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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1089.77203" ID-GBIF-Dataset="6d9fb3a1-d81a-4126-b7b5-41bae5b7173d" ID-PMC="PMC8948171" ID-Pensoft-Pub="1313-2970-1089-109" ID-Pensoft-UUID="C5F28A44BCB15640AD5D7B738E36C60A" ID-PubMed="35586598" ID-ZooBank="539FC2362F874F67AFA7A079C5027522" ModsDocID="1313-2970-1089-109" checkinTime="1647574147317" checkinUser="pensoft" docAuthor="Min, Rui, Zhao, Yahui, Shi, Jingsong &amp; Yang, Junxing" docDate="2022" docId="EEDA864C0C475F78A3EAD3865BBDB76D" docLanguage="en" docName="ZooKeys 1089: 109-124" docOrigin="ZooKeys 1089" docPubDate="2022-03-17" docSource="http://dx.doi.org/10.3897/zookeys.1089.77203" docTitle="Homatula robusta Min &amp; Zhao &amp; Shi &amp; Yang 2022, sp. nov." docType="treatment" docUuid="A27C86D0-FD58-448C-9A85-6129B1A18F66" docUuidSource="ZooBank" docVersion="5" id="C5F28A44BCB15640AD5D7B738E36C60A" lastPageNumber="109" masterDocId="C5F28A44BCB15640AD5D7B738E36C60A" masterDocTitle="A new species of Homatula (Teleostei, Cobitoidea, Nemacheilidae) from the Pearl River drainage, Yunnan, China" masterLastPageNumber="124" masterPageNumber="109" pageNumber="109" updateTime="1668151536174" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new species of Homatula (Teleostei, Cobitoidea, Nemacheilidae) from the Pearl River drainage, Yunnan, China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Min, Rui</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5893-4936</mods:nameIdentifier>
<mods:affiliation>Kunming Natural History Museum of Zoology, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan 650223, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhao, Yahui</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4615-596X</mods:nameIdentifier>
<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shi, Jingsong</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9168-1734</mods:nameIdentifier>
<mods:affiliation>Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing 100044, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yang, Junxing</mods:namePart>
<mods:affiliation>State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, The Innovative Academy of Seed Design, Chinese Academy of Sciences, Kunming, Yunnan 650223, China &amp; Yunnan Key Laboratory of Plateau Fish Breeding, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan 650223, China</mods:affiliation>
<mods:nameIdentifier type="email">yangjx@mail.kiz.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-03-17</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1089</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>109</mods:start>
<mods:end>124</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1089.77203</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1089.77203</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1089-109</mods:identifier>
<mods:identifier type="ZooBank">539FC2362F874F67AFA7A079C5027522</mods:identifier>
<mods:identifier type="Pensoft-UUID">C5F28A44BCB15640AD5D7B738E36C60A</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="194097467" LSID="urn:lsid:zoobank.org:act:A27C86D0-FD58-448C-9A85-6129B1A18F66" httpUri="http://treatment.plazi.org/id/EEDA864C0C475F78A3EAD3865BBDB76D" lastPageNumber="109" pageId="0" pageNumber="109">
<subSubSection pageId="0" pageNumber="109" type="nomenclature">
<paragraph pageId="0" pageNumber="109">
<taxonomicName LSID="http://zoobank.org/A27C86D0-FD58-448C-9A85-6129B1A18F66" authority="Min &amp; Zhao &amp; Shi &amp; Yang, 2022" authorityName="Min &amp; Zhao &amp; Shi &amp; Yang" authorityYear="2022" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Homatula robusta" order="Cypriniformes" pageId="0" pageNumber="109" phylum="Chordata" rank="species" species="robusta" status="sp. nov.">Homatula robusta</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="109">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="109" type="description">
