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<document id="85525594C68E6479A59BF90458CDB1C2" ID-CLB-Dataset="26049" ID-DOI="10.11646/zootaxa.4654.1.1" ID-GBIF-Dataset="9ce1d103-9234-4fbe-9544-f1c294bb8783" ID-ISSN="1175-5326" ID-Zenodo-Dep="3365803" ID-ZooBank="CFD82CC0-3110-472E-972B-7ADC0C523A04" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1565592061838" checkinUser="plazi" docAuthor="Hansen, Thomas" docDate="2019" docId="BF18F633A97FFF9A2B9CC213FACDF837" docLanguage="en" docName="zootaxa.4654.1.1.pdf" docOrigin="Zootaxa 4654 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Claviscala minor Hansen 2019, n. sp." docType="treatment" docUuid="37D25BF3-B53F-486F-BE1B-7E444C04D310" docUuidSource="ZooBank" docVersion="22" lastPageNumber="128" masterDocId="43218E4BA900FFE52B0BC14DFFA9FFE3" masterDocTitle="Gastropods from the Cretaceous-Palaeogene boundary in Denmark" masterLastPageNumber="196" masterPageNumber="1" pageNumber="128" updateTime="1698750564626" updateUser="ExternalLinkService">
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<mods:title id="A230B7FE2D3A1F177C350D8AB8D9BF34">Gastropods from the Cretaceous-Palaeogene boundary in Denmark</mods:title>
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<mods:namePart id="A23B0B91986F51701672C5D3D390A41F">Hansen, Thomas</mods:namePart>
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<mods:date id="C11BB8C0F8815734EF2DC34A53327239">2019</mods:date>
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<treatment id="BF18F633A97FFF9A2B9CC213FACDF837" ID-GBIF-Taxon="159074858" LSID="urn:lsid:zoobank.org:act:37D25BF3-B53F-486F-BE1B-7E444C04D310" httpUri="http://treatment.plazi.org/id/BF18F633A97FFF9A2B9CC213FACDF837" lastPageNumber="128" pageId="127" pageNumber="128">
<subSubSection id="7FAB14AEA97FFF9A2B9CC213FE06FC9A" box="[151,431,862,889]" pageId="127" pageNumber="128" type="nomenclature">
<paragraph id="370E4725A97FFF9A2B9CC213FE06FC9A" blockId="127.[151,431,862,924]" box="[151,431,862,889]" pageId="127" pageNumber="128">
<heading id="6C46F049A97FFF9A2B9CC213FE06FC9A" bold="true" box="[151,431,862,889]" fontSize="11" level="1" pageId="127" pageNumber="128" reason="1">
<emphasis id="05C59B37A97FFF9A2B9CC213FE06FC9A" bold="true" box="[151,431,862,889]" pageId="127" pageNumber="128">
<taxonomicName id="F0B13CA6A97FFF9A2B9CC213FECCFC9B" authority="Hansen, 2019" authorityName="Hansen" authorityYear="2019" box="[151,357,862,889]" class="Gastropoda" family="Epitoniidae" genus="Claviscala" kingdom="Animalia" pageId="127" pageNumber="128" phylum="Mollusca" rank="species" species="minor" status="sp. nov.">
<emphasis id="05C59B37A97FFF9A2B9CC213FECCFC9B" bold="true" box="[151,357,862,889]" italics="true" pageId="127" pageNumber="128">Claviscala minor</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1EF6264CA97FFF9A2A66C212FE06FC9A" box="[365,431,863,889]" pageId="127" pageNumber="128" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7FAB14AEA97FFF9A2B9CC2CFFE88FC7F" box="[151,289,898,924]" pageId="127" pageNumber="128" type="description">
<paragraph id="370E4725A97FFF9A2B9CC2CFFE88FC7F" blockId="127.[151,431,862,924]" box="[151,289,898,924]" pageId="127" pageNumber="128">
