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<document id="AA269DB19FDBB56DD2F22B6851809CAC" ID-CLB-Dataset="25134" ID-DOI="10.11646/zootaxa.4392.1.11" ID-GBIF-Dataset="6da59c8b-a336-4b16-84fb-66b22e06e129" ID-ISSN="1175-5326" ID-Zenodo-Dep="1194398" ID-ZooBank="E92FF3D8-B86B-4ED4-A074-5B266FE0711B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1520538817317" checkinUser="plazi" docAuthor="Carneiro, Eduardo, Dolibaina, Diego R., Grishin, Nick V. &amp; Warren, Andrew D." docDate="2018" docId="EB1F87ACDB58FFF4FF19A0323DF0D32A" docLanguage="en" docName="zootaxa.4392.1.11.pdf.imf" docOrigin="Zootaxa 4392 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Wahydra graslieae Carneiro &amp; Dolibaina &amp; Grishin &amp; Warren 2018, sp. nov." docType="treatment" docVersion="8" lastPageNumber="199" masterDocId="1726FFD4DB59FFF7FF8EA2103936D03F" masterDocTitle="A new species of Wahydra from Ecuador (Hesperiidae, Hesperiinae, Anthoptini)" masterLastPageNumber="200" masterPageNumber="196" pageNumber="197" updateTime="1698528438492" updateUser="ExternalLinkService">
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<mods:title id="6425AED2C4A1FD5BA14E3D6F1A8239A6">A new species of Wahydra from Ecuador (Hesperiidae, Hesperiinae, Anthoptini)</mods:title>
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<mods:namePart id="D139CC39018709835776EF5BC5B911A8">Carneiro, Eduardo</mods:namePart>
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<mods:namePart id="6403FD1525EE069EBA42503BB1E19E16">Dolibaina, Diego R.</mods:namePart>
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<mods:namePart id="4AA8A48BB20AFA3194560B948DC29E36">Grishin, Nick V.</mods:namePart>
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<mods:namePart id="CB4B961D75BF4CFFEC967FBBAE4CC242">Warren, Andrew D.</mods:namePart>
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<mods:title id="57B731124562DA3F6A6EED5514EB3BBF">Zootaxa</mods:title>
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<mods:date id="0C2A1B908C8F111C1FBFA7B59055B687">2018</mods:date>
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<mods:number id="BE0BD4D7936F7FCAA71C8FDCAAE7DED1">2018-03-08</mods:number>
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<mods:detail id="5269B6531131199415F0902B0539A18D" type="volume">
<mods:number id="356A952ABDF1BCDFA6CFE34C70941B6B">4392</mods:number>
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<mods:identifier id="10A1FB5E26045F7B71B82C218ABE8ADC" type="DOI">10.11646/zootaxa.4392.1.11</mods:identifier>
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<treatment id="EB1F87ACDB58FFF4FF19A0323DF0D32A" ID-DOI="http://doi.org/10.5281/zenodo.5973808" ID-GBIF-Taxon="159835467" ID-Zenodo-Dep="5973808" LSID="urn:lsid:plazi:treatment:EB1F87ACDB58FFF4FF19A0323DF0D32A" httpUri="http://treatment.plazi.org/id/EB1F87ACDB58FFF4FF19A0323DF0D32A" lastPageId="3" lastPageNumber="199" pageId="1" pageNumber="197">
<subSubSection id="2BAC6531DB58FFF6FF19A032383DD261" pageId="1" pageNumber="197" type="nomenclature">
<paragraph id="630936BADB58FFF6FF19A0323AA4D203" blockId="1.[151,914,546,606]" box="[151,914,546,572]" pageId="1" pageNumber="197">
<heading id="384181D6DB58FFF6FF19A0323AA4D203" bold="true" box="[151,914,546,572]" fontSize="11" level="1" pageId="1" pageNumber="197" reason="1">
<emphasis id="51C2EAA8DB58FFF6FF19A0323AA4D203" bold="true" box="[151,914,546,572]" pageId="1" pageNumber="197">
<taxonomicName id="A4B64D39DB58FFF6FF19A0323A11D203" authority="A. Warren, Carneiro &amp; Dolibaina" authorityName="Carneiro &amp; Dolibaina &amp; Grishin &amp; Warren" authorityYear="2018" box="[151,807,546,572]" class="Insecta" family="Hesperiidae" genus="Wahydra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="graslieae" status="sp. nov.">
<emphasis id="51C2EAA8DB58FFF6FF19A032384ED203" bold="true" box="[151,376,546,572]" italics="true" pageId="1" pageNumber="197">Wahydra graslieae</emphasis>
A. Warren, Carneiro &amp; Dolibaina
