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<document id="C88605D73374440EA3548B2C9E5CBD49" ID-DOI="10.11646/zootaxa.2160.1.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="5319514" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629626811990" checkinUser="felipe" docAuthor="Kjaerandsen, Jostein, Martinsson, Svante, Hedmark, Kjell &amp; Evenhuis, Neal L." docDate="2009" docId="D75E2A4D810DFFC2FF01FE1FFA8F6435" docLanguage="en" docName="zt02160p050.pdf" docOrigin="Zootaxa 2160 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2160.1.2" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Urytalpa galdes Kjaerandsen, Martinsson, Hedmark &amp; Evenhuis, 2009, sp. n." docType="treatment" docVersion="3" lastPageNumber="38" masterDocId="2B6752358104FFCBFF96FFFDFF806358" masterDocTitle="On the genus Urytalpa Edwards (Diptera: Keroplatidae) in the Nordic and Nearctic regions, with fixation of a new type species and a key to world males" masterLastPageNumber="50" masterPageNumber="29" pageNumber="38" updateTime="1699086153839" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="F898D63AAE2072568436BFC210F6D88F">On the genus Urytalpa Edwards (Diptera: Keroplatidae) in the Nordic and Nearctic regions, with fixation of a new type species and a key to world males</mods:title>
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<mods:namePart id="7363A6E1D027D50457E93AA3011D9BD6">Kjaerandsen, Jostein</mods:namePart>
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<mods:namePart id="5C2001912681122B2C935EFCC4E05AED">Hedmark, Kjell</mods:namePart>
<mods:affiliation id="E1DD7547F3004719FA8CA48BF6D1A4F3">Kyrkvägen 17, S- 960 30, Vuollerim, Sweden. E-mail: kjell. hedmark @ same. net</mods:affiliation>
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<mods:namePart id="C3E31181295C4AD01D97997FC7B01DE0">Evenhuis, Neal L.</mods:namePart>
<mods:affiliation id="9CE29218CC78DFC968FD72E8F8076C9F">Department of Natural Sciences, Bishop Museum, 1525 Bernice Street, Honolulu, Hawaii 96817, USA.</mods:affiliation>
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<taxonomicName id="98F7E0D8810DFFC2FF01FE1FFED462A4" ID-CoL="9HKQS" authorityName="Hedmark &amp; Kjaerandsen" authorityYear="2009" box="[151,340,482,508]" class="Insecta" family="Keroplatidae" genus="Urytalpa" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="galdes">
<emphasis id="6D834749810DFFC2FF01FE1FFED462A4" bold="true" box="[151,340,482,508]" italics="true" pageId="9" pageNumber="38">Urytalpa galdes</emphasis>
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&amp; Kjaerandsen
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.
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.
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(
<figureCitation id="C7CC87DE810DFFC2FF09FDF4FF7D617B" box="[159,253,521,547]" captionStart="FIGURE 2" captionStartId="5.[151,255,1931,1955]" captionTargetBox="[178,1414,190,1908]" captionTargetId="figure-66@5.[164,1429,173,1923]" captionTargetPageId="5" captionText="FIGURE 2. Habitus photos of males of Nordic species of the genus Urytalpa Edwards. — A. Urytalpa dorsalis (Staeger, 1840). — B. Urytalpa trivittata (Lundström, 1914). — C. Urytalpa atriceps (Edwards, 1913). — D. Urytalpa macrocera (Edwards, 1913). — E. Urytalpa galdes Hedmark &amp; Kjaerandsen sp. n." figureDoi="http://doi.org/10.5281/zenodo.5319518" httpUri="https://zenodo.org/record/5319518/files/figure.png" pageId="9" pageNumber="38">Figs 2E</figureCitation>
,
<figureCitation id="C7CC87DE810DFFC2FE9CFDF4FEAD617B" box="[266,301,521,547]" captionStart="FIGURE 4" captionStartId="8.[151,255,1928,1952]" captionTargetBox="[279,1315,198,1879]" captionTargetId="figure-66@8.[252,1334,181,1899]" captionTargetPageId="8" captionText="FIGURE 4. Wing photos of males of Nordic species of the genus Urytalpa Edwards. — A. Urytalpa dorsalis (Staeger, 1840). — B. Urytalpa trivittata (Lundström, 1914). — C. Urytalpa atriceps (Edwards, 1913). — D. Urytalpa macrocera (Edwards, 1913). — E. Urytalpa galdes Hedmark &amp; Kjaerandsen sp. n." figureDoi="http://doi.org/10.5281/zenodo.5319524" httpUri="https://zenodo.org/record/5319524/files/figure.png" pageId="9" pageNumber="38">4E</figureCitation>
