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<document id="74572FC1FDB4191D038A2D0F4301C0E3" ID-CLB-Dataset="55478" ID-DOI="10.11646/zootaxa.5138.3.3" ID-GBIF-Dataset="127cd1e7-81ce-4e94-baab-57a88d940d72" ID-ISSN="1175-5326" ID-Zenodo-Dep="6559653" ID-ZooBank="E3A9B988-A702-411A-A917-BB3760B9CECB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1652866326516" checkinUser="plazi" docAuthor="Anker, Arthur" docDate="2022" docId="039A8D0A8259FFE95593FF1D101AFBD1" docLanguage="en" docName="zootaxa.5138.3.3.pdf" docOrigin="Zootaxa 5138 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Salmoneus arabicus Anker 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="271" masterDocId="FFA3F572825FFFE35504FF8A124BFFC0" masterDocTitle="Three new species of the shrimp genus Salmoneus Holthuis, 1955 from the Red Sea, with a new record of S. poupini Anker, 2010 (Malacostraca: Decapoda Alpheidae)" masterLastPageNumber="282" masterPageNumber="261" pageNumber="267" updateTime="1699337770483" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="E0C6D09AD659AE67ADE5794CAE4FB6DD">
<mods:title id="C4B89F4CFBCD0D11E2FC69BDEB864033">Three new species of the shrimp genus Salmoneus Holthuis, 1955 from the Red Sea, with a new record of S. poupini Anker, 2010 (Malacostraca: Decapoda Alpheidae)</mods:title>
</mods:titleInfo>
<mods:name id="1E901B0AC6C5C1CF51D015D63FF7D82E" type="personal">
<mods:role id="9FD466ADE2B8168310156C7D28755B96">
<mods:roleTerm id="D1B8619AD8918BD6FBFDF0810058B7DF">Author</mods:roleTerm>
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<mods:namePart id="C1877433239E2FD351A220F73FA6A5D2">Anker, Arthur</mods:namePart>
</mods:name>
<mods:typeOfResource id="BB50C8C4CE3A2933645AC3E663D998C2">text</mods:typeOfResource>
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<mods:title id="5B287CBE7FCD6D9D3ECF7251D4946762">Zootaxa</mods:title>
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<mods:part id="A390F57D38D8AE6AA3F7645BDAF693FA">
<mods:date id="BE910AF6C97336E6C8C1D1EF08A312F7">2022</mods:date>
<mods:detail id="48BEC426D53788069482B3A0EBD571F8" type="pubDate">
<mods:number id="F3E515B45CD8A74CCDCA9AB92A175712">2022-05-18</mods:number>
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<mods:number id="F873DB426D3DFFB523F500D938A41EEC">5138</mods:number>
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<mods:number id="591441EAD137E64A191F87795452ADCD">3</mods:number>
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<mods:classification id="21D34EDB2EE6E3B77B7DEA6FB4749B05">journal article</mods:classification>
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<mods:identifier id="889BB1991593C00D1C365E78A639ADA7" type="DOI">10.11646/zootaxa.5138.3.3</mods:identifier>
<mods:identifier id="2C89070D4D7B4BAD53C6B8AFCFA54D4D" type="GBIF-Dataset">127cd1e7-81ce-4e94-baab-57a88d940d72</mods:identifier>
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<treatment id="039A8D0A8259FFE95593FF1D101AFBD1" ID-DOI="http://doi.org/10.5281/zenodo.6564887" ID-GBIF-Taxon="195535773" ID-Zenodo-Dep="6564887" LSID="urn:lsid:plazi:treatment:039A8D0A8259FFE95593FF1D101AFBD1" httpUri="http://treatment.plazi.org/id/039A8D0A8259FFE95593FF1D101AFBD1" lastPageId="10" lastPageNumber="271" pageId="6" pageNumber="267">
<subSubSection id="C3296F978259FFE55593FF1D13BAFF71" box="[151,497,151,178]" pageId="6" pageNumber="267" type="nomenclature">
<paragraph id="8B8C3C1C8259FFE55593FF1D13BAFF71" blockId="6.[151,497,151,214]" box="[151,497,151,178]" pageId="6" pageNumber="267">
<heading id="D0C48B708259FFE55593FF1D13BAFF71" bold="true" box="[151,497,151,178]" fontSize="11" level="1" pageId="6" pageNumber="267" reason="1">
<emphasis id="B947E00E8259FFE55593FF1D13BAFF71" bold="true" box="[151,497,151,178]" pageId="6" pageNumber="267">
<taxonomicName id="4C33479F8259FFE55593FF1D13C7FF72" authority="Anker, 2022" authorityName="Anker" authorityYear="2022" box="[151,396,151,178]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="267" phylum="Arthropoda" rank="species" species="arabicus" status="sp. nov.">
<emphasis id="B947E00E8259FFE55593FF1D13C7FF72" bold="true" box="[151,396,151,178]" italics="true" pageId="6" pageNumber="267">Salmoneus arabicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2745D758259FFE55490FF1D13BAFF71" box="[404,497,151,177]" pageId="6" pageNumber="267" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3296F978259FFE55593FF311359FF16" box="[151,274,187,214]" pageId="6" pageNumber="267" type="description">
<paragraph id="8B8C3C1C8259FFE55593FF311359FF16" blockId="6.[151,497,151,214]" box="[151,274,187,214]" pageId="6" pageNumber="267">
(
<figureCitation id="130820998259FFE5559BFF311341FF16" box="[159,266,187,214]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStartId-0="7.[151,250,1874,1898]" captionStartId-1="8.[151,250,1930,1954]" captionStartId-2="9.[151,250,1498,1522]" captionTargetBox-0="[215,1373,199,1839]" captionTargetBox-1="[166,1420,735,1894]" captionTargetBox-2="[151,1436,181,1474]" captionTargetId-0="figure-15@7.[201,1386,181,1851]" captionTargetId-1="figure-288@8.[151,1436,722,1906]" captionTargetId-2="figure-15@9.[151,1436,181,1474]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionText-0="FIGURE 4. Salmoneus arabicus sp. nov., holotype, ovigerous specimen (cl 2.8 mm) from Al Fahal Reef off Thuwal, Saudi Arabia (FLMNH UF 35964): a—frontal region, dorsal; b—same, lateral view; c—fourth, fifth and sixth pleonites, lateral; d—telson, lateral; e—ventromesial tooth of first article of antennular peduncle, lateral; f—third maxilliped, lateral; g—second pereiopod, lateral; h—third pereiopod, lateral; i—same, distal portion of propodus and dactylus, mesial; j—fifth pereiopod, lateral; k—same, propodus and dactylus, mesial." captionText-1="FIGURE 5. Salmoneus arabicus sp. nov., holotype, ovigerous specimen (cl 2.8 mm) from Al Fahal Reef off Thuwal, Saudi Arabia (FLMNH UF 35964): a—major (left) cheliped, mesial; b—same, lateral; c—same, chela, dorsomesial; d—minor (right) cheliped, lateral." captionText-2="FIGURE 6. Salmoneus arabicus sp. nov., holotype, ovigerous specimen (cl 2.8 mm) from Al Fahal Reef off Thuwal, Saudi Arabia (FLMNH UF 35964): a—shrimp in life, dorsal; b—same, lateral. Photographs by the author." figureDoi-0="http://doi.org/10.5281/zenodo.6559671" figureDoi-1="http://doi.org/10.5281/zenodo.6559673" figureDoi-2="http://doi.org/10.5281/zenodo.6559675" httpUri-0="https://zenodo.org/record/6559671/files/figure.png" httpUri-1="https://zenodo.org/record/6559673/files/figure.png" httpUri-2="https://zenodo.org/record/6559675/files/figure.png" pageId="6" pageNumber="267">Figs. 46</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3296F978259FFE55593FE891139FE11" pageId="6" pageNumber="267" type="materials_examined">
<paragraph id="8B8C3C1C8259FFE55593FE891139FE11" blockId="6.[151,1437,259,1978]" pageId="6" pageNumber="267">
<emphasis id="B947E00E8259FFE55593FE89130BFEDD" bold="true" box="[151,320,259,285]" pageId="6" pageNumber="267">Type material</emphasis>
.
<materialsCitation id="3B5B36418259FFE55454FE89106FFEA5" ID-GBIF-Occurrence="3775741304" collectingDate="2013-03-19" collectingDateMax="2013-03-22" collectingDateMin="2013-03-19" collectionCode="FLMNH" collectorName="A. Anker" country="Saudi Arabia" county="northern Farasan Banks" elevation="15" latitude="20.117" location="Red Sea" longLatPrecision="1" longitude="40.215" municipality="Al Fahal Reef" pageId="6" pageNumber="267" specimenCode="UF 35964, UF 35982, UF 35985" specimenCount="3" typeStatus="holotype">
<typeStatus id="548882BE8259FFE55454FE8913F0FEDD" box="[336,443,259,285]" pageId="6" pageNumber="267" type="holotype">Holotype</typeStatus>
, ovig. specimen (cl
<quantity id="4CCB91F98259FFE557A0FE89114AFEDE" box="[676,769,259,286]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="6" pageNumber="267" unit="mm" value="2.8">2.8 mm</quantity>
),
<collectionCode id="ED22A4D98259FFE55612FE8E1133FEDE" box="[790,888,260,286]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="6" pageNumber="267" type="Museum">FLMNH</collectionCode>
<specimenCode id="DB9594678259FFE55685FE8E11BCFEDE" box="[897,1015,259,286]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="6" pageNumber="267" type="Museum">UF 35964</specimenCode>
,
<location id="8EEC6AC78259FFE55100FE89162EFEDD" LSID="urn:lsid:plazi:treatment:039A8D0A8259FFE95593FF1D101AFBD1:8EEC6AC78259FFE55100FE89162EFEDD" box="[1028,1125,259,285]" country="Saudi Arabia" county="northern Farasan Banks" latitude="20.117" longLatPrecision="1" longitude="40.215" municipality="Al Fahal Reef" name="Red Sea" pageId="6" pageNumber="267">Red Sea</location>
,
<collectingCountry id="F3247C8C8259FFE55177FE891742FEDD" box="[1139,1289,259,285]" name="Saudi Arabia" pageId="6" pageNumber="267">Saudi Arabia</collectingCountry>
, off
<location id="8EEC6AC78259FFE55044FE8917D3FEDD" LSID="urn:lsid:plazi:treatment:039A8D0A8259FFE95593FF1D101AFBD1:8EEC6AC78259FFE55044FE8917D3FEDD" box="[1344,1432,259,285]" country="Saudi Arabia" county="northern Farasan Banks" latitude="20.117" longLatPrecision="1" longitude="40.215" municipality="Al Fahal Reef" name="Thuwal" pageId="6" pageNumber="267">Thuwal</location>
,
<collectingMunicipality id="6BE8A6668259FFE55593FEAD1372FE81" box="[151,313,295,321]" pageId="6" pageNumber="267">Al Fahal Reef</collectingMunicipality>
,
<geoCoordinate id="EE075ADB8259FFE55441FEAD13ABFE82" box="[325,480,295,322]" degrees="22" direction="north" minutes="16" orientation="latitude" pageId="6" pageNumber="267" precision="1" seconds="52.2" value="22.281166">22°16'52.2&quot;N</geoCoordinate>
,
<geoCoordinate id="EE075ADB8259FFE554E9FEAD10CEFE82" box="[493,645,295,322]" degrees="38" direction="east" minutes="57" orientation="longitude" pageId="6" pageNumber="267" precision="1" seconds="45.8" value="38.962723">38°57'45.8&quot;E</geoCoordinate>
, exposed side of offshore coral reef, depth:
<quantity id="4CCB91F98259FFE55172FEAD1697FE82" box="[1142,1244,295,322]" metricMagnitude="1" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="6" pageNumber="267" unit="m" value="15.0" valueMax="20.0" valueMin="10.0">
<elevation id="001EDB2F8259FFE55172FEAD1697FE82" box="[1142,1244,295,322]" metricMagnitude="1" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="6" pageNumber="267" unit="m" value="15.0" valueMax="20.0" valueMin="10.0">1020 m</elevation>
</quantity>
, leg.
