361">E. favonius</emphasis>
</taxonomicName>
, PARATYPE female, western Ecuador; C) 
<taxonomicName id="63F65986693ED6119CD3FA25FAE7FA46" box="[1214,1290,1471,1492]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="361" phylum="Arthropoda" rank="species" species="meta">
<emphasis id="9682FE17693ED6119CD3FA25FAE7FA46" box="[1214,1290,1471,1492]" italics="true" pageId="3" pageNumber="361">E. meta</emphasis>
</taxonomicName>
, male, eastern Ecuador; D) 
<taxonomicName id="63F65986693ED6119977FA44FE8AFA61" box="[282,359,1502,1523]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="361" phylum="Arthropoda" rank="species" species="meta">
<emphasis id="9682FE17693ED6119977FA44FE8AFA61" box="[282,359,1502,1523]" italics="true" pageId="3" pageNumber="361">E. meta</emphasis>
</taxonomicName>
, LECTOTYPE female, southeastern Peru.
</paragraph>
</caption>
<paragraph id="A4492205693ED61198AAF9A3FE9DF804" blockId="3.[151,1437,1593,2014]" pageId="3" pageNumber="361">
<emphasis id="9682FE17693ED61198AAF9A3FED8F9C0" bold="true" box="[199,309,1593,1618]" pageId="3" pageNumber="361">Genitalia</emphasis>
(
<figureCitation id="3CCD3E80693ED611992AF9A3FE7DF9C3" box="[327,400,1593,1618]" captionStart="FIGURE 2" captionStartId="5.[151,250,1929,1951]" captionTargetBox="[223,1366,206,1875]" captionTargetId="figure@5.[212,1374,193,1888]" captionTargetPageId="5" captionText="FIGURE 2. Genitalia of E. favonius: A) male genitalia, in lateral view; B) aedeagus, in lateral view; C) juxta, in posterior view; D) female genitalia, in posterior view; E) lamella antevaginalis, in ventral view; F) ductus bursae and corpus bursae, in dorsal view; G) signa (SN- 15 - 131; SN- 15 - 126) Scale bar = 1 mm." httpUri="https://zenodo.org/record/259867/files/figure.png" pageId="3" pageNumber="361">Figs 2</figureCitation>
A–C): Tegumen somewhat subtriangular in lateral view, dorsally flattened, anterior dorsal edge present, conspicuous posterior projection above uncus present, projecting from dorsal margin of tegumen and extending posteriorly; uncus narrow, long, without setae, slightly hooked, tapering posteriorly, slightly longer than tegumen in lateral view; gnathos absent; combination of ventral arms from tegumen and dorsal arms from saccus straight; appendices angulares absent; saccus approximately half length of uncus; juxta shallow ‘U-shaped’ in posterior view; valve sparsely setose, with distal half angled upward at approximately 30º to horizontal; basal half appears roughly trapezoidal in lateral view, ventral margin convex, dorsal margin slightly concave, distal half narrow with a rounded tip, basal half somewhat rectangular with costa projecting dorsally in lateral view; aedeagus slightly curved, similar in length to valve, tubular, open anterodorsally, cornuti absent; ventral surface of anal tube weakly sclerotized.
</paragraph>
<paragraph id="A4492205693ED61198AAF83BFCE3F82B" blockId="3.[151,1437,1593,2014]" box="[199,782,1953,1978]" pageId="3" pageNumber="361">
<emphasis id="9682FE17693ED61198AAF83BFEA5F828" bold="true" box="[199,328,1953,1978]" pageId="3" pageNumber="361">FEMALE:</emphasis>
forewing length: 
<quantity id="630E8FE0693ED6119A64F83BFD83F82B" box="[521,622,1953,1978]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" pageId="3" pageNumber="361" unit="mm" value="15.5">15.5 mm</quantity>
– 
<quantity id="630E8FE0693ED6119A10F83BFD24F82B" box="[637,713,1953,1977]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="361" unit="mm" value="18.0">18 mm</quantity>
(n=4)
</paragraph>
<paragraph id="A4492205693ED61698AAF85FFA71FF65" blockId="3.[151,1437,1593,2014]" lastBlockId="4.[151,1437,151,2015]" lastPageId="4" lastPageNumber="362" pageId="3" pageNumber="361">Similar to male except as follows: female foretarsus divided into five segments; forewing more rounded and broader, outer margin convex; ground colour of both wing surfaces white, apical half of dorsal forewing black, dark ventral bands and lines appearing as darkly scaled rather than just visible through wing; dorsal hindwing with dark marginal scaling stronger, submarginal and marginal lines visible to tornus, in addition to ocelli in cells M1 and Cu1.</paragraph>
<paragraph id="A44922056939D61698AAFE9CFD59FE3D" blockId="4.[151,1437,151,2015]" pageId="4" pageNumber="362">
<emphasis id="9682FE176939D61698AAFE9CFE75FE8D" bold="true" box="[199,408,262,287]" pageId="4" pageNumber="362">Female Genitalia</emphasis>
(
<figureCitation id="3CCD3E806939D61699C0FE9CFE14FE8D" box="[429,505,262,287]" captionStart="FIGURE 2" captionStartId="5.[151,250,1929,1951]" captionTargetBox="[223,1366,206,1875]" captionTargetId="figure@5.[212,1374,193,1888]" captionTargetPageId="5" captionText="FIGURE 2. Genitalia of E. favonius: A) male genitalia, in lateral view; B) aedeagus, in lateral view; C) juxta, in posterior view; D) female genitalia, in posterior view; E) lamella antevaginalis, in ventral view; F) ductus bursae and corpus bursae, in dorsal view; G) signa (SN- 15 - 131; SN- 15 - 126) Scale bar = 1 mm." httpUri="https://zenodo.org/record/259867/files/figure.png" pageId="4" pageNumber="362">Figs 2</figureCitation>
D–G): Lamella antevaginalis sclerotized, somewhat trapezoidal with very deep concavity at median region of posterior margin in ventral view, anterior portion appears darker in coloration than remainder of lamella antevaginalis; basal side of 8th abdominal segment sclerotized; ductus bursae membranous, origin of ductus seminalis close to ostium bursae; corpus bursae roughly oval in dorsal view, with two short signa, spines of signa developed and somewhat robust.
</paragraph>
<paragraph id="A44922056939D61698AAFE20FF14FD89" blockId="4.[151,1437,151,2015]" pageId="4" pageNumber="362">
<materialsCitation id="149E28586939D61698AAFE20FF14FD89" ID-GBIF-Occurrence="1324639628" collectingDate="1994-06-01" collectionCode="MGCL, MECN" collectorName="La Punta &amp; K. Willmott &amp; J. Hall &amp; Genitalic" country="Ecuador" county="Types" elevation="300" latitude="1.0652777" location="La Punta" longLatPrecision="925" longitude="-78.65" municipality="Lita-San Lorenzo rd." pageId="4" pageNumber="362" specimenCount="2" stateProvince="Esmeraldas" typeStatus="holotype">
<emphasis id="9682FE176939D61698AAFE20FD8FFE41" bold="true" box="[199,610,442,467]" pageId="4" pageNumber="362">
<typeStatus id="7B4D9CA76939D61698AAFE20FEFCFE41" box="[199,273,442,467]" pageId="4" pageNumber="362">
<collectingCounty id="4D285A896939D61698AAFE20FEFCFE41" box="[199,273,442,467]" pageId="4" pageNumber="362">Types</collectingCounty>
</typeStatus>
: 
<typeStatus id="7B4D9CA76939D6169971FE20FE52FE41" box="[284,447,442,467]" pageId="4" pageNumber="362" type="holotype">HOLOTYPE</typeStatus>
: 
<collectingCountry id="DCE162956939D61699A6FE20FDB0FE41" box="[459,605,442,467]" name="Ecuador" pageId="4" pageNumber="362">ECUADOR</collectingCountry>
:
</emphasis>
<emphasis id="9682FE176939D6169A05FE21FD01FE41" box="[616,748,443,467]" italics="true" pageId="4" pageNumber="362">
<collectingRegion id="6632ECE76939D6169A05FE21FD01FE41" box="[616,748,443,467]" country="Ecuador" name="Esmeraldas" pageId="4" pageNumber="362">Esmeraldas</collectingRegion>
</emphasis>
; km 44 