</taxonomicName>
,
<taxonomicNameLabel id="4AF157D3DB58FFF6FCBAA0323AA4D203" box="[820,914,546,572]" pageId="1" pageNumber="197" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="630936BADB58FFF6FF19A055383DD261" blockId="1.[151,914,546,606]" box="[151,267,581,606]" pageId="1" pageNumber="197">
(
<figureCitation id="FB8D2A3FDB58FFF6FF11A0553834D261" box="[159,258,581,606]" captionStart-0="FIGURES 1–2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="1.[151,264,1779,1802]" captionStartId-1="2.[151,250,1806,1828]" captionStartId-2="3.[151,250,1690,1712]" captionTargetBox-0="[151,1436,1240,1755]" captionTargetBox-1="[151,1436,744,1785]" captionTargetBox-2="[151,1436,844,1669]" captionTargetId-0="figure@1.[151,1436,1239,1755]" captionTargetId-1="figure@2.[151,1436,744,1785]" captionTargetId-2="figure@3.[151,1436,844,1669]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURES 1–2. Wahydra graslieae A. Warren, Carneiro, &amp; Dolibaina sp. nov., male holotype. 1—dorsal; 2—ventral. Scale bar = 1 cm." captionText-1="FIGURE 3. Left forewing of Wahydra graslieae A. Warren, Carneiro, &amp; Dolibaina sp. nov. (holotype), showing general aspect of brand, with positions of wing veins indicated." captionText-2="FIGURE 4. Male genitalia of Wahydra graslieae A. Warren, Carneiro &amp; Dolibaina sp. nov. (holotype). Scale bar = 1 mm." httpUri-0="https://zenodo.org/record/1194400/files/figure.png" httpUri-1="https://zenodo.org/record/1194402/files/figure.png" httpUri-2="https://zenodo.org/record/1194404/files/figure.png" pageId="1" pageNumber="197">Figs 1–4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="2BAC6531DB58FFF6FF19A09D3A4AD3D3" pageId="1" pageNumber="197" type="diagnosis">
<paragraph id="630936BADB58FFF6FF19A09D3A4AD3D3" blockId="1.[151,1437,653,1185]" pageId="1" pageNumber="197">
<emphasis id="51C2EAA8DB58FFF6FF19A09D3827D299" bold="true" box="[151,273,653,678]" pageId="1" pageNumber="197">Diagnosis.</emphasis>
DFW uniformly brown with a thin tripartite stigma between CuA1–2A. All other species of
<taxonomicName id="A4B64D39DB58FFF6FAB6A09D3CAAD299" authorityName="Steinhauser" authorityYear="1991" box="[1336,1436,653,678]" class="Insecta" family="Hesperiidae" genus="Wahydra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="51C2EAA8DB58FFF6FAB6A09D3CAAD299" box="[1336,1436,653,678]" italics="true" pageId="1" pageNumber="197">Wahydra</emphasis>
</taxonomicName>
have orange spots on DFW, though reduced in
<taxonomicName id="A4B64D39DB58FFF6FD2FA0A53AACD2F2" authority="(Evans, 1955)" authorityName="Evans" authorityYear="1955" baseAuthorityName="Evans" baseAuthorityYear="1955" box="[673,922,692,717]" class="Insecta" family="Hesperiidae" genus="Wahydra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="ekka">
<emphasis id="51C2EAA8DB58FFF6FD2FA0A53BC1D2F2" box="[673,759,692,717]" italics="true" pageId="1" pageNumber="197">W. ekka</emphasis>
(
<bibRefCitation id="07274B4BDB58FFF6FC8BA0A43AA4D2F2" box="[773,914,692,717]" pageId="1" pageNumber="197" refId="ref2242" refString="Evans, W. H. (1955) A Catalogue of the American Hesperiidae Indicating the Classification and Nomenclature Adopted in the British Museum (Natural History). British Museum (Natural History), London, vi + 499 pp., pls. 54 - 88." type="book chapter">Evans, 1955</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="A4B64D39DB58FFF6FC41A0A53C2AD2F3" authority="Steinhauser, 1991" authorityName="Steinhauser" authorityYear="1991" box="[975,1308,692,717]" class="Insecta" family="Hesperiidae" genus="Wahydra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="51C2EAA8DB58FFF6FC41A0A53D7AD2F2" box="[975,1100,692,717]" italics="true" pageId="1" pageNumber="197">W. obscura</emphasis>
<bibRefCitation id="07274B4BDB58FFF6FBDDA0A43C2AD2F3" box="[1107,1308,692,717]" pageId="1" pageNumber="197" refId="ref2543" refString="Steinhauser, S. R. (1991) Taxonomic notes and descriptions of new taxa in the Neotropical Hesperiidae. Part 2, Heteropterinae and Hesperiinae, Vinius group. Bulletin of the Allyn Museum, 132, 1 - 79." type="journal article">Steinhauser, 1991</bibRefCitation>