<collectorName id="26C659CA8259FFE5501DFEAD17CBFE81" box="[1305,1408,295,321]" pageId="6" pageNumber="267">A. Anker</collectorName>
<emphasis id="B947E00E8259FFE5508CFEA212FFFEA5" italics="true" pageId="6" pageNumber="267">et al.</emphasis>
,
<date id="FF8D1ADC8259FFE555C7FEC1130AFEA5" box="[195,321,331,357]" pageId="6" pageNumber="267" value="2013-03-19">
<collectingDate id="EFC9E3348259FFE555C7FEC1130AFEA5" box="[195,321,331,357]" pageId="6" pageNumber="267" value="2013-03-19">19.03.2013</collectingDate>
</date>
[fcn BDJRS-3038];
</materialsCitation>
<materialsCitation id="3B5B36418259FFE55729FEC6112FFE6D" ID-GBIF-Occurrence="3775741303" collectingDate="2013-03-19" collectingDateMax="2013-03-22" collectingDateMin="2013-03-19" collectionCode="FLMNH" collectorName="A. Anker" country="Saudi Arabia" county="northern Farasan Banks" elevation="15" latitude="20.117" location="Red Sea" longLatPrecision="1" longitude="40.215" municipality="Al Fahal Reef" pageId="6" pageNumber="267" specimenCode="UF 35964, UF 35982, UF 35985" specimenCount="3" typeStatus="paratype">
<typeStatus id="548882BE8259FFE55729FEC610D6FEA6" box="[557,669,332,358]" pageId="6" pageNumber="267" type="paratype">paratypes</typeStatus>
: 1 ovig. specimen (cl
<quantity id="4CCB91F98259FFE55692FEC111BAFEA6" box="[918,1009,331,358]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="6" pageNumber="267" unit="mm" value="3.1">3.1 mm</quantity>
),
<collectionCode id="ED22A4D98259FFE55101FEC6162CFEA6" box="[1029,1127,332,358]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="6" pageNumber="267" type="Museum">FLMNH</collectionCode>
<specimenCode id="DB9594678259FFE5516BFEC616AFFEA6" box="[1135,1252,331,358]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="6" pageNumber="267" type="Museum">UF 35982</specimenCode>
, Red Sea,
<collectingCountry id="F3247C8C8259FFE55058FEC112ADFE49" name="Saudi Arabia" pageId="6" pageNumber="267">Saudi Arabia</collectingCountry>
,
<collectingCounty id="62ED44908259FFE555F7FEE5104AFE49" box="[243,513,367,394]" pageId="6" pageNumber="267">northern Farasan Banks</collectingCounty>
off Al-Lith, Whale Shark Reef,
<geoCoordinate id="EE075ADB8259FFE55671FEE5165BFE49" box="[885,1040,367,393]" degrees="20" direction="north" minutes="07" orientation="latitude" pageId="6" pageNumber="267" precision="1" seconds="01.2" value="20.117">20°07'01.2&quot;N</geoCoordinate>
,
<geoCoordinate id="EE075ADB8259FFE5511AFEE516FDFE4A" box="[1054,1206,367,394]" degrees="40" direction="east" minutes="12" orientation="longitude" pageId="6" pageNumber="267" precision="1" seconds="54.0" value="40.215">40°12'54.0&quot;E</geoCoordinate>
, offshore coral reef, depth:
<quantity id="4CCB91F98259FFE555E7FE191355FE6E" box="[227,286,403,430]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="267" unit="m" value="10.0">
<elevation id="001EDB2F8259FFE555E7FE191355FE6E" box="[227,286,403,430]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="267" unit="m" value="10.0">10 m</elevation>
</quantity>
, leg.
<collectorName id="26C659CA8259FFE55452FE1913F7FE6D" box="[342,444,403,429]" pageId="6" pageNumber="267">A. Anker</collectorName>
<emphasis id="B947E00E8259FFE554C6FE1E13B1FE6D" box="[450,506,403,429]" italics="true" pageId="6" pageNumber="267">et al.</emphasis>
,
<date id="FF8D1ADC8259FFE55703FE1910CEFE6D" box="[519,645,403,429]" pageId="6" pageNumber="267" value="2013-03-22">
<collectingDate id="EFC9E3348259FFE55703FE1910CEFE6D" box="[519,645,403,429]" pageId="6" pageNumber="267" value="2013-03-22">22.03.2013</collectingDate>
</date>
[fcn BDJRS-3160];
</materialsCitation>
<materialsCitation id="3B5B36418259FFE5566FFE191139FE11" ID-GBIF-Occurrence="3775741314" collectingDate="2013-03-19" collectingDateMax="2013-03-22" collectingDateMin="2013-03-19" collectionCode="FLMNH" collectorName="A. Anker" country="Saudi Arabia" county="northern Farasan Banks" elevation="15" latitude="20.117" location="Red Sea" longLatPrecision="1" longitude="40.215" municipality="Al Fahal Reef" pageId="6" pageNumber="267" specimenCode="UF 35964, UF 35982, UF 35985" specimenCount="3" typeStatus="paratype">
1 ovig. specimen (cl
<quantity id="4CCB91F98259FFE55156FE1916E0FE6E" box="[1106,1195,403,430]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="6" pageNumber="267" unit="mm" value="2.9">2.9 mm</quantity>
),
<collectionCode id="ED22A4D98259FFE551B9FE1E1754FE6E" box="[1213,1311,404,430]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="6" pageNumber="267" type="Museum">FLMNH</collectionCode>
<specimenCode id="DB9594678259FFE55021FE1E17D3FE6E" box="[1317,1432,403,430]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="6" pageNumber="267" type="Museum">UF 35985</specimenCode>
, same collection data as for previous
<typeStatus id="548882BE8259FFE55734FE3210DBFE12" box="[560,656,440,466]" pageId="6" pageNumber="267" type="paratype">paratype</typeStatus>
[fcn BDJRS-3163].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3296F978259FFEA55C3FE511004F8D4" lastPageId="9" lastPageNumber="270" pageId="6" pageNumber="267" type="description">
<paragraph id="8B8C3C1C8259FFE555C3FE51165CFD0D" blockId="6.[151,1437,259,1978]" pageId="6" pageNumber="267">
<emphasis id="B947E00E8259FFE555C3FE51131AFE35" bold="true" box="[199,337,475,501]" pageId="6" pageNumber="267">Description</emphasis>
. Small-sized shrimp with moderately slender, non-compressed body. Carapace (
<figureCitation id="130820998259FFE551CFFE51177EFE36" box="[1227,1333,475,502]" captionStart="FIGURE 4" captionStartId="7.[151,250,1874,1898]" captionTargetBox="[215,1373,199,1839]" captionTargetId="figure-15@7.[201,1386,181,1851]" captionTargetPageId="7" captionText="FIGURE 4. Salmoneus arabicus sp. nov., holotype, ovigerous specimen (cl 2.8 mm) from Al Fahal Reef off Thuwal, Saudi Arabia (FLMNH UF 35964): a—frontal region, dorsal; b—same, lateral view; c—fourth, fifth and sixth pleonites, lateral; d—telson, lateral; e—ventromesial tooth of first article of antennular peduncle, lateral; f—third maxilliped, lateral; g—second pereiopod, lateral; h—third pereiopod, lateral; i—same, distal portion of propodus and dactylus, mesial; j—fifth pereiopod, lateral; k—same, propodus and dactylus, mesial." figureDoi="http://doi.org/10.5281/zenodo.6559671" httpUri="https://zenodo.org/record/6559671/files/figure.png" pageId="6" pageNumber="267">Fig. 4a, b</figureCitation>
) smooth, glabrous, completely covering eyes dorsally, only partly concealing them laterally; anterolateral suture present; pterygostomial angle broadly rounded; cardiac notch deep. Rostrum (
<figureCitation id="130820998259FFE5569AFDA91642FDFE" box="[926,1033,547,574]" captionStart="FIGURE 4" captionStartId="7.[151,250,1874,1898]" captionTargetBox="[215,1373,199,1839]" captionTargetId="figure-15@7.[201,1386,181,1851]" captionTargetPageId="7" captionText="FIGURE 4. Salmoneus arabicus sp. nov., holotype, ovigerous specimen (cl 2.8 mm) from Al Fahal Reef off Thuwal, Saudi Arabia (FLMNH UF 35964): a—frontal region, dorsal; b—same, lateral view; c—fourth, fifth and sixth pleonites, lateral; d—telson, lateral; e—ventromesial tooth of first article of antennular peduncle, lateral; f—third maxilliped, lateral; g—second pereiopod, lateral; h—third pereiopod, lateral; i—same, distal portion of propodus and dactylus, mesial; j—fifth pereiopod, lateral; k—same, propodus and dactylus, mesial." figureDoi="http://doi.org/10.5281/zenodo.6559671" httpUri="https://zenodo.org/record/6559671/files/figure.png" pageId="6" pageNumber="267">Fig. 4a, b</figureCitation>
) almost perfectly triangular, as long as broad, with acute tip, latter distinctly overreaching distal margin of first article of antennular peduncle; lateral margins straight to faintly convex; rostral carina short, low, not distinct in posterior half of rostrum; ventral margin unarmed. Orbital teeth (
<figureCitation id="130820998259FFE554AAFD051069FD6A" box="[430,546,655,682]" captionStart="FIGURE 4" captionStartId="7.[151,250,1874,1898]" captionTargetBox="[215,1373,199,1839]" captionTargetId="figure-15@7.[201,1386,181,1851]" captionTargetPageId="7" captionText="FIGURE 4. Salmoneus arabicus sp. nov., holotype, ovigerous specimen (cl 2.8 mm) from Al Fahal Reef off Thuwal, Saudi Arabia (FLMNH UF 35964): a—frontal region, dorsal; b—same, lateral view; c—fourth, fifth and sixth pleonites, lateral; d—telson, lateral; e—ventromesial tooth of first article of antennular peduncle, lateral; f—third maxilliped, lateral; g—second pereiopod, lateral; h—third pereiopod, lateral; i—same, distal portion of propodus and dactylus, mesial; j—fifth pereiopod, lateral; k—same, propodus and dactylus, mesial." figureDoi="http://doi.org/10.5281/zenodo.6559671" httpUri="https://zenodo.org/record/6559671/files/figure.png" pageId="6" pageNumber="267">Fig. 4a, b</figureCitation>
) well developed, about 0.2 of rostrum length, narrow, acute, extending well beyond eyes, directed slightly mesially in dorsal view, anteriorly in lateral view.