,
<figureCitation id="C7CC87DE810DFFC2FEAFFDF4FEFF617B" box="[313,383,521,547]" captionStart="FIGURE 7" captionStartId="12.[151,255,1865,1889]" captionTargetBox="[169,1435,214,1826]" captionTargetId="figure-82@12.[151,1436,194,1841]" captionTargetPageId="12" captionText="FIGURE 7. Male terminalia of Urytalpa galdes Hedmark &amp; Kjaerandsen sp. n. — A. Segment IX, hypandrial lobe and internal aedeagal apparatus in ventral view (pre-terminal segments not illustrated). — B. Tergite IX, proctiger and cerci in dorsal view. — C. Segment IX and beyond in lateral view. — D. Gonostylus in lateral view, enlarged. — E. Aedeagal apparatus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5319534" httpUri="https://zenodo.org/record/5319534/files/figure.png" pageId="9" pageNumber="38">7AE</figureCitation>
)
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<emphasis id="6D834749810DFFC2FF01FDABFE8F6128" bold="true" box="[151,271,598,624]" pageId="9" pageNumber="38">Diagnosis</emphasis>
. This species is closest to
<taxonomicName id="98F7E0D8810DFFC2FDD2FDAAFC546128" authority="Bechev &amp; Koc, 2008" authorityName="Bechev &amp; Koc" authorityYear="2008" box="[580,980,598,624]" class="Insecta" family="Keroplatidae" genus="Urytalpa" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="chandleri">
<emphasis id="6D834749810DFFC2FDD2FDAAFD576128" box="[580,727,599,624]" italics="true" pageId="9" pageNumber="38">U. chandleri</emphasis>
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Bechev &amp; Koç,
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in the shape of the male terminalia but is easily distinguished from all other species by the long club-shaped and gently incurved processes arising caudally from the gonocoxites (
<figureCitation id="C7CC87DE810DFFC2FD81FD59FDF661E6" box="[535,630,676,702]" captionStart="FIGURE 7" captionStartId="12.[151,255,1865,1889]" captionTargetBox="[169,1435,214,1826]" captionTargetId="figure-82@12.[151,1436,194,1841]" captionTargetPageId="12" captionText="FIGURE 7. Male terminalia of Urytalpa galdes Hedmark &amp; Kjaerandsen sp. n. — A. Segment IX, hypandrial lobe and internal aedeagal apparatus in ventral view (pre-terminal segments not illustrated). — B. Tergite IX, proctiger and cerci in dorsal view. — C. Segment IX and beyond in lateral view. — D. Gonostylus in lateral view, enlarged. — E. Aedeagal apparatus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5319534" httpUri="https://zenodo.org/record/5319534/files/figure.png" pageId="9" pageNumber="38">Fig. 7A</figureCitation>
) and by the large gonostylus with a single long and digitate apical process (
<figureCitation id="C7CC87DE810DFFC2FE90FD37FEEB61BC" box="[262,363,714,740]" captionStart="FIGURE 7" captionStartId="12.[151,255,1865,1889]" captionTargetBox="[169,1435,214,1826]" captionTargetId="figure-82@12.[151,1436,194,1841]" captionTargetPageId="12" captionText="FIGURE 7. Male terminalia of Urytalpa galdes Hedmark &amp; Kjaerandsen sp. n. — A. Segment IX, hypandrial lobe and internal aedeagal apparatus in ventral view (pre-terminal segments not illustrated). — B. Tergite IX, proctiger and cerci in dorsal view. — C. Segment IX and beyond in lateral view. — D. Gonostylus in lateral view, enlarged. — E. Aedeagal apparatus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5319534" httpUri="https://zenodo.org/record/5319534/files/figure.png" pageId="9" pageNumber="38">Fig. 7D</figureCitation>
). It appears distinctly smaller than the other Nordic species of
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<emphasis id="6D834749810DFFC2FB15FD37FB7261BB" box="[1155,1266,714,739]" italics="true" pageId="9" pageNumber="38">Urytalpa</emphasis>
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and the male abdomen is dorsoventrally rather than laterally compressed (
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).