<paragraph pageId="0" pageNumber="109">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Lateral A dorsal B and ventral views C of H. robusta sp. nov., holotype, KIZ 2009000125, 83.12 mm SL." figureDoi="10.3897/zookeys.1089.77203.figure1" httpUri="https://binary.pensoft.net/fig/659924" pageId="0" pageNumber="109">Figs 1</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Homatula robusta sp. nov., paratype, KIZ 2009000122, 81.32 mm SL, head in lateral A dorsal B and ventral C views." figureDoi="10.3897/zookeys.1089.77203.figure2" httpUri="https://binary.pensoft.net/fig/659925" pageId="0" pageNumber="109">, 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lateral view of H. robusta sp. nov. KIZ 2009000122 paratype A H. longidorsalis KIZ 1987003989 B H. nanpanjiangensis KIZ 1994000023 C H. potanini KIZ 2010000266 D." figureDoi="10.3897/zookeys.1089.77203.figure3" httpUri="https://binary.pensoft.net/fig/659926" pageId="0" pageNumber="109">, 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lateral CT scans of: A H. robusta sp. nov., KIZ 2009000125, lateral view and structure of caudal skeletons B H. nanpanjiangensis, KIZ 1994000029, lateral view and structure of caudal skeletons C H. wenshanensis, KIZ 2014005686, lateral view. Abbreviations: ph-partypurale, ns-neural spine, hs-haemal spine, h-hypural, epu-epurale, pst-pleurostyle." figureDoi="10.3897/zookeys.1089.77203.figure4" httpUri="https://binary.pensoft.net/fig/659927" pageId="0" pageNumber="109">, 4</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Distribution of Homatula from the Nanpanjiang River." figureDoi="10.3897/zookeys.1089.77203.figure5" httpUri="https://binary.pensoft.net/fig/659928" pageId="0" pageNumber="109">, 5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="109" type="materials_examined">
<paragraph pageId="0" pageNumber="109">Material.</paragraph>
<paragraph pageId="0" pageNumber="109">
<materialsCitation collectingDate="2009-03-14" collectorName="Wansheng Jiang, Weiying Wang" country="China" county="Luoping County" elevation="1210" latitude="25.033333" location="Changdi village" longLatPrecision="1248" longitude="104.5" municipality="upper Pearl River" specimenCount="1" stateProvince="Yunnan Province" typeStatus="Holotype">
<emphasis bold="true" pageId="0" pageNumber="109">
<emphasis italics="true" pageId="0" pageNumber="109">
<typeStatus>Holotype</typeStatus>
</emphasis>
.
</emphasis>
KIZ 2009000125,
<quantity metricMagnitude="-2" metricUnit="m" metricValue="8.312" unit="mm" value="83.12">83.12 mm</quantity>
SL; collected by
<collectorName>Wansheng Jiang</collectorName>
and
<collectorName>Weiying Wang</collectorName>
on
<collectingDate value="2009-03-14">14 March 2009</collectingDate>
at
<location LSID="urn:lsid:plazi:treatment:EEDA864C0C475F78A3EAD3865BBDB76D:E90A1BF0B2D09CCE65ADC65E287F2111" country="China" county="Luoping County" latitude="25.033333" longLatPrecision="1248" longitude="104.5" municipality="upper Pearl River" name="Changdi village" stateProvince="Yunnan Province">Changdi village</location>
,
<collectingCounty>Luoping County</collectingCounty>
,
<location LSID="urn:lsid:plazi:treatment:EEDA864C0C475F78A3EAD3865BBDB76D:29379086D9EF83390674A934946804B6" country="China" county="Luoping County" latitude="25.033333" longLatPrecision="1248" longitude="104.5" municipality="upper Pearl River" name="Qujing City" stateProvince="Yunnan Province">Qujing City</location>
,
<collectingRegion country="China" name="Yunnan">Yunnan Province</collectingRegion>
,
<collectingCountry name="China">China</collectingCountry>
;
<location LSID="urn:lsid:plazi:treatment:EEDA864C0C475F78A3EAD3865BBDB76D:31CFB36275CBEE1E8F25DAB2EE8D949A" country="China" county="Luoping County" latitude="25.033333" longLatPrecision="1248" longitude="104.5" municipality="upper Pearl River" name="Nanpanjiang River" stateProvince="Yunnan Province">Nanpanjiang River</location>
,
<collectingMunicipality>upper Pearl River</collectingMunicipality>
(
<geoCoordinate degrees="25" direction="north" minutes="02" orientation="latitude" precision="925" value="25.033333">25°02'N</geoCoordinate>
,
<geoCoordinate degrees="104" direction="east" minutes="30" orientation="longitude" precision="925" value="104.5">104°30'E</geoCoordinate>
; ca
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.21" unit="m" value="1210.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.21" unit="m" value="1210.0">1210 m</elevation>
</quantity>
)
</materialsCitation>
.
