903 lines
74 KiB
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903 lines
74 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/vz.71.e62787" ID-GBIF-Dataset="8a0fe91d-28b3-4736-90ae-456d38e8ed24" ID-GBIF-Taxon="191979928" ID-Pensoft-Pub="2625-8498-71-669" ID-Pensoft-UUID="560164B38FA75BB6B5B0BC67193D8AB3" ID-ZooBank="523BCE4C45C44E578B58282007807A97" ModsDocID="2625-8498-71-669" checkinTime="1641857276082" checkinUser="pensoft" docAuthor="Gowande, Gaurang, Pal, Saunak, Jablonski, Daniel, Masroor, Rafaqat, Phansalkar, Pushkar U., Dsouza, Princia, Jayarajan, Aditi & Shanker, Kartik" docDate="2021" docId="FF21F0287662550EB923206B9830BCA2" docLanguage="en" docName="VertZool 71: 669-696" docOrigin="Vertebrate Zoology 71" docPubDate="2021-10-20" docSource="http://dx.doi.org/10.3897/vz.71.e62787" docTitle="Calotes vultuosus comb. nov. (Harlan 1825" docType="treatment" docVersion="4" id="560164B38FA75BB6B5B0BC67193D8AB3" lastPageNumber="669" masterDocId="560164B38FA75BB6B5B0BC67193D8AB3" masterDocTitle="Molecular phylogenetics and taxonomic reassessment of the widespread agamid lizard Calotes versicolor (Daudin, 1802) (Squamata, Agamidae) across South Asia" masterLastPageNumber="696" masterPageNumber="669" pageId="0" pageNumber="669" updateTime="1641863297910" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>Molecular phylogenetics and taxonomic reassessment of the widespread agamid lizard Calotes versicolor (Daudin, 1802) (Squamata, Agamidae) across South Asia</mods:title>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Gowande, Gaurang</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8648-2957</mods:nameIdentifier>
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<mods:affiliation>Annasaheb Kulkarni Department of Biodiversity, Abasaheb Garware College, Pune - 411004, Maharashtra, India & Department of Biotechnology, Fergusson College, Pune - 411004, Maharashtra, India & Centre for Ecological Sciences, Indian Institute of Science, Bengaluru - 560012, Karnataka, India</mods:affiliation>
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<mods:nameIdentifier type="email">gaurang.gowande@gmail.com</mods:nameIdentifier>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Pal, Saunak</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8267-4708</mods:nameIdentifier>
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<mods:affiliation>Centre for Ecological Sciences, Indian Institute of Science, Bengaluru - 560012, Karnataka, India & Bombay Natural History Society, Hornbill House, Opposite Lion Gate, Fort, Mumbai - 400001, Maharashtra, India</mods:affiliation>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jablonski, Daniel</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5394-0114</mods:nameIdentifier>
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<mods:affiliation>Department of Zoology, Comenius University in Bratislava, Ilkovicova 6, Mlynska dolina, Bratislava - 84215, Slovakia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Masroor, Rafaqat</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-6248-546X</mods:nameIdentifier>
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<mods:affiliation>Zoological Sciences Division, Pakistan Museum of Natural History, Garden Avenue, Shakarparian, Islamabad - 440000, Pakistan</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Phansalkar, Pushkar U.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-3012-9481</mods:nameIdentifier>
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<mods:affiliation>A / 2, Ajinkyanagari, Karvenagar, Pune - 411052, Maharashtra, India</mods:affiliation>
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</mods:name>
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Dsouza, Princia</mods:namePart>
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<mods:affiliation>Centre for Ecological Sciences, Indian Institute of Science, Bengaluru - 560012, Karnataka, India</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jayarajan, Aditi</mods:namePart>
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<mods:affiliation>Centre for Ecological Sciences, Indian Institute of Science, Bengaluru - 560012, Karnataka, India</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Shanker, Kartik</mods:namePart>
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<mods:affiliation>Centre for Ecological Sciences, Indian Institute of Science, Bengaluru - 560012, Karnataka, India</mods:affiliation>
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<mods:title>Vertebrate Zoology</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-10-20</mods:number>
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<mods:detail type="volume">
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<mods:number>71</mods:number>
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<mods:extent unit="page">
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<mods:start>669</mods:start>
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<mods:end>696</mods:end>
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<mods:url>http://dx.doi.org/10.3897/vz.71.e62787</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/vz.71.e62787</mods:identifier>
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<mods:identifier type="Pensoft-Pub">2625-8498-71-669</mods:identifier>
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<mods:identifier type="ZooBank">523BCE4C45C44E578B58282007807A97</mods:identifier>
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<mods:identifier type="Pensoft-UUID">560164B38FA75BB6B5B0BC67193D8AB3</mods:identifier>
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<treatment ID-GBIF-Taxon="191979928" LSID="urn:lsid:plazi:treatment:FF21F0287662550EB923206B9830BCA2" httpUri="http://treatment.plazi.org/id/FF21F0287662550EB923206B9830BCA2" lastPageNumber="669" pageId="0" pageNumber="669">
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<subSubSection pageId="0" pageNumber="669" type="nomenclature">
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<paragraph pageId="0" pageNumber="669">
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<taxonomicName LSID="FF21F028-7662-550E-B923-206B9830BCA2" authority="comb. nov. (Harlan, 1825)" authorityName="comb. nov. (Harlan" authorityYear="1825" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calotes vultuosus" order="Squamata" pageId="0" pageNumber="669" phylum="Chordata" rank="species" species="vultuosus">Calotes vultuosus comb. nov. (Harlan, 1825)</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="669" type="description">
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<paragraph pageId="0" pageNumber="669">
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Holotype of Calotes vultuosus comb. nov. ANSP 7296. A. full body ventro-lateral, B. full body lateral, C. head dorsal, D. head ventral, E. head lateral. Photographs by Ned Gilmore." figureDoi="10.3897/vz.71.e62787.figure6" httpUri="https://binary.pensoft.net/fig/600933" pageId="0" pageNumber="669">Figs 6</figureCitation>
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. An uncollected male of Calotes vultuosus comb. nov., from Burdwan, West Bengal, India, in life. Photograph by Ayan Mondal." figureDoi="10.3897/vz.71.e62787.figure7" httpUri="https://binary.pensoft.net/fig/600934" pageId="0" pageNumber="669">, 7</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection pageId="0" pageNumber="669" type="reference_group">
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<paragraph pageId="0" pageNumber="669">
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<taxonomicName authorityName="Harlan" authorityYear="1825" class="Reptilia" family="Agamidae" genus="Agama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agama vultuosa" order="Squamata" pageId="0" pageNumber="669" phylum="Chordata" rank="species" species="vultuosa">Agama vultuosa</taxonomicName>
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<bibRefCitation author="Harlan, R" journalOrPublisher="Journal of the Academy of Natural Sciences of Philadelphia" pageId="0" pageNumber="669" pagination="296 - 305" refId="B27" refString="Harlan, R, 1825. Description of a new species of Agama. Journal of the Academy of Natural Sciences of Philadelphia 4 (1): 296 - 305" title="Description of a new species of Agama." volume="4" year="1825">Harlan 1825</bibRefCitation>