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: LU
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(
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&amp;
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) [
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, JKJ-SPM- 028550, on slide].
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<emphasis id="6D834749810DFFC2FECFFC76FE0260FD" bold="true" box="[345,386,907,933]" pageId="9" pageNumber="38">LU</emphasis>
,
<collectorName id="F202FE8D810DFFC2FE05FC71FD9560FE" box="[403,533,908,934]" pageId="9" pageNumber="38">Jokkmokk</collectorName>
,
<collectorName id="F202FE8D810DFFC2FDB4FC71FD4160FE" box="[546,705,908,934]" pageId="9" pageNumber="38">Kaltisbäcken</collectorName>
<location id="5A28CD80810DFFC2FD5CFC71FC4F60FE" LSID="urn:lsid:plazi:treatment:D75E2A4D810DFFC2FF01FE1FFA8F6435:5A28CD80810DFFC2FD5CFC71FC4F60FE" box="[714,975,908,934]" country="Sweden" latitude="66.69056" longLatPrecision="16" longitude="20.376945" municipality="Kaltisbacken" name="1 km NNE Messaure" pageId="9" pageNumber="38">
<quantity id="980F36BE810DFFC2FD5CFC71FC8860FE" box="[714,776,908,934]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="38" unit="km" value="1.0">1 km</quantity>
NNE Messaure
</location>
, same data as for holotype
</materialsCitation>
<materialsCitation id="EF9F9106810DFFC2FADFFC71FDD46094" collectionCode="MZLU" country="Sweden" pageId="9" pageNumber="38" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="49F150D2810DFFC2FADFFC71FA1C60FE" box="[1353,1436,908,934]" pageId="9" pageNumber="38" type="male">1 male</specimenCount>
[
<collectionCode id="39E6039E810DFFC2FF0BFC4FFF776094" box="[157,247,946,972]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34994" lsid="urn:lsid:biocol.org:col:34994" name="Lund University" pageId="9" pageNumber="38">MZLU</collectionCode>
, JKJ-SPM-016493, on slide]
</materialsCitation>
;
<materialsCitation id="EF9F9106810DFFC2FDF4FC4FFB116094" box="[610,1169,946,972]" collectionCode="MZLU" country="Sweden" pageId="9" pageNumber="38" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="49F150D2810DFFC2FDF4FC4FFD326094" box="[610,690,946,972]" pageId="9" pageNumber="38" type="male">1 male</specimenCount>
[
<collectionCode id="39E6039E810DFFC2FD57FC4FFC9B6094" box="[705,795,946,972]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34994" lsid="urn:lsid:biocol.org:col:34994" name="Lund University" pageId="9" pageNumber="38">MZLU</collectionCode>
, JKJ-SPM-033204, in alcohol]
</materialsCitation>
;
<materialsCitation id="EF9F9106810DFFC2FB08FC4FFBF460AB" collectingDate="1998-08-15" collectionCode="MZLU" collectorName="Leg. K." country="Sweden" location="Hedmark" pageId="9" pageNumber="38" specimenCount="1" specimenCount-male="1" stateProvince="Hedmark" typeStatus="paratype">
<specimenCount id="49F150D2810DFFC2FB08FC4FFB726094" box="[1182,1266,946,972]" pageId="9" pageNumber="38" type="male">1 male</specimenCount>
,
<date id="2B49BD9B810DFFC2FB69FC4FFA1B6094" box="[1279,1435,946,972]" pageId="9" pageNumber="38" value="1998-08-15">
<collectingDate id="3B0D4473810DFFC2FB69FC4FFA1B6094" box="[1279,1435,946,972]" pageId="9" pageNumber="38" value="1998-08-15">15 Aug 1998</collectingDate>
</date>
(
<collectorName id="F202FE8D810DFFC2FF09FC24FF7960AB" box="[159,249,985,1011]" pageId="9" pageNumber="38">Leg. K.</collectorName>
<collectingRegion id="9D3355B9810DFFC2FE97FC24FEF060AB" box="[257,368,985,1011]" country="Norway" name="Hedmark" pageId="9" pageNumber="38">Hedmark</collectingRegion>
) [
<collectionCode id="39E6039E810DFFC2FE10FC24FE5F60AB" box="[390,479,985,1011]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34994" lsid="urn:lsid:biocol.org:col:34994" name="Lund University" pageId="9" pageNumber="38">MZLU</collectionCode>