</paragraph>
<paragraph id="8B8C3C1C8259FFE555C3FD5D10FBFCD6" blockId="6.[151,1437,259,1978]" pageId="6" pageNumber="267">
Eyestalks with cornea occupying most of distal surface; anterodorsal margin without triangular process (
<figureCitation id="130820998259FFE5504CFD5D12E3FCD6" captionStart="FIGURE 4" captionStartId="7.[151,250,1874,1898]" captionTargetBox="[215,1373,199,1839]" captionTargetId="figure-15@7.[201,1386,181,1851]" captionTargetPageId="7" captionText="FIGURE 4. Salmoneus arabicus sp. nov., holotype, ovigerous specimen (cl 2.8 mm) from Al Fahal Reef off Thuwal, Saudi Arabia (FLMNH UF 35964): a—frontal region, dorsal; b—same, lateral view; c—fourth, fifth and sixth pleonites, lateral; d—telson, lateral; e—ventromesial tooth of first article of antennular peduncle, lateral; f—third maxilliped, lateral; g—second pereiopod, lateral; h—third pereiopod, lateral; i—same, distal portion of propodus and dactylus, mesial; j—fifth pereiopod, lateral; k—same, propodus and dactylus, mesial." figureDoi="http://doi.org/10.5281/zenodo.6559671" httpUri="https://zenodo.org/record/6559671/files/figure.png" pageId="6" pageNumber="267">Fig. 4a, b</figureCitation>
). Epistomial sclerites each with acute process.
</paragraph>
<paragraph id="8B8C3C1C8259FFE555C3FC95162EFC42" blockId="6.[151,1437,259,1978]" pageId="6" pageNumber="267">
Pleon (
<figureCitation id="130820998259FFE55412FC95132EFCF9" box="[278,357,799,825]" captionStart="FIGURE 4" captionStartId="7.[151,250,1874,1898]" captionTargetBox="[215,1373,199,1839]" captionTargetId="figure-15@7.[201,1386,181,1851]" captionTargetPageId="7" captionText="FIGURE 4. Salmoneus arabicus sp. nov., holotype, ovigerous specimen (cl 2.8 mm) from Al Fahal Reef off Thuwal, Saudi Arabia (FLMNH UF 35964): a—frontal region, dorsal; b—same, lateral view; c—fourth, fifth and sixth pleonites, lateral; d—telson, lateral; e—ventromesial tooth of first article of antennular peduncle, lateral; f—third maxilliped, lateral; g—second pereiopod, lateral; h—third pereiopod, lateral; i—same, distal portion of propodus and dactylus, mesial; j—fifth pereiopod, lateral; k—same, propodus and dactylus, mesial." figureDoi="http://doi.org/10.5281/zenodo.6559671" httpUri="https://zenodo.org/record/6559671/files/figure.png" pageId="6" pageNumber="267">Fig. 4c</figureCitation>
) with pleura of first to fourth pleonite rounded antero- and posteroventrally; fifth pleuron acutely produced posteroventrally; sixth pleonite with acute subtriangular projection flanking base of telson, oblique suture at posteroventral angle not distinct; preanal plate rounded, with depressed median part.
</paragraph>
<paragraph id="8B8C3C1C8259FFE555C3FC01163FFBD2" blockId="6.[151,1437,259,1978]" pageId="6" pageNumber="267">
Telson (
<figureCitation id="130820998259FFE55422FC011337FC66" box="[294,380,907,934]" captionStart="FIGURE 4" captionStartId="7.[151,250,1874,1898]" captionTargetBox="[215,1373,199,1839]" captionTargetId="figure-15@7.[201,1386,181,1851]" captionTargetPageId="7" captionText="FIGURE 4. Salmoneus arabicus sp. nov., holotype, ovigerous specimen (cl 2.8 mm) from Al Fahal Reef off Thuwal, Saudi Arabia (FLMNH UF 35964): a—frontal region, dorsal; b—same, lateral view; c—fourth, fifth and sixth pleonites, lateral; d—telson, lateral; e—ventromesial tooth of first article of antennular peduncle, lateral; f—third maxilliped, lateral; g—second pereiopod, lateral; h—third pereiopod, lateral; i—same, distal portion of propodus and dactylus, mesial; j—fifth pereiopod, lateral; k—same, propodus and dactylus, mesial." figureDoi="http://doi.org/10.5281/zenodo.6559671" httpUri="https://zenodo.org/record/6559671/files/figure.png" pageId="6" pageNumber="267">Fig. 4d</figureCitation>
) slender, somewhat elongate, subrectangular, tapering distally, about 2.6 times as long as proximal width; dorsal surface with two pairs of stout spiniform setae situated at about 0.50.6 and 0.70.8 telson length, respectively; posterior margin with two pairs of slender spiniform setae, mesial distinctly longer than lateral, and moderately deep, U-shaped median notch, latter flanked by two long plumose setae.
</paragraph>
<paragraph id="8B8C3C1C8259FFE555C3FB911338FAF1" blockId="6.[151,1437,259,1978]" pageId="6" pageNumber="267">
Antennular peduncle (
<figureCitation id="130820998259FFE554B9FB911075FBF6" box="[445,574,1051,1078]" captionStart="FIGURE 4" captionStartId="7.[151,250,1874,1898]" captionTargetBox="[215,1373,199,1839]" captionTargetId="figure-15@7.[201,1386,181,1851]" captionTargetPageId="7" captionText="FIGURE 4. Salmoneus arabicus sp. nov., holotype, ovigerous specimen (cl 2.8 mm) from Al Fahal Reef off Thuwal, Saudi Arabia (FLMNH UF 35964): a—frontal region, dorsal; b—same, lateral view; c—fourth, fifth and sixth pleonites, lateral; d—telson, lateral; e—ventromesial tooth of first article of antennular peduncle, lateral; f—third maxilliped, lateral; g—second pereiopod, lateral; h—third pereiopod, lateral; i—same, distal portion of propodus and dactylus, mesial; j—fifth pereiopod, lateral; k—same, propodus and dactylus, mesial." figureDoi="http://doi.org/10.5281/zenodo.6559671" httpUri="https://zenodo.org/record/6559671/files/figure.png" pageId="6" pageNumber="267">Fig. 4a, b, e</figureCitation>
) stout; dorsally visible portion of first article as long as wide; stylocerite stout, with subacute tip, latter reaching or slightly overreaching mid-length of second article; second article very stout, wider than long; lateral antennular flagellum biramous, with fused portion composed of two short subdivisions; shorter free ramus with at least three poorly demarcated subdivisions each furnished with bundle of aesthetascs. Antenna (
<figureCitation id="130820998259FFE5540EFB211334FB06" box="[266,383,1195,1222]" captionStart="FIGURE 4" captionStartId="7.[151,250,1874,1898]" captionTargetBox="[215,1373,199,1839]" captionTargetId="figure-15@7.[201,1386,181,1851]" captionTargetPageId="7" captionText="FIGURE 4. Salmoneus arabicus sp. nov., holotype, ovigerous specimen (cl 2.8 mm) from Al Fahal Reef off Thuwal, Saudi Arabia (FLMNH UF 35964): a—frontal region, dorsal; b—same, lateral view; c—fourth, fifth and sixth pleonites, lateral; d—telson, lateral; e—ventromesial tooth of first article of antennular peduncle, lateral; f—third maxilliped, lateral; g—second pereiopod, lateral; h—third pereiopod, lateral; i—same, distal portion of propodus and dactylus, mesial; j—fifth pereiopod, lateral; k—same, propodus and dactylus, mesial." figureDoi="http://doi.org/10.5281/zenodo.6559671" httpUri="https://zenodo.org/record/6559671/files/figure.png" pageId="6" pageNumber="267">Fig. 4a, b</figureCitation>
) with basicerite stout, armed with sharp distoventral tooth; scaphocerite not overreaching antennular peduncle, ovate, almost twice as long as broad; distolateral tooth not reaching anteriorly protruding distal margin of blade; carpocerite short, reaching half-length of scaphocerite; flagellum not particularly thickened, moderately slender.
