treatments-xml/data/D1/3B/87/D13B87B7FFC63645A7FB188CFED3AB0A.xml

<document id="603DF989CFEAEEF2EFBF8C1D27645D17" ID-CLB-Dataset="7185" ID-DOI="10.11646/zootaxa.4963.1.6" ID-GBIF-Dataset="e3c46cee-7d58-44ce-91f1-031dc312cd95" ID-ISSN="1175-5326" ID-Zenodo-Dep="4719970" ID-ZooBank="5AE81AAE-AF66-4973-B4E6-644134B7B67C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1618563571957" checkinUser="plazi" docAuthor="Cheng, Zhi-Fei, Pan, Zhao-Hui, Li, Li-Zhen &amp; Peng, Zhong" docDate="2021" docId="D13B87B7FFC63645A7FB188CFED3AB0A" docLanguage="en" docName="zootaxa.4963.1.6.pdf" docOrigin="Zootaxa 4963 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hygrodromicus carbonarius Cheng, Li &amp; Peng 2021, sp. n." docType="treatment" docVersion="7" lastPageNumber="142" masterDocId="2D02FFCFFFC73641A76C1A12FF9EAA4B" masterDocTitle="Three new species of Hygrodromicus Tronquet, 1981 (Coleoptera: Staphylinidae Omaliinae: Anthophagini) from China" masterLastPageNumber="148" masterPageNumber="138" pageNumber="139" updateTime="1698937162313" updateUser="ExternalLinkService">
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<mods:title id="BC75E4A2DA041990B2C35F6B7E2FDF04">Three new species of Hygrodromicus Tronquet, 1981 (Coleoptera: Staphylinidae Omaliinae: Anthophagini) from China</mods:title>
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<mods:namePart id="3D2221FBDBAB7E38F1CDEE3A6682C42D">Cheng, Zhi-Fei</mods:namePart>
<mods:affiliation id="E5C9C1F8C05F08554603016FEE496536">Lab of Environmental Entomology, College of Life Sciences, Shanghai Normal University, 100 Guilin Road, Shanghai, 200234, P. R. China.</mods:affiliation>
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<mods:namePart id="6333E1D2FD6BE47820C046E3EC4E6354">Pan, Zhao-Hui</mods:namePart>
<mods:affiliation id="077AAFA4A0D6380BC147A44199909EFC">XiZang agricultural and Animal Husbandry College, Linzhi 860000, P. R. China.</mods:affiliation>
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<mods:namePart id="5CF5E517CC389A223893DA8A27C554C2">Li, Li-Zhen</mods:namePart>
<mods:affiliation id="3B15F2F1354DD930F6CD98E52F1434A0">Lab of Environmental Entomology, College of Life Sciences, Shanghai Normal University, 100 Guilin Road, Shanghai, 200234, P. R. China.</mods:affiliation>
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<mods:namePart id="2E48DDEAD438893664CB41080B2C58A7">Peng, Zhong</mods:namePart>
<mods:affiliation id="8EE616B0A38534253D76D78275177B8A">Lab of Environmental Entomology, College of Life Sciences, Shanghai Normal University, 100 Guilin Road, Shanghai, 200234, P. R. China.</mods:affiliation>
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<taxonomicName id="9E924D22FFC63640A7FB188CFD47A8F2" ID-CoL="8Y4TG" authority="Cheng, Li &amp; Peng" authorityName="Cheng, Li &amp; Peng" authorityYear="2021" box="[151,729,670,697]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="139" phylum="Arthropoda" rank="species" species="carbonarius" status="sp. nov.">
<emphasis id="6BE6EAB3FFC63640A7FB188CFE71A8F2" bold="true" box="[151,495,670,697]" italics="true" pageId="1" pageNumber="139">Hygrodromicus carbonarius</emphasis>
Cheng, Li &amp; Peng
</taxonomicName>
, 
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(
<figureCitation id="C1A92A24FFC63640A7F318D1FE9AA896" box="[159,260,707,733]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,250,1872,1896]" captionStartId-1="3.[151,250,1494,1518]" captionTargetBox-0="[160,1431,284,1815]" captionTargetBox-1="[195,1418,224,1455]" captionTargetId-0="figure-61@2.[151,1436,273,1849]" captionTargetId-1="figure-27@3.[151,1436,181,1470]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Hygrodromicus carbonarius sp. n., male. Habitus of male in dorsal view (A); head in dorsal view (B); left antenna (C); labrum in dorsal view (D); left maxilla in ventral view (G); right (E) and left (F) mandibles in dorsal view; labium in ventral view (H); abdominal sternite VIII (I) and tergite VIII (J). Scale bar: 2.0 mm in A; 1.0 mm in C; 0.30 mm in B; 0.10 mm in D–J." captionText-1="FIGURE 2. Aedeagus features of Hygrodromicus carbonarius sp. n.: dorsal view (A), lateral view (B), and ventral view (C). Scale bar: 0.20 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4719973" figureDoi-1="http://doi.org/10.5281/zenodo.4719975" httpUri-0="https://zenodo.org/record/4719973/files/figure.png" httpUri-1="https://zenodo.org/record/4719975/files/figure.png" pageId="1" pageNumber="139">Figs 1–2</figureCitation>
, 
<figureCitation id="C1A92A24FFC63640A67C18D1FEBEA896" box="[272,288,707,733]" captionStart="FIGURE 7" captionStartId="9.[151,250,973,997]" captionTargetBox="[151,1436,181,949]" captionTargetId="figure-27@9.[151,1436,181,949]" captionTargetPageId="9" captionText="FIGURE 7. Distribution of Chinese species of the genus Hygrodromicus. H. carbonarius sp. n.: red circle; H. danlangi sp. n.: yellow triangles; H. shaanxiensis sp. n.: pink star." figureDoi="http://doi.org/10.5281/zenodo.4719983" httpUri="https://zenodo.org/record/4719983/files/figure.png" pageId="1" pageNumber="139">7</figureCitation>
, 
<figureCitation id="C1A92A24FFC63640A64018D1FED3A896" box="[300,333,707,733]" captionStart="FIGURE 8" captionStartId="9.[151,250,1936,1960]" captionTargetBox="[151,1436,1057,1912]" captionTargetId="figure-65@9.[151,1436,1057,1912]" captionTargetPageId="9" captionText="FIGURE 8. Type localities of Chinese species of the genus Hygrodromicus. A Chongzhe village, Bayi District, alt. 3230 m (Hygrodromicus danlangi sp. n.) B Gaduogang village, Gongbujiangda County, alt. 3620 m (Hygrodromicus danlangi sp. n. and Hygrodromicus carbonarius sp. n.)" figureDoi="http://doi.org/10.5281/zenodo.4719985" httpUri="https://zenodo.org/record/4719985/files/figure.png" pageId="1" pageNumber="139">8B</figureCitation>
)
</paragraph>
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<emphasis id="6BE6EAB3FFC63640A7FB1911FEA1A956" bold="true" box="[151,319,771,797]" pageId="1" pageNumber="139">Type material</emphasis>
(
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, 
<specimenCount id="4F94FD28FFC63640A6CC1911FE40A955" box="[416,478,771,798]" pageId="1" pageNumber="139" type="female">6 ♀♀</specimenCount>
). 