<figureCitation id="AF8A5BA0A97FFF9A2B9CC2CFFF40FC7F" box="[151,233,898,924]" captionStart="FIGURE 29" captionStartId="125.[151,250,188,213]" captionTargetId="figure@124.[151,1435,179,1918]" captionTargetPageId="124" captionText="FIGURE 29. AB. Ageria skeldervigensis n. sp., Silicone cast of holotype MGUH 33226. Width 3.8, 4.5 mm respectively. CE. Ageria laxa n. sp. CD. Silicone cast of holotype MGUH 33227. Width 4.6 mm for teleoconch, 0.3 mm for protoconch. E. Silicone cast of paratype MGUH 33228. Width 6.7 mm. FG. Ageria sp. F. Silicone cast of seemingly mislaid specimen MGUH 33229. Width 3.7 mm. G. Silicone cast of MGUH 33230. Width 0.6 mm. HK. Coniscala umbilica n. sp. H. Silicone cast of paratype MGUH 33231. Width 1.4 mm. IJ. Holotype MGUH 33232. Width 19.6 mm. K. Paratype MGUH 33233. Width 12.4 mm. LN. Opalia sp. LM. Silicone cast of MGUH 33234. Width 1.3 mm. N. Silicone cast of MGUH 33235. Width 0.5 mm. OQ. Claviscala minor n. sp. OP. Silicone cast of holotype MGUH 33236. Width of teleoconch 0.9 mm. Width of protoconch 0.4 mm. Q. Silicone cast of paratype MGUH 33237. Width 1.3 mm. R. Claviscala sp., Silicone cast of MGUH 33238. Width 1.4 mm. SV. Amaea aliceae n. sp. SU. Silicone cast and inner mould of holotype MGUH 33239. Width 2.7, 2.0 and 3.1 mm respectively. V. Silicone cast of paratype MGUH 33240. Width 1.7 mm. W. Amaea stevniensis n. sp., Silicone cast of holotype MGUH 33241. Width 1.0 mm. X. Amaea sp. A. Silicone cast of MGUH 33242. Width of teleoconch 1.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3442114" httpUri="https://zenodo.org/record/3442114/files/figure.png" pageId="127" pageNumber="128">Figs 29</figureCitation>
OQ
</paragraph>
</subSubSection>
<subSubSection id="7FAB14AEA97FFF9A2B9CC284FACDF837" pageId="127" pageNumber="128" type="materials_examined">
<paragraph id="370E4725A97FFF9A2B9CC284FA9FFBEB" blockId="127.[151,1437,969,2005]" pageId="127" pageNumber="128">
<emphasis id="05C59B37A97FFF9A2B9CC284FEBBFC07" bold="true" box="[151,274,969,996]" pageId="127" pageNumber="128">Diagnosis.</emphasis>
Shell small; conical protoconch with around 35 opisthocyrt and strong collabral threads per whorl; teleoconch whorls rather weakly convex with weakly opisthocline transverse ribs and no spiral ribs or threads.
</paragraph>
<paragraph id="370E4725A97FFF9A2BCCC55CFB62FBCE" blockId="127.[151,1437,969,2005]" box="[199,1227,1041,1069]" pageId="127" pageNumber="128">
<emphasis id="05C59B37A97FFF9A2BCCC55CFE1DFBCF" bold="true" box="[199,436,1041,1068]" pageId="127" pageNumber="128">Derivation of name.</emphasis>
Refers to the small size compared to the typical species of
<taxonomicName id="F0B13CA6A97FFF9A2F47C55FFB6DFBCF" authorityName="Boury" authorityYear="1909" box="[1100,1220,1042,1068]" class="Gastropoda" family="Epitoniidae" genus="Claviscala" kingdom="Animalia" pageId="127" pageNumber="128" phylum="Mollusca" rank="genus">
<emphasis id="05C59B37A97FFF9A2F47C55FFB6DFBCF" box="[1100,1220,1042,1068]" italics="true" pageId="127" pageNumber="128">Claviscala</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="370E4725A97FFF9A2BCCC57BFDC7FB7B" blockId="127.[151,1437,969,2005]" pageId="127" pageNumber="128">
<emphasis id="05C59B37A97FFF9A2BCCC57BFEDFFBB3" bold="true" box="[199,374,1077,1104]" pageId="127" pageNumber="128">Type material.</emphasis>
The external mould
<collectionCode id="51A0DFE0A97FFF9A2968C57BFD10FBB3" box="[611,697,1078,1104]" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="127" pageNumber="128">MGUH</collectionCode>
33236 consisting of the entire protoconch and first several teleoconch whorls is selected as
<typeStatus id="E80AF987A97FFF9A2AD0C517FDE8FB97" box="[475,577,1114,1140]" pageId="127" pageNumber="128" type="holotype">holotype</typeStatus>
.
<typeStatus id="E80AF987A97FFF9A2946C516FD06FB96" box="[589,687,1115,1141]" pageId="127" pageNumber="128" type="paratype">Paratype</typeStatus>
<collectionCode id="51A0DFE0A97FFF9A29B2C517FCA6FB97" box="[697,783,1114,1140]" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="127" pageNumber="128">MGUH</collectionCode>
33237 is an external mould from the lithified top of the Højerup Member at Rødvig, Stevns Klint.
