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<document id="17EC7EDBBD61FBC3D362C37C0BBCA90B" ID-CLB-Dataset="26049" ID-DOI="10.11646/zootaxa.4654.1.1" ID-GBIF-Dataset="9ce1d103-9234-4fbe-9544-f1c294bb8783" ID-ISSN="1175-5326" ID-Zenodo-Dep="3365803" ID-ZooBank="CFD82CC0-3110-472E-972B-7ADC0C523A04" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1565592061838" checkinUser="plazi" docAuthor="Hansen, Thomas" docDate="2019" docId="BF18F633A98DFF6B2B9CC558FCC8FF34" docLanguage="en" docName="zootaxa.4654.1.1.pdf" docOrigin="Zootaxa 4654 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Columbarium heberti D. D. Cr. D. D. 1880" docType="treatment" docVersion="21" lastPageNumber="142" masterDocId="43218E4BA900FFE52B0BC14DFFA9FFE3" masterDocTitle="Gastropods from the Cretaceous-Palaeogene boundary in Denmark" masterLastPageNumber="196" masterPageNumber="1" pageNumber="142" updateTime="1698750564626" updateUser="ExternalLinkService">
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<mods:title id="02432D0D9FABAC3264AD62047C2DA822">Gastropods from the Cretaceous-Palaeogene boundary in Denmark</mods:title>
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<treatment id="BF18F633A98DFF6B2B9CC558FCC8FF34" ID-GBIF-Taxon="159074686" LSID="urn:lsid:plazi:treatment:BF18F633A98DFF6B2B9CC558FCC8FF34" httpUri="http://treatment.plazi.org/id/BF18F633A98DFF6B2B9CC558FCC8FF34" lastPageId="142" lastPageNumber="142" pageId="141" pageNumber="142">
<subSubSection id="7FAB14AEA98DFF682B9CC558FD67FBCC" box="[151,718,1045,1072]" pageId="141" pageNumber="142" type="nomenclature">
<paragraph id="370E4725A98DFF682B9CC558FD67FBCC" blockId="141.[151,718,1045,1107]" box="[151,718,1045,1072]" pageId="141" pageNumber="142">
<heading id="6C46F049A98DFF682B9CC558FD67FBCC" bold="true" box="[151,718,1045,1072]" fontSize="11" level="1" pageId="141" pageNumber="142" reason="1">
<taxonomicName id="F0B13CA6A98DFF682B9CC558FD67FBCC" authority="(Briart &amp; Cornet, 1880)" authorityName="D. D. Cr. D. D." authorityYear="1880" baseAuthorityName="Briart &amp; Cornet" baseAuthorityYear="1880" box="[151,718,1045,1072]" class="Gastropoda" family="Turbinellidae" genus="Columbarium" kingdom="Animalia" order="Neogastropoda" pageId="141" pageNumber="142" phylum="Mollusca" rank="species" species="heberti">
<emphasis id="05C59B37A98DFF682B9CC558FD67FBCC" bold="true" box="[151,718,1045,1072]" pageId="141" pageNumber="142">
<emphasis id="05C59B37A98DFF682B9CC558FE34FBD3" bold="true" box="[151,413,1045,1072]" italics="true" pageId="141" pageNumber="142">Columbarium heberti</emphasis>
(
<bibRefCitation id="53203AD4A98DFF682AA7C558FD6FFBCC" author="Briart, A. &amp; Cornet, F. L." box="[428,710,1045,1071]" pageId="141" pageNumber="142" pagination="1 - 73" refId="ref126042" refString="Briart, A. &amp; Cornet, F. L. (1880) Descriptions des fossiles du calcaire grossier de Mons. Troisieme partie. Memoires courronnes et Memoires des Savants Etrangers, Academie Royale des Sciences, des Lettres et des Beaux-Arts de Belgique, 43 (1), i-viii + 1 - 73, pls. 13 - 18." type="journal article" year="1880">Briart &amp; Cornet, 1880</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7FAB14AEA98DFF682B9CC574FD75FB75" pageId="141" pageNumber="142" type="description">