<materialsCitation id="E9FA3CFCFFC63640A6951911FA03A90B" ID-GBIF-Occurrence="3091101302" collectingDate="2018-07-18" collectorName="Cheng, Peng &amp; Shen" country="China" county="Gongbujiangda County" elevation="3620" latitude="29.9" location="Gaduogang Village" longLatPrecision="20" longitude="93.23944" municipality="Linzhi City" pageId="1" pageNumber="139" specimenCount="1" specimenCount-male="1" stateProvince="Xizang" typeStatus="holotype">
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<typeStatus id="86298803FFC63640A6951911FDF4A956" box="[505,618,771,797]" pageId="1" pageNumber="139" type="holotype">Holotype</typeStatus>
:
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<specimenCount id="4F94FD28FFC63640A51B1916FD10A955" box="[631,654,772,798]" pageId="1" pageNumber="139" type="male">♂</specimenCount>
, ‘ 
<collectingCountry id="21857631FFC63640A5CC1911FD72A956" box="[672,748,771,797]" name="China" pageId="1" pageNumber="139">China</collectingCountry>
: 
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, 
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, 
<collectingCounty id="B04C4E2DFFC63640A48B1911FB70A956" box="[999,1262,771,797]" pageId="1" pageNumber="139">Gongbujiangda County</collectingCounty>
[工布江ffi县], near 
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[ḋľÑ村], 
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, 
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, alt. 
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, 
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<collectingDate id="3D68E989FFC63640A49E1934FBEAA90B" box="[1010,1140,806,832]" pageId="1" pageNumber="139" value="2018-07-18">18.vii.2018</collectingDate>
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, 
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, 
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&amp; 
<collectorName id="F4675377FFC63640A2581934FAF2A90B" box="[1332,1388,806,832]" pageId="1" pageNumber="139">Shen</collectorName>
leg.
</materialsCitation>
/ 
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<typeStatus id="86298803FFC63640A7C51958FEA3A92F" box="[169,317,842,868]" pageId="1" pageNumber="139" type="holotype">HOLOTYPE</typeStatus>
&lt;red rectangular label, printed&gt; 
<specimenCount id="4F94FD28FFC63640A5D71958FD4CA92F" box="[699,722,842,868]" pageId="1" pageNumber="139" type="male">♂</specimenCount>
, 
<taxonomicName id="9E924D22FFC63640A58C1958FC10A928" authority="Cheng &amp; Pan &amp; Li &amp; Peng, 2021" authorityName="Cheng &amp; Pan &amp; Li &amp; Peng" authorityYear="2021" box="[736,910,841,867]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="139" phylum="Arthropoda" rank="species" species="carbonarius" status="sp. nov.">
<emphasis id="6BE6EAB3FFC63640A58C1958FC10A928" box="[736,910,841,867]" italics="true" pageId="1" pageNumber="139">H. carbonarius</emphasis>
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<taxonomicNameLabel id="70D557C8FFC63640A4F41958FC44A92F" box="[920,986,842,868]" pageId="1" pageNumber="139" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, det. 
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, 
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&amp; 
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, 2021, SNUC’ (
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)
</materialsCitation>
. 