<collectingMunicipality id="442DB87F6939D6169B25FE20FBD9FE41" box="[840,1076,442,467]" pageId="4" pageNumber="362">Lita-San Lorenzo rd.</collectingMunicipality>
, 
<location id="A12974DE6939D6169C52FE21FB4BFE40" LSID="urn:lsid:plazi:treatment:2C5F9313693FD61598FAFF0DFE2AFEF7:A12974DE6939D6169C52FE21FB4BFE40" box="[1087,1190,443,466]" country="Ecuador" county="Types" latitude="1.0652777" longLatPrecision="925" longitude="-78.65" municipality="Lita-San Lorenzo rd." name="La Punta" pageId="4" pageNumber="362" stateProvince="Esmeraldas">
<collectorName id="090347D36939D6169C52FE21FB4BFE40" box="[1087,1190,443,466]" pageId="4" pageNumber="362">La Punta</collectorName>
</location>
, [
<geoCoordinate id="C1C244C26939D6169CD5FE20FACDFE41" box="[1208,1312,442,467]" direction="north" orientation="latitude" pageId="4" pageNumber="362" precision="15" value="1.0652777">1º3'55''N</geoCoordinate>
, 
<geoCoordinate id="C1C244C26939D6169D41FE21FA7CFE41" box="[1324,1425,443,467]" direction="west" orientation="longitude" pageId="4" pageNumber="362" precision="925" value="-78.65">78º39'W</geoCoordinate>
], 
<quantity id="630E8FE06939D61698FAFE45FF34FE65" box="[151,217,479,503]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="362" unit="m" value="300.0">
<elevation id="2FDBC5366939D61698FAFE45FF34FE65" box="[151,217,479,503]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="362" unit="m" value="300.0">300m</elevation>
</quantity>
, (
<collectorName id="090347D36939D616989FFE45FE90FE65" box="[242,381,478,503]" pageId="4" pageNumber="362">K. Willmott</collectorName>
&amp; 
<collectorName id="090347D36939D61699C7FE7AFE11FE65" box="[426,508,478,503]" pageId="4" pageNumber="362">J. Hall</collectorName>
), 21–22 
<date id="D04804C56939D6169A0BFE7AFD17FE65" box="[614,762,479,503]" pageId="4" pageNumber="362" value="1994-06-01">
<collectingDate id="C00CFD2D6939D6169A0BFE7AFD17FE65" box="[614,762,479,503]" pageId="4" pageNumber="362" value="1994-06-01">June 1994, 1</collectingDate>
</date>
♂, 
<collectorName id="090347D36939D6169B48FE44FC66FE65" box="[805,907,478,503]" pageId="4" pageNumber="362">Genitalic</collectorName>
vial SN-15-131, (
<collectionCode id="C2E7BAC06939D6169C0EFE45FB55FE65" box="[1123,1208,479,503]" pageId="4" pageNumber="362">MGCL</collectionCode>
, to be deposited in 
<collectionCode id="C2E7BAC06939D61698FAFD99FF00FD89" box="[151,237,515,539]" country="Ecuador" httpUri="http://grbio.org/cool/rm44-by7m" name="Museo Ecuadoriano de Ciencias Naturales" pageId="4" pageNumber="362">MECN</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="A44922056939D61698AAFDBCFE24FBED" blockId="4.[151,1437,151,2015]" pageId="4" pageNumber="362">
<materialsCitation id="149E28586939D61698AAFDBCFF13FDF1" ID-GBIF-Occurrence="1324639625" collectingDate="1975-04-01" collectingDateMax="61-09-19" collectingDateMin="1975-04-01" collectionCode="MGCL" collectorName="Finca La Selva" country="Costa Rica" location="Finca La Selva" pageId="4" pageNumber="362" specimenCount="1" stateProvince="Heredia" typeStatus="paratype">
<emphasis id="9682FE176939D61698AAFDBCFDD3FDAD" bold="true" box="[199,574,550,575]" pageId="4" pageNumber="362">
<typeStatus id="7B4D9CA76939D61698AAFDBCFE82FDAD" box="[199,367,550,575]" pageId="4" pageNumber="362" type="paratype">PARATYPES</typeStatus>
: 
<collectingCountry id="DCE162956939D61699ECFDBCFDD4FDAD" box="[385,569,550,575]" name="Costa Rica" pageId="4" pageNumber="362">COSTA RICA</collectingCountry>
:
</emphasis>
<emphasis id="9682FE176939D6169A26FDBDFD4BFDAD" box="[587,678,551,575]" italics="true" pageId="4" pageNumber="362">
<collectingRegion id="6632ECE76939D6169A26FDBDFD4BFDAD" box="[587,678,551,575]" country="Costa Rica" name="Heredia" pageId="4" pageNumber="362">Heredia</collectingRegion>
</emphasis>
; 
<collectorName id="090347D36939D6169AD6FDBDFC9FFDAD" box="[699,882,551,575]" pageId="4" pageNumber="362">
<location id="A12974DE6939D6169AD6FDBDFC9FFDAD" LSID="urn:lsid:plazi:treatment:2C5F9313693FD61598FAFF0DFE2AFEF7:A12974DE6939D6169AD6FDBDFC9FFDAD" box="[699,882,551,575]" country="Costa Rica" name="Finca La Selva" pageId="4" pageNumber="362" stateProvince="Heredia">Finca La Selva</location>
</collectorName>
, 12 
<collectingDate id="C00CFD2D6939D6169BC0FDBDFBADFDAD" box="[941,1088,551,575]" pageId="4" pageNumber="362" value="1975-04-01">
<date id="D04804C56939D6169BC0FDBDFBADFDAD" box="[941,1088,551,575]" pageId="4" pageNumber="362" value="1975-04-01">Apr 1975, 1</date>
</collectingDate>
♀, [FLMNH-MGCL- 
<date id="D04804C56939D6169D52FDBDFA7FFDAD" box="[1343,1426,551,575]" pageId="4" pageNumber="362" value="61-09-19">
<collectingDate id="C00CFD2D6939D6169D52FDBDFA7FFDAD" box="[1343,1426,551,575]" pageId="4" pageNumber="362" value="61-09-19">190961</collectingDate>
</date>
], (
<collectionCode id="C2E7BAC06939D61698F2FDD1FF1AFDF1" box="[159,247,587,611]" pageId="4" pageNumber="362">MGCL</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="149E28586939D6169967FDD1FE29FD15" ID-GBIF-Occurrence="1324639634" collectingDate="60-09-19" collectionCode="MGCL" collectorName="Fratello, S. &amp; Genitalic" country="Costa Rica" elevation="700" location="Cerro Brujo" municipality="Osa Peninsula" pageId="4" pageNumber="362" specimenCount="1" specimenCount-female="1" stateProvince="Puntarenas" typeStatus="paratype">
<emphasis id="9682FE176939D6169967FDD1FE67FDF0" box="[266,394,587,610]" italics="true" pageId="4" pageNumber="362">
<collectingRegion id="6632ECE76939D6169967FDD1FE67FDF0" box="[266,394,587,610]" country="Costa Rica" name="Puntarenas" pageId="4" pageNumber="362">Puntarenas</collectingRegion>
</emphasis>
; 
<collectingMunicipality id="442DB87F6939D61699F4FDD1FDD0FDF1" box="[409,573,587,611]" pageId="4" pageNumber="362">Osa Peninsula</collectingMunicipality>
, 
<location id="A12974DE6939D6169A25FDD1FD3EFDF1" LSID="urn:lsid:plazi:treatment:2C5F9313693FD61598FAFF0DFE2AFEF7:A12974DE6939D6169A25FDD1FD3EFDF1" box="[584,723,587,611]" country="Costa Rica" municipality="Osa Peninsula" name="Cerro Brujo" pageId="4" pageNumber="362" stateProvince="Puntarenas">Cerro Brujo</location>
, 
<quantity id="630E8FE06939D6169AB3FDD1FCCCFDF1" box="[734,801,587,611]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="362" unit="m" value="700.0">
<elevation id="2FDBC5366939D6169AB3FDD1FCCCFDF1" box="[734,801,587,611]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="362" unit="m" value="700.0">700m</elevation>
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, (
<collectorName id="090347D36939D6169B59FDD1FC5FFDF1" box="[820,946,587,611]" pageId="4" pageNumber="362">Fratello, S.</collectorName>
), 
<specimenCount id="B2F0E98C6939D6169BA7FDD1FC00FDF1" box="[970,1005,587,611]" pageId="4" pageNumber="362" type="female">1♀</specimenCount>
, 
<collectorName id="090347D36939D6169B91FDD1FB8FFDF1" box="[1020,1122,587,611]" pageId="4" pageNumber="362">Genitalic</collectorName>
vial SN-15-126, [FLMNH- MGCL- 
<date id="D04804C56939D6169898FDF4FEA6FD15" box="[245,331,622,647]" pageId="4" pageNumber="362" value="60-09-19">
<collectingDate id="C00CFD2D6939D6169898FDF4FEA6FD15" box="[245,331,622,647]" pageId="4" pageNumber="362" value="60-09-19">190960</collectingDate>
</date>
], (
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)
</materialsCitation>
. 