<paragraph id="370E4725A98DFF682B9CC574FF59FBB0" blockId="141.[151,718,1045,1107]" box="[151,240,1081,1107]" pageId="141" pageNumber="142">
<figureCitation id="AF8A5BA0A98DFF682B9CC574FF59FBB0" box="[151,240,1081,1107]" captionStart="FIGURE 31" captionStartId="141.[151,250,188,213]" captionTargetId="figure@140.[151,1435,179,1918]" captionTargetPageId="140" captionText="FIGURE 31. A–C. Vexillum tenestolidum n. sp. A–B. Silicone cast of holotype MGUH 33259. Width of protoconch 0.7 mm. Width of last whorl 3.4 mm. C. Silicone cast of paratype MGUH 33260. Width 5.8 mm. D–G. Vexillum? rosenkrantzi n. sp. D–E. Silicone cast and inner mould of holotype MGUH 33261. Width 1.9 and 1.6 mm respectively. F. Silicone cast of paratype MGUH 33262. Width 3.2 mm. G. Silicone cast of paratype MGUH 33263. Width 2.9 mm. H. Vexillum? sp. Silicone cast of MGUH 33264. Width 3.5 mm. I. Columbarium heberti (Briart &amp; Cornet, 1880). Silicone cast of last whorl MGUH 33265. Width 7.7 mm. Photograph by S.L. Jakobsen. J. Columbarium sp. Silicone cast of MGUH 33266. Width 13.3 mm. K–N. Conomitra glabra (Ravn, 1902). K. Silicone cast of MGUH 33267. Width 3.6 mm. L–M. Silicone cast of MGUH 33268. Width 3.6 and 3.9 mm respectively. N. External cast of syntype MGUH 116. Height 8.2 mm. Photograph by S.L. Jakobsen. O. Volutomitra cf. quinqueplicata Ravn, 1902. Silicone cast of MGUH 33269. Width 11.6 mm. P. Volutomitra quinqueplicata Ravn, 1902. External cast representing last whorl of type specimen MGUH 117. Middle Danien Faxe Formation, Denmark. Height 38.6 mm. Photograph by S.L. Jakobsen. Q–R. Spirancilla milthersii (Ravn, 1902). Q. Silicone cast of holotype MGUH 33270. Width 3.3 mm. R. Silicone cast of MGUH 33271. Width 3.1 mm. S–T. Exilia sp., Silicone cast of MGUH 33272. Width 1.6 mm. U–W. Latiromitra? sp. External cast and inner mould of MGUH 33273. Width 4.6 mm for casts and 4.0 mm for mould." figureDoi="http://doi.org/10.5281/zenodo.3442118" httpUri="https://zenodo.org/record/3442118/files/figure.png" pageId="141" pageNumber="142">Fig. 31I</figureCitation>
</paragraph>
<paragraph id="370E4725A98DFF682B9CC533FD75FB75" blockId="141.[151,1246,1149,1297]" box="[151,732,1149,1174]" pageId="141" pageNumber="142">
1880
<taxonomicName id="F0B13CA6A98DFF682BDBC533FDAFFB75" authority="Briart &amp; Cornet" authorityName="Briart &amp; Cornet-Gorbach" box="[208,518,1149,1174]" class="Gastropoda" family="Columbariidae" genus="Fusus" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="141" pageNumber="142" phylum="Mollusca" rank="species" species="heberti">
<emphasis id="05C59B37A98DFF682BDBC533FEF2FB75" box="[208,347,1150,1174]" italics="true" pageId="141" pageNumber="142">Fusus heberti</emphasis>
Briart &amp; Cornet
</taxonomicName>
: 20, pl. 14, fig. 8a–d.