</taxonomicName>
. The VHW ground color is ferruginous red, with a metallic silver discal spot and postdiscal band. These features immediately identify
<taxonomicName id="A4B64D39DB58FFF6FF76A0ED38B0D32A" authority="Carneiro &amp; Dolibaina &amp; Grishin &amp; Warren, 2018" authorityName="Carneiro &amp; Dolibaina &amp; Grishin &amp; Warren" authorityYear="2018" box="[248,390,764,789]" class="Insecta" family="Hesperiidae" genus="Wahydra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="graslieae" status="sp. nov.">
<emphasis id="51C2EAA8DB58FFF6FF76A0ED38B0D32A" box="[248,390,764,789]" italics="true" pageId="1" pageNumber="197">W. graslieae</emphasis>
</taxonomicName>
,
<emphasis id="51C2EAA8DB58FFF6FE16A0EC38C3D32A" bold="true" box="[408,501,764,789]" pageId="1" pageNumber="197">
<taxonomicNameLabel id="4AF157D3DB58FFF6FE16A0EC38C3D32A" box="[408,501,764,789]" pageId="1" pageNumber="197" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
as unique both among the species of
<taxonomicName id="A4B64D39DB58FFF6FC3CA0EC3D20D32A" authorityName="Steinhauser" authorityYear="1991" box="[946,1046,764,789]" class="Insecta" family="Hesperiidae" genus="Wahydra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="51C2EAA8DB58FFF6FC3CA0EC3D20D32A" box="[946,1046,764,789]" italics="true" pageId="1" pageNumber="197">Wahydra</emphasis>
</taxonomicName>
and all other known members of
<taxonomicName id="A4B64D39DB58FFF6FF19A1303822D306" authorityName="A.Warren" authorityYear="2009" box="[151,276,800,825]" class="Insecta" family="Hesperiidae" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="tribe" tribe="Anthoptini">Anthoptini</taxonomicName>
. Two potentially sympatric species belonging to
<taxonomicName id="A4B64D39DB58FFF6FCB7A1303AADD306" authorityName="A.Warren" authorityYear="2008" box="[825,923,800,825]" class="Insecta" family="Hesperiidae" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="tribe" tribe="Moncini">Moncini</taxonomicName>
,
<taxonomicName id="A4B64D39DB58FFF6FC29A1313C39D307" authority="(Weeks, 1901)" authorityName="Weeks" authorityYear="1901" baseAuthorityName="Weeks" baseAuthorityYear="1901" box="[935,1295,800,825]" class="Insecta" family="Hesperiidae" genus="Tigasis" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="viridenex">
<emphasis id="51C2EAA8DB58FFF6FC29A1313D53D306" box="[935,1125,800,825]" italics="true" pageId="1" pageNumber="197">Tigasis viridenex</emphasis>
(Weeks, 1901)
</taxonomicName>
and
<taxonomicName id="A4B64D39DB58FFF6FAC9A1313859D363" authority="(Bell, 1942)" authorityName="Bell" authorityYear="1942" baseAuthorityName="Bell" baseAuthorityYear="1942" class="Insecta" family="Hesperiidae" genus="Lerema" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="viridis">
<emphasis id="51C2EAA8DB58FFF6FAC9A13139E9D362" italics="true" pageId="1" pageNumber="197">Lerema viridis</emphasis>
(Bell, 1942)
</taxonomicName>
have a somewhat similar VHW, however, they are easily distinguished from
<taxonomicName id="A4B64D39DB58FFF6FB51A1553C5DD362" authority="Carneiro &amp; Dolibaina &amp; Grishin &amp; Warren, 2018" authorityName="Carneiro &amp; Dolibaina &amp; Grishin &amp; Warren" authorityYear="2018" box="[1247,1387,836,861]" class="Insecta" family="Hesperiidae" genus="Wahydra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="graslieae" status="sp. nov.">
<emphasis id="51C2EAA8DB58FFF6FB51A1553C5DD362" box="[1247,1387,836,861]" italics="true" pageId="1" pageNumber="197">W. graslieae</emphasis>
</taxonomicName>
,
<emphasis id="51C2EAA8DB58FFF6FAF4A15439FED3BE" bold="true" pageId="1" pageNumber="197">
<taxonomicNameLabel id="4AF157D3DB58FFF6FAF4A15439FED3BE" pageId="1" pageNumber="197" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by their greenish ground color and the highly reduced metallic silver markings. In addition, the bifurcated median apophysis of the tegumen of