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Wansheng Jiang &amp; Weiying Wang" country="China" county="Luoping County" elevation="1210" latitude="25.033333" location="Changdi village" longLatPrecision="1248" longitude="104.5" municipality="upper Pearl River" specimenCount="3" stateProvince="Yunnan" typeStatus="Paratypes">
<emphasis bold="true" pageId="0" pageNumber="109">
<emphasis italics="true" pageId="0" pageNumber="109">
<typeStatus>Paratypes</typeStatus>
</emphasis>
.
</emphasis>
KIZ 2009000122, 144, 146,
<specimenCount type="generic">3 ex.</specimenCount>
<quantity metricMagnitude="-2" metricUnit="m" metricValue="7.124" metricValueMax="8.132" metricValueMin="6.116" unit="mm" value="71.24" valueMax="81.32" valueMin="61.16">61.16-81.32 mm</quantity>
SL, same data as for holotype
</materialsCitation>
.
</paragraph>
<caption doi="10.3897/zookeys.1089.77203.figure1" httpUri="https://binary.pensoft.net/fig/659924" pageId="0" pageNumber="109" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="109">
<emphasis bold="true" pageId="0" pageNumber="109">Figure 1.</emphasis>
Lateral
<emphasis bold="true" pageId="0" pageNumber="109">A</emphasis>
dorsal
<emphasis bold="true" pageId="0" pageNumber="109">B</emphasis>
and ventral views
<emphasis bold="true" pageId="0" pageNumber="109">C</emphasis>
of
<taxonomicName class="Actinopterygii" kingdom="Animalia" lsidName="H. robusta" order="Cypriniformes" pageId="0" pageNumber="109" phylum="Chordata" rank="species" species="robusta">
<emphasis italics="true" pageId="0" pageNumber="109">H. robusta</emphasis>
</taxonomicName>
sp. nov.,
<typeStatus>holotype</typeStatus>
, KIZ 2009000125,
<quantity metricMagnitude="-2" metricUnit="m" metricValue="8.312" unit="mm" value="83.12">83.12 mm</quantity>
SL.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="109" type="diagnosis">
<paragraph pageId="0" pageNumber="109">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="109">
The new species can be distinguished from all other species of
<taxonomicName authorityName="Nichols" authorityYear="1925" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Homatula" order="Cypriniformes" pageId="0" pageNumber="109" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="109">Homatula</emphasis>
</taxonomicName>
by having the following combination of characters: naked and robust body with well-developed crests (caudal peduncle depth as a percentage of its length: 70.5-78.5%), lateral line complete, median notch on lower jaw, median gap on lower lip, three pairs of short barbels, with maxillary barbels extending posteriorly to the anterior edge of eyes, branched dorsal-fin rays
<normalizedToken originalValue="8½">81/2</normalizedToken>
, emarginated caudal fin, and vertebrae 37-39.
</paragraph>
<caption doi="10.3897/zookeys.1089.77203.figure2" httpUri="https://binary.pensoft.net/fig/659925" pageId="0" pageNumber="109" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="109">
<emphasis bold="true" pageId="0" pageNumber="109">Figure 2.</emphasis>
<taxonomicName authorityName="Min &amp; Zhao &amp; Shi &amp; Yang" authorityYear="2022" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Homatula robusta" order="Cypriniformes" pageId="0" pageNumber="109" phylum="Chordata" rank="species" species="robusta">
<emphasis italics="true" pageId="0" pageNumber="109">Homatula robusta</emphasis>
</taxonomicName>
sp. nov., paratype, KIZ 2009000122, 81.32 mm SL, head in lateral
<emphasis bold="true" pageId="0" pageNumber="109">A</emphasis>
dorsal
<emphasis bold="true" pageId="0" pageNumber="109">B</emphasis>
and ventral
<emphasis bold="true" pageId="0" pageNumber="109">C</emphasis>
views.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="109" type="description">
<paragraph pageId="0" pageNumber="109">Description.</paragraph>
<paragraph pageId="0" pageNumber="109">Anterior body cylindrical, posterior body laterally compressed; robust, depth 5.8-6.3 times in length. Caudal peduncle stout, depth 1.27-1.42 times in its length. Crests on dorsal and ventral midlines present and supported by rudimentary procurrent caudal-fin rays; dorsal crest starting immediately posterior of dorsal-fin base, ventral crest starting immediately posterior of anal-fin base.</paragraph>
<caption doi="10.3897/zookeys.1089.77203.figure3" httpUri="https://binary.pensoft.net/fig/659926" pageId="0" pageNumber="109" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="109">
<emphasis bold="true" pageId="0" pageNumber="109">Figure 3.</emphasis>
Lateral view of
<taxonomicName class="Actinopterygii" kingdom="Animalia" lsidName="H. robusta" order="Cypriniformes" pageId="0" pageNumber="109" phylum="Chordata" rank="species" species="robusta">
<emphasis italics="true" pageId="0" pageNumber="109">H. robusta</emphasis>
</taxonomicName>
sp. nov. KIZ 2009000122 paratype
<emphasis bold="true" pageId="0" pageNumber="109">A</emphasis>
<taxonomicName class="Actinopterygii" kingdom="Animalia" lsidName="H. longidorsalis" order="Cypriniformes" pageId="0" pageNumber="109" phylum="Chordata" rank="species" species="longidorsalis">
<emphasis italics="true" pageId="0" pageNumber="109">H. longidorsalis</emphasis>
</taxonomicName>
KIZ 1987003989
<emphasis bold="true" pageId="0" pageNumber="109">B</emphasis>
<taxonomicName class="Actinopterygii" kingdom="Animalia" lsidName="H. nanpanjiangensis" order="Cypriniformes" pageId="0" pageNumber="109" phylum="Chordata" rank="species" species="nanpanjiangensis">
<emphasis italics="true" pageId="0" pageNumber="109">H. nanpanjiangensis</emphasis>
</taxonomicName>
KIZ 1994000023
<emphasis bold="true" pageId="0" pageNumber="109">C</emphasis>
<taxonomicName class="Actinopterygii" kingdom="Animalia" lsidName="H. potanini" order="Cypriniformes" pageId="0" pageNumber="109" phylum="Chordata" rank="species" species="potanini">
<emphasis italics="true" pageId="0" pageNumber="109">H. potanini</emphasis>
</taxonomicName>
KIZ 2010000266
<emphasis bold="true" pageId="0" pageNumber="109">D</emphasis>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="109">Snout blunt in lateral view, cheeks inflated. Eyes elliptical horizontally, dorsolaterally positioned. Mouth inferior, slightly arched. Anterior nostril in flap, next to posterior nostril. Lips moderately thick, upper lip smooth, slightly notched medially, lower lip with shallow furrows and median gap. Processus dentiformis on upper jaw present with circular arc edge; lower jaw spoon-like with a median notch. Three pairs of barbels, maxillary barbel reaching anterior margin of eye, outer rostral barbel reaching inner corner of mouth and inner rostral barbel not.</paragraph>
<caption doi="10.3897/zookeys.1089.77203.figure4" httpUri="https://binary.pensoft.net/fig/659927" pageId="0" pageNumber="109" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="109">
<emphasis bold="true" pageId="0" pageNumber="109">Figure 4.</emphasis>
Lateral CT scans of:
<emphasis bold="true" pageId="0" pageNumber="109">A</emphasis>
<taxonomicName class="Actinopterygii" kingdom="Animalia" lsidName="H. robusta" order="Cypriniformes" pageId="0" pageNumber="109" phylum="Chordata" rank="species" species="robusta">
<emphasis italics="true" pageId="0" pageNumber="109">H. robusta</emphasis>
</taxonomicName>
sp. nov., KIZ 2009000125, lateral view and structure of caudal skeletons
<emphasis bold="true" pageId="0" pageNumber="109">B</emphasis>
<taxonomicName class="Actinopterygii" kingdom="Animalia" lsidName="H. nanpanjiangensis" order="Cypriniformes" pageId="0" pageNumber="109" phylum="Chordata" rank="species" species="nanpanjiangensis">
<emphasis italics="true" pageId="0" pageNumber="109">H. nanpanjiangensis</emphasis>
</taxonomicName>
, KIZ 1994000029, lateral view and structure of caudal skeletons
<emphasis bold="true" pageId="0" pageNumber="109">C</emphasis>
<taxonomicName class="Actinopterygii" kingdom="Animalia" lsidName="H. wenshanensis" order="Cypriniformes" pageId="0" pageNumber="109" phylum="Chordata" rank="species" species="wenshanensis">
<emphasis italics="true" pageId="0" pageNumber="109">H. wenshanensis</emphasis>
</taxonomicName>
, KIZ 2014005686, lateral view. Abbreviations: ph-partypurale, ns-neural spine, hs-haemal spine, h-hypural, epu-epurale, pst-pleurostyle.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="109">
Dorsal-fin rays iv,
<normalizedToken originalValue="8½">81/2</normalizedToken>
, distal margin slightly convex. Pectoral-fin rays 11, reaching about halfway from insertion of pectoral fin to insertion of pelvic fin. Pelvic-fin rays 6-8, reaching close to anus, inserted opposite of the first branched dorsal-fin ray. Anus located 1.53-2.17 times eye diameter in front of anal-fin origin. Anal-fin rays iii,
<normalizedToken originalValue="5½">51/2</normalizedToken>
. Caudal-fin rays 9+8, distal margin of caudal fin emarginated with upper and lower lobes almost equal in length. Moderate axillary pelvic lobe with free tip.
</paragraph>
<caption ID-Table-UUID="EE40A515BAF2D86CDB5C021112581663" httpUri="http://table.plazi.org/id/EE40A515BAF2D86CDB5C021112581663" pageId="0" pageNumber="109" start="Table 2" startId="T2">
<paragraph pageId="0" pageNumber="109">
<emphasis bold="true" pageId="0" pageNumber="109">Table 2.</emphasis>
Measurements of four type specimens of
<taxonomicName authorityName="Min &amp; Zhao &amp; Shi &amp; Yang" authorityYear="2022" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Homatula robusta" order="Cypriniformes" pageId="0" pageNumber="109" phylum="Chordata" rank="species" species="robusta">