, JKJ-SPM-036830, pinned with terminalia in glycerine]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="17EDC8D0810DFFC2FF53FC02FB886783" pageId="9" pageNumber="38" type="etymology">
<paragraph id="5F489B5B810DFFC2FF53FC02FB886783" blockId="9.[151,1436,598,1901]" pageId="9" pageNumber="38">
<emphasis id="6D834749810DFFC2FF53FC02FECD6741" bold="true" box="[197,333,1023,1049]" pageId="9" pageNumber="38">Etymology</emphasis>
. Based on the Sámi word &quot;gáldes&quot;, the first element of &quot;Gáldesjåhkå&quot; which is the original Sámi name for the
<typeStatus id="804C25F9810DFFC2FE16FBDBFE346718" box="[384,436,1062,1088]" pageId="9" pageNumber="38">type</typeStatus>
locality, a small stream close to Messaure. The meaning and origin of &quot;gáldes&quot; is unknown to the science of language. The word is apparently very old. The second element, &quot;jåhkå&quot;, means stream. We consider &quot;gáldes&quot; to be an unlatinized Sámi word and thus to be retained with original gender and termination unchanged (ICZN Article 31b). The name &quot;Gáldesjåhkå&quot; is usually transcribed to &quot;Kaltisjokk&quot; or &quot;Kaltisbäcken&quot; on maps and references to this locality in biological papers.
</paragraph>
</subSubSection>
<subSubSection id="17EDC8D0810DFFC2FF53FB1AFE14641E" pageId="9" pageNumber="38" type="description">
<paragraph id="5F489B5B810DFFC2FF53FB1AFDC76447" blockId="9.[151,1436,598,1901]" pageId="9" pageNumber="38">
<emphasis id="6D834749810DFFC2FF53FB1AFED66659" bold="true" box="[197,342,1255,1281]" pageId="9" pageNumber="38">Description</emphasis>
.
<emphasis id="6D834749810DFFC2FEF2FB1AFE236659" bold="true" box="[356,419,1255,1281]" pageId="9" pageNumber="38">Male</emphasis>
(n=2). Body length 4.55.5 (n=3) mm. Wing length
<quantity id="980F36BE810DFFC2FB99FB15FB2D665A" box="[1039,1197,1256,1282]" metricMagnitude="-3" metricUnit="m" metricValue="3.91" metricValueMax="3.92" metricValueMin="3.9" pageId="9" pageNumber="38" unit="mm" value="3.91" valueMax="3.92" valueMin="3.9">3.93.92 mm</quantity>
, or 3.53.55 as long as profemur. Antenna length
<quantity id="980F36BE810DFFC2FE6EFAF3FD2B6670" box="[504,683,1294,1320]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.26" metricValueMin="2.24" pageId="9" pageNumber="38" unit="mm" value="2.25" valueMax="2.26" valueMin="2.24">2.242.26 mm</quantity>
.
<emphasis id="6D834749810DFFC2FD21FAF3FCC16670" bold="true" box="[695,833,1294,1320]" pageId="9" pageNumber="38">Coloration</emphasis>
(specimen in alcohol,
<figureCitation id="C7CC87DE810DFFC2FBC1FAF3FB346670" box="[1111,1204,1294,1320]" captionStart="FIGURE 2" captionStartId="5.[151,255,1931,1955]" captionTargetBox="[178,1414,190,1908]" captionTargetId="figure-66@5.[164,1429,173,1923]" captionTargetPageId="5" captionText="FIGURE 2. Habitus photos of males of Nordic species of the genus Urytalpa Edwards. — A. Urytalpa dorsalis (Staeger, 1840). — B. Urytalpa trivittata (Lundström, 1914). — C. Urytalpa atriceps (Edwards, 1913). — D. Urytalpa macrocera (Edwards, 1913). — E. Urytalpa galdes Hedmark &amp; Kjaerandsen sp. n." figureDoi="http://doi.org/10.5281/zenodo.5319518" httpUri="https://zenodo.org/record/5319518/files/figure.png" pageId="9" pageNumber="38">Fig. 2E</figureCitation>
). Head brown with pale yellowish mouth parts. Antenna dark brown with basal part of first flagellomere pale. Thorax mainly brown; prescutum pale brown on anterior half; preepisternum pale on dorsal half; laterotergite pale on anterior part; mediotergite gradually darker towards posterior end; thoracic stripes dark blackish brown narrowly surrounded by pale yellow ground; humeral area pale, narrow pale stripe extending laterally along entire length of mesoscutum. Wings clear, yellow tinged, veins brown. Halter whitish. Legs whitish. Abdomen mainly dark brown; tergites IIV with apical half forming yellow bands; sternites yellow with thin dark bands basally.