<taxonomicName id="A4B64D39DB58FFF6FDBDA19D3B88D39A" authority="Carneiro &amp; Dolibaina &amp; Grishin &amp; Warren, 2018" authorityName="Carneiro &amp; Dolibaina &amp; Grishin &amp; Warren" authorityYear="2018" box="[563,702,908,933]" class="Insecta" family="Hesperiidae" genus="Wahydra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="graslieae" status="sp. nov.">
<emphasis id="51C2EAA8DB58FFF6FDBDA19D3B88D39A" box="[563,702,908,933]" italics="true" pageId="1" pageNumber="197">W. graslieae</emphasis>
</taxonomicName>
<emphasis id="51C2EAA8DB58FFF6FD4BA19C3A28D39A" bold="true" box="[709,798,908,933]" pageId="1" pageNumber="197">
<taxonomicNameLabel id="4AF157D3DB58FFF6FD4BA19C3A28D39A" box="[709,798,908,933]" pageId="1" pageNumber="197" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is not found in any known species of
<taxonomicName id="A4B64D39DB58FFF6FB48A19C3C1CD39A" authorityName="Steinhauser" authorityYear="1991" box="[1222,1322,908,933]" class="Insecta" family="Hesperiidae" genus="Wahydra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="51C2EAA8DB58FFF6FB48A19C3C1CD39A" box="[1222,1322,908,933]" italics="true" pageId="1" pageNumber="197">Wahydra</emphasis>
</taxonomicName>
nor in the somewhat similar
<taxonomicName id="A4B64D39DB58FFF6FEE1A1A138CED3F6" baseAuthorityName="Weeks" baseAuthorityYear="1901" box="[367,504,944,969]" class="Insecta" family="Hesperiidae" genus="Tigasis" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="viridenex">
<emphasis id="51C2EAA8DB58FFF6FEE1A1A138CED3F6" box="[367,504,944,969]" italics="true" pageId="1" pageNumber="197">T. viridenex</emphasis>
</taxonomicName>
and
<taxonomicName id="A4B64D39DB58FFF6FDB5A1A13B93D3F6" baseAuthorityName="Bell" baseAuthorityYear="1942" box="[571,677,944,969]" class="Insecta" family="Hesperiidae" genus="Lerema" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="viridis">
<emphasis id="51C2EAA8DB58FFF6FDB5A1A13B93D3F6" box="[571,677,944,969]" italics="true" pageId="1" pageNumber="197">L. viridis</emphasis>
</taxonomicName>
.
<taxonomicName id="A4B64D39DB58FFF6FD37A1A03D42D3F7" authority="Steinhauser, 1991" authorityName="Steinhauser" authorityYear="1991" box="[697,1140,944,969]" class="Insecta" family="Hesperiidae" genus="Wahydra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="nieblensis">
<emphasis id="51C2EAA8DB58FFF6FD37A1A03AACD3F6" box="[697,922,944,969]" italics="true" pageId="1" pageNumber="197">Wahydra nieblensis</emphasis>
<bibRefCitation id="07274B4BDB58FFF6FC29A1A03D42D3F7" box="[935,1140,944,969]" pageId="1" pageNumber="197" refId="ref2543" refString="Steinhauser, S. R. (1991) Taxonomic notes and descriptions of new taxa in the Neotropical Hesperiidae. Part 2, Heteropterinae and Hesperiinae, Vinius group. Bulletin of the Allyn Museum, 132, 1 - 79." type="journal article">Steinhauser, 1991</bibRefCitation>
</taxonomicName>
also has a large median apophysis of the tegumen, but without developed bifurcated arms.
</paragraph>
</subSubSection>
<subSubSection id="2BAC6531DB58FFF4FF49A1E838C1D193" lastPageId="3" lastPageNumber="199" pageId="1" pageNumber="197" type="description">
<paragraph id="630936BADB58FFF6FF49A1E83D75D49E" blockId="1.[151,1437,653,1185]" pageId="1" pageNumber="197">
<emphasis id="51C2EAA8DB58FFF6FF49A1E8386FD42E" bold="true" box="[199,345,1016,1041]" pageId="1" pageNumber="197">Description.</emphasis>
Male.
<emphasis id="51C2EAA8DB58FFF6FE29A1E838D5D42E" box="[423,483,1016,1041]" italics="true" pageId="1" pageNumber="197">Head</emphasis>
: Eyes red. Vertex dark brown scattered with red ferruginous scales. Antennae longer than 2/3 length of forewing costa; antennal club short (1/4 shaft), ventral shaft yellowish in basal portion, dark brownish in apical portion; nudum of 14 segments, covering all the apiculus and extended to the club. Palpus quadrate (inner edge equal to transverse width), first and second segments ventrally yellowish, third segment of medium length (around half the length of second segment), cylindrical, dark brown.