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: 296
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</paragraph>
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<paragraph pageId="0" pageNumber="669">
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<taxonomicName authorityName="Gray" authorityYear="1827" class="Reptilia" family="Agamidae" genus="Agama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agama indica" order="Squamata" pageId="0" pageNumber="669" phylum="Chordata" rank="species" species="indica">Agama indica</taxonomicName>
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<bibRefCitation author="Hardwicke, T" journalOrPublisher="Zoological Journal" pageId="0" pageNumber="669" pagination="213 - 229" refId="B26" refString="Hardwicke, T, Gray, J, 1827. A synopsis of the species of saurian reptiles, collected in India by Major-General Hardwicke. Zoological Journal 3: 213 - 229" title="A synopsis of the species of saurian reptiles, collected in India by Major-General Hardwicke." volume="3" year="1827">Hardwicke and Gray 1827</bibRefCitation>
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: 217
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</paragraph>
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<paragraph pageId="0" pageNumber="669">
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<taxonomicName authority="versicolor" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calotes versicolor subsp. versicolor" order="Squamata" pageId="0" pageNumber="669" phylum="Chordata" rank="subSpecies" species="versicolor" subSpecies="versicolor">Calotes cf. versicolor versicolor</taxonomicName>
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<bibRefCitation author="Manthey, U" journalOrPublisher="Herpetozoa" pageId="0" pageNumber="669" refId="B37" refString="Manthey, U, 2008. Agamid Lizards of Southern Asia - Draconinae 1: Terralog Vol. 7a. Chimaira Buchhandlunggesellschaft mbH, Frankfurt." title="Agamid Lizards of Southern Asia - Draconinae 1: Terralog Vol. 7 a. Chimaira Buchhandlunggesellschaft mbH, Frankfurt." year="2008">Manthey 2008</bibRefCitation>
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: 88 (in part)
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</paragraph>
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<paragraph pageId="0" pageNumber="669">
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<taxonomicName baseAuthorityName="Daudin" baseAuthorityYear="1802" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calotes versicolor" order="Squamata" pageId="0" pageNumber="669" phylum="Chordata" rank="species" species="versicolor">Calotes cf. versicolor</taxonomicName>
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-
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<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4482.3.1" author="Pal, S" journalOrPublisher="Zootaxa" pageId="0" pageNumber="669" pagination="401 - 450" refId="B47" refString="Pal, S, Vijayakumar, S, Shanker, K, Jayarajan, A, Deepak, V, 2018. A systematic revision of Calotes Cuvier, 1817 (Squamata: Agamidae) from the Western Ghats adds two new genera and reveals two new species. Zootaxa 4482 (3): 401 - 450, DOI: https://doi.org/10.11646/zootaxa.4482.3.1" title="A systematic revision of Calotes Cuvier, 1817 (Squamata: Agamidae) from the Western Ghats adds two new genera and reveals two new species." url="https://doi.org/10.11646/zootaxa.4482.3.1" volume="4482" year="2018">Pal et al. (2018)</bibRefCitation>
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(in part)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="0" pageNumber="669" type="holotype">
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<paragraph pageId="0" pageNumber="669">Holotype.</paragraph>
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<paragraph pageId="0" pageNumber="669">
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INDIA • 1 ♂; West Bengal, Kolkata;
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<geoCoordinate degrees="20.55" direction="north" orientation="latitude" precision="555" value="20.55">20.55°N</geoCoordinate>
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,
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<geoCoordinate degrees="88.36" direction="east" orientation="longitude" precision="555" value="88.36">88.36°E</geoCoordinate>
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, 10 m a. s. l.; Dr R. Coates leg.; ANSP 7296
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</paragraph>
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<paragraph pageId="0" pageNumber="669">
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<emphasis bold="true" pageId="0" pageNumber="669">Other material (morphological vouchers).</emphasis>
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All from INDIA • 1 ♂, 1 ♀; Maharashtra, Satara, Koyna WLS; Soman and Thakar leg; BNHS 356 (both); • 1 ♂; Maharashtra, Solapur, Kurduwadi;
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<geoCoordinate degrees="18.09" direction="north" orientation="latitude" precision="555" value="18.09">18.09°N</geoCoordinate>
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,
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<geoCoordinate degrees="75.43" direction="east" orientation="longitude" precision="555" value="75.43">75.43°E</geoCoordinate>
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, 516 m a. s. l.; BNHS 368; • 2 ♂; Maharashtra, Satara, Thoseghar;
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<geoCoordinate degrees="17.58880" direction="north" orientation="latitude" precision="1" value="17.5888">17.58880°N</geoCoordinate>
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,
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<geoCoordinate degrees="73.83805" direction="east" orientation="longitude" precision="1" value="73.83805">73.83805°E</geoCoordinate>
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, 1119 m a. s. l.; 06 Jun. 2015; Gaurang Gowande and Pushkar Phansalkar leg.; GenBank MW901288-89 (16S); CESL 1004-05; • 1 ♂; Maharashtra, Satara, Patan;
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<geoCoordinate degrees="17.35397" direction="north" orientation="latitude" precision="1" value="17.35397">17.35397°N</geoCoordinate>
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,
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<geoCoordinate degrees="73.80288" direction="east" orientation="longitude" precision="1" value="73.80288">73.80288°E</geoCoordinate>
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, 587 m a. s. l.; 10 Jun. 2015; Gaurang Gowande and Pushkar Phansalkar leg.; GenBank MW901290 (16S); CESL 1007; • 1 ♂; Maharashtra, Pune, Taleghar;
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<geoCoordinate degrees="19.04670" direction="north" orientation="latitude" precision="1" value="19.0467">19.04670°N</geoCoordinate>
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,
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<geoCoordinate degrees="73.55370" direction="east" orientation="longitude" precision="1" value="73.5537">73.55370°E</geoCoordinate>
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, 925 m a. s. l.; 13 Jun. 2015; Gaurang Gowande and Pushkar Phansalkar leg.; GenBank MW901255 (16S); CESL1009; • 2 ♂; Maharashtra, Pune, Waghapur;
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<geoCoordinate degrees="18.39990" direction="north" orientation="latitude" precision="1" value="18.3999">18.39990°N</geoCoordinate>
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,
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<geoCoordinate degrees="74.12400" direction="east" orientation="longitude" precision="1" value="74.124">74.12400°E</geoCoordinate>
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, 762 m a. s. l.; 11 Oct. 2015; Gaurang Gowande and Pushkar Phansalkar leg.; CESL 1042-43; • 1 ♂; Maharashtra, Chiplun, Guhagar;
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<geoCoordinate degrees="17.49495" direction="north" orientation="latitude" precision="1" value="17.49495">17.49495°N</geoCoordinate>
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,
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<geoCoordinate degrees="73.18475" direction="east" orientation="longitude" precision="1" value="73.18475">73.18475°E</geoCoordinate>