<emphasis id="9682FE176939D61699A2FDF4FDBFFD15" bold="true" box="[463,594,622,647]" pageId="4" pageNumber="362">PANAMA:</emphasis>
<materialsCitation id="149E28586939D6169A37FDF5FE97FD39" ID-GBIF-Occurrence="1324639631" collectingDate="1977-04-01" collectionCode="USNM" collectorName="Altos de Pacora &amp; Small, G. B." country="Panama" elevation="2000" latitude="9.233334" location="Altos de Pacora" longLatPrecision="1300" longitude="-79.333336" pageId="4" pageNumber="362" specimenCount="1" typeStatus="paratype">
<emphasis id="9682FE176939D6169A37FDF5FD5AFD14" box="[602,695,623,646]" italics="true" pageId="4" pageNumber="362">
<collectingCountry id="DCE162956939D6169A37FDF5FD5AFD14" box="[602,695,623,646]" name="Panama" pageId="4" pageNumber="362">Panamá</collectingCountry>
</emphasis>
; 
<location id="A12974DE6939D6169AAAFDF4FC91FD15" LSID="urn:lsid:plazi:treatment:2C5F9313693FD61598FAFF0DFE2AFEF7:A12974DE6939D6169AAAFDF4FC91FD15" box="[711,892,622,647]" country="Panama" latitude="9.233334" longLatPrecision="1300" longitude="-79.333336" name="Altos de Pacora" pageId="4" pageNumber="362">
<collectorName id="090347D36939D6169AAAFDF4FC91FD15" box="[711,892,622,647]" pageId="4" pageNumber="362">Altos de Pacora</collectorName>
</location>
, [
<geoCoordinate id="C1C244C26939D6169BE2FDF5FC31FD15" box="[911,988,623,647]" direction="north" orientation="latitude" pageId="4" pageNumber="362" precision="925" value="9.233334">9º14'N</geoCoordinate>
, 
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], 
<quantity id="630E8FE06939D6169C31FDF5FB40FD15" box="[1116,1197,623,647]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="362" unit="m" value="2000.0">
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, (
<collectorName id="090347D36939D6169CADFDF4FAA8FD15" box="[1216,1349,622,647]" pageId="4" pageNumber="362">Small, G.B.</collectorName>
), 5 
<date id="D04804C56939D6169D1DFDF5FF07FD39" pageId="4" pageNumber="362" value="1977-04-01">
<collectingDate id="C00CFD2D6939D6169D1DFDF5FF07FD39" pageId="4" pageNumber="362" value="1977-04-01">Apr 1977, 1</collectingDate>
</date>
♂, (
<collectionCode id="C2E7BAC06939D6169976FD09FE9FFD39" box="[283,370,659,683]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="362">USNM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="149E28586939D61699E5FD09FB19FD39" ID-GBIF-Occurrence="1324639629" box="[392,1268,659,683]" collectingDate="1977-04-01" collectionCode="USNM" collectorName="Small, G. B." country="Panama" elevation="900" latitude="9.233334" location="Cerro Jefe" longLatPrecision="1300" longitude="-79.36667" pageId="4" pageNumber="362" specimenCount="1" typeStatus="paratype">
<location id="A12974DE6939D61699E5FD09FDECFD39" LSID="urn:lsid:plazi:treatment:2C5F9313693FD61598FAFF0DFE2AFEF7:A12974DE6939D61699E5FD09FDECFD39" box="[392,513,659,683]" country="Panama" latitude="9.233334" longLatPrecision="1300" longitude="-79.36667" name="Cerro Jefe" pageId="4" pageNumber="362">Cerro Jefe</location>
, [
<geoCoordinate id="C1C244C26939D6169A78FD09FD8EFD39" box="[533,611,659,683]" direction="north" orientation="latitude" pageId="4" pageNumber="362" precision="925" value="9.233334">9º14'N</geoCoordinate>
, 
<geoCoordinate id="C1C244C26939D6169A0AFD09FD22FD39" box="[615,719,659,683]" direction="west" orientation="longitude" pageId="4" pageNumber="362" precision="925" value="-79.36667">79º22'W</geoCoordinate>
], 
<quantity id="630E8FE06939D6169A88FD09FCC5FD39" box="[741,808,659,683]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="362" unit="m" value="900.0">
<elevation id="2FDBC5366939D6169A88FD09FCC5FD39" box="[741,808,659,683]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="362" unit="m" value="900.0">900m</elevation>
</quantity>
, (
<collectorName id="090347D36939D6169B50FD09FC28FD39" box="[829,965,659,683]" pageId="4" pageNumber="362">Small, G.B.</collectorName>
), 
<date id="D04804C56939D6169BB1FD09FB8AFD39" box="[988,1127,659,683]" pageId="4" pageNumber="362" value="1977-04-01">
<collectingDate id="C00CFD2D6939D6169BB1FD09FB8AFD39" box="[988,1127,659,683]" pageId="4" pageNumber="362" value="1977-04-01">Apr 1977, 1</collectingDate>
</date>
♀, (
<collectionCode id="C2E7BAC06939D6169CF8FD09FB01FD39" box="[1173,1260,659,683]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="362">USNM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="149E28586939D6169D6FFD09FBD7FD5D" ID-GBIF-Occurrence="1324639637" collectingDate="1978-03-01" collectionCode="USNM" collectorName="Altos de Pacora &amp; Small, G. B." country="Panama" elevation="750" latitude="9.233334" location="Altos de Pacora" longLatPrecision="1300" longitude="-79.333336" pageId="4" pageNumber="362" specimenCount="1" typeStatus="paratype">
near 
<collectorName id="090347D36939D6169D56FD09FF0EFD5D" pageId="4" pageNumber="362">
<location id="A12974DE6939D6169D56FD09FF0EFD5D" LSID="urn:lsid:plazi:treatment:2C5F9313693FD61598FAFF0DFE2AFEF7:A12974DE6939D6169D56FD09FF0EFD5D" country="Panama" latitude="9.233334" longLatPrecision="1300" longitude="-79.333336" name="Altos de Pacora" pageId="4" pageNumber="362">Altos de Pacora</location>
</collectorName>
, [
<geoCoordinate id="C1C244C26939D6169968FD2DFEBEFD5D" box="[261,339,695,719]" direction="north" orientation="latitude" pageId="4" pageNumber="362" precision="925" value="9.233334">9º14'N</geoCoordinate>
, 
<geoCoordinate id="C1C244C26939D616993AFD2DFE52FD5D" box="[343,447,695,719]" direction="west" orientation="longitude" pageId="4" pageNumber="362" precision="925" value="-79.333336">79º20'W</geoCoordinate>
], 
<quantity id="630E8FE06939D61699B8FD2CFDF5FD5D" box="[469,536,694,719]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="4" pageNumber="362" unit="m" value="750.0">
<elevation id="2FDBC5366939D61699B8FD2CFDF5FD5D" box="[469,536,694,719]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="4" pageNumber="362" unit="m" value="750.0">750m</elevation>
</quantity>
, (
<collectorName id="090347D36939D6169A43FD2CFD58FD5D" box="[558,693,694,719]" pageId="4" pageNumber="362">Small, G.B.</collectorName>
), 15–19 
<date id="D04804C56939D6169B70FD22FC40FD5D" box="[797,941,695,719]" pageId="4" pageNumber="362" value="1978-03-01">
<collectingDate id="C00CFD2D6939D6169B70FD22FC40FD5D" box="[797,941,695,719]" pageId="4" pageNumber="362" value="1978-03-01">Mar 1978, 1</collectingDate>
</date>
♀, (
<collectionCode id="C2E7BAC06939D6169BB6FD2DFBDFFD5D" box="[987,1074,695,719]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="362">USNM</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="149E28586939D6169C2AFD2CFD16FD61" ID-GBIF-Occurrence="1324639632" collectingDate="1945-12-01" collectionCode="AMNH" collectorName="La Borrascosa" country="Colombia" elevation="850" location="La Borrascosa" pageId="4" pageNumber="362" specimenCount="1" stateProvince="Santander" typeStatus="paratype">
<emphasis id="9682FE176939D6169C2AFD2CFB02FD5D" bold="true" box="[1095,1263,694,719]" pageId="4" pageNumber="362">
<collectingCountry id="DCE162956939D6169C2AFD2CFB07FD5D" box="[1095,1258,694,719]" name="Colombia" pageId="4" pageNumber="362">COLOMBIA</collectingCountry>
:
</emphasis>
<emphasis id="9682FE176939D6169C94FD2DFA81FD5D" box="[1273,1388,695,719]" italics="true" pageId="4" pageNumber="362">
<collectingRegion id="6632ECE76939D6169C94FD2DFA81FD5D" box="[1273,1388,695,719]" country="Colombia" name="Santander" pageId="4" pageNumber="362">Santander</collectingRegion>
</emphasis>
; 
<location id="A12974DE6939D6169D13FD22FEFBFD60" LSID="urn:lsid:plazi:treatment:2C5F9313693FD61598FAFF0DFE2AFEF7:A12974DE6939D6169D13FD22FEFBFD60" country="Colombia" name="La Borrascosa" pageId="4" pageNumber="362" stateProvince="Santander">
<collectorName id="090347D36939D6169D13FD22FEFBFD60" pageId="4" pageNumber="362">La Borrascosa</collectorName>
</location>
, 