</paragraph>
<caption id="37C96632DB58FFF6FF19A4E3383DD717" httpUri="https://zenodo.org/record/1194400/files/figure.png" pageId="1" pageNumber="197" startId="1.[151,264,1779,1802]" targetBox="[151,1436,1240,1755]" targetPageId="1">
<paragraph id="630936BADB58FFF6FF19A4E3383DD717" blockId="1.[151,1436,1779,1832]" pageId="1" pageNumber="197">
<emphasis id="51C2EAA8DB58FFF6FF19A4E3380AD736" bold="true" box="[151,316,1779,1802]" pageId="1" pageNumber="197">FIGURES 1–2.</emphasis>
<taxonomicName id="A4B64D39DB58FFF6FECAA4E43BD7D735" authority="A. Warren, Carneiro" authorityName="A. Warren, Carneiro" box="[324,737,1780,1802]" class="Insecta" family="Hesperiidae" genus="Wahydra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="graslieae">
<emphasis id="51C2EAA8DB58FFF6FECAA4E43B33D736" box="[324,517,1780,1801]" italics="true" pageId="1" pageNumber="197">Wahydra graslieae</emphasis>
A. Warren, Carneiro
</taxonomicName>
, &amp; Dolibaina
<emphasis id="51C2EAA8DB58FFF6FCFBA4E43AFED736" bold="true" box="[885,968,1780,1801]" pageId="1" pageNumber="197">sp. nov.</emphasis>
, male holotype. 1—dorsal; 2—ventral. Scale bar = 1 cm.
</paragraph>
</caption>
<paragraph id="630936BADB58FFF5FF49A55F3DD6D177" blockId="1.[151,1436,1871,2012]" lastBlockId="2.[151,1436,151,692]" lastPageId="2" lastPageNumber="198" pageId="1" pageNumber="197">
<emphasis id="51C2EAA8DB58FFF6FF49A55F382ED757" box="[199,280,1871,1896]" italics="true" pageId="1" pageNumber="197">Thorax</emphasis>
: dorsally and ventrally covered by long brown and red ferruginous scales; midtibiae spined; hindtibiae with two pairs of spurs. Forewing length
<quantity id="A44E9B5FDB58FFF6FDE4A5643BE4D7B3" box="[618,722,1908,1932]" metricMagnitude="-2" metricUnit="m" metricValue="1.33" pageId="1" pageNumber="197" unit="mm" value="13.3">13.3 mm</quantity>
. DFW homogeneous brown, with sparse red ferruginous scales on costal area. Stigma black, thin and tripartite, consisting of an elongate portion in CuA1–CuA2, following CuA, slightly angled towards CuA2, and two smaller spots in CuA2–2A, the anterior quadrate, surrounding CuA2, the posterior reduced, drop-shaped, below the anal fold; one subapical hyaline spot in R5–M1; fringe brown. DHW homogeneous brown; fringe brown. VFW ground color dark brown; costal and outer area ferruginous red; CuA2- 2A area paler; subapical hyaline spot as on DFW; fringe as on DFW. VHW ground color red ferruginous, with a metallic silver spot in the inferior half of the discal cell; a postdiscal metallic silver band from Sc+R1 to CuA2, zigzag patterned, and a circular postdiscal spot in CuA2-2A proximally displaced; fringe as on DHW.
</paragraph>
<paragraph id="630936BADB5BFFF5FF49A3473BDCD14F" blockId="2.[151,1436,151,692]" box="[199,746,343,368]" pageId="2" pageNumber="198">
<emphasis id="51C2EAA8DB5BFFF5FF49A3473807D14F" box="[199,305,343,368]" italics="true" pageId="2" pageNumber="198">Abdomen</emphasis>
: dorsally brown; ventrally ferruginous.