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, 11 m a. s. l.; 24 Oct. 2016; Gaurang Gowande and Pushkar Phansalkar leg.; CESL 1036; • 1 ♂; Rajasthan, Pali;
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<geoCoordinate degrees="26.82485" direction="north" orientation="latitude" precision="1" value="26.82485">26.82485°N</geoCoordinate>
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,
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<geoCoordinate degrees="71.85335" direction="east" orientation="longitude" precision="1" value="71.85335">71.85335°E</geoCoordinate>
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, 230 m a. s. l.; 2 Jul. 2017; Vishal Varma leg.; GenBank MW901284 (16S); CESL 1041 • 1 ♂; Maharashtra, Mumbai, Bandra; 8 May 1959; H. Abdulali leg.; BNHS 350 • 1 ♂; Karnataka, Kadra; Dec. 1914; S. Prater leg.; BNHS 354 • 1 ♂; Maharashtra, Mumbai; Sir Norman Kinnear leg.; BNHS 343 • 1 ♂; Gujarat, Kutch; P. Soman leg.; BNHS 316 • 1 ♂; Maharashtra, Aurangabad; Gaurang Gowande and Pushkar Phansalkar leg.; CESL 1065 • 3 ♂; Telangana, Hyderabad; 14 Apr. 2017; Gaurang Gowande and Pushkar Phansalkar leg.; GenBank MW901282 (16S); CESL 1068-70 • 1 ♂; Maharashtra, Latur; 16 Apr. 2017; Gaurang Gowande and Pushkar Phansalkar leg.; CESL 1071 • 1 ♀; Maharashtra, Mumbai, Borivali; 6 Jun. 1971; BNHS 1027.
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</paragraph>
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<caption doi="10.3897/vz.71.e62787.figure6" httpUri="https://binary.pensoft.net/fig/600933" pageId="0" pageNumber="669" start="Figure 6" startId="F6">
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<paragraph pageId="0" pageNumber="669">
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<emphasis bold="true" pageId="0" pageNumber="669">Figure 6.</emphasis>
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Holotype of
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<taxonomicName authorityName="comb. nov. (Harlan" authorityYear="1825" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calotes vultuosus" order="Squamata" pageId="0" pageNumber="669" phylum="Chordata" rank="species" species="vultuosus">
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<emphasis italics="true" pageId="0" pageNumber="669">Calotes vultuosus</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="669">comb. nov.</emphasis>
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ANSP 7296.
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<emphasis bold="true" pageId="0" pageNumber="669">A.</emphasis>
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full body ventro-lateral,
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<emphasis bold="true" pageId="0" pageNumber="669">B.</emphasis>
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full body lateral,
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<emphasis bold="true" pageId="0" pageNumber="669">C.</emphasis>
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head dorsal,
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<emphasis bold="true" pageId="0" pageNumber="669">D.</emphasis>
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head ventral,
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<emphasis bold="true" pageId="0" pageNumber="669">E.</emphasis>
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head lateral. Photographs by Ned Gilmore.
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</paragraph>
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</caption>
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||
<caption doi="10.3897/vz.71.e62787.figure7" httpUri="https://binary.pensoft.net/fig/600934" pageId="0" pageNumber="669" start="Figure 7" startId="F7">
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||
<paragraph pageId="0" pageNumber="669">
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<emphasis bold="true" pageId="0" pageNumber="669">Figure 7.</emphasis>
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An uncollected male of
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<taxonomicName authorityName="comb. nov. (Harlan" authorityYear="1825" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calotes vultuosus" order="Squamata" pageId="0" pageNumber="669" phylum="Chordata" rank="species" species="vultuosus">
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<emphasis italics="true" pageId="0" pageNumber="669">Calotes vultuosus</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="669">comb. nov.</emphasis>
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, from Burdwan, West Bengal, India, in life. Photograph by Ayan Mondal.
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</paragraph>
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</caption>
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<caption ID-Table-UUID="2180D7EC720D2D52B3FF2C140ADD46FF" httpUri="http://table.plazi.org/id/2180D7EC720D2D52B3FF2C140ADD46FF" pageId="0" pageNumber="669" start="Table 4" startId="T4">
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<paragraph pageId="0" pageNumber="669">
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<emphasis bold="true" pageId="0" pageNumber="669">Table 4.</emphasis>
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Mensural and meristic characters of the type and non-type specimens of
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<taxonomicName authorityName="comb. nov. (Harlan" authorityYear="1825" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calotes vultuosus" order="Squamata" pageId="0" pageNumber="669" phylum="Chordata" rank="species" species="vultuosus">
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<emphasis italics="true" pageId="0" pageNumber="669">Calotes vultuosus</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="669">comb. nov.</emphasis>
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and
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<taxonomicName authorityName="Auffenberg and Rehman" authorityYear="1993" baseAuthorityName="Gowande & Pal & Jablonski & Masroor & Phansalkar & Dsouza & Jayarajan & Shanker" baseAuthorityYear="2021" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calotes farooqi" order="Squamata" pageId="0" pageNumber="669" phylum="Chordata" rank="species" species="farooqi">
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<emphasis italics="true" pageId="0" pageNumber="669">Calotes farooqi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="669">stat. nov.</emphasis>
|
||
, mensural characters recorded as a ratio of SVL. Note that the morphometric data for the types of
|
||
<taxonomicName genus="A." lsidName="A. indica" pageId="0" pageNumber="669" rank="species" species="indica">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">A. indica</emphasis>
|
||
</taxonomicName>
|
||
were not available, whereas those for
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. farooqi" order="Squamata" pageId="0" pageNumber="669" rank="species" species="farooqi">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. farooqi</emphasis>
|
||
</taxonomicName>
|
||
have been sourced from
|
||
<bibRefCitation author="Auffenberg, W" journalOrPublisher="Asiatic Herpetological Research" pageId="0" pageNumber="669" pagination="14 - 30" refId="B7" refString="Auffenberg, W, Rehman, H, 1993. Studies on Pakistan Reptiles. Pt. 3. Calotes versicolor. Asiatic Herpetological Research 5: 14 - 30" title="Studies on Pakistan Reptiles. Pt. 3. Calotes versicolor." volume="5" year="1993">Auffenberg and Rehman (1993)</bibRefCitation>
|
||
. - indicates that measurements/counts could not be taken.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="669">
|
||
<table pageId="0" pageNumber="669">
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="669">Species</emphasis>
|
||
</td>
|
||
<td colspan="2" pageId="0" pageNumber="669" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="669">
|
||
<taxonomicName authorityName="comb. nov. (Harlan" authorityYear="1825" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calotes vultuosus" order="Squamata" pageId="0" pageNumber="669" phylum="Chordata" rank="species" species="vultuosus">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">Calotes vultuosus</emphasis>
|
||
</taxonomicName>
|
||
comb. nov.