<quantity id="630E8FE06939D6169949FD40FE45FD60" box="[292,424,730,754]" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="12.0" metricValueMin="5.0" pageId="4" pageNumber="362" unit="m" value="850.0" valueMax="1200.0" valueMin="500.0">
<elevation id="2FDBC5366939D6169949FD40FE45FD60" box="[292,424,730,754]" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="12.0" metricValueMin="5.0" pageId="4" pageNumber="362" unit="m" value="850.0" valueMax="1200.0" valueMin="500.0">500-1200m</elevation>
</quantity>
, 13 
<date id="D04804C56939D61699B1FD41FD8BFD61" box="[476,614,730,755]" pageId="4" pageNumber="362" value="1945-12-01">
<collectingDate id="C00CFD2D6939D61699B1FD41FD8BFD61" box="[476,614,730,755]" pageId="4" pageNumber="362" value="1945-12-01">Dec 1945, 1</collectingDate>
</date>
♂, (
<collectionCode id="C2E7BAC06939D6169AFAFD41FD1EFD61" box="[663,755,731,755]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="4" pageNumber="362">AMNH</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="149E28586939D6169B64FD41FEDEFC85" ID-GBIF-Occurrence="1324639624" collectingDate="1945-12-01" collectionCode="AMNH" collectorName="La Lechera" country="Colombia" elevation="2000" latitude="7.1244445" location="La Lechera" longLatPrecision="21" longitude="-73.309166" pageId="4" pageNumber="362" specimenCount="1" stateProvince="Santander" typeStatus="paratype">
<collectorName id="090347D36939D6169B64FD41FC60FD61" box="[777,909,731,755]" pageId="4" pageNumber="362">
<location id="A12974DE6939D6169B64FD41FC60FD61" LSID="urn:lsid:plazi:treatment:2C5F9313693FD61598FAFF0DFE2AFEF7:A12974DE6939D6169B64FD41FC60FD61" box="[777,909,731,755]" country="Colombia" latitude="7.1244445" longLatPrecision="21" longitude="-73.309166" name="La Lechera" pageId="4" pageNumber="362" stateProvince="Santander">La Lechera</location>
</collectorName>
, [
<geoCoordinate id="C1C244C26939D6169BCFFD41FBE7FD61" box="[930,1034,731,755]" direction="north" orientation="latitude" pageId="4" pageNumber="362" precision="15" value="7.1244445">7º7'28''N</geoCoordinate>
, 
<geoCoordinate id="C1C244C26939D6169C63FD41FB76FD61" box="[1038,1179,731,755]" direction="west" orientation="longitude" pageId="4" pageNumber="362" precision="15" value="-73.309166">73º18'33''W</geoCoordinate>
], 
<quantity id="630E8FE06939D6169CDDFD41FAECFD61" box="[1200,1281,731,755]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="362" unit="m" value="2000.0">
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, 1 
<date id="D04804C56939D6169D4BFD41FF4FFC85" pageId="4" pageNumber="362" value="1945-12-01">
<collectingDate id="C00CFD2D6939D6169D4BFD41FF4FFC85" pageId="4" pageNumber="362" value="1945-12-01">Dec 1945, 1</collectingDate>
</date>
♂, (
<collectionCode id="C2E7BAC06939D61698A2FD65FEC6FC85" box="[207,299,767,791]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="4" pageNumber="362">AMNH</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="149E28586939D6169950FD65FC4CFC85" ID-GBIF-Occurrence="1324639630" box="[317,929,767,791]" collectionCode="ZMHU" country="Colombia" latitude="5.2166667" location="Rio San Juan" longLatPrecision="1305" longitude="-76.65" pageId="4" pageNumber="362" specimenCount="4" specimenCount-male="4" stateProvince="Choco" typeStatus="paratype">
<emphasis id="9682FE176939D6169950FD65FE6BFC85" box="[317,390,767,791]" italics="true" pageId="4" pageNumber="362">
<collectingRegion id="6632ECE76939D6169950FD65FE6BFC85" box="[317,390,767,791]" country="Panama" name="Embera-Wounaan" pageId="4" pageNumber="362">Chocó</collectingRegion>
</emphasis>
; 
<location id="A12974DE6939D61699F9FD65FDC7FC84" LSID="urn:lsid:plazi:treatment:2C5F9313693FD61598FAFF0DFE2AFEF7:A12974DE6939D61699F9FD65FDC7FC84" box="[404,554,767,791]" country="Colombia" latitude="5.2166667" longLatPrecision="1305" longitude="-76.65" name="Rio San Juan" pageId="4" pageNumber="362" stateProvince="Choco">Río San Juan</location>
, [
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, 
<geoCoordinate id="C1C244C26939D6169AE1FD65FD1BFC85" box="[652,758,767,791]" direction="west" orientation="longitude" pageId="4" pageNumber="362" precision="925" value="-76.65">76º39'W</geoCoordinate>
], 
<specimenCount id="B2F0E98C6939D6169B6AFD65FCC1FC85" box="[775,812,767,791]" pageId="4" pageNumber="362" type="male">4♂</specimenCount>
, (
<collectionCode id="C2E7BAC06939D6169B2DFD65FC74FC85" box="[832,921,767,791]" country="Germany" httpUri="http://grbio.org/cool/99a5-d0z2" name="Zoologisches Museum der Humboldt Universitaet" pageId="4" pageNumber="362">ZMHU</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="149E28586939D6169BC7FD64FA75FCA9" ID-GBIF-Occurrence="1324639635" collectingDate="2003-10-01" collectionCode="MGCL" collectorName="K. Willmott" country="Ecuador" elevation="875" latitude="0.8808333" location="Rio Chuchuvi" longLatPrecision="21" longitude="-78.515" municipality="Lita-San Lorenzo rd." pageId="4" pageNumber="362" specimenCount="1" stateProvince="Esmeraldas" typeStatus="paratype">
<emphasis id="9682FE176939D6169BC7FD64FBACFC85" bold="true" box="[938,1089,766,791]" pageId="4" pageNumber="362">
<collectingCountry id="DCE162956939D6169BC7FD64FBD1FC85" box="[938,1084,766,791]" name="Ecuador" pageId="4" pageNumber="362">ECUADOR</collectingCountry>
:
</emphasis>
<emphasis id="9682FE176939D6169C25FD65FB21FC85" box="[1096,1228,767,791]" italics="true" pageId="4" pageNumber="362">
<collectingRegion id="6632ECE76939D6169C25FD65FB21FC85" box="[1096,1228,767,791]" country="Ecuador" name="Esmeraldas" pageId="4" pageNumber="362">Esmeraldas</collectingRegion>
</emphasis>
; km 12.5 
<collectingMunicipality id="442DB87F6939D6169D51FD64FEF3FCA9" pageId="4" pageNumber="362">Lita-San Lorenzo rd.</collectingMunicipality>
, 
<location id="A12974DE6939D6169946FCB9FE20FCA9" LSID="urn:lsid:plazi:treatment:2C5F9313693FD61598FAFF0DFE2AFEF7:A12974DE6939D6169946FCB9FE20FCA9" box="[299,461,802,827]" country="Ecuador" latitude="0.8808333" longLatPrecision="21" longitude="-78.515" municipality="Lita-San Lorenzo rd." name="Rio Chuchuvi" pageId="4" pageNumber="362" stateProvince="Esmeraldas">Río Chuchuví</location>
, [
<geoCoordinate id="C1C244C26939D616998DFCB8FDB8FCA9" box="[480,597,802,827]" direction="north" orientation="latitude" pageId="4" pageNumber="362" precision="15" value="0.8808333">0º52'51''N</geoCoordinate>
, 
<geoCoordinate id="C1C244C26939D6169A37FCB8FD0AFCA9" box="[602,743,802,827]" direction="west" orientation="longitude" pageId="4" pageNumber="362" precision="15" value="-78.515">78º30'54''W</geoCoordinate>
], 
<quantity id="630E8FE06939D6169A96FCB9FC9BFCA8" box="[763,886,802,827]" metricMagnitude="2" metricUnit="m" metricValue="8.75" metricValueMax="9.5" metricValueMin="8.0" pageId="4" pageNumber="362" unit="m" value="875.0" valueMax="950.0" valueMin="800.0">
<elevation id="2FDBC5366939D6169A96FCB9FC9BFCA8" box="[763,886,802,827]" metricMagnitude="2" metricUnit="m" metricValue="8.75" metricValueMax="9.5" metricValueMin="8.0" pageId="4" pageNumber="362" unit="m" value="875.0" valueMax="950.0" valueMin="800.0">800–950m</elevation>
</quantity>
, (
<collectorName id="090347D36939D6169BE1FCB9FBF4FCA9" box="[908,1049,803,827]" pageId="4" pageNumber="362">K. Willmott</collectorName>
), 29 
<date id="D04804C56939D6169C3EFCB9FAE1FCA9" box="[1107,1292,803,827]" pageId="4" pageNumber="362" value="2003-10-01">
<collectingDate id="C00CFD2D6939D6169C3EFCB9FAE1FCA9" box="[1107,1292,803,827]" pageId="4" pageNumber="362" value="2003-10-01">October 2003, 1</collectingDate>
</date>
♀, (
<collectionCode id="C2E7BAC06939D6169D54FCB9FA7DFCA9" box="[1337,1424,803,827]" pageId="4" pageNumber="362">MGCL</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="149E28586939D61698FAFCDDFAF7FCCD" ID-GBIF-Occurrence="1324639633" box="[151,1306,839,863]" collectingDate="2000-05-01" collectionCode="MGCL" collectorName="K. Willmott" country="Ecuador" elevation="800" latitude="0.8977778" location="Lita-San Lorenzo rd." longLatPrecision="21" longitude="-78.52473" pageId="4" pageNumber="362" specimenCount="1" stateProvince="Esmeraldas" typeStatus="paratype">