</paragraph>
<paragraph id="630936BADB5BFFF5FF49A36B38A1D28C" blockId="2.[151,1436,151,692]" pageId="2" pageNumber="198">
<emphasis id="51C2EAA8DB5BFFF5FF49A36B3841D1AB" box="[199,375,379,404]" italics="true" pageId="2" pageNumber="198">Male genitalia:</emphasis>
tegumen rectangular, about twice as wide as long, distally narrowed; median apophysis of tegumen bifurcated, longer than fenestra, larger than half of fenestra; lateral apophysis of tegumen symmetrically pointed. Fenestra rectangular longer than wide. Saccus shorter than tegumen, lobed. Uncus as long as tegumen (including its median apophysis), distally narrowed, shallowly bifid, with two short, divergent arms. Gnathos divided and narrow. Valva somewhat ovoid, narrowed distally; harpe triangular, broad, with a narrow, curved and upturned projection, distally serrated and partially covered by several small spines; sacculus, costa and ampulla narrow. Aedeagus longer than valva; coecum short and undifferentiated, distal opening of aedeagus dorsal, anteriorly contiguous with the opening of the ejaculatory bulb; dorsal triangular lateral projections on distal part of aedeagus; no cornutus.
</paragraph>
<caption id="37C96632DB5BFFF5FF19A51E3BB4D77C" httpUri="https://zenodo.org/record/1194402/files/figure.png" pageId="2" pageNumber="198" startId="2.[151,250,1806,1828]" targetBox="[151,1436,744,1785]" targetPageId="2">
<paragraph id="630936BADB5BFFF5FF19A51E3BB4D77C" blockId="2.[151,1436,1806,1859]" pageId="2" pageNumber="198">
<emphasis id="51C2EAA8DB5BFFF5FF19A51E3824D71C" bold="true" box="[151,274,1806,1828]" pageId="2" pageNumber="198">FIGURE 3.</emphasis>
Left forewing of
<taxonomicName id="A4B64D39DB5BFFF5FE4AA51E3A6ED71B" authority="A. Warren, Carneiro" authorityName="A. Warren, Carneiro" box="[452,856,1806,1828]" class="Insecta" family="Hesperiidae" genus="Wahydra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="198" phylum="Arthropoda" rank="species" species="graslieae">
<emphasis id="51C2EAA8DB5BFFF5FE4AA51E3BB4D71C" box="[452,642,1806,1827]" italics="true" pageId="2" pageNumber="198">Wahydra graslieae</emphasis>
A. Warren, Carneiro
</taxonomicName>
, &amp; Dolibaina
<emphasis id="51C2EAA8DB5BFFF5FC6BA51F3D03D71B" bold="true" box="[997,1077,1807,1828]" pageId="2" pageNumber="198">sp. nov.</emphasis>
(holotype), showing general aspect of brand, with positions of wing veins indicated.
</paragraph>
</caption>
<paragraph id="630936BADB5BFFF5FF49A57A38A1D7BC" blockId="2.[151,1422,1898,2031]" box="[199,407,1898,1923]" pageId="2" pageNumber="198">Female. unknown.</paragraph>
<paragraph id="630936BADB5BFFF5FF49A59F385DD798" blockId="2.[151,1422,1898,2031]" box="[199,363,1934,1959]" pageId="2" pageNumber="198">DNA barcode.</paragraph>
<paragraph id="630936BADB5BFFF4FF49A5A338C1D193" blockId="2.[151,1422,1898,2031]" lastBlockId="3.[151,1437,152,789]" lastPageId="3" lastPageNumber="199" pageId="2" pageNumber="198">ACTTTATATTTTATTTTTGGTATTTGAGCAGGAATATTAGGAACTTCCCTAAGTTTATTAATTCGTAC AGAATTAGGTAATCCAGGATCTTTAATTGGAGATGATCAAATTTATAATACTATTGTTACAGCTCATGCT TTTATTATAATTTTTTTTATAGTTATACCTATTATAATTGGAGGATTCGGAAATTGATTAATTCCTTTAATAC TAGGTGCTCCTGATATAGCTTTCCCTCGAATAAATAATATAAGATTTTGAATATTACCCCCTTCTTTAATA TTACTAGTCTCTAGAAGAATTGTAGAAAATGGTGCAGGAACTGGTTGAACAGTTTACCCCCCCCTTTC ATCTAATATTGCTCATCAAGGATCCTCTGTTGATTTAGCAATCTTTTCTCTTCATTTAGCTGGAATTTCCT CTATTTTAGGAGCTATTAATTTTATTACTACAATTATTAATATACGAATTAAAAACATATCATTTGATCAAA TACCTTTATTTGTATGATCAGTAGGAATTACAGCTTTACTTTTACTTTTATCATTACCAGTACTAGCTGGA GCCATCACTATACTTTTAACTGATCGAAATTTAAATACATCTTTTTTTGATCCTGCAGGAGGAGGAGATC CAATCTTATATCAACATTTA</paragraph>
</subSubSection>
<subSubSection id="2BAC6531DB5AFFF4FF49A3A73BA6D298" pageId="3" pageNumber="199" type="materials_examined">
<paragraph id="630936BADB5AFFF4FF49A3A73D98D25F" blockId="3.[151,1437,152,789]" pageId="3" pageNumber="199">
<emphasis id="51C2EAA8DB5AFFF4FF49A3A7384ED1EF" bold="true" box="[199,376,439,464]" pageId="3" pageNumber="199">
<typeStatus id="BC0D8818DB5AFFF4FF49A3A73834D1EF" box="[199,258,439,464]" pageId="3" pageNumber="199">Type</typeStatus>
material.