|
||
</emphasis>
|
||
</td>
|
||
<td colspan="2" pageId="0" pageNumber="669" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="669">
|
||
<taxonomicName authorityName="Auffenberg and Rehman" authorityYear="1993" baseAuthorityName="Gowande & Pal & Jablonski & Masroor & Phansalkar & Dsouza & Jayarajan & Shanker" baseAuthorityYear="2021" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calotes farooqi" order="Squamata" pageId="0" pageNumber="669" phylum="Chordata" rank="species" species="farooqi">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">Calotes farooqi</emphasis>
|
||
</taxonomicName>
|
||
stat. nov.
|
||
</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="669">Specimens measured</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="669">Holotype ANSP 7296</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="669">
|
||
Other morphological vouchers (
|
||
<emphasis italics="true" pageId="0" pageNumber="669">n</emphasis>
|
||
=20)
|
||
</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="669">Holotype FLMNH/UF 79470</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="669">
|
||
Other morphological vouchers (
|
||
<emphasis italics="true" pageId="0" pageNumber="669">n</emphasis>
|
||
=3)
|
||
</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="669">Mensural characters</emphasis>
|
||
</td>
|
||
<td colspan="1" isEmpty="true" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="669">Range</emphasis>
|
||
</td>
|
||
<td colspan="1" isEmpty="true" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="669">Range</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">HeadL/SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.19</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.16-0.30</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.27</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.23-0.25</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">HeadW/SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.16-0.26</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.24</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.19-0.21</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">JawW/SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.17-0.21</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.17-0.20</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">HeadH/SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.16-0.21</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.20</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.14-0.16</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">SnEye/SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.08-0.11</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.09-0.11</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">NarEye/SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.04-0.06</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.04-0.05</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">EyeEar/SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.06-0.09</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.07</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">Snw/SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.05-0.08</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.06</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">InterOrb/SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.09-0.13</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.12-0.15</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">TrunkL/SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.49</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.37-0.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.48-0.52</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">PectW/SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.12-0.20</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.14-0.20</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">PelvW/SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.09-0.17</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.11-0.13</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">SnForeL/SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.34-0.45</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.31-0.35</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">UpArmL/SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.14-0.23</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.20</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.13-0.18</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">LoArmL/SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.16-0.23</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.16-0.17</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">ForefL/SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.14-0.20</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.17-0.18</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">4fingLng/SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.10-0.15</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.12-0.13</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">UpLegL/SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.18-0.29</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.21-0.22</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">CrusL/SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.19-0.25</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.22</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">HindfL/SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.28-0.34</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.30-0.35</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">4toeLng/SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.15-0.23</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.13-0.20</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">ForeLimbL/SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.47-0.59</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">HindLimbL/SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.68-0.85</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">ForeLimbL/HindLimbL</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.65-0.81</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">PelvW/PectW</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.74-1.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">0.60-0.86</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="669">Meristic characters</emphasis>
|
||
</td>
|
||
<td colspan="1" isEmpty="true" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" isEmpty="true" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" isEmpty="true" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" isEmpty="true" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">SnS</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">7</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">5-7</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">6</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">6</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">HeadSTr</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">14</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">10-16</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">14</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">11</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">HeadSLn</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">12</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">12-17</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">12</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">13-14</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">CanthR</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">7</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">7-8</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">8</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">8</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">Eyelid</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">13</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">11-14</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">9-11</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">Suplab</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">10</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">9-13</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">13</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">11-13</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">Inflab</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">10</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">9-13</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">13</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">12</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">TempSp</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">2</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">2</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">2</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">2</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">Dorsal</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">35-62</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">40-44</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">Midbody</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">37-45</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">51</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">41-44</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">4FingLm</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">19-22</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">21</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">17-18</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="669">
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">4ToeLm</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">23-28</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">23</td>
|
||
<td colspan="1" pageId="0" pageNumber="669" rowspan="1">24-25</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="669" type="genetic diagnosis">
|
||
<paragraph pageId="0" pageNumber="669">Genetic diagnosis.</paragraph>
|
||
<paragraph pageId="0" pageNumber="669">
|
||
The least within species divergence at 16S and COI was 0%, while the greatest within species distance recorded was 2.0% and 9.5% at 16S and COI respectively. The species was at least 3.3% and 13.0% divergent from