</paragraph>
<paragraph id="8B8C3C1C8259FFE555C3FAB1136FFA25" blockId="6.[151,1437,259,1978]" pageId="6" pageNumber="267">
Third maxilliped (
<figureCitation id="130820998259FFE55495FAB11396FA96" box="[401,477,1339,1366]" captionStart="FIGURE 4" captionStartId="7.[151,250,1874,1898]" captionTargetBox="[215,1373,199,1839]" captionTargetId="figure-15@7.[201,1386,181,1851]" captionTargetPageId="7" captionText="FIGURE 4. Salmoneus arabicus sp. nov., holotype, ovigerous specimen (cl 2.8 mm) from Al Fahal Reef off Thuwal, Saudi Arabia (FLMNH UF 35964): a—frontal region, dorsal; b—same, lateral view; c—fourth, fifth and sixth pleonites, lateral; d—telson, lateral; e—ventromesial tooth of first article of antennular peduncle, lateral; f—third maxilliped, lateral; g—second pereiopod, lateral; h—third pereiopod, lateral; i—same, distal portion of propodus and dactylus, mesial; j—fifth pereiopod, lateral; k—same, propodus and dactylus, mesial." figureDoi="http://doi.org/10.5281/zenodo.6559671" httpUri="https://zenodo.org/record/6559671/files/figure.png" pageId="6" pageNumber="267">Fig. 4f</figureCitation>
) slender; coxa with strap-like epipod and rounded lateral plate; antepenultimate article about 3.2 times as long as penultimate article; penultimate article moderately long, about 2.5 times as long as wide; ultimate article relatively short (compared to antepenultimate article), tapering distally, ending in corneous apical tooth, without spiniform setae; exopod well developed, reaching 0.8 length of antepenultimate article; arthrobranch rather small.
</paragraph>
<paragraph id="8B8C3C1C8259FFE555C3FA65119DF879" blockId="6.[151,1437,259,1978]" pageId="6" pageNumber="267">
First pereiopods = chelipeds (
<figureCitation id="130820998259FFE55712FA651012F9CA" box="[534,601,1519,1546]" captionStart="FIGURE 5" captionStartId="8.[151,250,1930,1954]" captionTargetBox="[166,1420,735,1894]" captionTargetId="figure-288@8.[151,1436,722,1906]" captionTargetPageId="8" captionText="FIGURE 5. Salmoneus arabicus sp. nov., holotype, ovigerous specimen (cl 2.8 mm) from Al Fahal Reef off Thuwal, Saudi Arabia (FLMNH UF 35964): a—major (left) cheliped, mesial; b—same, lateral; c—same, chela, dorsomesial; d—minor (right) cheliped, lateral." figureDoi="http://doi.org/10.5281/zenodo.6559673" httpUri="https://zenodo.org/record/6559673/files/figure.png" pageId="6" pageNumber="267">Fig. 5</figureCitation>
) very different in shape, asymmetrical in size. Major cheliped (
<figureCitation id="130820998259FFE5502DFA6517DFF9CA" box="[1321,1428,1519,1546]" captionStart="FIGURE 5" captionStartId="8.[151,250,1930,1954]" captionTargetBox="[166,1420,735,1894]" captionTargetId="figure-288@8.[151,1436,722,1906]" captionTargetPageId="8" captionText="FIGURE 5. Salmoneus arabicus sp. nov., holotype, ovigerous specimen (cl 2.8 mm) from Al Fahal Reef off Thuwal, Saudi Arabia (FLMNH UF 35964): a—major (left) cheliped, mesial; b—same, lateral; c—same, chela, dorsomesial; d—minor (right) cheliped, lateral." figureDoi="http://doi.org/10.5281/zenodo.6559673" httpUri="https://zenodo.org/record/6559673/files/figure.png" pageId="6" pageNumber="267">Fig. 5ac</figureCitation>
) moderately robust, carried flexed under body at rest; ischium slender, about three times as long as wide, armed with one robust spiniform seta on ventrolateral surface; merus long, slender, almost straight, slightly widening distally, about 4.7 times as long as distal width, shallowly depressed ventrally; carpus cup-shaped, short, with rounded distal lobes, without acute or subacute process; chela somewhat enlarged and swollen, slightly longer than merus, with smooth surfaces; palm about 2.2 times as long as maximal width, with deep transverse groove proximo-ventrally, near carpus; fingers about 0.8 length of palm, not gaping when closed, not noticeably twisted, with gently crossing fingertips; cutting edges with about 1012 rounded teeth in form of almost continuous serration (except for edges near fingertips), proximal-most and distal-most teeth smallest, those near mid-length largest. Minor cheliped (
<figureCitation id="130820998259FFE5506BF88512FFF88E" captionStart="FIGURE 5" captionStartId="8.[151,250,1930,1954]" captionTargetBox="[166,1420,735,1894]" captionTargetId="figure-288@8.[151,1436,722,1906]" captionTargetPageId="8" captionText="FIGURE 5. Salmoneus arabicus sp. nov., holotype, ovigerous specimen (cl 2.8 mm) from Al Fahal Reef off Thuwal, Saudi Arabia (FLMNH UF 35964): a—major (left) cheliped, mesial; b—same, lateral; c—same, chela, dorsomesial; d—minor (right) cheliped, lateral." figureDoi="http://doi.org/10.5281/zenodo.6559673" httpUri="https://zenodo.org/record/6559673/files/figure.png" pageId="6" pageNumber="267">Fig. 5d</figureCitation>
) much smaller and weaker than major cheliped; ischium about 3.5 times as long as wide, armed with two stout spiniform setae on ventrolateral margin; merus slightly longer than ischium, slightly swollen, about 3.2 times as long as wide; carpus subcylindrical, slightly shorter than merus, widening distally; chela of about same length as carpus; palm as long as fingers, latter simple, with unarmed cutting edges.
</paragraph>
<caption id="DF4C6C948258FFE45593F8D810CEF83B" ID-DOI="http://doi.org/10.5281/zenodo.6559671" ID-Zenodo-Dep="6559671" httpUri="https://zenodo.org/record/6559671/files/figure.png" pageId="7" pageNumber="268" startId="7.[151,250,1874,1898]" targetBox="[215,1373,199,1839]" targetPageId="7">
<paragraph id="8B8C3C1C8258FFE45593F8D810CEF83B" blockId="7.[151,1437,1874,2043]" pageId="7" pageNumber="268">
<emphasis id="B947E00E8258FFE45593F8D8135EF8AB" bold="true" box="[151,277,1874,1899]" pageId="7" pageNumber="268">FIGURE 4.</emphasis>
<taxonomicName id="4C33479F8258FFE45419F8D913A0F8AB" authority="Anker, 2022" authorityName="Anker" authorityYear="2022" box="[285,491,1875,1899]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="268" phylum="Arthropoda" rank="species" species="arabicus" status="sp. nov.">
<emphasis id="B947E00E8258FFE45419F8D913A0F8AB" box="[285,491,1875,1899]" italics="true" pageId="7" pageNumber="268">Salmoneus arabicus</emphasis>
</taxonomicName>
<emphasis id="B947E00E8258FFE454F7F8D9100EF8AB" bold="true" box="[499,581,1875,1899]" pageId="7" pageNumber="268">
<taxonomicNameLabel id="A2745D758258FFE454F7F8D9100EF8AB" box="[499,581,1875,1899]" pageId="7" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, ovigerous specimen (cl 2.8 mm) from Al Fahal Reef off Thuwal, Saudi Arabia (FLMNH UF 35964): a—frontal region, dorsal; b—same, lateral view; c—fourth, fifth and sixth pleonites, lateral; d—telson, lateral; e—ventromesial tooth of first article of antennular peduncle, lateral; f—third maxilliped, lateral; g—second pereiopod, lateral; h—third pereiopod, lateral; i—same, distal portion of propodus and dactylus, mesial; j—fifth pereiopod, lateral; k—same, propodus and dactylus, mesial.
</paragraph>
</caption>
<paragraph id="8B8C3C1C8257FFEB55C3FF1D1366FED9" blockId="8.[151,1437,151,699]" pageId="8" pageNumber="269">
Second pereiopod (
<figureCitation id="130820998257FFEB54A4FF1D13B9FF71" box="[416,498,151,177]" captionStart="FIGURE 4" captionStartId="7.[151,250,1874,1898]" captionTargetBox="[215,1373,199,1839]" captionTargetId="figure-15@7.[201,1386,181,1851]" captionTargetPageId="7" captionText="FIGURE 4. Salmoneus arabicus sp. nov., holotype, ovigerous specimen (cl 2.8 mm) from Al Fahal Reef off Thuwal, Saudi Arabia (FLMNH UF 35964): a—frontal region, dorsal; b—same, lateral view; c—fourth, fifth and sixth pleonites, lateral; d—telson, lateral; e—ventromesial tooth of first article of antennular peduncle, lateral; f—third maxilliped, lateral; g—second pereiopod, lateral; h—third pereiopod, lateral; i—same, distal portion of propodus and dactylus, mesial; j—fifth pereiopod, lateral; k—same, propodus and dactylus, mesial." figureDoi="http://doi.org/10.5281/zenodo.6559671" httpUri="https://zenodo.org/record/6559671/files/figure.png" pageId="8" pageNumber="269">
Fig.
<quantity id="4CCB91F98257FFEB54D1FF1D13B9FF71" box="[469,498,151,177]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="8" pageNumber="269" unit="g" value="4.0">4g</quantity>
</figureCitation>
) slender; ischium elongate, almost seven times as long as wide, with one spiniform seta on ventrolateral surface, at about 0.30.4 of ischial length; merus slightly longer than ischium; carpus with five subarticles, proximal equal to sum of remaining four; length ratio of carpal subarticles approximately equal to 6: 2: 1: 1.5: 2.2.