</emphasis>
The male
<typeStatus id="BC0D8818DB5AFFF4FE77A3A73B6DD1EF" box="[505,603,439,464]" pageId="3" pageNumber="199" type="holotype">holotype</typeStatus>
of
<taxonomicName id="A4B64D39DB5AFFF4FD08A3A93A22D1EF" authority="Carneiro &amp; Dolibaina &amp; Grishin &amp; Warren, 2018" authorityName="Carneiro &amp; Dolibaina &amp; Grishin &amp; Warren" authorityYear="2018" box="[646,788,439,464]" class="Insecta" family="Hesperiidae" genus="Wahydra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="199" phylum="Arthropoda" rank="species" species="graslieae" status="sp. nov.">
<emphasis id="51C2EAA8DB5AFFF4FD08A3A93A22D1EF" box="[646,788,439,464]" italics="true" pageId="3" pageNumber="199">W. graslieae</emphasis>
</taxonomicName>
<emphasis id="51C2EAA8DB5AFFF4FC91A3A73A4AD1EF" bold="true" box="[799,892,439,464]" pageId="3" pageNumber="199">
<taxonomicNameLabel id="4AF157D3DB5AFFF4FC91A3A73A4AD1EF" box="[799,892,439,464]" pageId="3" pageNumber="199" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has the following labels: white, handprinted: /
<collectingCountry id="1BA1762ADB5AFFF4FF19A3CC3815D1CB" box="[151,291,476,500]" name="Ecuador" pageId="3" pageNumber="199">ECUADOR</collectingCountry>
:
<collectingRegion id="A172F858DB5AFFF4FEA0A3CC3849D1CB" box="[302,383,476,500]" country="Ecuador" name="Napo" pageId="3" pageNumber="199">
<collectionCode id="05A7AE7FDB5AFFF4FEA0A3CC3849D1CB" box="[302,383,476,500]" pageId="3" pageNumber="199">NAPO</collectionCode>
</collectingRegion>
: /
<quantity id="A44E9B5FDB5AFFF4FE15A3CC38D5D1CA" box="[411,483,476,501]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="199" unit="km" value="14.0">14 km</quantity>
E of Yanayacu / Biological Station / along Cosanga River /
<quantity id="A44E9B5FDB5AFFF4FB1BA3CC3DD0D1CB" box="[1173,1254,476,500]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="3" pageNumber="199" unit="m" value="2400.0">2400m</quantity>
,
<date id="1708107ADB5AFFF4FB7CA3CC3CBBD1CB" box="[1266,1421,476,500]" pageId="3" pageNumber="199" value="2004-06-17">17-June-2004</date>
/ Harold Greeney [
<emphasis id="51C2EAA8DB5AFFF4FED3A3EF38B1D227" box="[349,391,511,536]" italics="true" pageId="3" pageNumber="199">leg.</emphasis>
] / H09-2030, 11:30 hrs. /; white, printed and handprinted: / EC022 /
<taxonomicName id="A4B64D39DB5AFFF4FB13A3EF3DC9D227" authorityName="Steinhauser" authorityYear="1991" box="[1181,1279,511,536]" class="Insecta" family="Hesperiidae" genus="Wahydra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="199" phylum="Arthropoda" rank="genus">
<emphasis id="51C2EAA8DB5AFFF4FB13A3EF3DC9D227" box="[1181,1279,511,536]" italics="true" pageId="3" pageNumber="199">Wahydra</emphasis>
</taxonomicName>
/ E. Carneiro det. 2015 /; white, printed: /
<collectionCode id="05A7AE7FDB5AFFF4FE64A0343B11D203" box="[490,551,548,572]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="3" pageNumber="199">DNA</collectionCode>
sample ID: / NVG-5287 / c/o Nick Grishin /; red, printed: /
<typeStatus id="BC0D8818DB5AFFF4FB78A0343CBCD203" box="[1270,1418,548,572]" pageId="3" pageNumber="199" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="A4B64D39DB5AFFF4FF19A057385CD25F" authorityName="Carneiro &amp; Dolibaina &amp; Grishin &amp; Warren" authorityYear="2018" box="[151,362,583,608]" class="Insecta" family="Hesperiidae" genus="Wahydra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="199" phylum="Arthropoda" rank="species" species="graslieae">
<emphasis id="51C2EAA8DB5AFFF4FF19A057385CD25F" box="[151,362,583,608]" italics="true" pageId="3" pageNumber="199">Wahydra graslieae</emphasis>
</taxonomicName>
/ A. Warren, Carneiro &amp; Dolibaina /. The
<typeStatus id="BC0D8818DB5AFFF4FCCBA0573A91D25F" box="[837,935,583,608]" pageId="3" pageNumber="199" type="holotype">holotype</typeStatus>
is deposited at
<collectionCode id="05A7AE7FDB5AFFF4FBD8A0583D9CD25F" box="[1110,1194,584,608]" pageId="3" pageNumber="199">MGCL</collectionCode>
.