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. versicolor" order="Squamata" pageId="0" pageNumber="669" rank="species" species="versicolor">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. versicolor</emphasis>
|
||
</taxonomicName>
|
||
, 2.7% and 12.8% divergent from
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. irawadi" order="Squamata" pageId="0" pageNumber="669" rank="species" species="irawadi">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. irawadi</emphasis>
|
||
</taxonomicName>
|
||
, and 3.9% and 15.5% divergent from Clade4 at 16S and COI respectively (Table
|
||
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Table representing the least and the greatest un-corrected percent pairwise genetic p-distances within and between species recovered in this study. Values above the diagonal represent divergence on COI, whereas those below the diagonal represent divergence on 16 S. Values in bold along the diagonal represent distance within species, written as COI / 16 S. * Does not include Calotes aff. farooqi." httpUri="http://table.plazi.org/id/E5FE76105C59120F4551F581BC8CD722" pageId="0" pageNumber="669" tableUuid="E5FE76105C59120F4551F581BC8CD722">2</tableCitation>
|
||
). The species differed from
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. calotes" order="Squamata" pageId="0" pageNumber="669" rank="species" species="calotes">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. calotes</emphasis>
|
||
</taxonomicName>
|
||
by sequence divergence of> 5% at 16S and 16.3% at COI from
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. calotes" order="Squamata" pageId="0" pageNumber="669" rank="species" species="calotes">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. calotes</emphasis>
|
||
</taxonomicName>
|
||
. Phylogenetically, the species was recovered as sister to a clade comprising of
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. calotes" order="Squamata" pageId="0" pageNumber="669" rank="species" species="calotes">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. calotes</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. versicolor" order="Squamata" pageId="0" pageNumber="669" rank="species" species="versicolor">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. versicolor</emphasis>
|
||
</taxonomicName>
|
||
, and Clade4 (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Maximum Likelihood 16 S gene tree based on the ~ 416 base pair sequence data revealing the position of the species of the Calotes versicolor complex within the genus Calotes. Numbers at the nodes indicate bootstrap values. The tree was rooted using the Mantheyus phuwuanensis (Manthey and Nabhitabhata, 1991) as the out group. The tree was analysed under the TIM 2 + F + I + G 4 model of sequence evolution, implementing 1000 ultrafast bootstraps on the web platform of IQ-TREE. Species delimited in this study are colour coded: Green - C. farooqi stat. nov.; Yellow - C. irawadi; Red - C. versicolor; Blue - C. vultuosus comb. nov." figureDoi="10.3897/vz.71.e62787.figure3" httpUri="https://binary.pensoft.net/fig/600930" pageId="0" pageNumber="669">3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="669" type="diagnosis">
|
||
<paragraph pageId="0" pageNumber="669">Diagnosis and comparison.</paragraph>
|
||
<paragraph pageId="0" pageNumber="669">
|
||
A medium to large sized species of
|
||
<taxonomicName class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calotes" order="Squamata" pageId="0" pageNumber="669" phylum="Chordata" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">Calotes</emphasis>
|
||
</taxonomicName>
|
||
, adult males averaging 106 mm in SVL, females averaging 78 mm in SVL; body compressed; head relatively short; dorso-lateral scales posterodorsally oriented, large, weakly to strongly keeled, homogeneous; ventral scales smaller than the dorso-lateral scales, strongly keeled; 37-45 scales around the mid-body; anti-humeral fold absent; two distinct spines in the supratympanic region, posterior spine as long as the anterior spine, at times longer, more prominent; nuchal and dorsal crest continuous, distinct, slightly recurved; scales of the nuchal crest large, those of dorsal crest reduced to mere denticulation towards the base of the tail, generally ending in the region between the mid-body and the tail, rarely continues to the base of the tail; nuchal, dorsal and supratympanic spines more pronounced in males; limbs slender, dorsal surface of the limbs strongly keeled, ventral surface weakly keeled, that of the thighs smooth.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="669">
|
||
The species can be separated from the members of
|
||
<normalizedToken originalValue="Smith’s">Smith's</normalizedToken>
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. versicolor" order="Squamata" pageId="0" pageNumber="669" rank="species" species="versicolor">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. versicolor</emphasis>
|
||
</taxonomicName>
|
||
group (as defined above), by a combination of characters: absence of crescent-shaped patch of granular scales at the insertion of the forelimbs (vs. present in
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. emma" order="Squamata" pageId="0" pageNumber="669" rank="species" species="emma">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. emma</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. grandisquamis" order="Squamata" pageId="0" pageNumber="669" rank="species" species="grandisquamis">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. grandisquamis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. jerdoni" order="Squamata" pageId="0" pageNumber="669" rank="species" species="jerdoni">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. jerdoni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. mystaceus" order="Squamata" pageId="0" pageNumber="669" rank="species" species="mystaceus">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. mystaceus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. nemoricola" order="Squamata" pageId="0" pageNumber="669" rank="species" species="nemoricola">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. nemoricola</emphasis>
|
||
</taxonomicName>
|
||
), 37-45 Mid-body scale rows (vs. 49-65 in
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. emma" order="Squamata" pageId="0" pageNumber="669" rank="species" species="emma">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. emma</emphasis>
|
||
</taxonomicName>
|
||
, 27-35 in
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. grandisquamis" order="Squamata" pageId="0" pageNumber="669" rank="species" species="grandisquamis">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. grandisquamis</emphasis>
|
||
</taxonomicName>
|
||
, 45-57 in
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. jerdoni" order="Squamata" pageId="0" pageNumber="669" rank="species" species="jerdoni">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. jerdoni</emphasis>
|
||
</taxonomicName>
|
||
, 58-63 in
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. maria" order="Squamata" pageId="0" pageNumber="669" rank="species" species="maria">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. maria</emphasis>
|
||
</taxonomicName>
|
||
Gray, 48-60 in
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. minor" order="Squamata" pageId="0" pageNumber="669" rank="species" species="minor">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. minor</emphasis>
|
||
</taxonomicName>
|
||
, and 45-58 in
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. mystaceus" order="Squamata" pageId="0" pageNumber="669" rank="species" species="mystaceus">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. mystaceus</emphasis>
|
||
</taxonomicName>
|
||
); nuchal and dorsal crest scales well developed, nuchal crest scales slightly larger than the dorsal crest scales, dorsal crest scales become progressively smaller towards the base of the tail (vs. nuchal spines much longer, dorsal spines reduced in
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. maria" order="Squamata" pageId="0" pageNumber="669" rank="species" species="maria">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. maria</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. nemoricola" order="Squamata" pageId="0" pageNumber="669" rank="species" species="nemoricola">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. nemoricola</emphasis>
|
||
</taxonomicName>
|
||
; nuchal spines much longer than dorsal spines in
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. calotes" order="Squamata" pageId="0" pageNumber="669" rank="species" species="calotes">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. calotes</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. emma" order="Squamata" pageId="0" pageNumber="669" rank="species" species="emma">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. emma</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. grandisquamis" order="Squamata" pageId="0" pageNumber="669" rank="species" species="grandisquamis">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. grandisquamis</emphasis>
|
||
</taxonomicName>
|
||
); two well-separated supratympanic clusters of spine-like scales, one from each cluster enlarged, prominent to form a spine (vs. row of 3-4 compressed supratympanic spines in
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. grandisquamis" order="Squamata" pageId="0" pageNumber="669" rank="species" species="grandisquamis">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. grandisquamis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. nemoricola" order="Squamata" pageId="0" pageNumber="669" rank="species" species="nemoricola">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. nemoricola</emphasis>
|
||
</taxonomicName>
|
||
, 8-9 compressed spines above tympanum in
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. calotes" order="Squamata" pageId="0" pageNumber="669" rank="species" species="calotes">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. calotes</emphasis>
|
||
</taxonomicName>
|
||
; two parallel rows of supratympanic scales in
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. jerdoni" order="Squamata" pageId="0" pageNumber="669" rank="species" species="jerdoni">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. jerdoni</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. maria" order="Squamata" pageId="0" pageNumber="669" rank="species" species="maria">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. maria</emphasis>