<emphasis id="6D834749810DFFC2FF6BF9E0FE0C656F" bold="true" box="[253,396,1565,1591]" pageId="9" pageNumber="38">Terminalia</emphasis>
(
<figureCitation id="C7CC87DE810DFFC2FE09F9E0FE6A656F" box="[415,490,1565,1591]" captionStart="FIGURE 7" captionStartId="12.[151,255,1865,1889]" captionTargetBox="[169,1435,214,1826]" captionTargetId="figure-82@12.[151,1436,194,1841]" captionTargetPageId="12" captionText="FIGURE 7. Male terminalia of Urytalpa galdes Hedmark &amp; Kjaerandsen sp. n. — A. Segment IX, hypandrial lobe and internal aedeagal apparatus in ventral view (pre-terminal segments not illustrated). — B. Tergite IX, proctiger and cerci in dorsal view. — C. Segment IX and beyond in lateral view. — D. Gonostylus in lateral view, enlarged. — E. Aedeagal apparatus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5319534" httpUri="https://zenodo.org/record/5319534/files/figure.png" pageId="9" pageNumber="38">Fig. 7</figureCitation>
). Gonocoxites divided ventrally, with long club-shaped and gently incurved process apicolaterally and short setose lobe medially. Hypandrial lobe not discernible, apparently absent or very hyaline. Gonostylus large, with a rounded lateral knob and a long digitate apical process. Aedeagal apparatus of medium length, medial compressed plate not high but extending distinctly beyond apex of the stout lateral apodeme (
<figureCitation id="C7CC87DE810DFFC2FE35F945FE7F658A" box="[419,511,1720,1746]" captionStart="FIGURE 7" captionStartId="12.[151,255,1865,1889]" captionTargetBox="[169,1435,214,1826]" captionTargetId="figure-82@12.[151,1436,194,1841]" captionTargetPageId="12" captionText="FIGURE 7. Male terminalia of Urytalpa galdes Hedmark &amp; Kjaerandsen sp. n. — A. Segment IX, hypandrial lobe and internal aedeagal apparatus in ventral view (pre-terminal segments not illustrated). — B. Tergite IX, proctiger and cerci in dorsal view. — C. Segment IX and beyond in lateral view. — D. Gonostylus in lateral view, enlarged. — E. Aedeagal apparatus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5319534" httpUri="https://zenodo.org/record/5319534/files/figure.png" pageId="9" pageNumber="38">Fig. 7E</figureCitation>
). Tergite IX without outgrowths, apical corner rounded, without patch of short internal setae; with deep posterior and shallow anterior U-shaped incision. Proctiger relatively long; cercus parallelogram shaped in lateral view.
</paragraph>
<paragraph id="5F489B5B810DFFC2FF53F8D6FE14641E" blockId="9.[151,1436,598,1901]" box="[197,404,1835,1862]" pageId="9" pageNumber="38">
<emphasis id="6D834749810DFFC2FF53F8D6FEA0641D" bold="true" box="[197,288,1835,1861]" pageId="9" pageNumber="38">Female</emphasis>
unknown
</paragraph>
</subSubSection>
<subSubSection id="17EDC8D0810DFFC2FF53F8AFFA8F6435" box="[197,1295,1874,1901]" pageId="9" pageNumber="38" type="distribution">
<paragraph id="5F489B5B810DFFC2FF53F8AFFA8F6435" blockId="9.[151,1436,598,1901]" box="[197,1295,1874,1901]" pageId="9" pageNumber="38">
<emphasis id="6D834749810DFFC2FF53F8AFFEDE6434" bold="true" box="[197,350,1874,1900]" pageId="9" pageNumber="38">Distribution</emphasis>
. Northern Europe; so far known only from the
<typeStatus id="804C25F9810DFFC2FC1FF8AEFC3B6435" box="[905,955,1875,1901]" pageId="9" pageNumber="38">type</typeStatus>
locality in northern
<collectingCountry id="27E0DBCB810DFFC2FB3CF8AEFA8B6435" box="[1194,1291,1875,1901]" name="Sweden" pageId="9" pageNumber="38">Sweden</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>