</paragraph>
<paragraph id="8B8C3C1C8257FFEB55C3FEAB16FAFD7A" blockId="8.[151,1437,151,699]" pageId="8" pageNumber="269">
Third pereiopod (
<figureCitation id="130820998257FFEB5488FEAB13B8FEFC" box="[396,499,289,316]" captionStart="FIGURE 4" captionStartId="7.[151,250,1874,1898]" captionTargetBox="[215,1373,199,1839]" captionTargetId="figure-15@7.[201,1386,181,1851]" captionTargetPageId="7" captionText="FIGURE 4. Salmoneus arabicus sp. nov., holotype, ovigerous specimen (cl 2.8 mm) from Al Fahal Reef off Thuwal, Saudi Arabia (FLMNH UF 35964): a—frontal region, dorsal; b—same, lateral view; c—fourth, fifth and sixth pleonites, lateral; d—telson, lateral; e—ventromesial tooth of first article of antennular peduncle, lateral; f—third maxilliped, lateral; g—second pereiopod, lateral; h—third pereiopod, lateral; i—same, distal portion of propodus and dactylus, mesial; j—fifth pereiopod, lateral; k—same, propodus and dactylus, mesial." figureDoi="http://doi.org/10.5281/zenodo.6559671" httpUri="https://zenodo.org/record/6559671/files/figure.png" pageId="8" pageNumber="269">Fig. 4h, i</figureCitation>
) moderately slender; ischium about 3.5 times as long as wide, with three spiniform setae on ventrolateral surface; merus about 1.4 times as long as ischium, about six times as long as maximal width; carpus distinctly slenderer and shorter than merus, with small spiniform seta on distoventral margin; propodus subequal to carpus in length, with four slender spiniform setae on ventral margin and one pair of longer spiniform setae on distoventral margin, near dactylar base; dactylus moderately slender, gently curved, conical, about 0.4 length of propodus. Fourth pereiopod generally similar to third pereiopod, slenderer; ischium with two or three spiniform setae on distoventral margin. Fifth pereiopod (
<figureCitation id="130820998257FFEB5794FE7B10BFFDCC" box="[656,756,497,524]" captionStart="FIGURE 4" captionStartId="7.[151,250,1874,1898]" captionTargetBox="[215,1373,199,1839]" captionTargetId="figure-15@7.[201,1386,181,1851]" captionTargetPageId="7" captionText="FIGURE 4. Salmoneus arabicus sp. nov., holotype, ovigerous specimen (cl 2.8 mm) from Al Fahal Reef off Thuwal, Saudi Arabia (FLMNH UF 35964): a—frontal region, dorsal; b—same, lateral view; c—fourth, fifth and sixth pleonites, lateral; d—telson, lateral; e—ventromesial tooth of first article of antennular peduncle, lateral; f—third maxilliped, lateral; g—second pereiopod, lateral; h—third pereiopod, lateral; i—same, distal portion of propodus and dactylus, mesial; j—fifth pereiopod, lateral; k—same, propodus and dactylus, mesial." figureDoi="http://doi.org/10.5281/zenodo.6559671" httpUri="https://zenodo.org/record/6559671/files/figure.png" pageId="8" pageNumber="269">Fig. 4j, k</figureCitation>
) slenderer than third and fourth pereiopods; ischium about 3.5 times as long as wide, unarmed; merus about seven times as long as maximal width; carpus about as long as merus, much slenderer, with unarmed distoventral margin; propodus about 1.3 times as long as carpus, with four widely spaced spiniform setae along ventral margin and two longer spiniform setae on distoventral margin, near dactylar base; propodal cleaning brush well developed, composed of dozen or so transverse rows of microserrulate setae occupying distal third of propodus; dactylus slender, slightly curved, conical, about 0.3 length of propodus.
</paragraph>
<caption id="DF4C6C948257FFEB5593F800130BF82A" ID-DOI="http://doi.org/10.5281/zenodo.6559673" ID-Zenodo-Dep="6559673" httpUri="https://zenodo.org/record/6559673/files/figure.png" pageId="8" pageNumber="269" startId="8.[151,250,1930,1954]" targetBox="[166,1420,735,1894]" targetPageId="8">
<paragraph id="8B8C3C1C8257FFEB5593F800130BF82A" blockId="8.[151,1437,1930,2026]" pageId="8" pageNumber="269">
<emphasis id="B947E00E8257FFEB5593F800135EF862" bold="true" box="[151,277,1930,1954]" pageId="8" pageNumber="269">FIGURE 5.</emphasis>
<taxonomicName id="4C33479F8257FFEB5419F80013A0F862" authority="Anker, 2022" authorityName="Anker" authorityYear="2022" box="[285,491,1930,1954]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="269" phylum="Arthropoda" rank="species" species="arabicus" status="sp. nov.">
<emphasis id="B947E00E8257FFEB5419F80013A0F862" box="[285,491,1930,1954]" italics="true" pageId="8" pageNumber="269">Salmoneus arabicus</emphasis>
</taxonomicName>
<emphasis id="B947E00E8257FFEB54F7F800100EF862" bold="true" box="[499,581,1930,1954]" pageId="8" pageNumber="269">
<taxonomicNameLabel id="A2745D758257FFEB54F7F800100EF862" box="[499,581,1930,1954]" pageId="8" pageNumber="269" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, ovigerous specimen (cl 2.8 mm) from Al Fahal Reef off Thuwal, Saudi Arabia (FLMNH UF 35964): a—major (left) cheliped, mesial; b—same, lateral; c—same, chela, dorsomesial; d—minor (right) cheliped, lateral.
</paragraph>
</caption>
<caption id="DF4C6C948256FFEA5593FA5016C5F9D6" ID-DOI="http://doi.org/10.5281/zenodo.6559675" ID-Zenodo-Dep="6559675" httpUri="https://zenodo.org/record/6559675/files/figure.png" pageId="9" pageNumber="270" startId="9.[151,250,1498,1522]" targetBox="[151,1436,181,1474]" targetPageId="9">
<paragraph id="8B8C3C1C8256FFEA5593FA5016C5F9D6" blockId="9.[151,1437,1498,1559]" pageId="9" pageNumber="270">
<emphasis id="B947E00E8256FFEA5593FA50135FFA32" bold="true" box="[151,276,1498,1522]" pageId="9" pageNumber="270">FIGURE 6.</emphasis>
<taxonomicName id="4C33479F8256FFEA5419FA5013A0FA32" authority="Anker, 2022" authorityName="Anker" authorityYear="2022" box="[285,491,1498,1522]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="270" phylum="Arthropoda" rank="species" species="arabicus" status="sp. nov.">
<emphasis id="B947E00E8256FFEA5419FA5013A0FA32" box="[285,491,1498,1522]" italics="true" pageId="9" pageNumber="270">Salmoneus arabicus</emphasis>
</taxonomicName>
<emphasis id="B947E00E8256FFEA54F7FA51100EFA33" bold="true" box="[499,581,1499,1523]" pageId="9" pageNumber="270">
<taxonomicNameLabel id="A2745D758256FFEA54F7FA51100EFA33" box="[499,581,1499,1523]" pageId="9" pageNumber="270" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, ovigerous specimen (cl 2.8 mm) from Al Fahal Reef off Thuwal, Saudi Arabia (FLMNH UF 35964): a—shrimp in life, dorsal; b—same, lateral. Photographs by the author.
</paragraph>
</caption>
<paragraph id="8B8C3C1C8256FFEA55C3F9CF114BF90B" blockId="9.[151,1436,1605,2027]" pageId="9" pageNumber="270">
Second pleopod with appendix masculina of about same length as appendix interna. Uropod (
<figureCitation id="130820998256FFEA51CCF9CF1759F9A0" box="[1224,1298,1605,1632]" captionStart="FIGURE 4" captionStartId="7.[151,250,1874,1898]" captionTargetBox="[215,1373,199,1839]" captionTargetId="figure-15@7.[201,1386,181,1851]" captionTargetPageId="7" captionText="FIGURE 4. Salmoneus arabicus sp. nov., holotype, ovigerous specimen (cl 2.8 mm) from Al Fahal Reef off Thuwal, Saudi Arabia (FLMNH UF 35964): a—frontal region, dorsal; b—same, lateral view; c—fourth, fifth and sixth pleonites, lateral; d—telson, lateral; e—ventromesial tooth of first article of antennular peduncle, lateral; f—third maxilliped, lateral; g—second pereiopod, lateral; h—third pereiopod, lateral; i—same, distal portion of propodus and dactylus, mesial; j—fifth pereiopod, lateral; k—same, propodus and dactylus, mesial." figureDoi="http://doi.org/10.5281/zenodo.6559671" httpUri="https://zenodo.org/record/6559671/files/figure.png" pageId="9" pageNumber="270">Fig. 4l</figureCitation>
) with lateral lobe of protopod produced into sharp tooth; exopod moderately broad, ovate, with long spiniform seta flanked by small subacute distolateral tooth and subtriangular lateral tooth of somewhat sinuous diaeresis; endopod as long as exopod, noticeably narrower, without specific features.
</paragraph>
<paragraph id="8B8C3C1C8256FFEA55C3F95E1004F8D4" blockId="9.[151,1436,1605,2027]" pageId="9" pageNumber="270">
<emphasis id="B947E00E8256FFEA55C3F95E132CF92E" bold="true" box="[199,359,1748,1774]" pageId="9" pageNumber="270">Colour in life</emphasis>
. Semi-transparent whitish; yellow-orange inner organs visible due to translucence of carapace; freshly laid eggs bright orange (
<figureCitation id="130820998256FFEA54F9F973100AF8D4" box="[509,577,1785,1812]" captionStart="FIGURE 6" captionStartId="9.[151,250,1498,1522]" captionTargetBox="[151,1436,181,1474]" captionTargetId="figure-15@9.[151,1436,181,1474]" captionTargetPageId="9" captionText="FIGURE 6. Salmoneus arabicus sp. nov., holotype, ovigerous specimen (cl 2.8 mm) from Al Fahal Reef off Thuwal, Saudi Arabia (FLMNH UF 35964): a—shrimp in life, dorsal; b—same, lateral. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.6559675" httpUri="https://zenodo.org/record/6559675/files/figure.png" pageId="9" pageNumber="270">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3296F978256FFEA55C3F8971021F8F7" box="[199,618,1820,1847]" pageId="9" pageNumber="270" type="materials_examined">
<paragraph id="8B8C3C1C8256FFEA55C3F8971021F8F7" blockId="9.[151,1436,1605,2027]" box="[199,618,1820,1847]" pageId="9" pageNumber="270">
<materialsCitation id="3B5B36418256FFEA55C3F897102DF8F7" ID-GBIF-Occurrence="3775741312" box="[199,614,1820,1847]" country="Saudi Arabia" location="Thuwal" pageId="9" pageNumber="270" specimenCount="1" typeStatus="holotype">
<emphasis id="B947E00E8256FFEA55C3F897132BF8F6" bold="true" box="[199,352,1820,1847]" pageId="9" pageNumber="270">
<typeStatus id="548882BE8256FFEA55C3F8971349F8F7" box="[199,258,1821,1847]" pageId="9" pageNumber="270">Type</typeStatus>
locality
</emphasis>
.