km 15 
<location id="A12974DE6939D6169889FCDDFE22FCCD" LSID="urn:lsid:plazi:treatment:2C5F9313693FD61598FAFF0DFE2AFEF7:A12974DE6939D6169889FCDDFE22FCCD" box="[228,463,839,863]" country="Ecuador" latitude="0.8977778" longLatPrecision="21" longitude="-78.52473" name="Lita-San Lorenzo rd." pageId="4" pageNumber="362" stateProvince="Esmeraldas">Lita-San Lorenzo rd.</location>
, [
<geoCoordinate id="C1C244C26939D616998FFCDDFDB4FCCD" box="[482,601,839,863]" direction="north" orientation="latitude" pageId="4" pageNumber="362" precision="15" value="0.8977778">0º53'52''N</geoCoordinate>
, 
<geoCoordinate id="C1C244C26939D6169A30FCDDFD05FCCD" box="[605,744,839,863]" direction="west" orientation="longitude" pageId="4" pageNumber="362" precision="15" value="-78.52473">78º31'29''W</geoCoordinate>
], 
<quantity id="630E8FE06939D6169A97FCDDFCD0FCCD" box="[762,829,839,863]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="362" unit="m" value="800.0">
<elevation id="2FDBC5366939D6169A97FCDDFCD0FCCD" box="[762,829,839,863]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="362" unit="m" value="800.0">800m</elevation>
</quantity>
, (
<collectorName id="090347D36939D6169B3CFCDDFC31FCCD" box="[849,988,839,863]" pageId="4" pageNumber="362">K. Willmott</collectorName>
), 6 
<collectingDate id="C00CFD2D6939D6169C6EFCDDFB7DFCCD" box="[1027,1168,839,863]" pageId="4" pageNumber="362" value="2000-05-01">
<date id="D04804C56939D6169C6EFCDDFB7DFCCD" box="[1027,1168,839,863]" pageId="4" pageNumber="362" value="2000-05-01">May 2000, 1</date>
</collectingDate>
♀, (
<collectionCode id="C2E7BAC06939D6169CD6FCDDFAFFFCCD" box="[1211,1298,839,863]" pageId="4" pageNumber="362">MGCL</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="149E28586939D6169D48FCDDFAAFFC35" ID-GBIF-Occurrence="1324639626" collectingDate="2015-07-01" collectionCode="MGCL" collectorName="K. R., J. C. R, J. I. R. Willmott" country="Ecuador" elevation="225" latitude="1.1072222" location="San Francisco" longLatPrecision="21" longitude="-78.69861" municipality="San Lorenzo-Ibarra rd." pageId="4" pageNumber="362" specimenCount="1" stateProvince="Esmeraldas" typeStatus="paratype">
km 17 
<collectingMunicipality id="442DB87F6939D6169D1FFCDDFE84FC11" pageId="4" pageNumber="362">San Lorenzo-Ibarra rd.</collectingMunicipality>
, 
<location id="A12974DE6939D6169918FCF1FDF9FC11" LSID="urn:lsid:plazi:treatment:2C5F9313693FD61598FAFF0DFE2AFEF7:A12974DE6939D6169918FCF1FDF9FC11" box="[373,532,874,899]" country="Ecuador" latitude="1.1072222" longLatPrecision="21" longitude="-78.69861" municipality="San Lorenzo-Ibarra rd." name="San Francisco" pageId="4" pageNumber="362" stateProvince="Esmeraldas">San Francisco</location>
ridge, [
<geoCoordinate id="C1C244C26939D6169A06FCF0FD3DFC11" box="[619,720,874,899]" direction="north" orientation="latitude" pageId="4" pageNumber="362" precision="15" value="1.1072222">1º6'26''N</geoCoordinate>
, 
<geoCoordinate id="C1C244C26939D6169AB9FCF0FC89FC11" box="[724,868,874,899]" direction="west" orientation="longitude" pageId="4" pageNumber="362" precision="15" value="-78.69861">78º41'55''W</geoCoordinate>
], 
<quantity id="630E8FE06939D6169B1AFCF0FC1CFC11" box="[887,1009,874,899]" metricMagnitude="2" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="4" pageNumber="362" unit="m" value="225.0" valueMax="250.0" valueMin="200.0">
<elevation id="2FDBC5366939D6169B1AFCF0FC1CFC11" box="[887,1009,874,899]" metricMagnitude="2" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="4" pageNumber="362" unit="m" value="225.0" valueMax="250.0" valueMin="200.0">200–250m</elevation>
</quantity>
, (
<collectorName id="090347D36939D6169C6AFCF1FAA9FC11" box="[1031,1348,874,899]" pageId="4" pageNumber="362">K.R., J.C.R, J.I.R. Willmott</collectorName>
), 6 
<date id="D04804C56939D6169D00FCF0FF01FC35" pageId="4" pageNumber="362" value="2015-07-01">
<collectingDate id="C00CFD2D6939D6169D00FCF0FF01FC35" pageId="4" pageNumber="362" value="2015-07-01">July 2015, 1</collectingDate>
</date>
♂, [FLMNH-MGCL-217573], (
<collectionCode id="C2E7BAC06939D6169A31FC15FD5EFC35" box="[604,691,911,935]" pageId="4" pageNumber="362">MGCL</collectionCode>
), 8 
<date id="D04804C56939D6169A8EFC15FC9FFC35" box="[739,882,911,935]" pageId="4" pageNumber="362" value="2015-07-01">
<collectingDate id="C00CFD2D6939D6169A8EFC15FC9FFC35" box="[739,882,911,935]" pageId="4" pageNumber="362" value="2015-07-01">July 2015, 1</collectingDate>
</date>
♂, [FLMNH-MGCL-217574], (
<collectionCode id="C2E7BAC06939D6169C8EFC15FAD7FC35" box="[1251,1338,911,935]" pageId="4" pageNumber="362">MGCL</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="149E28586939D6169D3CFC15FA75FC59" ID-GBIF-Occurrence="1324639627" collectionCode="MGCL" collectorName="K. Willmott &amp; J. Hall &amp; July" country="Ecuador" elevation="350" latitude="1.0408334" location="Lita-San Lorenzo rd." longLatPrecision="21" longitude="-78.634445" pageId="4" pageNumber="362" specimenCount="1" specimenCount-female="1" stateProvince="Durango" typeStatus="paratype">
km 40 
<location id="A12974DE6939D61698FAFC29FE69FC59" LSID="urn:lsid:plazi:treatment:2C5F9313693FD61598FAFF0DFE2AFEF7:A12974DE6939D61698FAFC29FE69FC59" box="[151,388,947,971]" country="Ecuador" latitude="1.0408334" longLatPrecision="21" longitude="-78.634445" name="Lita-San Lorenzo rd." pageId="4" pageNumber="362" stateProvince="Durango">Lita-San Lorenzo rd.</location>
, El 
<collectingRegion id="6632ECE76939D61699DCFC29FDFBFC59" box="[433,534,947,971]" country="Mexico" name="Durango" pageId="4" pageNumber="362">Durango</collectingRegion>
, [
<geoCoordinate id="C1C244C26939D6169A44FC29FD60FC59" box="[553,653,947,971]" direction="north" orientation="latitude" pageId="4" pageNumber="362" precision="15" value="1.0408334">1º2'27''N</geoCoordinate>
, 
<geoCoordinate id="C1C244C26939D6169AFFFC29FCFEFC59" box="[658,787,947,971]" direction="west" orientation="longitude" pageId="4" pageNumber="362" precision="15" value="-78.634445">78º38'4''W</geoCoordinate>
], 
<quantity id="630E8FE06939D6169B4BFC29FC4CFC59" box="[806,929,947,971]" metricMagnitude="2" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="4" pageNumber="362" unit="m" value="350.0" valueMax="400.0" valueMin="300.0">
<elevation id="2FDBC5366939D6169B4BFC29FC4CFC59" box="[806,929,947,971]" metricMagnitude="2" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="4" pageNumber="362" unit="m" value="350.0" valueMax="400.0" valueMin="300.0">300–400m</elevation>
</quantity>
, (
<collectorName id="090347D36939D6169BD8FC29FBD0FC59" box="[949,1085,947,971]" pageId="4" pageNumber="362">K. Willmott</collectorName>
&amp; 
<collectorName id="090347D36939D6169C0FFC29FB42FC59" box="[1122,1199,947,971]" pageId="4" pageNumber="362">J. Hall</collectorName>
), 
<collectorName id="090347D36939D6169CA9FC29FB19FC59" box="[1220,1268,947,971]" pageId="4" pageNumber="362">July</collectorName>
, 
<specimenCount id="B2F0E98C6939D6169C92FC29FACFFC59" box="[1279,1314,947,971]" pageId="4" pageNumber="362" type="female">1♀</specimenCount>
, (
<collectionCode id="C2E7BAC06939D6169D54FC29FA7DFC59" box="[1337,1424,947,971]" pageId="4" pageNumber="362">MGCL</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="149E28586939D61698FAFC4CFDB9FB81" ID-GBIF-Occurrence="1324639623" collectingDate="1995-03-01" collectionCode="MGCL" collectorName="La Punta &amp; K. Willmott &amp; J. Hall &amp; Genitalic" country="Ecuador" elevation="300" latitude="1.0652777" location="La Punta" longLatPrecision="925" longitude="-78.65" municipality="Lita-San Lorenzo rd." pageId="4" pageNumber="362" specimenCount="1" stateProvince="Durango" typeStatus="paratype">
km 44 
<collectingMunicipality id="442DB87F6939D6169885FC4CFE35FC7D" box="[232,472,982,1007]" pageId="4" pageNumber="362">Lita-San Lorenzo rd.</collectingMunicipality>
, 
<location id="A12974DE6939D6169988FC42FDA3FC7D" LSID="urn:lsid:plazi:treatment:2C5F9313693FD61598FAFF0DFE2AFEF7:A12974DE6939D6169988FC42FDA3FC7D" box="[485,590,984,1007]" country="Ecuador" latitude="1.0652777" longLatPrecision="925" longitude="-78.65" municipality="Lita-San Lorenzo rd." name="La Punta" pageId="4" pageNumber="362" stateProvince="Durango">
<collectorName id="090347D36939D6169988FC42FDA3FC7D" box="[485,590,984,1007]" pageId="4" pageNumber="362">La Punta</collectorName>
</location>
, [
<geoCoordinate id="C1C244C26939D6169A0FFC4CFD29FC7D" box="[610,708,982,1007]" direction="north" orientation="latitude" pageId="4" pageNumber="362" precision="15" value="1.0652777">1º3'55''N</geoCoordinate>
, 
<geoCoordinate id="C1C244C26939D6169AA4FC4CFCDEFC7D" box="[713,819,982,1007]" direction="west" orientation="longitude" pageId="4" pageNumber="362" precision="925" value="-78.65">78º39'W</geoCoordinate>
], 
<quantity id="630E8FE06939D6169B25FC4DFC66FC7D" box="[840,907,983,1007]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="362" unit="m" value="300.0">
<elevation id="2FDBC5366939D6169B25FC4DFC66FC7D" box="[840,907,983,1007]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="362" unit="m" value="300.0">300m</elevation>
</quantity>
, (
<collectorName id="090347D36939D6169BCCFC4DFBC4FC7D" box="[929,1065,982,1007]" pageId="4" pageNumber="362">K. Willmott</collectorName>
&amp; 
<collectorName id="090347D36939D6169C3DFC42FB72FC7D" box="[1104,1183,982,1007]" pageId="4" pageNumber="362">J. Hall</collectorName>
), 24 
<date id="D04804C56939D6169CB5FC4CFA90FC7D" box="[1240,1405,982,1007]" pageId="4" pageNumber="362" value="1995-03-01">
<collectingDate id="C00CFD2D6939D6169CB5FC4CFA90FC7D" box="[1240,1405,982,1007]" pageId="4" pageNumber="362" value="1995-03-01">March 1995, 1</collectingDate>
</date>
♂, 
<collectorName id="090347D36939D61698FAFC60FF13FB81" box="[151,254,1018,1043]" pageId="4" pageNumber="362">Genitalic</collectorName>
vial KW-14-32, (
<collectionCode id="C2E7BAC06939D6169999FC61FDA6FB81" box="[500,587,1019,1043]" pageId="4" pageNumber="362">MGCL</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="149E28586939D6169A02FC60FA75FBA5" ID-GBIF-Occurrence="1324639622" collectingDate="2015-07-01" collectionCode="MGCL" collectorName="NE San Francisco &amp; N La Ceiba &amp; K. R. Willmott &amp; R. Aldaz" country="Ecuador" elevation="250" latitude="1.1325" location="Lita-San Lorenzo rd." longLatPrecision="21" longitude="-78.65889" pageId="4" pageNumber="362" specimenCount="1" stateProvince="Durango" typeStatus="paratype">
<location id="A12974DE6939D6169A02FC60FC91FB81" LSID="urn:lsid:plazi:treatment:2C5F9313693FD61598FAFF0DFE2AFEF7:A12974DE6939D6169A02FC60FC91FB81" box="[623,892,1018,1043]" country="Ecuador" latitude="1.1325" longLatPrecision="21" longitude="-78.65889" name="Lita-San Lorenzo rd." pageId="4" pageNumber="362" stateProvince="Durango">Lita-San Lorenzo rd.</location>
, 
<collectorName id="090347D36939D6169BF4FC61FB6AFB81" box="[921,1159,1018,1043]" pageId="4" pageNumber="362">NE San Francisco</collectorName>
, ridge 
<collectorName id="090347D36939D6169C9EFC61FA75FB81" box="[1267,1432,1018,1043]" pageId="4" pageNumber="362">N La Ceiba</collectorName>
, [
<geoCoordinate id="C1C244C26939D61698F3FB85FEECFBA5" box="[158,257,1055,1079]" direction="north" orientation="latitude" pageId="4" pageNumber="362" precision="15" value="1.1325">1º7'57''N</geoCoordinate>
, 
<geoCoordinate id="C1C244C26939D6169968FB85FE7BFBA5" box="[261,406,1055,1079]" direction="west" orientation="longitude" pageId="4" pageNumber="362" precision="15" value="-78.65889">78º39'32''W</geoCoordinate>
], 
<quantity id="630E8FE06939D61699C7FB85FE00FBA5" box="[426,493,1055,1079]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="362" unit="m" value="250.0">
<elevation id="2FDBC5366939D61699C7FB85FE00FBA5" box="[426,493,1055,1079]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="362" unit="m" value="250.0">250m</elevation>
</quantity>
, (
<collectorName id="090347D36939D6169A6CFB85FD4BFBA5" box="[513,678,1055,1079]" pageId="4" pageNumber="362">K.R. Willmott</collectorName>
, 
<collectorName id="090347D36939D6169ADFFB85FCF6FBA5" box="[690,795,1055,1079]" pageId="4" pageNumber="362">R. Aldaz</collectorName>
), 5 
<date id="D04804C56939D6169B2BFB85FC3CFBA5" box="[838,977,1055,1079]" pageId="4" pageNumber="362" value="2015-07-01">
<collectingDate id="C00CFD2D6939D6169B2BFB85FC3CFBA5" box="[838,977,1055,1079]" pageId="4" pageNumber="362" value="2015-07-01">July 2015, 1</collectingDate>
</date>
♀, [FLMNH-MGCL-195691], (
<collectionCode id="C2E7BAC06939D6169D54FB85FA7DFBA5" box="[1337,1424,1055,1079]" pageId="4" pageNumber="362">MGCL</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="149E28586939D61698FAFBD9FE28FBED" ID-GBIF-Occurrence="1324639636" collectingDate="1997-09-01" collectionCode="MGCL, BMNH" collectorName="El Placer &amp; K. Willmott" country="Ecuador" elevation="750" latitude="0.9083333" location="El Placer" longLatPrecision="925" longitude="-78.6" municipality="San Lorenzo-Lita" pageId="4" pageNumber="362" specimenCount="1" stateProvince="Durango" typeStatus="paratype">
<collectingMunicipality id="442DB87F6939D61698FAFBD9FEB0FBC9" box="[151,349,1090,1115]" pageId="4" pageNumber="362">San Lorenzo-Lita</collectingMunicipality>
railway, 
<location id="A12974DE6939D61699A8FBD8FDC3FBC9" LSID="urn:lsid:plazi:treatment:2C5F9313693FD61598FAFF0DFE2AFEF7:A12974DE6939D61699A8FBD8FDC3FBC9" box="[453,558,1090,1115]" country="Ecuador" latitude="0.9083333" longLatPrecision="925" longitude="-78.6" municipality="San Lorenzo-Lita" name="El Placer" pageId="4" pageNumber="362" stateProvince="Durango">
<collectorName id="090347D36939D61699A8FBD8FDC3FBC9" box="[453,558,1090,1115]" pageId="4" pageNumber="362">El Placer</collectorName>
</location>
, [
<geoCoordinate id="C1C244C26939D6169A2CFBD8FD5AFBC9" box="[577,695,1090,1115]" direction="north" orientation="latitude" pageId="4" pageNumber="362" precision="15" value="0.9083333">0º54'30''N</geoCoordinate>
, 
<geoCoordinate id="C1C244C26939D6169AD1FBD8FCCFFBC9" box="[700,802,1090,1115]" direction="west" orientation="longitude" pageId="4" pageNumber="362" precision="925" value="-78.6">78º36'W</geoCoordinate>
], 
<quantity id="630E8FE06939D6169B58FBD8FC95FBC9" box="[821,888,1090,1115]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="4" pageNumber="362" unit="m" value="750.0">
<elevation id="2FDBC5366939D6169B58FBD8FC95FBC9" box="[821,888,1090,1115]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="4" pageNumber="362" unit="m" value="750.0">750m</elevation>
</quantity>
, (
<collectorName id="090347D36939D6169BE1FBD9FBF4FBC9" box="[908,1049,1090,1115]" pageId="4" pageNumber="362">K. Willmott</collectorName>
), 9 
<date id="D04804C56939D6169C2FFBD8FAFAFBC9" box="[1090,1303,1090,1115]" pageId="4" pageNumber="362" value="1997-09-01">
<collectingDate id="C00CFD2D6939D6169C2FFBD8FAFAFBC9" box="[1090,1303,1090,1115]" pageId="4" pageNumber="362" value="1997-09-01">September 1997, 1</collectingDate>
</date>
♀, (
<collectionCode id="C2E7BAC06939D6169D2FFBD9FA7AFBC9" box="[1346,1431,1091,1115]" pageId="4" pageNumber="362">MGCL</collectionCode>
, to be deposited in 
<collectionCode id="C2E7BAC06939D6169909FBFDFE50FBED" box="[356,445,1127,1151]" pageId="4" pageNumber="362">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="A44922056939D61698AAFB10FC07FB55" blockId="4.[151,1437,151,2015]" pageId="4" pageNumber="362">