<materialsCitation id="E9FA3CFCFFC63640A66F197FFC00A9CC" ID-GBIF-Occurrence="3091101304" box="[259,926,877,903]" collectionCode="SNUC" country="China" determinerName="Cheng &amp; Li &amp; Peng" pageId="1" pageNumber="139" specimenCount="7" specimenCount-female="6" specimenCount-male="1" stateProvince="Xizang" typeStatus="paratype">
<emphasis id="6BE6EAB3FFC63640A66F197FFE1AA9CC" bold="true" box="[259,388,877,903]" pageId="1" pageNumber="139">
<typeStatus id="86298803FFC63640A66F197FFEE1A9CC" box="[259,383,877,903]" pageId="1" pageNumber="139" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="4F94FD28FFC63640A6E0197FFE27A9CC" box="[396,441,877,903]" pageId="1" pageNumber="139" type="male">1 ♂</specimenCount>
, 
<specimenCount id="4F94FD28FFC63640A6A9197FFD98A9CC" box="[453,518,877,903]" pageId="1" pageNumber="139" type="female">6 ♀♀</specimenCount>
, same data as the holotype (
<collectionCode id="3F83AE64FFC63640A429197FFC08A9CC" box="[837,918,877,903]" pageId="1" pageNumber="139">SNUC</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="E9FA3CFCFFC63640A4C6197FFCB5A986" ID-GBIF-Occurrence="3091101310" collectingDate="2005-08-08" collectionCode="SNUC" collectorName="Liang H. B. &amp; Zhang J. F. " country="China" county="Fugong" elevation="3095" latitude="27.18408" location="Shibali" longLatPrecision="1" longitude="98.71882" municipality="Lishadi Town" pageId="1" pageNumber="139" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
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, ‘ 
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: 
<collectingRegion id="9B56F843FFC63640A352197FFB47A9CC" box="[1086,1241,877,903]" country="China" name="Yunnan" pageId="1" pageNumber="139">Yunnan Prov.</collectingRegion>
, 
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, 
<collectingMunicipality id="B949ACDBFFC63640A224197FFF47A9E0" pageId="1" pageNumber="139">Lishadi Town</collectingMunicipality>
, 
<quantity id="9E6A9B44FFC63640A78B1982FEA6A9E1" box="[231,312,912,939]" metricMagnitude="3" metricUnit="m" metricValue="8.5" pageId="1" pageNumber="139" unit="km" value="8.5">8.5 km</quantity>
up 
<location id="5C4D607AFFC63640A60B1982FE27A9E1" LSID="urn:lsid:plazi:treatment:D13B87B7FFC63645A7FB188CFED3AB0A:5C4D607AFFC63640A60B1982FE27A9E1" box="[359,441,912,938]" country="China" county="Fugong" latitude="27.18408" longLatPrecision="1" longitude="98.71882" municipality="Lishadi Town" name="Shibali" pageId="1" pageNumber="139" stateProvince="Yunnan">Shibali</location>
, riverside, 
<geoCoordinate id="3CA65066FFC63640A5551982FD5AA9E1" box="[569,708,912,938]" degrees="27.18408" direction="north" orientation="latitude" pageId="1" pageNumber="139" precision="1" value="27.18408">27.18408°N</geoCoordinate>
, 
<geoCoordinate id="3CA65066FFC63640A5BE1982FCC4A9E1" box="[722,858,912,938]" degrees="98.71882" direction="east" orientation="longitude" pageId="1" pageNumber="139" precision="1" value="98.71882">98.71882°E</geoCoordinate>
, 
<quantity id="9E6A9B44FFC63640A4041982FC5FA9E0" box="[872,961,912,939]" metricMagnitude="3" metricUnit="m" metricValue="3.0949999999999998" pageId="1" pageNumber="139" unit="m" value="3095.0">
<elevation id="D2BFD192FFC63640A4041982FC5FA9E0" box="[872,961,912,939]" metricMagnitude="3" metricUnit="m" metricValue="3.0949999999999998" pageId="1" pageNumber="139" unit="m" value="3095.0">3095 m</elevation>
</quantity>
, 
<date id="2D2C1061FFC63640A4BC1982FBCDA9E0" box="[976,1107,912,939]" pageId="1" pageNumber="139" value="2005-08-08">
<collectingDate id="3D68E989FFC63640A4BC1982FBCDA9E0" box="[976,1107,912,939]" pageId="1" pageNumber="139" value="2005-08-08">2005.VIII.8</collectingDate>
</date>
day, 
<collectorName id="F4675377FFC63640A3F91982FA88A9E0" box="[1173,1302,912,939]" pageId="1" pageNumber="139">Liang H.B.</collectorName>
, 
<collectorName id="F4675377FFC63640A2491982FEE7A986" pageId="1" pageNumber="139">Zhang J.F. California Academy</collectorName>
&amp; IOZ., 
<collectorName id="F4675377FFC63640A68D19A1FD20A986" box="[481,702,947,973]" pageId="1" pageNumber="139">Chinese. Acad. Sci.</collectorName>
’ (
<collectionCode id="3F83AE64FFC63640A5BE19A1FCBDA986" box="[722,803,947,973]" pageId="1" pageNumber="139">SNUC</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="E9FA3CFCFFC63640A45919A1FBD8A986" ID-GBIF-Occurrence="3091101306" box="[821,1094,947,974]" country="China" location="All" pageId="1" pageNumber="139" specimenCount="1" stateProvince="Yunnan" typeStatus="paratype">
<location id="5C4D607AFFC63640A45919A1FCC7A986" LSID="urn:lsid:plazi:treatment:D13B87B7FFC63645A7FB188CFED3AB0A:5C4D607AFFC63640A45919A1FCC7A986" box="[821,857,947,973]" country="China" name="All" pageId="1" pageNumber="139" stateProvince="Yunnan">All</location>
<typeStatus id="86298803FFC63640A40C19A6FC55A985" box="[864,971,948,974]" pageId="1" pageNumber="139" type="paratype">paratypes</typeStatus>
with label
</materialsCitation>
: ‘ 
<typeStatus id="86298803FFC63640A33819A1FB7AA986" box="[1108,1252,947,973]" pageId="1" pageNumber="139" type="paratype">PARATYPE</typeStatus>
&lt;yellow rectan- gular, printed&gt; 
<specimenCount id="4F94FD28FFC63640A62819C5FEC7A9BA" box="[324,345,983,1009]" pageId="1" pageNumber="139" type="male">♂</specimenCount>
[or 
<specimenCount id="4F94FD28FFC63640A6EB19C4FE02A9BA" box="[391,412,982,1009]" pageId="1" pageNumber="139" type="female">♀</specimenCount>
], 
<taxonomicName id="9E924D22FFC63640A6D819C5FDC1A9BB" authority="Cheng &amp; Pan &amp; Li &amp; Peng, 2021" authorityName="Cheng &amp; Pan &amp; Li &amp; Peng" authorityYear="2021" box="[436,607,982,1008]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="139" phylum="Arthropoda" rank="species" species="carbonarius" status="sp. nov.">
<emphasis id="6BE6EAB3FFC63640A6D819C5FDC1A9BB" box="[436,607,982,1008]" italics="true" pageId="1" pageNumber="139">H. carbonarius</emphasis>
</taxonomicName>
<emphasis id="6BE6EAB3FFC63640A50A19C4FD3BA9BB" bold="true" box="[614,677,982,1008]" pageId="1" pageNumber="139">
<taxonomicNameLabel id="70D557C8FFC63640A50A19C4FD3BA9BB" box="[614,677,982,1008]" pageId="1" pageNumber="139" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, det. Cheng, Li &amp; Peng, 2021, SNUC’.