|
||
</taxonomicName>
|
||
). The species differs from
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. paulus" order="Squamata" pageId="0" pageNumber="669" rank="species" species="paulus">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. paulus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. zolaiking" order="Squamata" pageId="0" pageNumber="669" rank="species" species="zolaiking">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. zolaiking</emphasis>
|
||
</taxonomicName>
|
||
primarily by the homogeneous scalation on the dorsolateral region (vs. heterogeneous) and a comparatively well-developed dorsal crest. From the dubious species
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. bhutanensis" order="Squamata" pageId="0" pageNumber="669" rank="species" species="bhutanensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. bhutanensis</emphasis>
|
||
</taxonomicName>
|
||
, the species differs in possessing longer head, concave orbital region, and by the absence of a row of erect scales on the sides of the neck. From
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. chincollium" order="Squamata" pageId="0" pageNumber="669" rank="species" species="chincollium">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. chincollium</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. nigriplicatus" order="Squamata" pageId="0" pageNumber="669" rank="species" species="nigriplicatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. nigriplicatus</emphasis>
|
||
</taxonomicName>
|
||
, and other members of the
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. mystaceus" order="Squamata" pageId="0" pageNumber="669" rank="species" species="mystaceus">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. mystaceus</emphasis>
|
||
</taxonomicName>
|
||
complex (
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. bachae" order="Squamata" pageId="0" pageNumber="669" rank="species" species="bachae">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. bachae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. geissleri" order="Squamata" pageId="0" pageNumber="669" rank="species" species="geissleri">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. geissleri</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. goetzi" order="Squamata" pageId="0" pageNumber="669" rank="species" species="goetzi">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. goetzi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. mystaceus" order="Squamata" pageId="0" pageNumber="669" rank="species" species="mystaceus">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. mystaceus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. vindumbarbatus" order="Squamata" pageId="0" pageNumber="669" rank="species" species="vindumbarbatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. vindumbarbatus</emphasis>
|
||
</taxonomicName>
|
||
, sensu
|
||
<bibRefCitation author="Wagner, P" journalOrPublisher="Bonn Zoological Bulletin" pageId="0" pageNumber="669" pagination="141 - 171" refId="B68" refString="Wagner, P, Ihlow, F, Hartmann, T, Flecks, M, Schmitz, A, Boehme, W, 2021. Integrative approach to resolve Calotes mystaceus Dumeril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn Zoological Bulletin 70 (1): 141 - 171" title="Integrative approach to resolve Calotes mystaceus Dumeril & Bibron, 1837 species complex (Squamata: Agamidae)." volume="70" year="2021">Wagner et al. (2021)</bibRefCitation>
|
||
) the species differs by the absence of an oblique fold of skin in front of forelimbs or shoulder (vs. present). From the Sri Lankan congeners (
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. ceylonensis" order="Squamata" pageId="0" pageNumber="669" rank="species" species="ceylonensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. ceylonensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. desilvai" order="Squamata" pageId="0" pageNumber="669" rank="species" species="desilvai">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. desilvai</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. liocephalus" order="Squamata" pageId="0" pageNumber="669" rank="species" species="liocephalus">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. liocephalus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. liolepis" order="Squamata" pageId="0" pageNumber="669" rank="species" species="liolepis">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. liolepis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. manamendrai" order="Squamata" pageId="0" pageNumber="669" rank="species" species="manamendrai">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. manamendrai</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. nigrilabris" order="Squamata" pageId="0" pageNumber="669" rank="species" species="nigrilabris">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. nigrilabris</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. pethiyagodai" order="Squamata" pageId="0" pageNumber="669" rank="species" species="pethiyagodai">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. pethiyagodai</emphasis>
|
||
</taxonomicName>
|
||
) the species differs by its posterodorsal orientation of lateral body scales (vs. posteroventral) and absence of shoulder pit (vs. present). The species differs from
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. irawadi" order="Squamata" pageId="0" pageNumber="669" rank="species" species="irawadi">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. irawadi</emphasis>
|
||
</taxonomicName>
|
||
by its much larger adult male body size (average SVL 106 mm, vs. 82.4 mm), lesser number of scales around the mid-body (average 42, vs. 47 in
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. irawadi" order="Squamata" pageId="0" pageNumber="669" rank="species" species="irawadi">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. irawadi</emphasis>
|
||
</taxonomicName>
|
||
); from
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. htunwini" order="Squamata" pageId="0" pageNumber="669" rank="species" species="htunwini">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. htunwini</emphasis>
|
||
</taxonomicName>
|
||
by the posterodorsal or vertical orientation of scale rows on the sides of the neck and supra-axillary area (vs. horizontal in
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. htunwini" order="Squamata" pageId="0" pageNumber="669" rank="species" species="htunwini">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. htunwini</emphasis>
|
||
</taxonomicName>
|
||
). For comparison with the subspecies elevated to species rank in this communication, see the diagnosis and comparison section for that species.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="669">
|
||
The species can be further differentiated by its southern congener
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. versicolor" order="Squamata" pageId="0" pageNumber="669" rank="species" species="versicolor">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. versicolor</emphasis>
|
||
</taxonomicName>
|
||
by its slightly smaller adult body size (average male SVL 106 mm vs. 108 mm in
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. versicolor" order="Squamata" pageId="0" pageNumber="669" rank="species" species="versicolor">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. versicolor</emphasis>
|
||
</taxonomicName>
|
||
, female SVL 77.5 vs. 92.2 mm in
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. versicolor" order="Squamata" pageId="0" pageNumber="669" rank="species" species="versicolor">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. versicolor</emphasis>
|
||
</taxonomicName>
|
||
), dorsal crest composed of comparatively smaller scales, which become progressively smaller to the base of the tail in both sexes (vs. dorsal crest composed of large scales, which continues to the base of the tail in
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. versicolor" order="Squamata" pageId="0" pageNumber="669" rank="species" species="versicolor">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. versicolor</emphasis>
|
||
</taxonomicName>
|
||
), supratympanic spines shorter in both sexes (vs. longer in
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. versicolor" order="Squamata" pageId="0" pageNumber="669" rank="species" species="versicolor">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. versicolor</emphasis>
|
||
</taxonomicName>
|
||
). The species has shorter crus than
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. versicolor" order="Squamata" pageId="0" pageNumber="669" rank="species" species="versicolor">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. versicolor</emphasis>
|
||
</taxonomicName>
|
||
(average male CrusL 22.7 vs. 26.6 in
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. versicolor" order="Squamata" pageId="0" pageNumber="669" rank="species" species="versicolor">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. versicolor</emphasis>
|
||
</taxonomicName>
|
||
). The species differs in the overall shape of the trunk, which tapers to a lesser extent in
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. vultuosus" order="Squamata" pageId="0" pageNumber="669" rank="species" species="vultuosus">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. vultuosus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="669">comb. nov.</emphasis>
|
||
(average PectW/PelvW 0.81) vs. trunk tapers to a greater extent in
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. versicolor" order="Squamata" pageId="0" pageNumber="669" rank="species" species="versicolor">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. versicolor</emphasis>
|
||
</taxonomicName>
|
||
(average PectW/PelvW 0.70 in
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. versicolor" order="Squamata" pageId="0" pageNumber="669" rank="species" species="versicolor">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. versicolor</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="669">
|
||
Besides, the species differs from
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. versicolor" order="Squamata" pageId="0" pageNumber="669" rank="species" species="versicolor">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. versicolor</emphasis>
|
||
</taxonomicName>
|
||
in adult male coloration during the breeding season.