<location id="8EEC6AC78256FFEA5469F897138EF8F7" LSID="urn:lsid:plazi:treatment:039A8D0A8259FFE95593FF1D101AFBD1:8EEC6AC78256FFEA5469F897138EF8F7" box="[365,453,1821,1847]" country="Saudi Arabia" name="Thuwal" pageId="9" pageNumber="270">Thuwal</location>
,
<collectingCountry id="F3247C8C8256FFEA54D5F897102DF8F7" box="[465,614,1821,1847]" name="Saudi Arabia" pageId="9" pageNumber="270">Saudi Arabia</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3296F978256FFEA55C3F8CA137BF8BF" pageId="9" pageNumber="270" type="distribution">
<paragraph id="8B8C3C1C8256FFEA55C3F8CA137BF8BF" blockId="9.[151,1436,1605,2027]" pageId="9" pageNumber="270">
<emphasis id="B947E00E8256FFEA55C3F8CA1312F89A" bold="true" box="[199,345,1856,1882]" pageId="9" pageNumber="270">Distribution</emphasis>
. Red Sea: presently known only from the reefs off Thuwal and Al-Lith (northern Farasan Banks),
<collectingCountry id="F3247C8C8256FFEA5593F8EF1367F8BF" box="[151,300,1893,1919]" name="Saudi Arabia" pageId="9" pageNumber="270">Saudi Arabia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3296F978256FFEA55C3F80317DEF863" box="[199,1429,1929,1955]" pageId="9" pageNumber="270" type="biology_ecology">
<paragraph id="8B8C3C1C8256FFEA55C3F80317DEF863" blockId="9.[151,1436,1605,2027]" box="[199,1429,1929,1955]" pageId="9" pageNumber="270">
<emphasis id="B947E00E8256FFEA55C3F803136DF863" bold="true" box="[199,294,1929,1955]" pageId="9" pageNumber="270">Ecology</emphasis>
. The
<typeStatus id="548882BE8256FFEA5462F80313DDF863" box="[358,406,1929,1955]" pageId="9" pageNumber="270">type</typeStatus>
specimens were collected under coral rubble on offshore coral reefs, at depths of
<quantity id="4CCB91F98256FFEA502EF80317DBF863" box="[1322,1424,1929,1955]" metricMagnitude="1" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="9" pageNumber="270" unit="m" value="15.0" valueMax="20.0" valueMin="10.0">1020 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C3296F978256FFEA55C3F8271348F82B" pageId="9" pageNumber="270" type="etymology">
<paragraph id="8B8C3C1C8256FFEA55C3F8271348F82B" blockId="9.[151,1436,1605,2027]" pageId="9" pageNumber="270">
<emphasis id="B947E00E8256FFEA55C3F8271303F807" bold="true" box="[199,328,1965,1991]" pageId="9" pageNumber="270">Etymology</emphasis>
. The new species name alludes to
<collectingCountry id="F3247C8C8256FFEA57F0F82711C6F807" box="[756,909,1965,1991]" name="Saudi Arabia" pageId="9" pageNumber="270">Saudi Arabia</collectingCountry>
, the country of the
<typeStatus id="548882BE8256FFEA5179F82716E6F807" box="[1149,1197,1965,1991]" pageId="9" pageNumber="270">type</typeStatus>
locality; used as an adjective.
</paragraph>
</subSubSection>
<subSubSection id="C3296F978255FFE955C3FF1D101AFBD1" pageId="10" pageNumber="271" type="discussion">
<paragraph id="8B8C3C1C8255FFE955C3FF1D165EFC41" blockId="10.[151,1437,151,1041]" pageId="10" pageNumber="271">
<emphasis id="B947E00E8255FFE955C3FF1D137FFF71" bold="true" box="[199,308,151,177]" pageId="10" pageNumber="271">Remarks</emphasis>
.
<taxonomicName id="4C33479F8255FFE95445FF1D1068FF71" authority="Anker, 2022" authorityName="Anker" authorityYear="2022" box="[321,547,151,177]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="arabicus" status="sp. nov.">
<emphasis id="B947E00E8255FFE95445FF1D1068FF71" box="[321,547,151,177]" italics="true" pageId="10" pageNumber="271">Salmoneus arabicus</emphasis>
</taxonomicName>
<emphasis id="B947E00E8255FFE9572DFF1D10CAFF71" bold="true" box="[553,641,151,177]" pageId="10" pageNumber="271">
<taxonomicNameLabel id="A2745D758255FFE9572DFF1D10CAFF71" box="[553,641,151,177]" pageId="10" pageNumber="271" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
appears to be morphologically closest to
<taxonomicName id="4C33479F8255FFE95149FF1D169EFF71" baseAuthorityName="Coutiere" baseAuthorityYear="1897" box="[1101,1237,151,177]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="latirostris">
<emphasis id="B947E00E8255FFE95149FF1D169EFF71" box="[1101,1237,151,177]" italics="true" pageId="10" pageNumber="271">S. latirostris</emphasis>
</taxonomicName>
,
<taxonomicName id="4C33479F8255FFE951E6FF1D13DBFF15" authority="Anker &amp; Marin, 2006" authorityName="Anker &amp; Marin" authorityYear="2006" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="nhatrangensis">
<emphasis id="B947E00E8255FFE951E6FF1D17D6FF71" box="[1250,1437,151,177]" italics="true" pageId="10" pageNumber="271">S. nhatrangensis</emphasis>
<bibRefCitation id="EFA241ED8255FFE95593FF3113DBFF15" author="Anker, A. &amp; Marin, I. N." box="[151,400,187,213]" pageId="10" pageNumber="271" pagination="295 - 319" refId="ref10562" refString="Anker, A. &amp; Marin, I. N. (2006) New records and species of Alpheidae (Crustacea, Decapoda) from Vietnam. Part I. Genus Salmoneus Holthuis, 1955. Raffles Bulletin of Zoology, 54, 295 - 319." type="journal article" year="2006">Anker &amp; Marin, 2006</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C33479F8255FFE95498FF31101BFF15" box="[412,592,187,213]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="serratidigitus">
<emphasis id="B947E00E8255FFE95498FF31101BFF15" box="[412,592,187,213]" italics="true" pageId="10" pageNumber="271">S. serratidigitus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C33479F8255FFE9575AFF3110FBFF14" authorityName="Manning &amp; Chace" authorityYear="1990" box="[606,688,187,213]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="teres">
<emphasis id="B947E00E8255FFE9575AFF3110FBFF14" box="[606,688,187,213]" italics="true" pageId="10" pageNumber="271">S. teres</emphasis>
</taxonomicName>
,
<taxonomicName id="4C33479F8255FFE957BBFF31113FFF14" box="[703,884,187,213]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="inconspucuus">
<emphasis id="B947E00E8255FFE957BBFF31113FFF14" box="[703,884,187,213]" italics="true" pageId="10" pageNumber="271">S. inconspucuus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C33479F8255FFE95687FF3116F4FF15" authority="Anker, 2010" authorityName="Anker" authorityYear="2010" box="[899,1215,187,213]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="camaroncito">
<emphasis id="B947E00E8255FFE95687FF311667FF14" box="[899,1068,187,213]" italics="true" pageId="10" pageNumber="271">S. camaroncito</emphasis>
<bibRefCitation id="EFA241ED8255FFE95137FF3116F4FF15" author="Anker, A." box="[1075,1215,187,213]" pageId="10" pageNumber="271" pagination="177 - 205" refId="ref10192" refString="Anker, A. (2010) The shrimp genus Salmoneus Holthuis, 1955 (Crustacea, Decapoda, Alpheidae) in the tropical western Atlantic, with description of five new species. Zootaxa, 2372 (1), 177 - 205. https: // doi. org / 10.11646 / zootaxa. 2372.1.18" type="journal article" year="2010">Anker, 2010</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C33479F8255FFE951F3FF31136CFF39" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="saotomensis">
<emphasis id="B947E00E8255FFE951F3FF3117D7FF14" box="[1271,1436,187,213]" italics="true" pageId="10" pageNumber="271">S. saotomensis</emphasis>
Anker, 2020
</taxonomicName>
, especially in the presence of a basally broad rostrum with a short, not posteriorly extending carina, the dorsally completely or almost completely covered eyes, and the general shape of the chelipeds (
<bibRefCitation id="EFA241ED8255FFE951F7FE8917DCFEDD" author="Coutiere, H." box="[1267,1431,259,285]" pageId="10" pageNumber="271" pagination="1 - 559" refId="ref10857" refString="Coutiere, H. (1899) Les Alpheidae. Morphologie externe et interne, formes larvaires, bionomie. Annales des Sciences Naturelles, Zoologie et Paleontologie, 8 (9), 1 - 559, 6 pls. Masson, Paris. https: // doi. org / 10.5962 / bhl. title. 13143" type="journal article" year="1899">Coutière 1899</bibRefCitation>
;
<bibRefCitation id="EFA241ED8255FFE95593FEAD13E5FE81" author="Manning, R. B. &amp; Chace, F. A. Jr." box="[151,430,295,321]" pageId="10" pageNumber="271" pagination="1 - 91" refId="ref11355" refString="Manning, R. B. &amp; Chace, F. A. Jr. (1990) Decapod and stomatopod Crustacea from Ascension Island, South Atlantic Ocean. Smithsonian Contributions to Zoology, 503, 1 - 91. https: // doi. org / 10.5479 / si. 00810282.503" type="journal article" year="1990">Manning &amp; Chace 1990</bibRefCitation>
;
<bibRefCitation id="EFA241ED8255FFE954BFFEAD10F9FE81" author="Anker, A. &amp; Marin, I. N." box="[443,690,295,321]" pageId="10" pageNumber="271" pagination="295 - 319" refId="ref10562" refString="Anker, A. &amp; Marin, I. N. (2006) New records and species of Alpheidae (Crustacea, Decapoda) from Vietnam. Part I. Genus Salmoneus Holthuis, 1955. Raffles Bulletin of Zoology, 54, 295 - 319." type="journal article" year="2006">Anker &amp; Marin 2006</bibRefCitation>
;
<bibRefCitation id="EFA241ED8255FFE957BAFEAD1103FE81" author="Anker, A." box="[702,840,295,321]" pageId="10" pageNumber="271" pagination="21 - 39" refId="ref10136" refString="Anker, A. (2007) New species and records of alpheid shrimps, genera Salmoneus Holthuis and Parabetaeus Coutiere, from the tropical western Atlantic (Decapoda, Caridea). Zootaxa, 1653 (1), 21 - 39. https: // doi. org / 10.11646 / zootaxa. 1653.1.2" type="journal article" year="2007">Anker 2007</bibRefCitation>
,
<bibRefCitation id="EFA241ED8255FFE95651FEAD11DBFE81" author="Anker, A." box="[853,912,295,321]" pageId="10" pageNumber="271" pagination="177 - 205" refId="ref10192" refString="Anker, A. (2010) The shrimp genus Salmoneus Holthuis, 1955 (Crustacea, Decapoda, Alpheidae) in the tropical western Atlantic, with description of five new species. Zootaxa, 2372 (1), 177 - 205. https: // doi. org / 10.11646 / zootaxa. 2372.1.18" type="journal article" year="2010">2010</bibRefCitation>