</paragraph>
</subSubSection>
<subSubSection id="7FAB14AEA98DFF682B9CC5D1FC2CFB30" pageId="141" pageNumber="142" type="reference_group">
<paragraph id="370E4725A98DFF682B9CC5D1FB77FB57" blockId="141.[151,1246,1149,1297]" box="[151,1246,1180,1205]" pageId="141" pageNumber="142">
1969
<taxonomicName id="F0B13CA6A98DFF682BDBC5D1FD33FB56" authority="(Briart &amp; Cornet 1880)" authorityName="D. D. Cr. D. D." authorityYear="1880" baseAuthorityName="Briart &amp; Cornet" baseAuthorityYear="1880" box="[208,666,1180,1205]" class="Gastropoda" family="Turbinellidae" genus="Columbarium" kingdom="Animalia" order="Neogastropoda" pageId="141" pageNumber="142" phylum="Mollusca" rank="species" species="heberti">
<emphasis id="05C59B37A98DFF682BDBC5D1FE02FB57" box="[208,427,1180,1204]" italics="true" pageId="141" pageNumber="142">Columbarium heberti</emphasis>
(
<bibRefCitation id="53203AD4A98DFF682AB3C5D1FD3AFB56" author="Briart, A. &amp; Cornet, F. L." box="[440,659,1180,1205]" pageId="141" pageNumber="142" pagination="1 - 73" refId="ref126042" refString="Briart, A. &amp; Cornet, F. L. (1880) Descriptions des fossiles du calcaire grossier de Mons. Troisieme partie. Memoires courronnes et Memoires des Savants Etrangers, Academie Royale des Sciences, des Lettres et des Beaux-Arts de Belgique, 43 (1), i-viii + 1 - 73, pls. 13 - 18." type="journal article" year="1880">Briart &amp; Cornet 1880</bibRefCitation>
)
</taxonomicName>
—Darragh: 72–73, pl. 2, figs 19–21, 26–28, textfigs 3, 9.
</paragraph>
<paragraph id="370E4725A98DFF682B9CC5F6FC2CFB30" blockId="141.[151,1246,1149,1297]" box="[151,901,1210,1235]" pageId="141" pageNumber="142">
1972
<taxonomicName id="F0B13CA6A98DFF682BDBC5F1FDDEFB30" authority="Briart &amp; Cornet-Gorbach" authorityName="Briart &amp; Cornet-Gorbach" box="[208,631,1210,1235]" class="Gastropoda" family="Columbariidae" genus="Fusus" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="141" pageNumber="142" phylum="Mollusca" rank="species" species="heberti">
<emphasis id="05C59B37A98DFF682BDBC5F1FEF2FB30" box="[208,347,1211,1235]" italics="true" pageId="141" pageNumber="142">Fusus heberti</emphasis>
Briart &amp; Cornet—Gorbach
</taxonomicName>
: 102–103, pl. 15, figs 2–4.
</paragraph>
</subSubSection>
<subSubSection id="7FAB14AEA98DFF682B9CC597FDDFFAF3" pageId="141" pageNumber="142" type="description">
<paragraph id="370E4725A98DFF682B9CC597FC76FB11" blockId="141.[151,1246,1149,1297]" box="[151,991,1241,1266]" pageId="141" pageNumber="142">
1973
<taxonomicName id="F0B13CA6A98DFF682BDBC597FD09FB11" authority="(Briart &amp; Cornet, 1870)" baseAuthorityName="Briart &amp; Cornet" baseAuthorityYear="1870" box="[208,672,1241,1266]" class="Gastropoda" family="Turbinellidae" genus="Columbarium" kingdom="Animalia" order="Neogastropoda" pageId="141" pageNumber="142" phylum="Mollusca" rank="species" species="heberti">
<emphasis id="05C59B37A98DFF682BDBC597FE02FB11" box="[208,427,1242,1266]" italics="true" pageId="141" pageNumber="142">Columbarium heberti</emphasis>
(Briart &amp; Cornet, 1870)
</taxonomicName>
—Glibert: 64–65, pl. 7, fig. 19.
</paragraph>
<paragraph id="370E4725A98DFF682B9CC5B4FDDFFAF3" blockId="141.[151,1246,1149,1297]" box="[151,630,1272,1297]" pageId="141" pageNumber="142">
For more synonymy see
<bibRefCitation id="53203AD4A98DFF682A99C5B5FD83FAF2" author="Darragh, T. A." box="[402,554,1272,1297]" pageId="141" pageNumber="142" pagination="63 - 119" refId="ref128270" refString="Darragh, T. A. (1969) A Revision of the Family Columbariidae (Mollusca: Gastropoda). Proceedings of the Royal Society of Victoria, 83 (1), 63 - 119, 5 pls." type="book chapter" year="1969">Darragh (1969</bibRefCitation>
: p. 72).