|
||
<taxonomicName authorityName="comb. nov. (Harlan" authorityYear="1825" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calotes vultuosus" order="Squamata" pageId="0" pageNumber="669" phylum="Chordata" rank="species" species="vultuosus">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">Calotes vultuosus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="669">comb. nov.</emphasis>
|
||
males generally attain a cream to brown body coloration, the head and the anterior two-thirds of the trunk attain orange color, which at times extends to the forelimbs; the posterior parts of the trunk and the hind limbs remain dull, whereas
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. versicolor" order="Squamata" pageId="0" pageNumber="669" rank="species" species="versicolor">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. versicolor</emphasis>
|
||
</taxonomicName>
|
||
males attain yellowish overall coloration, the trunk and the orbital region turns bright orange, forelimbs and hind limbs turn dark to black. Further, the black patches under the throat extend anteriorly onto the jaw musculature, at times running along the lower jaw margin on each side, before terminating posterior to the post-mental scales, whereas, in
|
||
<taxonomicName family="Agamidae" genus="C." kingdom="Animalia" lsidName="C. versicolor" order="Squamata" pageId="0" pageNumber="669" rank="species" species="versicolor">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">C. versicolor</emphasis>
|
||
</taxonomicName>
|
||
, the black patches under the throat do not extend anteriorly onto the jaw muscles.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="669" type="description">
|
||
<paragraph pageId="0" pageNumber="669">Description of the holotype ANSP 7296 (based on photographic data, Fig. 6).</paragraph>
|
||
<paragraph pageId="0" pageNumber="669">A medium sized male, SVL ~77 mm. Tail complete, TailL ~160 mm. The specimen is in a damaged condition, there are multiple openings on the mid-body and near the vent, all artefacts of preservation. Further, the specimen has developed randomly distributed white patches on the scales of the head and the body. The description is based on the images of the specimen and modified from the original description; hence the mensural details should be treated with caution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="669">Head moderately large (HL/SVL ~0.19), snout tip blunt, rounded in dorsal perspective; loreal region between the nasals and the orbit slightly concave, acute, covered by heterogeneous, juxtaposed scales, some of which are ridged; the area comprising the loreal region and nasal shield triangular from the lateral perspective; eight CanthR scales, elongate, with their ends overlapping; supraciliary and canthal edge sharp, giving the head a flat appearance laterally from the dorsal perspective; large supraocular scales do not form shields, become parallel to convergent at the supraocular region; edges of the head divergent bordered by scales of the canthus rostralis and supraciliary; rostral broader than high; nasal shields single on each side, subtriangular, pointed anteriorly, rounded posteriorly, separated from the first Suplab by one prenasal scale, from rostral by two scales, from each other by seven SnS; nostrils round, in a single large nasal shield each, centrally placed; scales between rostral and SnS small, juxtaposed; SnS heterogeneous, the median SnS smallest, the pair bordering the median scale elongate; scales of the forehead posterior to SnS sub-imbricate, very irregular in shape and size, some rugose; HeadSLn 12 bordered anteriorly by the rostral scale, posteriorly by a single, large interparietal shield; orbit encased in a sock of granular scales, separated from the nasal shield by 6-8 scale rows, from Suplab by 3-4 scale rows; HeadSTr 14, between the posteriormost supraciliary scales on each side, just anterior to the interparietal; eye opening bordered dorsally and ventrally by two rows of non-granular scales, outer row composed of larger, square-shaped scales, inner row similar in shape, slightly smaller; eyelid scales (Eyelid) 13; pupil round, large; region between the orbit and the tympanum covered by rows of 6-9 smooth, roughly hexagonal scales; tympanum large, round, naked; its greatest diameter roughly 42% of horizontal diameter of the orbit; supratympanic scales weakly keeled; two enlarged supratympanic spines, subequal in size, separated from each other by four scales; anterior spine slender, bent parallel to the horizontal plane, posterior spine erect, prominent, its apex bent; scales in a row connecting the enlarged supratympanic spines give the posterior region of the head a serrated appearance; posterior region of the jaws swollen, bulging out, covered by subtriangular, mucronate, imbricate, postero-ventrally directed scales, upper border of the jaw muscles conceal the lower portion of the tympanum; labial scales large, sub-rectangular; Suplab 10; Inflab 10; two parallel rows of scales border the upper margin of the Suplab, lower originates above the second Suplab, separating the nasal shield and the second Suplab, terminates near the last Suplab; the upper row originates slightly posteriorly, terminates abruptly above the sixth Suplab; interparietal large, irregularly pentagonal, posterior border straight, tapers toward the anteriority, anterior border nearly pointed; interparietal bordered by 10 smooth, heterogeneous scales; nuchal crest starts 3-4 scales behind the interparietal; mental shield large, single, subtriangular, narrower than rostral; two pairs of elongate postmentals, anterior pair joint, in contact with mental, narrow; posterior pair broader, separated by two small chin scales; chin scales behind the postmental scales small, progressively become larger towards the throat, median row of gular scales enlarged, mucronate, and form a longitudinal fold.</paragraph>
|
||
<paragraph pageId="0" pageNumber="669">First nuchal scale smallest, size of the nuchal scale progressively increases towards the median nuchal scale, nuchal scales beyond the median nuchal scale roughly the same size as, or slightly smaller than the median nuchal scale; nuchal crest composed of ~10 long, conical spines, scales recurved, continue into dorsal crest scales; dorsal crest composed of spines that progressively become smaller towards the middle of the back; continues as an obsolete row of vertebral scales to the base of the tail, beyond which the mid-dorsal crest row terminates at the keeled tail scale rows; paravertebral scales recurved, strongly keeled, mucronate; dorso-lateral scales sub-triangular, imbricate, moderately keeled, posterodorsally oriented; those in the supra-axillary region mostly dorsally oriented, slightly smaller than those of the mid-body; body at the mid-body to the pelvic region damaged; ventrals strongly keeled, smaller than the dorso-lateral scales, mucronate, imbricate; ventral scales between the insertion of the forelimbs weakly keeled, some with rounded apices.</paragraph>