</paragraph>
<paragraph id="370E4725A97FFF9A2BCCC5ECFAF0FB03" blockId="127.[151,1437,969,2005]" pageId="127" pageNumber="128">
<emphasis id="05C59B37A97FFF9A2BCCC5ECFE17FB5F" bold="true" box="[199,446,1185,1212]" pageId="127" pageNumber="128">Additional material.</emphasis>
<collectionCode id="51A0DFE0A97FFF9A2AC2C5ECFDA0FB5F" box="[457,521,1185,1212]" country="USA" httpUri="http://grbio.org/cool/2e2p-880n" name="Ohio State University Museum" pageId="127" pageNumber="128">ØSM</collectionCode>
.10042-354, six specimens with the informal sample numbers SR.808, SR.906, SR.996.AB, SR.997.BC, SR.1048 and SR.1065.BC, and likely also SR.982, SR.1050 and SR.1051.AB.
</paragraph>
<paragraph id="370E4725A97FFF9A2BCCC5A7FDA5FACB" blockId="127.[151,1437,969,2005]" pageId="127" pageNumber="128">
<materialsCitation id="87D94D78A97FFF9A2BCCC5A7FDAEFACB" ID-GBIF-Occurrence="2331574345" county="Stevns Klint" location="Lithified" municipality="Hojerup Church" pageId="127" pageNumber="128" specimenCount="2" typeStatus="holotype">
<emphasis id="05C59B37A97FFF9A2BCCC5A7FD91FAE7" bold="true" box="[199,568,1257,1284]" pageId="127" pageNumber="128">
<typeStatus id="E80AF987A97FFF9A2BCCC5A7FEABFAE7" box="[199,258,1258,1284]" pageId="127" pageNumber="128">Type</typeStatus>
stratum and
<typeStatus id="E80AF987A97FFF9A2AAAC5A7FE7DFAE7" box="[417,468,1258,1284]" pageId="127" pageNumber="128">type</typeStatus>
locality.
</emphasis>
<location id="326E11FEA97FFF9A2935C5A7FD09FAE7" LSID="urn:lsid:plazi:treatment:BF18F633A97FFF9A2B9CC213FACDF837:326E11FEA97FFF9A2935C5A7FD09FAE7" box="[574,672,1258,1284]" county="Stevns Klint" municipality="Hojerup Church" name="Lithified" pageId="127" pageNumber="128">Lithified</location>
top of the
<location id="326E11FEA97FFF9A2813C5A7FC70FAE7" LSID="urn:lsid:plazi:treatment:BF18F633A97FFF9A2B9CC213FACDF837:326E11FEA97FFF9A2813C5A7FC70FAE7" box="[792,985,1258,1284]" county="Stevns Klint" municipality="Hojerup Church" name="Hojerup Member" pageId="127" pageNumber="128">Højerup Member</location>
of the
<location id="326E11FEA97FFF9A2F2DC5A6FB61FAE7" LSID="urn:lsid:plazi:treatment:BF18F633A97FFF9A2B9CC213FACDF837:326E11FEA97FFF9A2F2DC5A6FB61FAE7" box="[1062,1224,1258,1285]" county="Stevns Klint" municipality="Hojerup Church" name="Tor Formation" pageId="127" pageNumber="128">Tor Formation</location>
exposed just south of
<collectingMunicipality id="D76ADD5FA97FFF9A2BBDC443FEC4FACB" box="[182,365,1294,1320]" pageId="127" pageNumber="128">Højerup Church</collectingMunicipality>
,
<collectingCounty id="DE6F3FA9A97FFF9A2A72C443FDAEFACB" box="[377,519,1294,1320]" pageId="127" pageNumber="128">Stevns Klint</collectingCounty>
</materialsCitation>
.
</paragraph>
<paragraph id="370E4725A97FFF9A2BCCC47CFAF3FA93" blockId="127.[151,1437,969,2005]" pageId="127" pageNumber="128">
<emphasis id="05C59B37A97FFF9A2BCCC47CFEF3FAAF" bold="true" box="[199,346,1329,1356]" pageId="127" pageNumber="128">Occurrence.</emphasis>
This species is known exclusively from the lithified top of the upper Maastrichtian Højerup Member of the Tor Formation at Rødvig, just south of Højerup Church, and north of Kulsti Rende at Stevns Klint.