<emphasis italics="true" pageId="0" pageNumber="109">Homatula robusta</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="109">
<table pageId="0" pageNumber="109">
<tr pageId="0" pageNumber="109">
<th colspan="1" pageId="0" pageNumber="109" rowspan="2">Measurements (mm)</th>
<th colspan="4" pageId="0" pageNumber="109" rowspan="1">
<taxonomicName class="Actinopterygii" kingdom="Animalia" lsidName="H. robusta" order="Cypriniformes" pageId="0" pageNumber="109" phylum="Chordata" rank="species" species="robusta">
<emphasis italics="true" pageId="0" pageNumber="109">H. robusta</emphasis>
</taxonomicName>
</th>
</tr>
<tr pageId="0" pageNumber="109">
<th colspan="1" pageId="0" pageNumber="109" rowspan="1">2009000125</th>
<th colspan="1" pageId="0" pageNumber="109" rowspan="1">2009000122</th>
<th colspan="1" pageId="0" pageNumber="109" rowspan="1">2009000144</th>
<th colspan="1" pageId="0" pageNumber="109" rowspan="1">2009000146</th>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">SL</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">83.12</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">81.32</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">73.70</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">61.16</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">Head length</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">18.76</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">18.28</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">16.58</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">13.28</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">Predorsal length</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">38.78</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">37.34</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">35.16</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">29.06</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">Preventral length</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">41.80</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">40.94</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">37.06</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">29.98</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">Preanal length</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">60.42</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">60.46</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">54.52</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">45.60</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">Preanus length</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">55.92</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">56.00</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">50.16</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">41.96</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">Body depth</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">14.40</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">13.82</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">11.72</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">10.28</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">Caudal peduncle length (CPL)</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">16.56</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">14.78</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">13.32</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">9.96</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">Caudal peduncle depth (CPD)</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">11.68</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">11.60</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">10.20</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">7.04</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">Body width</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">9.36</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">9.38</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">8.66</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">7.06</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">Dorsal-fin length</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">11.28</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">11.48</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">10.24</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">7.12</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">Anal-fin length</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">12.32</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">11.94</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">11.58</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">9.14</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">Pelvic-fin length</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">12.12</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">12.28</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">11.22</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">8.72</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">Pectoral-fin length</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">14.82</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">14.76</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">13.50</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">11.12</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">Head depth at neck</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">11.84</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">11.72</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">10.70</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">8.56</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">Snout length</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">8.64</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">8.48</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">7.48</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">6.10</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">Head width at eye</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">13.12</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">13.34</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">12.40</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">9.46</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">Max head width</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">14.08</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">13.90</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">12.88</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">10.78</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">Interorbital width</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">4.30</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">4.44</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">4.24</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">3.74</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">eye diameter</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">2.62</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">2.34</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">2.28</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">2.10</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="109">Body scaleless, or sparse scales scattered along lateral line after posterior end of anal-fiin base, embedded beneath skin. Lateral line completed with 85-89 pores. Supraorbital pores 7, postorbital pores 3, sub- and preorbital pores 12, preoperculo-mandibular pores 10, supratemporal pores 3.</paragraph>
<paragraph pageId="0" pageNumber="109">Vertebrae (three specimens), 4+37-39; four hypural elements with h-1 &amp; h-2 fused, epurale present, last four neural spines (ns-1 to ns-4) and last three haemal spines (hs-1 to hs-3) on the caudal vertebrae are significantly enlarged. U-shaped stomach; intestine almost straight, with small bend next to stomach posterior. Longest recorded length is 83.1 mm SL, 95.7 mm total length (KIZ 2009000125, holotype).</paragraph>
<caption ID-Table-UUID="2B3F0EEBD95CF5A2003EF44C633C58A4" httpUri="http://table.plazi.org/id/2B3F0EEBD95CF5A2003EF44C633C58A4" pageId="0" pageNumber="109" start="Table 3" startId="T3">
<paragraph pageId="0" pageNumber="109">
<emphasis bold="true" pageId="0" pageNumber="109">Table 3.</emphasis>
Morphometrics of
<taxonomicName authorityName="Min &amp; Zhao &amp; Shi &amp; Yang" authorityYear="2022" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Homatula robusta" order="Cypriniformes" pageId="0" pageNumber="109" phylum="Chordata" rank="species" species="robusta">
<emphasis italics="true" pageId="0" pageNumber="109">Homatula robusta</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName baseAuthorityName="Min, Chen &amp; Yang" baseAuthorityYear="2010" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Homatula nanpanjiangensis" order="Cypriniformes" pageId="0" pageNumber="109" phylum="Chordata" rank="species" species="nanpanjiangensis">
<emphasis italics="true" pageId="0" pageNumber="109">Homatula nanpanjiangensis</emphasis>
</taxonomicName>
. SD, standard deviation.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="109">
<table pageId="0" pageNumber="109">
<tr pageId="0" pageNumber="109">
<th colspan="1" isEmpty="true" pageId="0" pageNumber="109" rowspan="1">-</th>
<th colspan="5" pageId="0" pageNumber="109" rowspan="1">
<taxonomicName class="Actinopterygii" kingdom="Animalia" lsidName="H. robusta" order="Cypriniformes" pageId="0" pageNumber="109" phylum="Chordata" rank="species" species="robusta">
<emphasis italics="true" pageId="0" pageNumber="109">H. robusta</emphasis>
</taxonomicName>
</th>
<th colspan="5" pageId="0" pageNumber="109" rowspan="1">
<taxonomicName class="Actinopterygii" kingdom="Animalia" lsidName="H. nanpanjiangensis" order="Cypriniformes" pageId="0" pageNumber="109" phylum="Chordata" rank="species" species="nanpanjiangensis">
<emphasis italics="true" pageId="0" pageNumber="109">H. nanpanjiangensis</emphasis>
</taxonomicName>
</th>
</tr>
<tr pageId="0" pageNumber="109">
<th colspan="1" pageId="0" pageNumber="109" rowspan="1">Measurements</th>
<th colspan="1" pageId="0" pageNumber="109" rowspan="1">N</th>
<th colspan="1" pageId="0" pageNumber="109" rowspan="1">Min</th>
<th colspan="1" pageId="0" pageNumber="109" rowspan="1">Max</th>
<th colspan="1" pageId="0" pageNumber="109" rowspan="1">Mean</th>
<th colspan="1" pageId="0" pageNumber="109" rowspan="1">SD</th>
<th colspan="1" pageId="0" pageNumber="109" rowspan="1">N</th>
<th colspan="1" pageId="0" pageNumber="109" rowspan="1">Min</th>
<th colspan="1" pageId="0" pageNumber="109" rowspan="1">Max</th>
<th colspan="1" pageId="0" pageNumber="109" rowspan="1">Mean</th>
<th colspan="1" pageId="0" pageNumber="109" rowspan="1">SD</th>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">SL (mm)</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">61.1</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">83.12</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">74.83</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">8.65</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">19</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">63.82</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">88.74</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">78.37</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">7.56</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="11" pageId="0" pageNumber="109" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="109">As percent of SL</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">Head length</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">13.28</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">18.76</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">16.73</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">2.15</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">19</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">15.46</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">20.36</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">18.63</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">1.53</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">Predorsal length</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">45.92</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">47.71</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">46.95</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">0.72</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">19</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">45.62</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">50.47</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">47.98</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">1.37</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">Preventral length</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">49.02</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">50.34</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">49.98</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">0.56</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">19</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">49.58</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">54.03</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">51.14</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">1.14</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">Preanal length</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">72.69</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">74.56</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">73.89</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">0.73</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">19</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">71.25</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">76.94</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">74.37</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">1.47</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">Preanus length</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">67.28</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">68.86</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">68.20</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">0.61</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">19</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">66.55</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">70.46</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">69.03</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">1.01</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">Body depth</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">15.9</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">17.32</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">16.76</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">0.53</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">19</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">12.64</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">15.30</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">13.77</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">0.73</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">Caudal peduncle length (CPL)</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">16.29</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">19.92</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">18.11</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">1.29</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">19</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">16.49</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">20.92</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">18.64</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">1.06</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">Caudal peduncle depth (CPD)</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">11.51</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">14.26</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">13.42</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">1.11</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">19</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">9.45</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">12.06</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">10.83</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">0.66</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">Body width</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">11.26</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">11.75</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">11.52</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">0.17</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">19</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">9.24</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">13.65</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">10.73</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">1.10</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">Dorsal-fin length</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">11.64</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">14.12</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">13.31</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">0.98</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">7</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">12.00</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">15.83</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">14.21</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">1.19</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">Anal-fin length</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">14.68</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">15.71</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">15.04</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">0.40</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">6</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">14.47</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">16.99</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">15.56</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">0.82</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">Pelvic-fin length</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">14.26</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">15.22</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">14.79</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">0.39</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">6</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">13.47</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">15.67</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">14.40</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">0.68</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">Pectoral-fin length</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">17.83</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">18.32</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">18.12</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">0.18</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">6</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">15.35</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">19.30</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">17.49</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">1.16</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="11" pageId="0" pageNumber="109" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="109">As percent of head length</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">Head depth at neck</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">63.11</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">64.54</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">64.06</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">0.57</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">19</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">49.88</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">61.19</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">55.01</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">3.79</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">Snout