|
||
<paragraph pageId="0" pageNumber="669">Limbs long and slender, covered with keeled scales, similar to dorso-laterals in shape and size, forming parallel longitudinal rows; scales on the dorsal surfaces of the thigh weakly keeled, ventral surfaces smooth, those on the crus and sole strongly keeled; hindlimbs longer, almost 80% of SVL; relative length of fingers 4>3>2>5>1, fourth slightly longer than third; relative length of toes 4>3>5>2>1; subdigital lamellae bicarinate, keels sharp; digits and subdigital lamellae slender, digits swollen at base; tail damaged ventrally near the vent, exposing the hemipenis; tail scales large, imbricate, strongly keeled on dorsal and ventral aspects, mucronate; TailL ~160 mm.</paragraph>
|
||
<paragraph pageId="0" pageNumber="669">Coloration in preservation: Dorsum generally dark brown, venter paler, light yellow; head dark brown dorsally, throat paler; orbit whitish, nasal shield, canthals and supraciliary region almost white; irregularly distributed white patches on the labials, throat, jaw muscles, axillary region, on the scales between the tympanum and the ear, the supratympanic region, and on the nuchal crest scales; the white patches continue on to the upper arm, lower arm, dorso-laterals, supraxillary region, and on the dorsal crest; white patches also present on the venter, the hindlimbs and the tail; tail scales show marks of corrosion due to preservative near the vent.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="669" type="description">
|
||
<paragraph pageId="0" pageNumber="669">Original description.</paragraph>
|
||
<paragraph pageId="0" pageNumber="669">Harlan R (1825). Journal of the Academy of Natural Sciences of Philadelphia 4(1): 296-305.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="669" type="etymology">
|
||
<paragraph pageId="0" pageNumber="669">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="669">
|
||
The specific epithet '
|
||
<emphasis italics="true" pageId="0" pageNumber="669">vultuosa</emphasis>
|
||
' could refer to the grim or frowning look of the species when viewed frontally, which it gets due to the flattened scales of the CanthR and the supercilium. The specific epithet
|
||
<taxonomicName class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calotes vultuosa" order="Squamata" pageId="0" pageNumber="669" phylum="Chordata" rank="species" species="vultuosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">Calotes vultuosa</emphasis>
|
||
</taxonomicName>
|
||
is feminine singular and is here changed to the masculine gender
|
||
<taxonomicName authorityName="comb. nov. (Harlan" authorityYear="1825" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calotes vultuosus" order="Squamata" pageId="0" pageNumber="669" phylum="Chordata" rank="species" species="vultuosus">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">Calotes vultuosus</emphasis>
|
||
</taxonomicName>
|
||
in agreement with the generic epithet,
|
||
<taxonomicName class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calotes" order="Squamata" pageId="0" pageNumber="669" phylum="Chordata" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="669">Calotes</emphasis>
|
||
</taxonomicName>
|
||
, which is masculine in gender.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="669" type="materials_examined">
|
||
<paragraph pageId="0" pageNumber="669">Variation in the material examined.</paragraph>
|
||
<paragraph pageId="0" pageNumber="669">
|
||
Eighteen male specimens were examined. The means for the mensural characters in mm are: HL 21.2; HW 25.3; HH 19.9; JawW 19.9; SnEye 10.9; NarEye 5.7; EyeEar 8.6; SnW 6.5; Interorb 11.1; SVL 106.0; TrunkL 47.1; TailH 16.4; TailW 13.1; PectW 18.4; PelvW 15.1; SnForeL 41.2; UpArmL 19.8; LoArmL 19.5; ForefL 19.2; 4FingLng 12.7; UpLegL 25.1; CrusL 23.1; HindfL 30.8; 4ToeLng 19.0; ForeLimbL 58.5; HindLimbL 79.1. The means for meristic characters are: SnS 7; HeadSTr 13; HeadSLn 14; CanthR 7; Eyelid 12; Suplab 11; Inflab 11; TempSp 2; Dorsal 48; Mid-body 42; 4FingLm 21; 4ToeLm 26. Additionally,
|
||
<specimenCount type="female">two female</specimenCount>
|
||
specimens were examined. The means for the mensural characters in mm are: HL 17.4; HW 13; HH 12.4; JawW 13; SnEye 8.1; NarEye 4.1; EyeEar 4.8; SnW 4.5; Interorb 8.7; SVL 77; TrunkL 36.8; TailH 6.3; TailW 5.9; PectW 11.5; PelvW 9.6; SnForeL 28.9; UpArmL 16.6; LoArmL 13.5; ForefL 14.7; 4FingLng 10.7; UpLegL 17.6; CrusL 17.9; HindfL 23.7; 4ToeLng 16.3; ForeLimbL 44.3; HindLimbL 59.3. The means for meristic characters are: SnS 6; HeadSTr 13; HeadSLn 13; CanthR 7; Eyelid 10; Suplab 10; Inflab 10; TempSp 2; Dorsal 45; Mid-body 40; 4FingLm 18; 4ToeLm 25. The ranges for each of these characters are given in Table
|
||
<tableCitation captionStart="Table 4" captionStartId="T4" captionText="Table 4. Mensural and meristic characters of the type and non-type specimens of Calotes vultuosus comb. nov. and Calotes farooqi stat. nov., mensural characters recorded as a ratio of SVL. Note that the morphometric data for the types of A. indica were not available, whereas those for C. farooqi have been sourced from Auffenberg and Rehman (1993). - indicates that measurements / counts could not be taken." httpUri="http://table.plazi.org/id/2180D7EC720D2D52B3FF2C140ADD46FF" pageId="0" pageNumber="669" tableUuid="2180D7EC720D2D52B3FF2C140ADD46FF">4</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="669" type="distribution">
|
||
<paragraph pageId="0" pageNumber="669">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="669">
|
||
The species appears to be widely distributed, occupying parts of the Dharwar Craton till Brahmagiri, Deccan Volcanic Province, the Central Highlands, the Gangetic Plains, and the Indian Deserts (Fig.
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. A map of south and Southeast Asia representing the localities from which morphological and / or primary or secondary genetic data were collected. Red circles indicate the localities from which C. versicolor sensu stricto (Clade 1) originate, yellow samples indicate those of C. irawadi sensu lato (Clade 2), blue circles represent the localities of C. vultuosus comb. nov. (Clade 3), green circles indicate Clade 4, with C. farooqi stat. nov. (77, 83, 85, 86) and C. aff. farooqi (78, 79). A single sequence of Calotes irawadi from Huanan Province, China (locality 73) is not marked on the map, since it was very distant to all other localities, as also those whose precise localities were unknown. Triangles indicate type localities of the respective species. See supplementary Table S 1 for details on molecular and / or morphological vouchers used, and their respective localities." figureDoi="10.3897/vz.71.e62787.figure1" httpUri="https://binary.pensoft.net/fig/600928" pageId="0" pageNumber="669">1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |