treatments-xml/data/B5/6A/87/B56A8789FFEFE50D0D02FCB5FC50FC41.xml
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<document id="C33607FDF28ADD373870B24A73A64389" ID-CLB-Dataset="2656" ID-DOI="10.5169/seals-787080" ID-GBIF-Dataset="42d9e06a-40e4-456d-bdac-8503804a01ce" ID-ISSN="2624-9170" ID-Zenodo-Dep="5838544" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1637933905077" checkinUser="carolina" docAuthor="Neubert, Eike &amp; Damme, Dirk van" docDate="2012" docId="B56A8789FFEFE50D0D02FCB5FC50FC41" docLanguage="en" docName="ContribNatHist.20.1-28.pdf.imf" docOrigin="Contributions to Natural History 20" docStyle="DocumentStyle:D319E9290ED630111D2A11900DAFC11F.4:ContribNatHist.2011-.journal_article" docStyleId="D319E9290ED630111D2A11900DAFC11F" docStyleName="ContribNatHist.2011-.journal_article" docStyleVersion="4" docTitle="Succinea omanensis Neubert &amp; Damme 2012, sp. nov." docType="treatment" docVersion="6" lastPageNumber="13" masterDocId="4953FFF1FFE4E5000D7BFFD8FFB4FFEB" masterDocTitle="Palaeogene continental molluscs of Oman" masterLastPageNumber="28" masterPageNumber="1" pageNumber="12" updateTime="1699470429660" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="21815D91EE01855F5A515489CB71234D">Palaeogene continental molluscs of Oman</mods:title>
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<mods:namePart id="074387168AAE86E7D34E04510DEEDE16">Neubert, Eike</mods:namePart>
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<mods:namePart id="B0EA373320970E49FB3453125D2E6DBD">Damme, Dirk van</mods:namePart>
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<mods:title id="4404BC602E5D00E92476791E99798FD0">Contributions to Natural History</mods:title>
</mods:titleInfo>
<mods:part id="3D6B0D47F51331671F92DE8859D36943">
<mods:date id="AE2DD1A73D95CD3DE1B3F8CE8F8E0433">2012</mods:date>
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<mods:number id="4E0D69BC79A8196FB33080A2214C167B">2012-12-21</mods:number>
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<mods:identifier id="C377C0CB169D14C60DB4FBE1AB4380B3" type="DOI">10.5169/seals-787080</mods:identifier>
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<treatment id="B56A8789FFEFE50D0D02FCB5FC50FC41" ID-DOI="http://doi.org/10.5281/zenodo.5838583" ID-GBIF-Taxon="191991078" ID-Zenodo-Dep="5838583" LSID="urn:lsid:plazi:treatment:B56A8789FFEFE50D0D02FCB5FC50FC41" httpUri="http://treatment.plazi.org/id/B56A8789FFEFE50D0D02FCB5FC50FC41" lastPageId="13" lastPageNumber="13" pageId="11" pageNumber="12">
<subSubSection id="75D96514FFEFE50B0D02FCB5FE0AFC6E" box="[121,446,877,902]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="3D7C369FFFEFE50B0D02FCB5FE0AFC6E" blockId="11.[121,597,877,902]" box="[121,446,877,902]" pageId="11" pageNumber="12">
<heading id="663481F3FFEFE50B0D02FCB5FE0AFC6E" box="[121,446,877,902]" centered="true" fontSize="36" level="2" pageId="11" pageNumber="12" reason="4">
<taxonomicName id="FAC34D1CFFEFE50B0D02FCB5FED5FC6D" ID-CoL="53DYN" authority="Neubert &amp; Damme, 2012" authorityName="Neubert &amp; Damme" authorityYear="2012" box="[121,353,877,902]" class="Gastropoda" family="Succineidae" genus="Succinea" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="omanensis" status="sp. nov.">
<emphasis id="0FB7EA8DFFEFE50B0D02FCB5FED5FC6D" bold="true" box="[121,353,877,902]" italics="true" pageId="11" pageNumber="12">Succinea omanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="148457F6FFEFE50B0C1DFCB5FE0AFC6E" box="[358,446,877,901]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="75D96514FFEFE50B0CB8FCB5FDE1FC6E" box="[451,597,877,901]" pageId="11" pageNumber="12" type="description">
<paragraph id="3D7C369FFFEFE50B0CB8FCB5FDE1FC6E" blockId="11.[121,597,877,902]" box="[451,597,877,901]" pageId="11" pageNumber="12">
<heading id="663481F3FFEFE50B0CB8FCB5FDE1FC6E" box="[451,597,877,901]" centered="true" fontSize="36" level="2" pageId="11" pageNumber="12" reason="4">
(
<figureCitation id="A5F82A1AFFEFE50B0CB1FCB5FDF9FC6E" box="[458,589,877,901]" captionStart="Figs" captionStartId="12.[159,198,1292,1310]" captionTargetBox="[159,1043,91,1250]" captionTargetId="figure-75@12.[159,1043,91,1250]" captionTargetPageId="12" captionText="Figs. 1112: Succinea omanensis sp. nov. Fig. 11: Wadi Darbat, holotype NMBE 5018975 in frontal and ventral view. Fig. 12: Paratype NMBE 5018976, same locality." figureDoi="http://doi.org/10.5281/zenodo.5838564" httpUri="https://zenodo.org/record/5838564/files/figure.png" pageId="11" pageNumber="12">Figs. 1112</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="75D96514FFEFE50B0D02FC60FDCCFBF1" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="3D7C369FFFEFE50B0D02FC60FD3DFC1E" blockId="11.[121,1006,952,1573]" pageId="11" pageNumber="12">
Type specimens:
<materialsCitation id="8DAB3CC2FFEFE50B0C4EFC60FDF3FC3B" ID-GBIF-Occurrence="3434796324" box="[309,583,952,976]" collectionCode="NMBE" collectionRegion="Dhofar" country="Oman" location="Wadi Darbat" pageId="11" pageNumber="12" specimenCode="NMBE 5018975" specimenCount="1" typeStatus="holotype">
<typeStatus id="E278883DFFEFE50B0C4EFC60FE2DFC3B" box="[309,409,952,976]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
<specimenCode id="6D659EE4FFEFE50B0CE5FC60FDF3FC3B" box="[414,583,952,976]" collectionCode="NMBE" country="Switzerland" httpUri="http://grbio.org/cool/5hwi-0wgz" name="Naturhistorisches Museum der Burgergemeinde Bern" pageId="11" pageNumber="12">NMBE 5018975</specimenCode>
</materialsCitation>
;
<materialsCitation id="8DAB3CC2FFEFE50B0F2EFC60FC5EFC3B" ID-GBIF-Occurrence="3434796314" box="[597,1002,952,976]" collectionCode="NMBE" collectionRegion="Dhofar" country="Oman" location="Wadi Darbat" pageId="11" pageNumber="12" specimenCode="NMBE 5018976, 5019044" specimenCount="2" typeStatus="paratype">
<typeStatus id="E278883DFFEFE50B0F2EFC60FD77FC3B" box="[597,707,952,976]" pageId="11" pageNumber="12" type="paratype">paratypes</typeStatus>
<specimenCode id="6D659EE4FFEFE50B0FB2FC60FCC1FC3B" box="[713,885,952,976]" collectionCode="NMBE" country="Switzerland" httpUri="http://grbio.org/cool/5hwi-0wgz" name="Naturhistorisches Museum der Burgergemeinde Bern" pageId="11" pageNumber="12">NMBE 5018976</specimenCode>
,
<specimenCode id="6D659EE4FFEFE50B0E05FC60FC5EFC3B" box="[894,1002,952,976]" collectionCode="NMBE" country="Switzerland" httpUri="http://grbio.org/cool/5hwi-0wgz" name="Naturhistorisches Museum der Burgergemeinde Bern" pageId="11" pageNumber="12">5019044</specimenCode>
</materialsCitation>
;
<materialsCitation id="8DAB3CC2FFEFE50B0D02FC05FD3DFC1E" ID-GBIF-Occurrence="3434796302" box="[121,649,989,1013]" collectionCode="NMBE" collectionRegion="Dhofar" country="Oman" location="Thaytiniti" pageId="11" pageNumber="12" specimenCode="NMBE 5019045-5019047" specimenCount="3" typeStatus="paratype">
<typeStatus id="E278883DFFEFE50B0D02FC05FF58FC1E" box="[121,236,989,1013]" pageId="11" pageNumber="12" type="paratype">paratypes</typeStatus>
<location id="381C6044FFEFE50B0D89FC05FEE9FC1E" LSID="urn:lsid:plazi:treatment:B56A8789FFEFE50D0D02FCB5FC50FC41:381C6044FFEFE50B0D89FC05FEE9FC1E" box="[242,349,989,1013]" name="Thaytiniti" pageId="11" pageNumber="12">Thaytiniti</location>
<specimenCode id="6D659EE4FFEFE50B0C18FC05FD36FC1E" box="[355,642,989,1013]" collectionCode="NMBE" country="Switzerland" httpUri="http://grbio.org/cool/5hwi-0wgz" name="Naturhistorisches Museum der Burgergemeinde Bern" pageId="11" pageNumber="12">NMBE 50190455019047</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="3D7C369FFFEFE50B0DE5FBDAFDCCFBF1" blockId="11.[121,1006,952,1573]" box="[158,632,1026,1050]" pageId="11" pageNumber="12">
<materialsCitation id="8DAB3CC2FFEFE50B0DE5FBDAFDC0FBF1" ID-GBIF-Occurrence="3434796315" box="[158,628,1026,1050]" country="Oman" location="Wadi Darbat" pageId="11" pageNumber="12" specimenCount="1" stateProvince="Zufar" typeStatus="holotype">
<typeStatus id="E278883DFFEFE50B0DE5FBDAFF67FBF1" box="[158,211,1026,1050]" pageId="11" pageNumber="12">Type</typeStatus>
locality:
<location id="381C6044FFEFE50B0C40FBDAFE7FFBF1" LSID="urn:lsid:plazi:treatment:B56A8789FFEFE50D0D02FCB5FC50FC41:381C6044FFEFE50B0C40FBDAFE7FFBF1" box="[315,459,1026,1050]" country="Oman" name="Wadi Darbat" pageId="11" pageNumber="12" stateProvince="Zufar">Wadi Darbat</location>
,
<collectingRegion id="FF07F87DFFEFE50B0CAFFBDAFD90FBF1" box="[468,548,1026,1050]" country="Oman" name="Zufar" pageId="11" pageNumber="12">Dhofar</collectingRegion>
,
<collectingCountry id="45D4760FFFEFE50B0F56FBDAFDC0FBF1" box="[557,628,1026,1050]" name="Oman" pageId="11" pageNumber="12">Oman</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="75D96514FFEFE50B0DE5FBF0FE26FB8E" pageId="11" pageNumber="12" type="multiple">
<paragraph id="3D7C369FFFEFE50B0DE5FBF0FC59FBAB" blockId="11.[121,1006,952,1573]" box="[158,1005,1064,1088]" pageId="11" pageNumber="12">Stratum typicum: Paludal biomicritic limestones of the Zalumah Formation.</paragraph>
<paragraph id="3D7C369FFFEFE50B0DE5FB95FE26FB8E" blockId="11.[121,1006,952,1573]" box="[158,402,1101,1125]" pageId="11" pageNumber="12">Age: Late Priabonian.</paragraph>
</subSubSection>
<subSubSection id="75D96514FFEFE50B0DE5FBAAFD3BFB61" box="[158,655,1138,1162]" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="3D7C369FFFEFE50B0DE5FBAAFD3BFB61" blockId="11.[121,1006,952,1573]" box="[158,655,1138,1162]" pageId="11" pageNumber="12">
Material: Only known from the
<typeStatus id="E278883DFFEFE50B0C87FBAAFD9BFB61" box="[508,559,1138,1162]" pageId="11" pageNumber="12">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="75D96514FFEFE50B0DE5FB40FC71FB3E" pageId="11" pageNumber="12" type="diagnosis">
<paragraph id="3D7C369FFFEFE50B0DE5FB40FC71FB3E" blockId="11.[121,1006,952,1573]" pageId="11" pageNumber="12">
Diagnosis: A small shelled member of
<taxonomicName id="FAC34D1CFFEFE50B0F39FB40FD11FB5B" authorityName="Draparnaud" authorityYear="1801" box="[578,677,1176,1200]" class="Gastropoda" family="Succineidae" genus="Succinea" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">
<emphasis id="0FB7EA8DFFEFE50B0F39FB40FD11FB5B" box="[578,677,1176,1200]" italics="true" pageId="11" pageNumber="12">Succinea</emphasis>
</taxonomicName>
, with a rapidly growing teleoconch and the last whorl exceeding half of the complete height of the shell.
</paragraph>
</subSubSection>
<subSubSection id="75D96514FFEFE50B0DE5FB3AFC2BFB11" box="[158,927,1250,1274]" pageId="11" pageNumber="12" type="etymology">
<paragraph id="3D7C369FFFEFE50B0DE5FB3AFC2BFB11" blockId="11.[121,1006,952,1573]" box="[158,927,1250,1274]" pageId="11" pageNumber="12">
Etymology:
<taxonomicName id="FAC34D1CFFEFE50B0C5DFB3AFE15FB11" authorityName="Neubert &amp; Damme" authorityYear="2012" box="[294,417,1250,1274]" class="Gastropoda" family="Succineidae" genus="Succinea" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="omanensis">
<emphasis id="0FB7EA8DFFEFE50B0C5DFB3AFE15FB11" box="[294,417,1250,1274]" italics="true" pageId="11" pageNumber="12">omanensis</emphasis>
</taxonomicName>
refers to the provenance of this new species.
</paragraph>
</subSubSection>
<subSubSection id="75D96514FFEFE50B0DE5FAD0FE34F9CE" pageId="11" pageNumber="12" type="description">
<paragraph id="3D7C369FFFEFE50B0DE5FAD0FDDFFA31" blockId="11.[121,1006,952,1573]" pageId="11" pageNumber="12">
Description: Small shells with a rapidly growing teleoconch, shells probably reach a total shell length of about
<quantity id="FA3B9B7AFFEFE50B0F41FAF5FD39FAAE" box="[570,653,1325,1349]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="12" unit="mm" value="10.0">10 mm</quantity>
; preserved
<typeStatus id="E278883DFFEFE50B0E69FAF5FCCCFAAE" box="[786,888,1325,1349]" pageId="11" pageNumber="12" type="paratype">paratype</typeStatus>
shell with almost 4 whorls, protoconch eroded; suture shallow, well-marked and somewhat reinforced, subsuturally slightly crenulate; teleoconch covered by a regular pattern of fine, axial growth riblets; last whorl amply open, constituting more than half of the complete height of the shell.
</paragraph>
<paragraph id="3D7C369FFFEFE50B0DE5FA30FE34F9CE" blockId="11.[121,1006,952,1573]" pageId="11" pageNumber="12">
Measurements:
<typeStatus id="E278883DFFEFE50B0C29FA30FE0CF9EB" box="[338,440,1512,1536]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
(
<figureCitation id="A5F82A1AFFEFE50B0CBEFA30FDBAF9EB" box="[453,526,1512,1536]" captionStart="Figs" captionStartId="12.[159,198,1292,1310]" captionTargetBox="[159,1043,91,1250]" captionTargetId="figure-75@12.[159,1043,91,1250]" captionTargetPageId="12" captionText="Figs. 1112: Succinea omanensis sp. nov. Fig. 11: Wadi Darbat, holotype NMBE 5018975 in frontal and ventral view. Fig. 12: Paratype NMBE 5018976, same locality." figureDoi="http://doi.org/10.5281/zenodo.5838564" httpUri="https://zenodo.org/record/5838564/files/figure.png" pageId="11" pageNumber="12">Fig. 11</figureCitation>
): H =
<quantity id="FA3B9B7AFFEFE50B0F28FA30FD0EF9EB" box="[595,698,1512,1536]" metricMagnitude="-3" metricUnit="m" metricValue="4.94" pageId="11" pageNumber="12" unit="mm" value="4.94">4.94 mm</quantity>
; Wh
<emphasis id="0FB7EA8DFFEFE50B0F96FA30FD49FA14" bold="true" box="[749,765,1512,1535]" italics="true" pageId="11" pageNumber="12">&lt;</emphasis>
3;
<typeStatus id="E278883DFFEFE50B0E64FA30FC37F9EB" box="[799,899,1512,1536]" pageId="11" pageNumber="12" type="paratype">paratype</typeStatus>
(
<figureCitation id="A5F82A1AFFEFE50B0EEBFA30FC6FF9EB" box="[912,987,1512,1536]" captionStart="Figs" captionStartId="12.[159,198,1292,1310]" captionTargetBox="[159,1043,91,1250]" captionTargetId="figure-75@12.[159,1043,91,1250]" captionTargetPageId="12" captionText="Figs. 1112: Succinea omanensis sp. nov. Fig. 11: Wadi Darbat, holotype NMBE 5018975 in frontal and ventral view. Fig. 12: Paratype NMBE 5018976, same locality." figureDoi="http://doi.org/10.5281/zenodo.5838564" httpUri="https://zenodo.org/record/5838564/files/figure.png" pageId="11" pageNumber="12">Fig. 12</figureCitation>
): H =
<quantity id="FA3B9B7AFFEFE50B0DD3F9D5FEB8F9CE" box="[168,268,1549,1573]" metricMagnitude="-3" metricUnit="m" metricValue="7.67" pageId="11" pageNumber="12" unit="mm" value="7.67">7.67 mm</quantity>
; Wh
<emphasis id="0FB7EA8DFFEFE50B0C3BF9D5FEE4F9CF" bold="true" box="[320,336,1549,1572]" italics="true" pageId="11" pageNumber="12">&gt;</emphasis>
3.5.
</paragraph>
</subSubSection>
<caption id="69BC6617FFE8E50C0DE4FAD4FD01FAD3" ID-DOI="http://doi.org/10.5281/zenodo.5838564" ID-Zenodo-Dep="5838564" httpUri="https://zenodo.org/record/5838564/files/figure.png" pageId="12" pageNumber="13" startId="12.[159,198,1292,1310]" targetBox="[159,1043,91,1250]" targetPageId="12">
<paragraph id="3D7C369FFFE8E50C0DE4FAD4FD01FAD3" blockId="12.[159,1044,1292,1336]" pageId="12" pageNumber="13">
Figs. 1112:
<taxonomicName id="FAC34D1CFFE8E50C0C77FAD4FE08FAF5" authority="Neubert &amp; Damme, 2012" authorityName="Neubert &amp; Damme" authorityYear="2012" box="[268,444,1292,1310]" class="Gastropoda" family="Succineidae" genus="Succinea" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="omanensis" status="sp. nov.">
<emphasis id="0FB7EA8DFFE8E50C0C77FAD4FE08FAF5" bold="true" box="[268,444,1292,1310]" italics="true" pageId="12" pageNumber="13">Succinea omanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="148457F6FFE8E50C0CBAFAD4FDB0FAF5" box="[449,516,1292,1310]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
Fig. 11: Wadi Darbat, holotype NMBE 5018975 in frontal and ventral view. Fig. 12: Paratype NMBE 5018976, same locality.
</paragraph>
</caption>
<subSubSection id="75D96514FFE8E50D0DBFFAA0FC50FC41" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="discussion">
<paragraph id="3D7C369FFFE8E50D0DBFFAA0FC50FC41" blockId="12.[159,1043,1400,1573]" lastBlockId="13.[121,1005,765,938]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
Remarks: It has to be stressed that the affiliation of this species to the extant genus
<taxonomicName id="FAC34D1CFFE8E50C0C42FA45FE2AFA5E" authorityName="Draparnaud" authorityYear="1801" box="[313,414,1437,1461]" class="Gastropoda" family="Succineidae" genus="Succinea" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="0FB7EA8DFFE8E50C0C42FA45FE2AFA5E" box="[313,414,1437,1461]" italics="true" pageId="12" pageNumber="13">Succinea</emphasis>
</taxonomicName>
is debatable. However, the shells are in fact quite similar to a number of Modern species within
<taxonomicName id="FAC34D1CFFE8E50C0F04FA1AFD56FA31" authorityName="Draparnaud" authorityYear="1801" box="[639,738,1474,1498]" class="Gastropoda" family="Succineidae" genus="Succinea" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="0FB7EA8DFFE8E50C0F04FA1AFD56FA31" box="[639,738,1474,1498]" italics="true" pageId="12" pageNumber="13">Succinea</emphasis>
</taxonomicName>
and related genera, which today are defined by using anatomical and molecular data, inapplicable to fossils. Creating a fossil genus to harbour these shells seems not to be advisable as long as material is so meagre and good arguments in favour of such a genus are available. Today, there are only two records of an extant succineid species from the Arabian Peninsula, i.e.
<taxonomicName id="FAC34D1CFFE9E50D0CD6FC90FC9CFC8B" authority="(MORELET, 1848)" baseAuthorityName="MORELET" baseAuthorityYear="1848" box="[429,808,840,864]" class="Gastropoda" family="Succineidae" genus="Quickia" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="concisa">
<emphasis id="0FB7EA8DFFE9E50D0CD6FC90FDEAFC8B" box="[429,606,840,864]" italics="true" pageId="13" pageNumber="14">Quickia concisa</emphasis>
(MORELET, 1848)
</taxonomicName>
. The shell of this species is smaller, and it has a deep and simple suture, and thus cannot be identified with the fossil specimens from Wadi Darbat (
<bibRefCitation id="59524B6EFFE9E50D0FD2FC4AFC35FC41" author="Neubert, E." box="[681,897,914,938]" pageId="13" pageNumber="14" pagination="333 - 461" refId="ref8326" refString="Neubert, E. (1998): Annotated checklist of the terrestrial and freshwater Molluscs of the Arabian Peninsula. - Fauna of Arabia, 17: 333 - 461; Basel / Riyadh." type="journal article" year="1998">Neubert 1998: 370</bibRefCitation>
, fig. 63).
</paragraph>
</subSubSection>
</treatment>
</document>