<materialsCitation id="149E28586939D61698AAFB10FC07FB55" ID-GBIF-Occurrence="1324639621" collectingDate="2013-09-01" collectionCode="OTHER, RECORDS" collectorName="Playa de Oro &amp; Hardwick, R." country="Ecuador" elevation="70" latitude="0.87583333" location="Rio Santiago" longLatPrecision="21" longitude="-78.79444" pageId="4" pageNumber="362" specimenCount="1" stateProvince="Esmeraldas">
<emphasis id="9682FE176939D61698AAFB10FD89FB31" bold="true" box="[199,612,1162,1187]" pageId="4" pageNumber="362">
<collectionCode id="C2E7BAC06939D61698AAFB10FE2CFB31" box="[199,449,1162,1187]" pageId="4" pageNumber="362">OTHER RECORDS</collectionCode>
: 
<collectingCountry id="DCE162956939D61699A0FB10FDB2FB31" box="[461,607,1162,1187]" name="Ecuador" pageId="4" pageNumber="362">ECUADOR</collectingCountry>
:
</emphasis>
<emphasis id="9682FE176939D6169A01FB11FD1DFB31" box="[620,752,1163,1187]" italics="true" pageId="4" pageNumber="362">
<collectingRegion id="6632ECE76939D6169A01FB11FD1DFB31" box="[620,752,1163,1187]" country="Ecuador" name="Esmeraldas" pageId="4" pageNumber="362">Esmeraldas</collectingRegion>
</emphasis>
; 
<location id="A12974DE6939D6169B6DFB11FC78FB31" LSID="urn:lsid:plazi:treatment:2C5F9313693FD61598FAFF0DFE2AFEF7:A12974DE6939D6169B6DFB11FC78FB31" box="[768,917,1162,1187]" country="Ecuador" latitude="0.87583333" longLatPrecision="21" longitude="-78.79444" name="Rio Santiago" pageId="4" pageNumber="362" stateProvince="Esmeraldas">Río Santiago</location>
, 
<collectorName id="090347D36939D6169BCDFB10FBDBFB31" box="[928,1078,1162,1187]" pageId="4" pageNumber="362">Playa de Oro</collectorName>
, [
<geoCoordinate id="C1C244C26939D6169C24FB10FB53FB31" box="[1097,1214,1162,1187]" direction="north" orientation="latitude" pageId="4" pageNumber="362" precision="15" value="0.87583333">0º52'33''N</geoCoordinate>
, 
<geoCoordinate id="C1C244C26939D6169CAEFB10FABCFB31" box="[1219,1361,1162,1187]" direction="west" orientation="longitude" pageId="4" pageNumber="362" precision="15" value="-78.79444">78º47'40''W</geoCoordinate>
], 
<quantity id="630E8FE06939D6169D0EFB11FA7AFB31" box="[1379,1431,1163,1187]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="362" unit="m" value="70.0">
<elevation id="2FDBC5366939D6169D0EFB11FA7AFB31" box="[1379,1431,1163,1187]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="362" unit="m" value="70.0">70m</elevation>
</quantity>
, (
<collectorName id="090347D36939D61698F3FB34FED5FB55" box="[158,312,1198,1223]" pageId="4" pageNumber="362">Hardwick, R.</collectorName>
), 26–29 
<date id="D04804C56939D61699F6FB34FD80FB55" box="[411,621,1198,1223]" pageId="4" pageNumber="362" value="2013-09-01">
<collectingDate id="C00CFD2D6939D61699F6FB34FD80FB55" box="[411,621,1198,1223]" pageId="4" pageNumber="362" value="2013-09-01">September 2013, 1</collectingDate>
</date>
♀, (photograph of live specimen).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="ECEC718E6939D61698AAFB48FAACFA9D" pageId="4" pageNumber="362" type="etymology">
<paragraph id="A44922056939D61698AAFB48FAACFA9D" blockId="4.[151,1437,151,2015]" pageId="4" pageNumber="362">
<emphasis id="9682FE176939D61698AAFB48FEA3FB79" bold="true" box="[199,334,1234,1259]" pageId="4" pageNumber="362">Etymology.</emphasis>
This specific epithet is derived from the masculine Latin noun, 
<taxonomicName id="63F659866939D6169C51FB49FB76FB79" box="[1084,1179,1235,1259]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="362" phylum="Arthropoda" rank="species" species="favonius">
<emphasis id="9682FE176939D6169C51FB49FB76FB79" box="[1084,1179,1235,1259]" italics="true" pageId="4" pageNumber="362">favonius</emphasis>
</taxonomicName>
, used in apposition, a word meaning ‘west wind’, in reference to the fact that this species is only known from west of the Andes.
</paragraph>
</subSubSection>
<subSubSection id="ECEC718E6939D61698AAFA80FE79FAC5" pageId="4" pageNumber="362" type="distribution">
<paragraph id="A44922056939D61698AAFA80FE79FAC5" blockId="4.[151,1437,151,2015]" pageId="4" pageNumber="362">
<emphasis id="9682FE176939D61698AAFA80FE8DFAA1" bold="true" box="[199,352,1306,1331]" pageId="4" pageNumber="362">Distribution.</emphasis>
(
<figureCitation id="3CCD3E806939D616991CFA80FE58FAA1" box="[369,437,1306,1331]" captionStart="FIGURE 3" captionStartId="6.[151,250,1977,1999]" captionTargetBox="[163,1418,212,1938]" captionTargetId="figure@6.[151,1436,193,1956]" captionTargetPageId="6" captionText="FIGURE 3. Distribution map for E. favonius." httpUri="https://zenodo.org/record/164789/files/figure.png" pageId="4" pageNumber="362">Fig. 3</figureCitation>
): Based on the examined specimens, this new species is distributed from 
<collectingCountry id="DCE162956939D6169D6FFA81FA90FAA1" box="[1282,1405,1306,1331]" name="Costa Rica" pageId="4" pageNumber="362">Costa Rica</collectingCountry>
to northwestern 
<collectingCountry id="DCE162956939D616995DFAA4FE7DFAC5" box="[304,400,1342,1367]" name="Ecuador" pageId="4" pageNumber="362">Ecuador</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="ECEC718E6939D61698AAFAF8FC9BF84D" pageId="4" pageNumber="362" type="biology_ecology">
<paragraph id="A44922056939D61698AAFAF8FDB9F999" blockId="4.[151,1437,151,2015]" pageId="4" pageNumber="362">
<emphasis id="9682FE176939D61698AAFAF8FECAFAE9" bold="true" box="[199,295,1378,1403]" pageId="4" pageNumber="362">Biology.</emphasis>
In western 
<collectingCountry id="DCE162956939D61699C5FAF8FDEAFAE9" box="[424,519,1378,1403]" name="Ecuador" pageId="4" pageNumber="362">Ecuador</collectingCountry>
, 
<taxonomicName id="63F659866939D6169A7CFAF9FD62FAE9" box="[529,655,1379,1403]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="362" phylum="Arthropoda" rank="species" species="favonius">
<emphasis id="9682FE176939D6169A7CFAF9FD62FAE9" box="[529,655,1379,1403]" italics="true" pageId="4" pageNumber="362">E. favonius</emphasis>
</taxonomicName>
occurs in wet lowland rainforest from 
<quantity id="630E8FE06939D6169C53FAF9FB43FAE8" box="[1086,1198,1379,1403]" metricMagnitude="2" metricUnit="m" metricValue="4.85" metricValueMax="9.0" metricValueMin="0.7" pageId="4" pageNumber="362" unit="m" value="485.0" valueMax="900.0" valueMin="70.0">70–900 m</quantity>
elevation, where it is uncommon. Males were only encountered on prominent hilltops, where they perched and patrolled singly approximately 
<quantity id="630E8FE06939D6169952FA31FE87FA50" box="[319,362,1451,1475]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="362" unit="m" value="1.0">1 m</quantity>
above the ground at the edges of light gaps in the mid-morning. Females were observed both on hill and ridgetops as well as hillsides, flying along trails in the primary forest understorey about 
<quantity id="630E8FE06939D6169CB3FA55FACAFA75" box="[1246,1319,1487,1511]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="4" pageNumber="362" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">1–2 m</quantity>
above the ground from midday to early afternoon.
</paragraph>
<paragraph id="A44922056939D61698AAF98CFC9BF84D" blockId="4.[151,1437,151,2015]" pageId="4" pageNumber="362">
<emphasis id="9682FE176939D61698AAF98CFD1CF9BD" bold="true" box="[199,753,1558,1583]" pageId="4" pageNumber="362">
Taxonomy of 
<taxonomicName id="63F659866939D6169900F98CFDCEF9BC" box="[365,547,1558,1583]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="362" phylum="Arthropoda" rank="species" species="meta">
<emphasis id="9682FE176939D6169900F98CFDCEF9BC" bold="true" box="[365,547,1558,1583]" italics="true" pageId="4" pageNumber="362">Euptychia meta</emphasis>
</taxonomicName>
and 
<taxonomicName id="63F659866939D6169A09F98DFD04F9BD" box="[612,745,1558,1583]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="362" phylum="Arthropoda" rank="species" species="favonius">
<emphasis id="9682FE176939D6169A09F98DFD04F9BD" bold="true" box="[612,745,1558,1583]" italics="true" pageId="4" pageNumber="362">E. favonius</emphasis>
</taxonomicName>
.