</paragraph>
<paragraph id="370E4725A97FFF9A2BCCC434FE74F9E3" blockId="127.[151,1437,969,2005]" pageId="127" pageNumber="128">
<emphasis id="05C59B37A97FFF9A2BCCC434FEF0FA77" bold="true" box="[199,345,1401,1428]" pageId="127" pageNumber="128">Description.</emphasis>
Protoconch conical multispiral with four convex whorls,
<typeStatus id="E80AF987A97FFF9A28F3C437FBF3FA77" box="[1016,1114,1402,1428]" pageId="127" pageNumber="128" type="holotype">holotype</typeStatus>
<quantity id="F049EAC0A97FFF9A2F6FC434FB62FA76" box="[1124,1227,1401,1429]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="127" pageNumber="128" unit="mm" value="0.65">0.65 mm</quantity>
high and
<quantity id="F049EAC0A97FFF9A2E48C437FA34FA76" box="[1347,1437,1402,1429]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="127" pageNumber="128" unit="mm" value="0.4">0.4 mm</quantity>
wide. First protoconch whorl seemingly smooth, the following covered by up to around 35 closely spaced opisthocyrt collabral threads per whorl. Transition to teleoconch seemingly sharp, seen as a change in sculpture and a decrease in whorl expansion.
</paragraph>
<paragraph id="370E4725A97FFF9A2BCCC747FCE4F957" blockId="127.[151,1437,969,2005]" pageId="127" pageNumber="128">Teleoconch spire slender, conical with evenly increasing whorl width; initial whorl moderately convex, succeeding ones flattened, constricted just above narrow and relatively shallow abapical suture; whorl height corresponding to approximately 80 % of width. Transition to smooth flat base abrupt, marked by subperipheral carination. Aperture not known. Whorls carrying coarse, sharp and weakly opisthocline transverse ribs, their number increasing very slowly with growth from around 14 on first teleoconch whorl.</paragraph>
<paragraph id="370E4725A97FFF9A2BCCC7F3FD84F91F" blockId="127.[151,1437,969,2005]" pageId="127" pageNumber="128">
<emphasis id="05C59B37A97FFF9A2BCCC7F3FED7F93B" bold="true" box="[199,382,1726,1752]" pageId="127" pageNumber="128">Measurements.</emphasis>
The
<typeStatus id="E80AF987A97FFF9A2ABCC7F3FDB0F93B" box="[439,537,1726,1752]" pageId="127" pageNumber="128" type="holotype">holotype</typeStatus>
<collectionCode id="51A0DFE0A97FFF9A292BC7F3FDDFF93B" box="[544,630,1726,1752]" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="127" pageNumber="128">MGUH</collectionCode>
33236 is
<quantity id="F049EAC0A97FFF9A29EFC7F3FC93F93A" box="[740,826,1726,1753]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="127" pageNumber="128" unit="mm" value="2.9">2.9 mm</quantity>
high and at least
<quantity id="F049EAC0A97FFF9A28F4C7F3FBFCF93A" box="[1023,1109,1726,1753]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="127" pageNumber="128" unit="mm" value="0.9">0.9 mm</quantity>
wide, consisting of four protoconch and five teleoconch whorls.