</paragraph>
<paragraph id="630936BADB5AFFF4FF49A07C3BA6D298" blockId="3.[151,1437,152,789]" pageId="3" pageNumber="199">
<emphasis id="51C2EAA8DB5AFFF4FF49A07C385ED2BA" bold="true" box="[199,360,620,645]" pageId="3" pageNumber="199">Type locality.</emphasis>
The holotype of
<taxonomicName id="A4B64D39DB5AFFF4FDBCA07D3B8BD2BA" authority="Carneiro &amp; Dolibaina &amp; Grishin &amp; Warren, 2018" authorityName="Carneiro &amp; Dolibaina &amp; Grishin &amp; Warren" authorityYear="2018" box="[562,701,620,645]" class="Insecta" family="Hesperiidae" genus="Wahydra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="199" phylum="Arthropoda" rank="species" species="graslieae" status="sp. nov.">
<emphasis id="51C2EAA8DB5AFFF4FDBCA07D3B8BD2BA" box="[562,701,620,645]" italics="true" pageId="3" pageNumber="199">W. graslieae</emphasis>
</taxonomicName>
<emphasis id="51C2EAA8DB5AFFF4FD47A07C3A13D2BA" bold="true" box="[713,805,620,645]" pageId="3" pageNumber="199">
<taxonomicNameLabel id="4AF157D3DB5AFFF4FD47A07C3A13D2BA" box="[713,805,620,645]" pageId="3" pageNumber="199" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was collected along the edge of secondary flood plain forest dominated by
<taxonomicName id="A4B64D39DB5AFFF4FEF0A09F38CAD297" authority="Mill." authorityName="Mill." box="[382,508,655,680]" class="Magnoliopsida" family="Betulaceae" genus="Alnus" kingdom="Plantae" order="Fagales" pageId="3" pageNumber="199" phylum="Tracheophyta" rank="genus">
<emphasis id="51C2EAA8DB5AFFF4FEF0A09F3888D297" box="[382,446,655,680]" italics="true" pageId="3" pageNumber="199">Alnus</emphasis>
Mill.
</taxonomicName>
and
<taxonomicName id="A4B64D39DB5AFFF4FDBCA0813B46D297" authorityName="Linnaeus" authorityYear="1753" box="[562,624,657,680]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="3" pageNumber="199" phylum="Tracheophyta" rank="genus">
<emphasis id="51C2EAA8DB5AFFF4FDBCA0813B46D297" box="[562,624,657,680]" italics="true" pageId="3" pageNumber="199">Piper</emphasis>
</taxonomicName>
L.
</paragraph>
</subSubSection>
<subSubSection id="2BAC6531DB5AFFF4FF49A0A43DF0D32A" pageId="3" pageNumber="199" type="etymology">
<paragraph id="630936BADB5AFFF4FF49A0A43DF0D32A" blockId="3.[151,1437,152,789]" pageId="3" pageNumber="199">
<emphasis id="51C2EAA8DB5AFFF4FF49A0A43878D2F2" bold="true" box="[199,334,692,717]" pageId="3" pageNumber="199">Etymology.</emphasis>
We are delighted to name this species in honor of Emily Graslie, Chief Curiosity Correspondent at the Field Museum (Chicago,
<collectingRegion id="A172F858DB5AFFF4FE65A0C73B0BD2CF" box="[491,573,727,752]" country="United States of America" name="Illinois" pageId="3" pageNumber="199">Illinois</collectingRegion>
,
<collectingCountry id="1BA1762ADB5AFFF4FDC2A0C83BBED2CF" box="[588,648,728,752]" name="United States of America" pageId="3" pageNumber="199">USA</collectingCountry>
), in recognition of her efforts to promote natural history collections through her YouTube channel The Brain Scoop (https://www.youtube.com/user/thebrainscoop).
</paragraph>
</subSubSection>
</treatment>
</document>
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