length</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">45.11</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">46.39</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">45.87</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">0.47</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">19</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">40.92</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">47.69</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">44.16</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">2.28</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">Head width at eye</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">69.94</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">74.79</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">72.23</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">1.83</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">19</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">49.63</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">72.79</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">59.56</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">6.77</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">Max head width</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">75.05</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">81.17</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">77.49</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">2.33</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">19</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">63.22</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">77.27</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">69.29</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">3.72</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">Interorbital width</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">22.92</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">28.16</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">25.24</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">1.93</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">19</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">22.21</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">26.67</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">24.29</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">1.17</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">Eye diameter</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">12.8</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">15.81</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">14.08</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">1.09</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">19</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">13.79</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">17.94</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">15.95</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">1.05</td>
</tr>
<tr pageId="0" pageNumber="109">
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="109">CPD/CPL (%)</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">70.53</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">78.48</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">74.07</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">3.53</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">19</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">49.50</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">65.66</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">58.22</td>
<td colspan="1" pageId="0" pageNumber="109" rowspan="1">4.22</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="109" type="coloration of preserved specimens">
<paragraph pageId="0" pageNumber="109">Coloration of preserved specimens.</paragraph>
<paragraph pageId="0" pageNumber="109">Body light brown with vertical brown bars in formalin-fixed specimens. Bars on predorsal body usually blurred and indistinct, or countable and separated by extraordinary narrow interspaces just in KIZ 2009000122. Bars and interspaces getting wider towards caudal-fin base and approximately equal width on posterior body. Usually, bars regularly shaped and jointed on dorsal midline from opposite sides, or two bars met on upper body and last bar diffused or formed by two combined bars (KIZ 2009000122). Dark black bar on caudal-fin base, reaching dorsal and ventral extremities. All fin rays pale brown and covered by melanophores, series of stripes halfway up each dorsal-fin ray. Color in alcohol-fixed specimens is paler than those in formalin-fixed specimens.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="109" type="sexual dimorphism">
<paragraph pageId="0" pageNumber="109">Sexual dimorphism.</paragraph>
<paragraph pageId="0" pageNumber="109">No sexual dimorphism was observed.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="109" type="etymology">
<paragraph pageId="0" pageNumber="109">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="109">
Robusta is a Latin word meaning
<normalizedToken originalValue="strong">'strong'</normalizedToken>
, in reference to the stout body and caudal peduncle. The Chinese common name is suggested as
<normalizedToken originalValue="粗壮荷马条鳅">粗壮荷马条鳅</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="109" type="distribution">
<paragraph pageId="0" pageNumber="109">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="109">
Only known from the type locality, Changdi village, Luoping County, Qujing City, Yunnan Province, China (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Distribution of Homatula from the Nanpanjiang River." figureDoi="10.3897/zookeys.1089.77203.figure5" httpUri="https://binary.pensoft.net/fig/659928" pageId="0" pageNumber="109">5</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1089.77203.figure5" httpUri="https://binary.pensoft.net/fig/659928" pageId="0" pageNumber="109" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="109">
<emphasis bold="true" pageId="0" pageNumber="109">Figure 5.</emphasis>
Distribution of
<taxonomicName authorityName="Nichols" authorityYear="1925" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Homatula" order="Cypriniformes" pageId="0" pageNumber="109" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="109">Homatula</emphasis>
</taxonomicName>
from the Nanpanjiang River.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>