</paragraph>
</subSubSection>
<subSubSection id="1188652AFFC63642A7AB19EBFD54AD89" lastPageId="3" lastPageNumber="141" pageId="1" pageNumber="139" type="description">
<paragraph id="592D36A1FFC63640A7AB19EBFCA4AE11" blockId="1.[151,1437,771,2030]" pageId="1" pageNumber="139">
<emphasis id="6BE6EAB3FFC63640A7AB19EBFEC7AE58" bold="true" box="[199,345,1017,1043]" pageId="1" pageNumber="139">Description.</emphasis>
Measurements (in mm) and ratios: BL 4.45–5.95; FL 2.84‒3.56; HL 0.61–0.89; HW 0.83–0.95; PL 0.78–1.17; PWb 0.83–1.00; PWm 0.83‒1.17; EL 1.28–1.72; EW 1.45‒2.06; AnL 2.71‒3.41; AeL 0.93‒0.99; HL/HW 0.69–0.94; PL/PWm 0.76‒0.84; EL/EW 0.79–0.90.
</paragraph>
<paragraph id="592D36A1FFC63640A7AB1E71FF78AEEB" blockId="1.[151,1437,771,2030]" pageId="1" pageNumber="139">
Body (
<figureCitation id="C1A92A24FFC63640A67E1E71FEF9AE36" box="[274,359,1123,1149]" captionStart="FIGURE 1" captionStartId="2.[151,250,1872,1896]" captionTargetBox="[160,1431,284,1815]" captionTargetId="figure-61@2.[151,1436,273,1849]" captionTargetPageId="2" captionText="FIGURE 1. Hygrodromicus carbonarius sp. n., male. Habitus of male in dorsal view (A); head in dorsal view (B); left antenna (C); labrum in dorsal view (D); left maxilla in ventral view (G); right (E) and left (F) mandibles in dorsal view; labium in ventral view (H); abdominal sternite VIII (I) and tergite VIII (J). Scale bar: 2.0 mm in A; 1.0 mm in C; 0.30 mm in B; 0.10 mm in D–J." figureDoi="http://doi.org/10.5281/zenodo.4719973" httpUri="https://zenodo.org/record/4719973/files/figure.png" pageId="1" pageNumber="139">Fig. 1A</figureCitation>
) broad, black, antennae paler, basal portions of femora, apical portions of tibiae and tarsi reddishbrown.
</paragraph>
<paragraph id="592D36A1FFC63640A7AB1EBBFAA2AC9F" blockId="1.[151,1437,771,2030]" pageId="1" pageNumber="139">
Head (
<figureCitation id="C1A92A24FFC63640A67D1EBBFEF8AE8F" box="[273,358,1193,1220]" captionStart="FIGURE 1" captionStartId="2.[151,250,1872,1896]" captionTargetBox="[160,1431,284,1815]" captionTargetId="figure-61@2.[151,1436,273,1849]" captionTargetPageId="2" captionText="FIGURE 1. Hygrodromicus carbonarius sp. n., male. Habitus of male in dorsal view (A); head in dorsal view (B); left antenna (C); labrum in dorsal view (D); left maxilla in ventral view (G); right (E) and left (F) mandibles in dorsal view; labium in ventral view (H); abdominal sternite VIII (I) and tergite VIII (J). Scale bar: 2.0 mm in A; 1.0 mm in C; 0.30 mm in B; 0.10 mm in D–J." figureDoi="http://doi.org/10.5281/zenodo.4719973" httpUri="https://zenodo.org/record/4719973/files/figure.png" pageId="1" pageNumber="139">Fig. 1B</figureCitation>
) subtriangular and transverse, broadest at level of eyes; vertex strongly transverse, with deep Ushaped impression between eyes; frons between supra-anternnal tubercles slightly impressed; clypeus short and narrow, weakly convex; eyes moderately prominent, about 1.50 times as long as temples (
<typeStatus id="86298803FFC63640A3071EE2FB4DAF40" box="[1131,1235,1264,1291]" pageId="1" pageNumber="139" type="holotype">holotype</typeStatus>
); ocelli indistinct, distance between ocelli 1.1 times as long as distance between ocellus and posterior margin of eye (
<typeStatus id="86298803FFC63640A3B61F01FADDAF65" box="[1242,1347,1299,1326]" pageId="1" pageNumber="139" type="holotype">holotype</typeStatus>
). Vertex and frons covered with coarse and dense punctures, and reticular microsculpture. Pubescence on head long, dense and decumbent in anterior portion. Labrum (
<figureCitation id="C1A92A24FFC63640A5E51F4BFD41AF3F" box="[649,735,1369,1396]" captionStart="FIGURE 1" captionStartId="2.[151,250,1872,1896]" captionTargetBox="[160,1431,284,1815]" captionTargetId="figure-61@2.[151,1436,273,1849]" captionTargetPageId="2" captionText="FIGURE 1. Hygrodromicus carbonarius sp. n., male. Habitus of male in dorsal view (A); head in dorsal view (B); left antenna (C); labrum in dorsal view (D); left maxilla in ventral view (G); right (E) and left (F) mandibles in dorsal view; labium in ventral view (H); abdominal sternite VIII (I) and tergite VIII (J). Scale bar: 2.0 mm in A; 1.0 mm in C; 0.30 mm in B; 0.10 mm in D–J." figureDoi="http://doi.org/10.5281/zenodo.4719973" httpUri="https://zenodo.org/record/4719973/files/figure.png" pageId="1" pageNumber="139">Fig. 1D</figureCitation>
) subrectangular, with slightly concaved anterior margin. Mandibles (
<figureCitation id="C1A92A24FFC63640A7851F6FFEC1AFDC" box="[233,351,1405,1431]" captionStart="FIGURE 1" captionStartId="2.[151,250,1872,1896]" captionTargetBox="[160,1431,284,1815]" captionTargetId="figure-61@2.[151,1436,273,1849]" captionTargetPageId="2" captionText="FIGURE 1. Hygrodromicus carbonarius sp. n., male. Habitus of male in dorsal view (A); head in dorsal view (B); left antenna (C); labrum in dorsal view (D); left maxilla in ventral view (G); right (E) and left (F) mandibles in dorsal view; labium in ventral view (H); abdominal sternite VIII (I) and tergite VIII (J). Scale bar: 2.0 mm in A; 1.0 mm in C; 0.30 mm in B; 0.10 mm in D–J." figureDoi="http://doi.org/10.5281/zenodo.4719973" httpUri="https://zenodo.org/record/4719973/files/figure.png" pageId="1" pageNumber="139">Figs 1E–F</figureCitation>
) subtriangular, each with broad base, and curved distal tooth; right mandible armed with a small distinct subtriangular tooth on inner margin near middle; prostheca with short and dense pubescence from base to above middle of mandible. Maxilla (
<figureCitation id="C1A92A24FFC63640A5541FD1FD0EAF96" box="[568,656,1475,1501]" captionStart="FIGURE 1" captionStartId="2.[151,250,1872,1896]" captionTargetBox="[160,1431,284,1815]" captionTargetId="figure-61@2.[151,1436,273,1849]" captionTargetPageId="2" captionText="FIGURE 1. Hygrodromicus carbonarius sp. n., male. Habitus of male in dorsal view (A); head in dorsal view (B); left antenna (C); labrum in dorsal view (D); left maxilla in ventral view (G); right (E) and left (F) mandibles in dorsal view; labium in ventral view (H); abdominal sternite VIII (I) and tergite VIII (J). Scale bar: 2.0 mm in A; 1.0 mm in C; 0.30 mm in B; 0.10 mm in D–J." figureDoi="http://doi.org/10.5281/zenodo.4719973" httpUri="https://zenodo.org/record/4719973/files/figure.png" pageId="1" pageNumber="139">Fig. 1G</figureCitation>
) with smooth subtriangular stipes, subrectangular palpifer with long pubescence on external margin, elongate galea with short pubescence, and slender lacinia with dense pubescence and long teeth. Maxillary palpi composed of minute and elongate palpomere 1, slightly curved palpomere 2, broadened palpomere 3, and short subcylindrical palpomere 4. Labium (
<figureCitation id="C1A92A24FFC63640A4F51C3FFC6CAC0C" box="[921,1010,1581,1607]" captionStart="FIGURE 1" captionStartId="2.[151,250,1872,1896]" captionTargetBox="[160,1431,284,1815]" captionTargetId="figure-61@2.[151,1436,273,1849]" captionTargetPageId="2" captionText="FIGURE 1. Hygrodromicus carbonarius sp. n., male. Habitus of male in dorsal view (A); head in dorsal view (B); left antenna (C); labrum in dorsal view (D); left maxilla in ventral view (G); right (E) and left (F) mandibles in dorsal view; labium in ventral view (H); abdominal sternite VIII (I) and tergite VIII (J). Scale bar: 2.0 mm in A; 1.0 mm in C; 0.30 mm in B; 0.10 mm in D–J." figureDoi="http://doi.org/10.5281/zenodo.4719973" httpUri="https://zenodo.org/record/4719973/files/figure.png" pageId="1" pageNumber="139">Fig. 1H</figureCitation>
) elongate, with flattened prementum divided by longitudinal groove anteriorly. Labial palpi long, palpomere 1 moderately large and elongate; palpomere 2 robust, broadened apically; palpomere 3 slender, distinctly narrower than palpomere 2. Antennae (
<figureCitation id="C1A92A24FFC63640A2651C61FAC6ACC6" box="[1289,1368,1651,1677]" captionStart="FIGURE 1" captionStartId="2.[151,250,1872,1896]" captionTargetBox="[160,1431,284,1815]" captionTargetId="figure-61@2.[151,1436,273,1849]" captionTargetPageId="2" captionText="FIGURE 1. Hygrodromicus carbonarius sp. n., male. Habitus of male in dorsal view (A); head in dorsal view (B); left antenna (C); labrum in dorsal view (D); left maxilla in ventral view (G); right (E) and left (F) mandibles in dorsal view; labium in ventral view (H); abdominal sternite VIII (I) and tergite VIII (J). Scale bar: 2.0 mm in A; 1.0 mm in C; 0.30 mm in B; 0.10 mm in D–J." figureDoi="http://doi.org/10.5281/zenodo.4719973" httpUri="https://zenodo.org/record/4719973/files/figure.png" pageId="1" pageNumber="139">Fig 1C</figureCitation>
) with long antennomeres; length × width (in mm) of antennomeres 1–11 (
<typeStatus id="86298803FFC63640A4EF1C84FC72ACFA" box="[899,1004,1686,1713]" pageId="1" pageNumber="139" type="holotype">holotype</typeStatus>
): 0.26 × 0.13: 0.15 × 0.11: 0.19 × 0.11: 0.21 × 0.11: 0.22 × 0.11: 0.22 × 0.11: 0.25 × 0.11: 0.25 × 0.11: 0.25 × 0.11: 0.22 × 0.11: 0.34 × 0.11.
</paragraph>
<paragraph id="592D36A1FFC63640A7AB1CCFFAF7ADC8" blockId="1.[151,1437,771,2030]" pageId="1" pageNumber="139">Pronotum subtrapezoidal, broadest near anterior fourth; anterior margin widely rounded, lateral margins in anterior 2/3 rounded and nearly straight at posterior third, basal margin straight; hind angles obtuse; disc distinctly convex, with indistinct impression, stretching from middle to base. Punctures of disc finer and denser than those of head; pubescence moderately long, dense and decumbent in posterior portion; microsculpture distinct and evenly distributed. Scutellum triangular, covered with several punctures and pubescence, and distinct microsculpture.</paragraph>
<paragraph id="592D36A1FFC63640A7AB1D9FFE4FADA6" blockId="1.[151,1437,771,2030]" pageId="1" pageNumber="139">Elytra subtrapezoidal, short, distinctly wider than long, gradually dilated posteriorly, posterior angles broadly rounded; punctures and pubescence similar to those of pronotum; microsculpture absent. Hind wings brachypterous, but not completely reduced.</paragraph>
<paragraph id="592D36A1FFC53643A7AB1A85FC1DAAB2" blockId="2.[151,1436,151,249]" pageId="2" pageNumber="140">Abdomen distinctly transverse, widest at segment IV, evenly narrowing posteriorly. Tergites with fine punctures and microsculpture, and short decumbent pubescence, denser on apical tergites; tergites IV–V with a pair of tomentose spots in middle, spots on tergite V smaller and less transverse.</paragraph>
<caption id="0DED6629FFC53643A7FB1D42FF59AD9F" ID-DOI="http://doi.org/10.5281/zenodo.4719973" ID-Zenodo-Dep="4719973" httpUri="https://zenodo.org/record/4719973/files/figure.png" pageId="2" pageNumber="140" startId="2.[151,250,1872,1896]" targetBox="[160,1431,284,1815]" targetPageId="2">
<paragraph id="592D36A1FFC53643A7FB1D42FF59AD9F" blockId="2.[151,1436,1872,2004]" pageId="2" pageNumber="140">
<emphasis id="6BE6EAB3FFC53643A7FB1D42FE8DAD23" bold="true" box="[151,275,1872,1896]" pageId="2" pageNumber="140">FIGURE 1.</emphasis>
<taxonomicName id="9E924D22FFC53643A6751D42FDA6AD23" authority="Cheng &amp; Pan &amp; Li &amp; Peng, 2021" authorityName="Cheng &amp; Pan &amp; Li &amp; Peng" authorityYear="2021" box="[281,568,1872,1896]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="140" phylum="Arthropoda" rank="species" species="carbonarius" status="sp. nov.">
<emphasis id="6BE6EAB3FFC53643A6751D42FDA6AD23" box="[281,568,1872,1896]" italics="true" pageId="2" pageNumber="140">Hygrodromicus carbonarius</emphasis>
</taxonomicName>
<emphasis id="6BE6EAB3FFC53643A5521D43FDE8AD22" bold="true" box="[574,630,1873,1897]" pageId="2" pageNumber="140">
<taxonomicNameLabel id="70D557C8FFC53643A5521D43FDE8AD22" box="[574,630,1873,1897]" pageId="2" pageNumber="140" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, male. Habitus of male in dorsal view (A); head in dorsal view (B); left antenna (C); labrum in dorsal view (D); left maxilla in ventral view (G); right (E) and left (F) mandibles in dorsal view; labium in ventral view (H); abdominal sternite VIII (I) and tergite VIII (J). Scale bar: 2.0 mm in A; 1.0 mm in C; 0.30 mm in B; 0.10 mm in D–J.
</paragraph>
</caption>
<caption id="0DED6629FFC43642A7FB1FC4FEFDAC5A" ID-DOI="http://doi.org/10.5281/zenodo.4719975" ID-Zenodo-Dep="4719975" httpUri="https://zenodo.org/record/4719975/files/figure.png" pageId="3" pageNumber="141" startId="3.[151,250,1494,1518]" targetBox="[195,1418,224,1455]" targetPageId="3">
<paragraph id="592D36A1FFC43642A7FB1FC4FEFDAC5A" blockId="3.[151,1437,1494,1554]" pageId="3" pageNumber="141">
<emphasis id="6BE6EAB3FFC43642A7FB1FC4FE8AAFA5" bold="true" box="[151,276,1494,1518]" pageId="3" pageNumber="141">FIGURE 2.</emphasis>
Aedeagus features of 
<taxonomicName id="9E924D22FFC43642A6951FC4FC87AFA5" authority="Cheng &amp; Pan &amp; Li &amp; Peng, 2021" authorityName="Cheng &amp; Pan &amp; Li &amp; Peng" authorityYear="2021" box="[505,793,1494,1518]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="141" phylum="Arthropoda" rank="species" species="carbonarius" status="sp. nov.">
<emphasis id="6BE6EAB3FFC43642A6951FC4FC87AFA5" box="[505,793,1494,1518]" italics="true" pageId="3" pageNumber="141">Hygrodromicus carbonarius</emphasis>
</taxonomicName>
<emphasis id="6BE6EAB3FFC43642A44C1FC4FCC4AFA5" bold="true" box="[800,858,1494,1518]" pageId="3" pageNumber="141">
<taxonomicNameLabel id="70D557C8FFC43642A44C1FC4FCC4AFA5" box="[800,858,1494,1518]" pageId="3" pageNumber="141" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
: dorsal view (A), lateral view (B), and ventral view (C). Scale bar: 0.20 mm.
</paragraph>
</caption>
<paragraph id="592D36A1FFC43642A7AB1C52FD14ACA1" blockId="3.[151,1437,1600,2023]" pageId="3" pageNumber="141">
Male. Apical margin of abdominal tergite VIII (
<figureCitation id="C1A92A24FFC43642A5841C52FCABAC11" box="[744,821,1600,1626]" captionStart="FIGURE 1" captionStartId="2.[151,250,1872,1896]" captionTargetBox="[160,1431,284,1815]" captionTargetId="figure-61@2.[151,1436,273,1849]" captionTargetPageId="2" captionText="FIGURE 1. Hygrodromicus carbonarius sp. n., male. Habitus of male in dorsal view (A); head in dorsal view (B); left antenna (C); labrum in dorsal view (D); left maxilla in ventral view (G); right (E) and left (F) mandibles in dorsal view; labium in ventral view (H); abdominal sternite VIII (I) and tergite VIII (J). Scale bar: 2.0 mm in A; 1.0 mm in C; 0.30 mm in B; 0.10 mm in D–J." figureDoi="http://doi.org/10.5281/zenodo.4719973" httpUri="https://zenodo.org/record/4719973/files/figure.png" pageId="3" pageNumber="141">Fig. 1I</figureCitation>
) somewhat truncated, indistinctly emarginated; apical margin of abdominal sternite VIII (
<figureCitation id="C1A92A24FFC43642A55C1C76FD1FAC35" box="[560,641,1636,1662]" captionStart="FIGURE 1" captionStartId="2.[151,250,1872,1896]" captionTargetBox="[160,1431,284,1815]" captionTargetId="figure-61@2.[151,1436,273,1849]" captionTargetPageId="2" captionText="FIGURE 1. Hygrodromicus carbonarius sp. n., male. Habitus of male in dorsal view (A); head in dorsal view (B); left antenna (C); labrum in dorsal view (D); left maxilla in ventral view (G); right (E) and left (F) mandibles in dorsal view; labium in ventral view (H); abdominal sternite VIII (I) and tergite VIII (J). Scale bar: 2.0 mm in A; 1.0 mm in C; 0.30 mm in B; 0.10 mm in D–J." figureDoi="http://doi.org/10.5281/zenodo.4719973" httpUri="https://zenodo.org/record/4719973/files/figure.png" pageId="3" pageNumber="141">Fig. 1J</figureCitation>
) broadly emarginated. Aedeagus (
<figureCitation id="C1A92A24FFC43642A3611C76FB13AC35" box="[1037,1165,1636,1662]" captionStart="FIGURE 2" captionStartId="3.[151,250,1494,1518]" captionTargetBox="[195,1418,224,1455]" captionTargetId="figure-27@3.[151,1436,181,1470]" captionTargetPageId="3" captionText="FIGURE 2. Aedeagus features of Hygrodromicus carbonarius sp. n.: dorsal view (A), lateral view (B), and ventral view (C). Scale bar: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.4719975" httpUri="https://zenodo.org/record/4719975/files/figure.png" pageId="3" pageNumber="141">Figs 2A–C</figureCitation>
) elongate, median lobe slightly emarginated in lateral margins near anterior third; parameres slightly asymmetrical, slender, significantly exceeding apex of median lobe, each bearing four apical setae; internal sac with submedian elongate membranous structures and long flagellum between them.
</paragraph>
<paragraph id="592D36A1FFC43642A7AB1CE6FBB6AD45" blockId="3.[151,1437,1600,2023]" box="[199,1064,1780,1807]" pageId="3" pageNumber="141">Female. Apical margin of abdominal sternite VIII without concavity apically.</paragraph>
<paragraph id="592D36A1FFC43642A7AB1D0AFD54AD89" blockId="3.[151,1437,1600,2023]" pageId="3" pageNumber="141">
<emphasis id="6BE6EAB3FFC43642A7AB1D0AFE2DAD79" bold="true" box="[199,435,1816,1842]" pageId="3" pageNumber="141">Comparative notes.</emphasis>
Based on the general shape of the forebody, coloration, shapes of maxillary and labial palpi, the presence of U-shaped impression between eyes, and shapes of apical margins of male tergite and sternite VIII, 
<taxonomicName id="9E924D22FFC43642A7BB1D73FE1DAD31" authority="Cheng &amp; Pan &amp; Li &amp; Peng, 2021" authorityName="Cheng &amp; Pan &amp; Li &amp; Peng" authorityYear="2021" box="[215,387,1888,1914]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="141" phylum="Arthropoda" rank="species" species="carbonarius" status="sp. nov.">
<emphasis id="6BE6EAB3FFC43642A7BB1D73FE1DAD31" box="[215,387,1888,1914]" italics="true" pageId="3" pageNumber="141">H. carbonarius</emphasis>
</taxonomicName>
<emphasis id="6BE6EAB3FFC43642A6E71D72FE52AD31" bold="true" box="[395,460,1888,1914]" pageId="3" pageNumber="141">
<taxonomicNameLabel id="70D557C8FFC43642A6E71D72FE52AD31" box="[395,460,1888,1914]" pageId="3" pageNumber="141" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is most similar to 
<taxonomicName id="9E924D22FFC43642A5CD1D73FCB7AD31" authority="Cheng &amp; Pan &amp; Li &amp; Peng, 2021" authorityName="Cheng &amp; Pan &amp; Li &amp; Peng" authorityYear="2021" box="[673,809,1888,1915]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="141" phylum="Arthropoda" rank="species" species="danlangi" status="sp. nov.">
<emphasis id="6BE6EAB3FFC43642A5CD1D73FD2BAD31" box="[673,693,1889,1914]" italics="true" pageId="3" pageNumber="141">H</emphasis>
. 