</paragraph>
</subSubSection>
<subSubSection id="7FAB14AEA98DFF6B2B9CC473FCC8FF34" lastPageId="142" lastPageNumber="143" pageId="141" pageNumber="142" type="materials_examined">
<paragraph id="370E4725A98DFF682B9CC473FB19FA42" blockId="141.[151,1436,1342,2017]" pageId="141" pageNumber="142">
<emphasis id="05C59B37A98DFF682B9CC473FEBBFABA" bold="true" box="[151,274,1342,1369]" pageId="141" pageNumber="142">Diagnosis.</emphasis>
Protoconch of 1½ small whorls, embryonic whorl deviated slightly from axis. Teleoconch keel with scales. Sculpture of strong and low transverse costae and three sharp and strong spiral cords, one on keel, the second halfway between keel and abapical suture, and the third hidden by succeeding whorl on spire.
</paragraph>
<paragraph id="370E4725A98DFF682BCCC4E7FB13FA26" blockId="141.[151,1436,1342,2017]" box="[199,1210,1450,1477]" pageId="141" pageNumber="142">
<emphasis id="05C59B37A98DFF682BCCC4E7FE27FA26" bold="true" box="[199,398,1450,1477]" pageId="141" pageNumber="142">Danish material.</emphasis>
Specimen
<collectionCode id="51A0DFE0A98DFF682900C4E6FDC8FA26" box="[523,609,1451,1477]" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="141" pageNumber="142">MGUH</collectionCode>
33265 is a fragmentary mould of the last two whorls.
</paragraph>
<paragraph id="370E4725A98DFF682BCCC482FCEEF9EE" blockId="141.[151,1436,1342,2017]" pageId="141" pageNumber="142">
<emphasis id="05C59B37A98DFF682BCCC482FD9EFA0A" bold="true" box="[199,567,1486,1513]" pageId="141" pageNumber="142">Type stratum and type locality.</emphasis>
The
<typeStatus id="E80AF987A98DFF682964C482FD7FFA0A" box="[623,726,1487,1513]" pageId="141" pageNumber="142" type="lectotype">lectotype</typeStatus>
(
<bibRefCitation id="53203AD4A98DFF6829EFC482FC71FA0A" author="Briart, A. &amp; Cornet, F. L." box="[740,984,1487,1513]" pageId="141" pageNumber="142" pagination="1 - 73" refId="ref126042" refString="Briart, A. &amp; Cornet, F. L. (1880) Descriptions des fossiles du calcaire grossier de Mons. Troisieme partie. Memoires courronnes et Memoires des Savants Etrangers, Academie Royale des Sciences, des Lettres et des Beaux-Arts de Belgique, 43 (1), i-viii + 1 - 73, pls. 13 - 18." type="journal article" year="1880">Briart &amp; Cornet 1880</bibRefCitation>
: fig. 8d) as suggested by
<bibRefCitation id="53203AD4A98DFF682FE4C482FA35FA09" author="Darragh, T. A." box="[1263,1436,1487,1514]" pageId="141" pageNumber="142" pagination="63 - 119" refId="ref128270" refString="Darragh, T. A. (1969) A Revision of the Family Columbariidae (Mollusca: Gastropoda). Proceedings of the Royal Society of Victoria, 83 (1), 63 - 119, 5 pls." type="book chapter" year="1969">Darragh (1969)</bibRefCitation>
comes from the Danian Calcaire de Mons, Hainaut,
<collectingCountry id="4FA607B5A98DFF6829D5C4BEFCEBF9EE" box="[734,834,1523,1549]" name="Belgium" pageId="141" pageNumber="142">Belgium</collectingCountry>
.