</emphasis>
<bibRefCitation id="C0675FF46939D6169A97F982FC47F9BD" author="Weymer" box="[762,938,1559,1584]" pageId="4" pageNumber="362" refString="Weymer, G. (1911) 4. Familie: Satyridae. In: Seitz, A. (Ed.), Die Gross-Schmetterlinge der Erde. Vol. 5. A. Kernen, Stuttgart, pp. 185 - 225." type="book chapter" year="1911">Weymer (1911)</bibRefCitation>
described 
<taxonomicName id="63F659866939D6169C45F982FB6DF9BD" box="[1064,1152,1560,1583]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="362" phylum="Arthropoda" rank="species" species="meta">
<emphasis id="9682FE176939D6169C45F982FB6DF9BD" box="[1064,1152,1560,1583]" italics="true" pageId="4" pageNumber="362">E. meta</emphasis>
</taxonomicName>
based on an unspecified number of specimens from Marcapata, Peru. Although only one syntype in the ZMHU has so far been located (a female), examination of the original description indicates that Weymer believed that he had examined both sexes. In plate 46 the dorsal surface of the female is figured, but the sex is not mentioned and it is difficult to judge whether Weymer correctly realized that this species was sexually dimorphic. Therefore, additional syntypes (if they exist) might consist just of female specimen(s). The single known female syntype (
<figureCitation id="3CCD3E806939D6169C56F950FB97F971" box="[1083,1146,1738,1763]" captionStart="FIGURE 1" captionStartId="3.[151,250,1439,1461]" captionTargetBox="[175,1404,477,1408]" captionTargetId="figure@3.[151,1435,459,1415]" captionTargetPageId="3" captionText="FIGURE 1. Male and female specimens of E. favonius sp. nov. and E. meta, left dorsal and right ventral surfaces: A) E. favonius, HOLOTYPE male, western Ecuador; B) E. favonius, PARATYPE female, western Ecuador; C) E. meta, male, eastern Ecuador; D) E. meta, LECTOTYPE female, southeastern Peru." httpUri="https://zenodo.org/record/164787/files/figure.png" pageId="4" pageNumber="362">Fig. 1</figureCitation>
D) exhibits an undulating marginal band on both ventral wings, similar to 
<taxonomicName id="63F659866939D6169A8FF96AFC8BF895" box="[738,870,1775,1799]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="362" phylum="Arthropoda" rank="species" species="favonius">
<emphasis id="9682FE176939D6169A8FF96AFC8BF895" box="[738,870,1775,1799]" italics="true" pageId="4" pageNumber="362">E. favonius</emphasis>
</taxonomicName>
in this respect. However, the ventral hindwing submarginal band and outer ring of concentric submarginal ocelli are both typical of other east Ecuadorian 
<taxonomicName id="63F659866939D6169D2BF889FA76F8B8" box="[1350,1435,1811,1834]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="362" phylum="Arthropoda" rank="species" species="meta">
<emphasis id="9682FE176939D6169D2BF889FA76F8B8" box="[1350,1435,1811,1834]" italics="true" pageId="4" pageNumber="362">E. meta</emphasis>
</taxonomicName>
and are distinct from those of 
<taxonomicName id="63F659866939D616998BF8A2FD89F8DD" box="[486,612,1847,1871]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="362" phylum="Arthropoda" rank="species" species="favonius">
<emphasis id="9682FE176939D616998BF8A2FD89F8DD" box="[486,612,1847,1871]" italics="true" pageId="4" pageNumber="362">E. favonius</emphasis>
</taxonomicName>
(see diagnosis). Since cryptic species are frequently found in euptychiine butterflies (e.g. 
<bibRefCitation id="C0675FF46939D6169925F8C1FDD2F8E1" author="Cong" box="[328,575,1882,1907]" pageId="4" pageNumber="362" refString="Cong, Q. &amp; Grishin, N. V. (2014) A new Hermeuptychia (Lepidoptera, Nymphalidae, Satyrinae) is sympatric and synchronic with H. sosybius in southeast US coastal plains, while another new Hermeuptychia species - not hermes - inhabits south Texas and northeast Mexico. ZooKeys, 91, 43 - 91. http: // dx. doi. org / 10.3897 / zookeys. 379.6394" type="journal article" year="2014">Cong &amp; Grishin 2014</bibRefCitation>
), we designate this female syntype (
<figureCitation id="3CCD3E806939D6169BBFF8C0FBFCF8E1" box="[978,1041,1882,1907]" captionStart="FIGURE 1" captionStartId="3.[151,250,1439,1461]" captionTargetBox="[175,1404,477,1408]" captionTargetId="figure@3.[151,1435,459,1415]" captionTargetPageId="3" captionText="FIGURE 1. Male and female specimens of E. favonius sp. nov. and E. meta, left dorsal and right ventral surfaces: A) E. favonius, HOLOTYPE male, western Ecuador; B) E. favonius, PARATYPE female, western Ecuador; C) E. meta, male, eastern Ecuador; D) E. meta, LECTOTYPE female, southeastern Peru." httpUri="https://zenodo.org/record/164787/files/figure.png" pageId="4" pageNumber="362">Fig. 1</figureCitation>
D, forewing length = 
<quantity id="630E8FE06939D6169D6BF8C1FAA3F8E1" box="[1286,1358,1883,1907]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="4" pageNumber="362" unit="mm" value="17.0">17mm</quantity>
) in the ZMHU with the following labels as the lectotype of 
<taxonomicName id="63F659866939D6169B68F81AFC8DF805" box="[773,864,1920,1943]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="362" phylum="Arthropoda" rank="species" species="meta">
<emphasis id="9682FE176939D6169B68F81AFC8DF805" box="[773,864,1920,1943]" italics="true" pageId="4" pageNumber="362">E. meta</emphasis>
</taxonomicName>
in order to stabilize the nomenclature and allow confident identification of 
<taxonomicName id="63F659866939D61699A1F839FDA3F829" box="[460,590,1955,1979]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="362" phylum="Arthropoda" rank="species" species="favonius">
<emphasis id="9682FE176939D61699A1F839FDA3F829" box="[460,590,1955,1979]" italics="true" pageId="4" pageNumber="362">E. favonius</emphasis>
</taxonomicName>
: LECTOTYPE ♀ 
<taxonomicName id="63F659866939D6169B44F838FBA9F829" authority="Weymer" authorityName="Weymer" box="[809,1092,1954,1979]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="362" phylum="Arthropoda" rank="species" species="meta">Euptychia meta Weymer</taxonomicName>
designated by Lee D. Miller. 1989 
<emphasis id="9682FE176939D61698A2F85CFF32F84D" bold="true" box="[207,223,1990,2015]" pageId="4" pageNumber="362">//</emphasis>
Marcapata Cuzco 
<emphasis id="9682FE176939D61699C7F85CFE57F84D" bold="true" box="[426,442,1990,2015]" pageId="4" pageNumber="362">//</emphasis>
genitalia vial M-9055 ♀ Lee D. Miller.
</paragraph>
</subSubSection>
<caption id="F089728D6938D61798FAF813FCC2F84E" httpUri="https://zenodo.org/record/259867/files/figure.png" pageId="5" pageNumber="363" targetBox="[223,1366,206,1875]" targetPageId="5">
<paragraph id="A44922056938D61798FAF813FCC2F84E" blockId="5.[151,1436,1929,2012]" pageId="5" pageNumber="363">
<emphasis id="9682FE176938D61798FAF813FEF8F80C" bold="true" box="[151,277,1929,1951]" pageId="5" pageNumber="363">FIGURE 2.</emphasis>
Genitalia of 
<taxonomicName id="63F659866938D61799CCF810FDFBF80D" box="[417,534,1929,1951]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="363" phylum="Arthropoda" rank="species" species="favonius">
<emphasis id="9682FE176938D61799CCF810FDFBF80D" box="[417,534,1929,1951]" italics="true" pageId="5" pageNumber="363">E. favonius</emphasis>
</taxonomicName>
: A) male genitalia, in lateral view; B) aedeagus, in lateral view; C) juxta, in posterior view; D) female genitalia, in posterior view; E) lamella antevaginalis, in ventral view; F) ductus bursae and corpus bursae, in dorsal view; G) signa (SN-15-131; SN-15-126) Scale bar = 1mm.
</paragraph>
</caption>
<caption id="F089728D693BD61498FAF823FD8AF85D" box="[151,615,1977,1999]" httpUri="https://zenodo.org/record/164789/files/figure.png" pageId="6" pageNumber="364" targetBox="[163,1418,212,1938]" targetPageId="6">
<paragraph id="A4492205693BD61498FAF823FD8AF85D" blockId="6.[151,615,1977,1999]" box="[151,615,1977,1999]" pageId="6" pageNumber="364">
<emphasis id="9682FE17693BD61498FAF823FEFFF85C" bold="true" box="[151,274,1977,1999]" pageId="6" pageNumber="364">FIGURE 3.</emphasis>
Distribution map for 
<taxonomicName id="63F65986693BD6149982F820FD8CF85D" box="[495,609,1977,1999]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="364" phylum="Arthropoda" rank="species" species="favonius">
<emphasis id="9682FE17693BD6149982F820FD8CF85D" box="[495,609,1977,1999]" italics="true" pageId="6" pageNumber="364">E. favonius</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="ECEC718E693AD61598AAFF02FE2AFEF7" pageId="7" pageNumber="365" type="description">
<paragraph id="A4492205693AD61598AAFF02FE2AFEF7" blockId="7.[151,1437,151,357]" pageId="7" pageNumber="365">
<taxonomicName id="63F65986693AD61598AAFF02FE97FF22" box="[199,378,152,176]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="365" phylum="Arthropoda" rank="species" species="meta">
<emphasis id="9682FE17693AD61598AAFF02FE97FF22" box="[199,378,152,176]" italics="true" pageId="7" pageNumber="365">Euptychia meta</emphasis>
</taxonomicName>
and 
<taxonomicName id="63F65986693AD61599D1FF03FDD2FF22" box="[444,575,152,176]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="365" phylum="Arthropoda" rank="species" species="favonius">
<emphasis id="9682FE17693AD61599D1FF03FDD2FF22" box="[444,575,152,176]" italics="true" pageId="7" pageNumber="365">E. favonius</emphasis>
</taxonomicName>
are clearly sister taxa, based on both morphological and molecular data (Nakahara, unpublished data), and given their allopatry they could arguably be regarded as conspecific. However, the small but consistent wing pattern differences are similar in extent to those distinguishing other sympatric, closely related 
<taxonomicName id="63F65986693AD6159927FE9EFE51FE8E" box="[330,444,260,284]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="9682FE17693AD6159927FE9EFE51FE8E" box="[330,444,260,284]" italics="true" pageId="7" pageNumber="365">Euptychia</emphasis>
</taxonomicName>
species (Nakahara, unpublished data). Moreover, the significant divergence in COI, which is of an order of magnitude greater between the two taxa than within the two taxa, suggests that separate species status is warranted.
</paragraph>
</subSubSection>
</treatment>
</document>