, 2020). However,
<taxonomicName id="4C33479F8255FFE95160FEAD16AFFE81" authority="Anker, 2022" authorityName="Anker" authorityYear="2022" box="[1124,1252,295,321]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="arabicus" status="sp. nov.">
<emphasis id="B947E00E8255FFE95160FEAD16AFFE81" box="[1124,1252,295,321]" italics="true" pageId="10" pageNumber="271">S. arabicus</emphasis>
</taxonomicName>
<emphasis id="B947E00E8255FFE951E9FEAD1702FE81" bold="true" box="[1261,1353,295,321]" pageId="10" pageNumber="271">
<taxonomicNameLabel id="A2745D758255FFE951E9FEAD1702FE81" box="[1261,1353,295,321]" pageId="10" pageNumber="271" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be separated from
<taxonomicName id="4C33479F8255FFE9544CFEC11399FEA5" baseAuthorityName="Coutiere" baseAuthorityYear="1897" box="[328,466,331,357]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="latirostris">
<emphasis id="B947E00E8255FFE9544CFEC11399FEA5" box="[328,466,331,357]" italics="true" pageId="10" pageNumber="271">S. latirostris</emphasis>
</taxonomicName>
,
<taxonomicName id="4C33479F8255FFE954E5FEC1107FFEA4" authorityName="Manning &amp; Chace" authorityYear="1990" box="[481,564,331,357]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="teres">
<emphasis id="B947E00E8255FFE954E5FEC1107FFEA4" box="[481,564,331,357]" italics="true" pageId="10" pageNumber="271">S. teres</emphasis>
</taxonomicName>
and
<taxonomicName id="4C33479F8255FFE95769FEC11157FEA4" authorityName="Anker" authorityYear="2020" box="[621,796,331,357]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="inconspicuus">
<emphasis id="B947E00E8255FFE95769FEC11157FEA4" box="[621,796,331,357]" italics="true" pageId="10" pageNumber="271">S. inconspicuus</emphasis>
</taxonomicName>
by the more triangular rostrum, with the lateral margins almost straight (
<emphasis id="B947E00E8255FFE9544DFEE5132BFE48" box="[329,352,367,392]" italics="true" pageId="10" pageNumber="271">vs</emphasis>
. proximally convex in
<taxonomicName id="4C33479F8255FFE95758FEE510A8FE49" baseAuthorityName="Coutiere" baseAuthorityYear="1897" box="[604,739,367,393]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="latirostris">
<emphasis id="B947E00E8255FFE95758FEE510A8FE49" box="[604,739,367,393]" italics="true" pageId="10" pageNumber="271">S. latirostris</emphasis>
</taxonomicName>
); from
<taxonomicName id="4C33479F8255FFE95630FEE511F0FE49" baseAuthorityName="Coutiere" baseAuthorityYear="1897" box="[820,955,367,393]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="latirostris">
<emphasis id="B947E00E8255FFE95630FEE511F0FE49" box="[820,955,367,393]" italics="true" pageId="10" pageNumber="271">S. latirostris</emphasis>
</taxonomicName>
and
<taxonomicName id="4C33479F8255FFE956E9FEE516D5FE49" box="[1005,1182,367,393]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="serratidigitus">
<emphasis id="B947E00E8255FFE956E9FEE516D5FE49" box="[1005,1182,367,393]" italics="true" pageId="10" pageNumber="271">S. serratidigitus</emphasis>
</taxonomicName>
by the uniform whitish colouration (
<emphasis id="B947E00E8255FFE9542DFE19130BFE6C" box="[297,320,403,428]" italics="true" pageId="10" pageNumber="271">vs</emphasis>
. red banded in
<taxonomicName id="4C33479F8255FFE954F7FE191035FE6D" baseAuthorityName="Coutiere" baseAuthorityYear="1897" box="[499,638,403,429]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="latirostris">
<emphasis id="B947E00E8255FFE954F7FE191035FE6D" box="[499,638,403,429]" italics="true" pageId="10" pageNumber="271">S. latirostris</emphasis>
</taxonomicName>
and bright yellow-orange in
<taxonomicName id="4C33479F8255FFE956C8FE1916CBFE6D" box="[972,1152,403,429]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="serratidigitus">
<emphasis id="B947E00E8255FFE956C8FE1916CBFE6D" box="[972,1152,403,429]" italics="true" pageId="10" pageNumber="271">S. serratidigitus</emphasis>
</taxonomicName>
); from
<taxonomicName id="4C33479F8255FFE951DDFE1917DDFE6D" authorityName="Anker &amp; Marin" authorityYear="2006" box="[1241,1430,403,429]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="nhatrangensis">
<emphasis id="B947E00E8255FFE951DDFE1917DDFE6D" box="[1241,1430,403,429]" italics="true" pageId="10" pageNumber="271">S. nhatrangensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C33479F8255FFE95593FE3D1307FE11" box="[151,332,439,465]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="serratidigitus">
<emphasis id="B947E00E8255FFE95593FE3D1307FE11" box="[151,332,439,465]" italics="true" pageId="10" pageNumber="271">S. serratidigitus</emphasis>
</taxonomicName>
and possibly
<taxonomicName id="4C33479F8255FFE954EAFE3D1032FE11" baseAuthorityName="Coutiere" baseAuthorityYear="1897" box="[494,633,439,465]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="latirostris">
<emphasis id="B947E00E8255FFE954EAFE3D1032FE11" box="[494,633,439,465]" italics="true" pageId="10" pageNumber="271">S. latirostris</emphasis>
</taxonomicName>
by the ischia of the major and minor cheliped and the ischium of the second pereiopod armed with spiniform setae (
<emphasis id="B947E00E8255FFE957ACFE5110F4FE34" box="[680,703,475,500]" italics="true" pageId="10" pageNumber="271">vs</emphasis>
. unarmed in the other species, to be confirmed for
<taxonomicName id="4C33479F8255FFE95005FE5117C0FE35" baseAuthorityName="Coutiere" baseAuthorityYear="1897" box="[1281,1419,475,501]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="latirostris">
<emphasis id="B947E00E8255FFE95005FE5117C0FE35" box="[1281,1419,475,501]" italics="true" pageId="10" pageNumber="271">S. latirostris</emphasis>
</taxonomicName>
); from
<taxonomicName id="4C33479F8255FFE955D1FE75136DFDD8" authorityName="Manning &amp; Chace" authorityYear="1990" box="[213,294,511,537]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="teres">
<emphasis id="B947E00E8255FFE955D1FE75136DFDD8" box="[213,294,511,537]" italics="true" pageId="10" pageNumber="271">S. teres</emphasis>
</taxonomicName>
and
<taxonomicName id="4C33479F8255FFE9545AFE751046FDD8" authorityName="Anker" authorityYear="2020" box="[350,525,511,537]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="inconspicuus">
<emphasis id="B947E00E8255FFE9545AFE751046FDD8" box="[350,525,511,537]" italics="true" pageId="10" pageNumber="271">S. inconspicuus</emphasis>
</taxonomicName>
by the major cheliped armed with a spiniform seta (
<emphasis id="B947E00E8255FFE9515FFE751639FDD8" box="[1115,1138,511,536]" italics="true" pageId="10" pageNumber="271">vs</emphasis>
. unarmed in the other two species); from
<taxonomicName id="4C33479F8255FFE9543FFDA913A6FDFD" box="[315,493,547,573]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="serratidigitus">
<emphasis id="B947E00E8255FFE9543FFDA913A6FDFD" box="[315,493,547,573]" italics="true" pageId="10" pageNumber="271">S. serratidigitus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C33479F8255FFE954FEFDA91001FDFC" authorityName="Manning &amp; Chace" authorityYear="1990" box="[506,586,547,573]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="teres">
<emphasis id="B947E00E8255FFE954FEFDA91001FDFC" box="[506,586,547,573]" italics="true" pageId="10" pageNumber="271">S. teres</emphasis>
</taxonomicName>
and
<taxonomicName id="4C33479F8255FFE9577AFDA91160FDFC" authorityName="Anker" authorityYear="2020" box="[638,811,547,573]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="inconspicuus">
<emphasis id="B947E00E8255FFE9577AFDA91160FDFC" box="[638,811,547,573]" italics="true" pageId="10" pageNumber="271">S. inconspicuus</emphasis>
</taxonomicName>
by the shorter stylocerite, only slightly overreaching the mid-length of the second article of the antennular peduncle (
<emphasis id="B947E00E8255FFE9563AFDCD111EFDA0" box="[830,853,583,608]" italics="true" pageId="10" pageNumber="271">vs</emphasis>
. reaching to the end of the second article or beyond it in the other species); from
<taxonomicName id="4C33479F8255FFE954DCFDE110D8FD45" authorityName="Anker &amp; Marin" authorityYear="2006" box="[472,659,619,645]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="nhatrangensis">
<emphasis id="B947E00E8255FFE954DCFDE110D8FD45" box="[472,659,619,645]" italics="true" pageId="10" pageNumber="271">S. nhatrangensis</emphasis>
</taxonomicName>
by the much shallower notches between the orbital teeth and the base of the rostrum; from
<taxonomicName id="4C33479F8255FFE95482FD051393FD68" authorityName="Manning &amp; Chace" authorityYear="1990" box="[390,472,655,681]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="teres">
<emphasis id="B947E00E8255FFE95482FD051393FD68" box="[390,472,655,681]" italics="true" pageId="10" pageNumber="271">S. teres</emphasis>
</taxonomicName>
by the third pereiopod ischium armed with spiniform setae (
<emphasis id="B947E00E8255FFE9518FFD0516E9FD68" box="[1163,1186,655,680]" italics="true" pageId="10" pageNumber="271">vs</emphasis>
. unarmed in
<taxonomicName id="4C33479F8255FFE9503DFD0517C0FD68" authorityName="Manning &amp; Chace" authorityYear="1990" box="[1337,1419,655,681]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="teres">
<emphasis id="B947E00E8255FFE9503DFD0517C0FD68" box="[1337,1419,655,681]" italics="true" pageId="10" pageNumber="271">S. teres</emphasis>
</taxonomicName>
); from
<taxonomicName id="4C33479F8255FFE955D0FD391330FD0C" authorityName="Anker" authorityYear="2010" box="[212,379,691,717]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="camaroncito">
<emphasis id="B947E00E8255FFE955D0FD391330FD0C" box="[212,379,691,717]" italics="true" pageId="10" pageNumber="271">S. camaroncito</emphasis>