</paragraph>
<paragraph id="370E4725A97FFF9A2BCCC64BFACDF837" blockId="127.[151,1437,969,2005]" pageId="127" pageNumber="128">
<emphasis id="05C59B37A97FFF9A2BCCC64BFE92F8C3" bold="true" box="[199,315,1798,1824]" pageId="127" pageNumber="128">Remarks.</emphasis>
<taxonomicName id="F0B13CA6A97FFF9A2A49C64BFDAAF8C3" authority="Hansen, 2019" authorityName="Hansen" authorityYear="2019" box="[322,515,1798,1824]" class="Gastropoda" family="Epitoniidae" genus="Claviscala" kingdom="Animalia" pageId="127" pageNumber="128" phylum="Mollusca" rank="species" species="minor" status="sp. nov.">
<emphasis id="05C59B37A97FFF9A2A49C64BFDAAF8C3" box="[322,515,1798,1824]" italics="true" pageId="127" pageNumber="128">Claviscala minor</emphasis>
</taxonomicName>
<emphasis id="05C59B37A97FFF9A2902C64BFDE1F8C3" bold="true" box="[521,584,1798,1824]" pageId="127" pageNumber="128">
<taxonomicNameLabel id="1EF6264CA97FFF9A2902C64BFDE1F8C3" box="[521,584,1798,1824]" pageId="127" pageNumber="128" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is unusually small for this genus, but otherwise its features correspond quite well with the generic diagnosis. This species is easily distinguished from the Early Cretaceous
<taxonomicName id="F0B13CA6A97FFF9A2FBDC667FEDFF88B" authority="(Michelin, 1833)" baseAuthorityName="Michelin" baseAuthorityYear="1833" class="Gastropoda" family="Epitoniidae" genus="Claviscala" kingdom="Animalia" pageId="127" pageNumber="128" phylum="Mollusca" rank="species" species="clementina">
<emphasis id="05C59B37A97FFF9A2FBDC667FF1AF88B" italics="true" pageId="127" pageNumber="128">Claviscala clementina</emphasis>
(
<bibRefCitation id="53203AD4A97FFF9A2BCAC603FEC7F88B" author="Michelin, H." box="[193,366,1870,1896]" pageId="127" pageNumber="128" pagination="29" refId="ref136077" refString="Michelin, H. (1833) Classe V, Mollusques. In: Guerin-Meneville, F. - E. (Ed.), Magasin de Zoologie, Paris, Serie 1, 3, pls. 29. [pl. with description]" type="book chapter" year="1833">Michelin, 1833</bibRefCitation>
)
</taxonomicName>
from
<collectingCountry id="4FA607B5A97FFF9A2AB2C602FDACF88A" box="[441,517,1871,1897]" name="France" pageId="127" pageNumber="128">France</collectingCountry>
by the higher density of transverse ribs, which are also more transversely directed. It differs from the Late Cretaceous
<taxonomicName id="F0B13CA6A97FFF9A292FC63FFD35F86F" authorityName="Boury" authorityYear="1909" box="[548,668,1906,1932]" class="Gastropoda" family="Epitoniidae" genus="Claviscala" kingdom="Animalia" pageId="127" pageNumber="128" phylum="Mollusca" rank="genus">
<emphasis id="05C59B37A97FFF9A292FC63FFD35F86F" box="[548,668,1906,1932]" italics="true" pageId="127" pageNumber="128">Claviscala</emphasis>
</taxonomicName>
sp. of
<bibRefCitation id="53203AD4A97FFF9A29E2C63FFBADF86F" author="Squires, R. L. &amp; Saul, L. R." box="[745,1028,1906,1932]" pageId="127" pageNumber="128" pagination="50 - 63" refId="ref141206" refString="Squires, R. L. &amp; Saul, L. R. (2003 a) New Late Cretaceous (Campanian and Maastrichtian) Marine Gastropods from California. Journal of Paleontology, 77 (1), 50 - 63. https: // doi. org / 10.1666 / 0022 - 3360 (2003) 077 % 3 C 0050: NLCCAM % 3 E 2.0. CO; 2" type="journal article" year="2003">Squires and Saul (2003a)</bibRefCitation>
from California by a slightly higher whorl convexity, a sharper transition to the base of the last whorl and the opisthocline nature of the transverse ribs. It differs from the Cretaceous
<taxonomicName id="F0B13CA6A97FFF9A2AE2C6F7FCC1F837" authority="Abbass, 1963" authorityName="Abbass" authorityYear="1963" box="[489,872,1978,2004]" class="Gastropoda" family="Epitoniidae" genus="Claviscala" kingdom="Animalia" pageId="127" pageNumber="128" phylum="Mollusca" rank="species" species="darwishi">
<emphasis id="05C59B37A97FFF9A2AE2C6F7FD63F837" box="[489,714,1978,2004]" italics="true" pageId="127" pageNumber="128">Claviscala darwishi</emphasis>
<bibRefCitation id="53203AD4A97FFF9A29C4C6F7FCC1F837" author="Abbass, H. L." box="[719,872,1978,2004]" pageId="127" pageNumber="128" pagination="1 - 146" refId="ref123269" refString="Abbass, H. L. (1963) A Monograph on the Egyptian Cretaceous Gastropods. United Arab Republic, Geological Survey and Mineral Research Department, Palaeontological Series, Monograph 2, 1 - 146, pls. 1 - 12." type="journal article" year="1963">Abbass, 1963</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="4FA607B5A97FFF9A28A7C6F6FC46F836" box="[940,1007,1979,2005]" name="Egypt" pageId="127" pageNumber="128">Egypt</collectingCountry>
by the much less convex whorls.
</paragraph>
</subSubSection>
</treatment>
</document>