<emphasis id="6BE6EAB3FFC43642A5A91D72FCB7AD31" box="[709,809,1888,1914]" italics="true" pageId="3" pageNumber="141">danlangi</emphasis>
</taxonomicName>
<emphasis id="6BE6EAB3FFC43642A45D1D72FCEFAD31" bold="true" box="[817,881,1888,1914]" pageId="3" pageNumber="141">
<taxonomicNameLabel id="70D557C8FFC43642A45D1D72FCEFAD31" box="[817,881,1888,1914]" pageId="3" pageNumber="141" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, but differs by the wider pronotum, the presence of additional small tooth in inner margins of mandible, less concaved apical margin of labrum, and details of the internal and external morphology of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="1188652AFFC43645A7AB1DDEFF6EAAB2" lastPageId="4" lastPageNumber="142" pageId="3" pageNumber="141" type="distribution">
<paragraph id="592D36A1FFC43642A7AB1DDEFCF0ADAC" blockId="3.[151,1437,1600,2023]" box="[199,878,1996,2023]" pageId="3" pageNumber="141">
<emphasis id="6BE6EAB3FFC43642A7AB1DDEFEFFADAD" bold="true" box="[199,353,1996,2022]" pageId="3" pageNumber="141">Distribution.</emphasis>
Southwestern 
<collectingCountry id="21857631FFC43642A56B1DDEFDD0ADAD" box="[519,590,1996,2022]" name="China" pageId="3" pageNumber="141">China</collectingCountry>
: 
<collectingRegion id="9B56F843FFC43642A5351DDEFD33ADAD" box="[601,685,1996,2022]" country="China" name="Xizang" pageId="3" pageNumber="141">Xizang</collectingRegion>
, 
<collectingRegion id="9B56F843FFC43642A5DA1DDFFC92ADAC" box="[694,780,1997,2023]" country="China" name="Yunnan" pageId="3" pageNumber="141">Yunnan</collectingRegion>
(
<figureCitation id="C1A92A24FFC43642A4771DDEFCFEADAC" box="[795,864,1996,2023]" captionStart="FIGURE 7" captionStartId="9.[151,250,973,997]" captionTargetBox="[151,1436,181,949]" captionTargetId="figure-27@9.[151,1436,181,949]" captionTargetPageId="9" captionText="FIGURE 7. Distribution of Chinese species of the genus Hygrodromicus. H. carbonarius sp. n.: red circle; H. danlangi sp. n.: yellow triangles; H. shaanxiensis sp. n.: pink star." figureDoi="http://doi.org/10.5281/zenodo.4719983" httpUri="https://zenodo.org/record/4719983/files/figure.png" pageId="3" pageNumber="141">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="592D36A1FFC33645A7AB1A84FF6EAAB2" blockId="4.[151,1436,150,321]" pageId="4" pageNumber="142">
<emphasis id="6BE6EAB3FFC33645A7AB1A84FED5AAFB" bold="true" box="[199,331,150,176]" pageId="4" pageNumber="142">Bionomics.</emphasis>
The species inhabits wet mosses or mixed leaf litter on banks of small streams with very cold water (
<figureCitation id="C1A92A24FFC33645A7CC1AA9FF66AA9E" box="[160,248,187,213]" captionStart="FIGURE 8" captionStartId="9.[151,250,1936,1960]" captionTargetBox="[151,1436,1057,1912]" captionTargetId="figure-65@9.[151,1436,1057,1912]" captionTargetPageId="9" captionText="FIGURE 8. Type localities of Chinese species of the genus Hygrodromicus. A Chongzhe village, Bayi District, alt. 3230 m (Hygrodromicus danlangi sp. n.) B Gaduogang village, Gongbujiangda County, alt. 3620 m (Hygrodromicus danlangi sp. n. and Hygrodromicus carbonarius sp. n.)" figureDoi="http://doi.org/10.5281/zenodo.4719985" httpUri="https://zenodo.org/record/4719985/files/figure.png" pageId="4" pageNumber="142">Fig. 8B</figureCitation>
). Specimens were collected at high elevations more than 
<quantity id="9E6A9B44FFC33645A4111AA9FC4DAA9E" box="[893,979,187,213]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="142" unit="m" value="3000.0">3000 m</quantity>
a.s.l. in 
<collectingRegion id="9B56F843FFC33645A35E1AA9FB1DAA9E" box="[1074,1155,187,213]" country="China" name="Xizang" pageId="4" pageNumber="142">Xizang</collectingRegion>
in July and in 
<collectingRegion id="9B56F843FFC33645A2451AA9FAE1AA9E" box="[1321,1407,187,213]" country="China" name="Yunnan" pageId="4" pageNumber="142">Yunnan</collectingRegion>
in August.
</paragraph>
</subSubSection>
<subSubSection id="1188652AFFC33645A7AB1B11FED3AB0A" pageId="4" pageNumber="142" type="etymology">
<paragraph id="592D36A1FFC33645A7AB1B11FED3AB0A" blockId="4.[151,1436,150,321]" pageId="4" pageNumber="142">
<emphasis id="6BE6EAB3FFC33645A7AB1B11FED0AB56" bold="true" box="[199,334,259,285]" pageId="4" pageNumber="142">Etymology.</emphasis>
The specific epithet ‘ 
<taxonomicName id="9E924D22FFC33645A5271B11FD4AAB56" authorityName="Cheng &amp; Pan &amp; Li &amp; Peng" authorityYear="2021" box="[587,724,259,285]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="142" phylum="Arthropoda" rank="species" species="carbonarius">
<emphasis id="6BE6EAB3FFC33645A5271B11FD4AAB56" box="[587,724,259,285]" italics="true" pageId="4" pageNumber="142">carbonarius</emphasis>
</taxonomicName>
’ means ‘carbon’, referring to the coloration of a new species, black as carbon.
</paragraph>
</subSubSection>
</treatment>
</document>