</taxonomicName>
and
<taxonomicName id="4C33479F8255FFE954ABFD391018FD0C" authorityName="Anker" authorityYear="2020" box="[431,595,691,717]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="saotomensis">
<emphasis id="B947E00E8255FFE954ABFD391018FD0C" box="[431,595,691,717]" italics="true" pageId="10" pageNumber="271">S. saotomensis</emphasis>
</taxonomicName>
by the minor cheliped non-enlarged and with the chela fingers unarmed (
<emphasis id="B947E00E8255FFE9507AFD3917DEFD0C" box="[1406,1429,691,716]" italics="true" pageId="10" pageNumber="271">vs</emphasis>
. noticeably enlarged and with the chela fingers armed with small teeth in
<taxonomicName id="4C33479F8255FFE956CDFD5D1639FD30" authorityName="Anker" authorityYear="2010" box="[969,1138,727,753]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="camaroncito">
<emphasis id="B947E00E8255FFE956CDFD5D1639FD30" box="[969,1138,727,753]" italics="true" pageId="10" pageNumber="271">S. camaroncito</emphasis>
</taxonomicName>
and
<taxonomicName id="4C33479F8255FFE951ACFD5D1705FD30" authorityName="Anker" authorityYear="2020" box="[1192,1358,727,753]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="saotomensis">
<emphasis id="B947E00E8255FFE951ACFD5D1705FD30" box="[1192,1358,727,753]" italics="true" pageId="10" pageNumber="271">S. saotomensis</emphasis>
</taxonomicName>
); from
<taxonomicName id="4C33479F8255FFE95593FD71130BFCD4" authorityName="Anker" authorityYear="2010" box="[151,320,763,789]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="camaroncito">
<emphasis id="B947E00E8255FFE95593FD71130BFCD4" box="[151,320,763,789]" italics="true" pageId="10" pageNumber="271">S. camaroncito</emphasis>
</taxonomicName>
by the rostrum directed forwards, without subdistal tooth on its ventral margin (
<emphasis id="B947E00E8255FFE951C6FD711692FCD4" box="[1218,1241,763,788]" italics="true" pageId="10" pageNumber="271">vs</emphasis>
. directed slightly downwards, with a small subdistal tooth, in
<taxonomicName id="4C33479F8255FFE95787FC951161FCF8" authorityName="Anker" authorityYear="2010" box="[643,810,799,825]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="camaroncito">
<emphasis id="B947E00E8255FFE95787FC951161FCF8" box="[643,810,799,825]" italics="true" pageId="10" pageNumber="271">S. camaroncito</emphasis>
</taxonomicName>
); and from
<taxonomicName id="4C33479F8255FFE956ABFC95161FFCF8" authorityName="Anker" authorityYear="2020" box="[943,1108,799,825]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="saotomensis">
<emphasis id="B947E00E8255FFE956ABFC95161FFCF8" box="[943,1108,799,825]" italics="true" pageId="10" pageNumber="271">S. saotomensis</emphasis>
</taxonomicName>
by the major cheliped carpus unarmed ventrally (
<emphasis id="B947E00E8255FFE9547DFCC913DBFC9C" box="[377,400,835,860]" italics="true" pageId="10" pageNumber="271">vs</emphasis>
. with a strong ventromesial process in
<taxonomicName id="4C33479F8255FFE9565EFCC91649FC9C" authorityName="Anker" authorityYear="2020" box="[858,1026,835,861]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="saotomensis">
<emphasis id="B947E00E8255FFE9565EFCC91649FC9C" box="[858,1026,835,861]" italics="true" pageId="10" pageNumber="271">S. saotomensis</emphasis>
</taxonomicName>
) (
<bibRefCitation id="EFA241ED8255FFE95118FCC91688FC9D" author="Coutiere, H." box="[1052,1219,835,861]" pageId="10" pageNumber="271" pagination="1 - 559" refId="ref10857" refString="Coutiere, H. (1899) Les Alpheidae. Morphologie externe et interne, formes larvaires, bionomie. Annales des Sciences Naturelles, Zoologie et Paleontologie, 8 (9), 1 - 559, 6 pls. Masson, Paris. https: // doi. org / 10.5962 / bhl. title. 13143" type="journal article" year="1899">Coutière 1899</bibRefCitation>
;
<bibRefCitation id="EFA241ED8255FFE951D5FCC91299FC41" author="Banner, D. M. &amp; Banner, A. H." pageId="10" pageNumber="271" pagination="1 - 99" refId="ref10705" refString="Banner, D. M. &amp; Banner, A. H. (1981) Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, Leiden, 190, 1 - 99." type="journal article" year="1981">Banner &amp; Banner 1981</bibRefCitation>
;
<bibRefCitation id="EFA241ED8255FFE955DAFCED13A4FC41" author="Manning, R. B. &amp; Chace, F. A. Jr." box="[222,495,871,897]" pageId="10" pageNumber="271" pagination="1 - 91" refId="ref11355" refString="Manning, R. B. &amp; Chace, F. A. Jr. (1990) Decapod and stomatopod Crustacea from Ascension Island, South Atlantic Ocean. Smithsonian Contributions to Zoology, 503, 1 - 91. https: // doi. org / 10.5479 / si. 00810282.503" type="journal article" year="1990">Manning &amp; Chace 1990</bibRefCitation>
;
<bibRefCitation id="EFA241ED8255FFE954FDFCED10A1FC41" author="Anker, A. &amp; Marin, I. N." box="[505,746,871,897]" pageId="10" pageNumber="271" pagination="295 - 319" refId="ref10562" refString="Anker, A. &amp; Marin, I. N. (2006) New records and species of Alpheidae (Crustacea, Decapoda) from Vietnam. Part I. Genus Salmoneus Holthuis, 1955. Raffles Bulletin of Zoology, 54, 295 - 319." type="journal article" year="2006">Anker &amp; Marin 2006</bibRefCitation>
;
<bibRefCitation id="EFA241ED8255FFE957F7FCED1130FC41" author="Anker, A." box="[755,891,871,897]" pageId="10" pageNumber="271" pagination="21 - 39" refId="ref10136" refString="Anker, A. (2007) New species and records of alpheid shrimps, genera Salmoneus Holthuis and Parabetaeus Coutiere, from the tropical western Atlantic (Decapoda, Caridea). Zootaxa, 1653 (1), 21 - 39. https: // doi. org / 10.11646 / zootaxa. 1653.1.2" type="journal article" year="2007">Anker 2007</bibRefCitation>
,
<bibRefCitation id="EFA241ED8255FFE95682FCED118AFC41" author="Anker, A." box="[902,961,871,897]" pageId="10" pageNumber="271" pagination="177 - 205" refId="ref10192" refString="Anker, A. (2010) The shrimp genus Salmoneus Holthuis, 1955 (Crustacea, Decapoda, Alpheidae) in the tropical western Atlantic, with description of five new species. Zootaxa, 2372 (1), 177 - 205. https: // doi. org / 10.11646 / zootaxa. 2372.1.18" type="journal article" year="2010">2010</bibRefCitation>
, 2020).
</paragraph>
<paragraph id="8B8C3C1C8255FFE955C3FC01101AFBD1" blockId="10.[151,1437,151,1041]" pageId="10" pageNumber="271">
All other species of
<taxonomicName id="4C33479F8255FFE954ADFC011069FC65" authorityName="Holthuis" authorityYear="1955" box="[425,546,907,933]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="genus">
<emphasis id="B947E00E8255FFE954ADFC011069FC65" box="[425,546,907,933]" italics="true" pageId="10" pageNumber="271">Salmoneus</emphasis>
</taxonomicName>
differ from
<taxonomicName id="4C33479F8255FFE957A8FC011161FC65" authority="Anker, 2022" authorityName="Anker" authorityYear="2022" box="[684,810,907,933]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="arabicus" status="sp. nov.">
<emphasis id="B947E00E8255FFE957A8FC011161FC65" box="[684,810,907,933]" italics="true" pageId="10" pageNumber="271">S. arabicus</emphasis>
</taxonomicName>
<emphasis id="B947E00E8255FFE95636FC0111C0FC65" bold="true" box="[818,907,907,933]" pageId="10" pageNumber="271">
<taxonomicNameLabel id="A2745D758255FFE95636FC0111C0FC65" box="[818,907,907,933]" pageId="10" pageNumber="271" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by a greater number of morphological features and/or colour (where known). This includes all remaining species known from the Red Sea and adjacent areas of the Indian Ocean, e.g.
<taxonomicName id="4C33479F8255FFE9546DFC591393FC2D" authorityName="Ashrafi, Duris &amp; Naderloo" authorityYear="2020" box="[361,472,979,1005]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="rashedi">
<emphasis id="B947E00E8255FFE9546DFC591393FC2D" box="[361,472,979,1005]" italics="true" pageId="10" pageNumber="271">S. rashedi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C33479F8255FFE954E1FC59102AFC2C" authorityName="Anker" authorityYear="2019" box="[485,609,979,1005]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="venustus">
<emphasis id="B947E00E8255FFE954E1FC59102AFC2C" box="[485,609,979,1005]" italics="true" pageId="10" pageNumber="271">S. venustus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C33479F8255FFE9576BFC591145FC2D" authorityName="Duris &amp; Horka" authorityYear="2016" box="[623,782,979,1005]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="chadwickae">
<emphasis id="B947E00E8255FFE9576BFC591145FC2D" box="[623,782,979,1005]" italics="true" pageId="10" pageNumber="271">S. chadwickae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C33479F8255FFE95647FC5911F4FC2C" baseAuthorityName="Coutiere" baseAuthorityYear="1897" box="[835,959,979,1005]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="cristatus">
<emphasis id="B947E00E8255FFE95647FC5911F4FC2C" box="[835,959,979,1005]" italics="true" pageId="10" pageNumber="271">S. cristatus</emphasis>
</taxonomicName>
(all with different colourations), as well as the above-described
<taxonomicName id="4C33479F8255FFE9547BFC7D13BAFBD1" authority="Anker, 2022" authorityName="Anker" authorityYear="2022" box="[383,497,1015,1041]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="farasan" status="sp. nov.">
<emphasis id="B947E00E8255FFE9547BFC7D13BAFBD1" box="[383,497,1015,1041]" italics="true" pageId="10" pageNumber="271">S. farasan</emphasis>
</taxonomicName>
<emphasis id="B947E00E8255FFE954F3FC7D101AFBD1" bold="true" box="[503,593,1015,1041]" pageId="10" pageNumber="271">
<taxonomicNameLabel id="A2745D758255FFE954F3FC7D101AFBD1" box="[503,593,1015,1041]" pageId="10" pageNumber="271" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>