</paragraph>
<paragraph id="370E4725A98DFF682BCCC75BFE08F999" blockId="141.[151,1436,1342,2017]" pageId="141" pageNumber="142">
<emphasis id="05C59B37A98DFF682BCCC75BFEF3F9D2" bold="true" box="[199,346,1558,1585]" pageId="141" pageNumber="142">Occurrence.</emphasis>
In
<collectingCountry id="4FA607B5A98DFF682A75C75AFE4FF9D2" box="[382,486,1559,1585]" name="Denmark" pageId="141" pageNumber="142">Denmark</collectingCountry>
it has been found in the
<taxonomicName id="F0B13CA6A98DFF6829F3C75AFBE5F9D2" authority="Limestone Member" authorityName="Gilleaudeau et al." authorityYear="2018" box="[760,1100,1559,1585]" class="Gastropoda" family="Cerithiidae" genus="Cerithium" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="141" pageNumber="142" phylum="Mollusca" rank="genus">Cerithium Limestone Member</taxonomicName>
at Stevns Klint. Outside
<collectingCountry id="4FA607B5A98DFF682E6FC75AFF78F9B6" name="Denmark" pageId="141" pageNumber="142">Denmark</collectingCountry>
it is known from the Maastrichtian at Maastricht in
<collectingCountry id="4FA607B5A98DFF68282AC776FCD2F9B6" box="[801,891,1595,1621]" name="Netherlands" pageId="141" pageNumber="142">Holland</collectingCountry>
and from the Danian (Montian) in
<collectingCountry id="4FA607B5A98DFF682E00C776FAC2F9B6" box="[1291,1387,1595,1621]" name="Belgium" pageId="141" pageNumber="142">Belgium</collectingCountry>
and
<collectingCountry id="4FA607B5A98DFF682B9CC712FF4CF99A" box="[151,229,1631,1657]" name="Poland" pageId="141" pageNumber="142">Poland</collectingCountry>
(
<bibRefCitation id="53203AD4A98DFF682BF8C712FE3DF999" author="Darragh, T. A." box="[243,404,1631,1658]" pageId="141" pageNumber="142" pagination="63 - 119" refId="ref128270" refString="Darragh, T. A. (1969) A Revision of the Family Columbariidae (Mollusca: Gastropoda). Proceedings of the Royal Society of Victoria, 83 (1), 63 - 119, 5 pls." type="book chapter" year="1969">Darragh 1969</bibRefCitation>
).
</paragraph>
<paragraph id="370E4725A98DFF682BCCC7CFFC6FF8EA" blockId="141.[151,1436,1342,2017]" pageId="141" pageNumber="142">
<emphasis id="05C59B37A98DFF682BCCC7CFFDEFF97E" bold="true" box="[199,582,1666,1693]" pageId="141" pageNumber="142">Description of Danish material.</emphasis>
Whorls angular with flattened shoulder ramp and pronounced shoulder at periphery, succeeded abapically by two weaker carinas. Suture moderately deep, wavy, located immediately adapically of abapical carina. Aperture sub-ovoid with thick callus and outer lip, extending abapically into siphonal canal of unknown length. Columella straight. Teeth or columellar folds absent.
</paragraph>
<paragraph id="370E4725A98DFF682BCCC65EFC23F896" blockId="141.[151,1436,1342,2017]" pageId="141" pageNumber="142">Teleoconch sculpture consisting of perhaps 7 or 8 low and broad transverse costae fading out towards abapical carina and crossed by three lamella-like spiral cords on shoulder and carina. Growth lines fine and densely spaced, prosocline on shoulder ramp, becoming orthocline on whorl flanks.</paragraph>
<paragraph id="370E4725A98DFF682BCCC632FD0DF85E" blockId="141.[151,1436,1342,2017]" pageId="141" pageNumber="142">
<emphasis id="05C59B37A98DFF682BCCC632FED7F87A" bold="true" box="[199,382,1919,1945]" pageId="141" pageNumber="142">Measurements.</emphasis>
No clear size can be given for the present fragmentary specimen, but the preserved part of the two whorls are
<quantity id="F049EAC0A98DFF682A4EC6EEFE32F85E" box="[325,411,1955,1981]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="141" pageNumber="142" unit="mm" value="9.1">9.1 mm</quantity>
wide and
<quantity id="F049EAC0A98DFF682905C6EEFDCDF85E" box="[526,612,1955,1981]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="141" pageNumber="142" unit="mm" value="8.1">8.1 mm</quantity>
high.
</paragraph>
<paragraph id="370E4725A98DFF6B2BCCC68AFCC8FF34" blockId="141.[151,1436,1342,2017]" lastBlockId="142.[151,1436,153,215]" lastPageId="142" lastPageNumber="143" pageId="141" pageNumber="142">
<emphasis id="05C59B37A98DFF682BCCC68AFE92F802" bold="true" box="[199,315,1991,2017]" pageId="141" pageNumber="142">Remarks.</emphasis>
The characteristic shape of the shell makes for a fairly confident assignment to this species even though the specimen is very fragmentary. It seems to only have occurred in the Danish area in the form of strays from the more shallow-water environments further to the south.
</paragraph>
</subSubSection>
</treatment>
</document>
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