595 lines
79 KiB
XML
595 lines
79 KiB
XML
<document id="FC196F6447764D93A266114EE8791C38" ID-CLB-Dataset="36828" ID-DOI="10.11646/zootaxa.3824.1.1" ID-GBIF-Dataset="df227efc-13d9-421e-92ed-f9aa417b0cf7" ID-ISSN="1175-5326" ID-Zenodo-Dep="286404" ID-ZooBank="023D78D0-4182-48D2-BAEB-CDA6473CF585" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460443577100" checkinUser="plazi" docAuthor="Webster, Mark & Bohach, Lisa L." docDate="2014" docId="B10C8793FFE7FF9761B5F99DFBCB87C2" docLanguage="en" docName="zt03824p066.pdf" docOrigin="Zootaxa 3824 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Lochmanolenellus primus" docType="treatment" docVersion="6" lastPageNumber="32" masterDocId="4D35FFEBFFFDFF886122FFE2FFE18379" masterDocTitle="Systematic revision of the trilobite genera Laudonia and Lochmanolenellus (Olenelloidea) from the lower Dyeran (Cambrian Series 2) of western Laurentia" masterLastPageNumber="66" masterPageNumber="1" pageNumber="27" updateTime="1698333190807" updateUser="ExternalLinkService">
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<mods:mods id="937E92DF546C9C00813176B3206E85DE" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B55568872F7D273AFD97FF672FB05BA2">
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<mods:title id="ADCA6A35FBC038EFBEBDD005A6415CE7">Systematic revision of the trilobite genera Laudonia and Lochmanolenellus (Olenelloidea) from the lower Dyeran (Cambrian Series 2) of western Laurentia</mods:title>
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<mods:namePart id="BA103B0A3F272769E46A949B7246A77F">Webster, Mark</mods:namePart>
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<mods:name id="A1A812897F3C1E4596FC765EC998AE30" type="personal">
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<mods:roleTerm id="96E4009D22F78A14FB29763DA802E172">Author</mods:roleTerm>
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<mods:namePart id="424B1BAA8BB4EB92E5ED330F50552F60">Bohach, Lisa L.</mods:namePart>
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<mods:typeOfResource id="24CB5B73172EEC3EECCA1C2D277F3FE0">text</mods:typeOfResource>
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<mods:titleInfo id="E3F921A97D3CE98AB12A8D20C433121D">
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<mods:title id="30B3615338BA99B526E3274BCF7E683E">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="608EBB87D62B0765DACF49E9F90BA731">
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<mods:date id="CE204057E2E6D7990B5D343BD8AFD303">2014</mods:date>
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<mods:number id="13FD20E58206E29030B82244707F72DC">3824</mods:number>
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<mods:identifier id="2B2489FBA71FF3F86FF21EA22F24D8D3" type="DOI">10.11646/zootaxa.3824.1.1</mods:identifier>
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<treatment id="B10C8793FFE7FF9761B5F99DFBCB87C2" ID-DOI="http://doi.org/10.5281/zenodo.6129724" ID-GBIF-Taxon="127668554" ID-Zenodo-Dep="6129724" LSID="urn:lsid:plazi:treatment:B10C8793FFE7FF9761B5F99DFBCB87C2" httpUri="http://treatment.plazi.org/id/B10C8793FFE7FF9761B5F99DFBCB87C2" lastPageId="31" lastPageNumber="32" pageId="26" pageNumber="27">
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<subSubSection id="71BF650EFFE7FF9361B5F99DFCB38251" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="nomenclature">
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<paragraph id="391A3685FFE7FF9261B5F99DFC9585E3" blockId="26.[151,884,1663,1723]" box="[151,884,1663,1690]" pageId="26" pageNumber="27">
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<heading id="625281E9FFE7FF9261B5F99DFC9585E3" bold="true" box="[151,884,1663,1690]" fontSize="11" level="1" pageId="26" pageNumber="27" reason="1">
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||
<emphasis id="0BD1EA97FFE7FF9261B5F99DFC9585E3" bold="true" box="[151,884,1663,1690]" pageId="26" pageNumber="27">
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||
<taxonomicName id="FEA54D06FFE7FF9261B5F99DFE3585E0" box="[151,468,1663,1689]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="primus">
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<emphasis id="0BD1EA97FFE7FF9261B5F99DFE3585E0" bold="true" box="[151,468,1663,1689]" italics="true" pageId="26" pageNumber="27">Lochmanolenellus primus</emphasis>
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</taxonomicName>
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(Lochman in
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<bibRefCitation id="5D344B74FFE7FF9263A1F962FC8D85E3" author="Cooper" box="[643,876,1663,1690]" pageId="26" pageNumber="27" refString="Cooper, G. A., Arellano, A. R. V., Johnson, J. H., Okulitch, V. J., Stoyanow, A. & Lochman, C. (1952) Cambrian stratigraphy and paleontology near Caborca, northwestern Sonora, Mexico. Smithsonian Miscellaneous Collections, 119 (1), 1 - 184." type="journal article" year="1952">
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Cooper
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<emphasis id="0BD1EA97FFE7FF9263C4F99DFCFB85E0" bold="true" box="[742,794,1663,1689]" italics="true" pageId="26" pageNumber="27">et al</emphasis>
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., 1952
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</bibRefCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="391A3685FFE7FF9261B5F940FF3885C2" blockId="26.[151,884,1663,1723]" box="[151,217,1698,1723]" pageId="26" pageNumber="27">
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<figureCitation id="A19E2A00FFE7FF9261B5F940FF3885C2" box="[151,217,1698,1723]" captionStart="FIGURE 8" captionStartId="29.[151,250,890,912]" captionTargetBox="[164,1432,197,860]" captionTargetId="figure@29.[151,1436,181,869]" captionTargetPageId="29" captionText="FIGURE 8. Holotype cephalon of Lochmanolenellus primus (Lochman in Cooper et al., 1952); USNM 115681. 1, Dorsal view. 2, Anterior view. 3, Left lateral view. 4, Oblique anterolateral view. All x 4. From upper portion of Puerto Blanco Formation at USNM locality 801 c, Cerros de la Proveedora, Caborca area, Sonora, Mexico." httpUri="https://zenodo.org/record/286412/files/figure.png" pageId="26" pageNumber="27">Fig. 8</figureCitation>
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</paragraph>
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<paragraph id="391A3685FFE7FF9261B5F908FC348466" blockId="26.[151,1436,1770,2038]" pageId="26" pageNumber="27">
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1952
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<taxonomicName id="FEA54D06FFE7FF9261F7F909FDB88479" authority="Lochman" authorityName="Lochman" box="[213,601,1770,1792]" class="Trilobita" family="Holmiidae" genus="Wanneria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="26" pageNumber="27" phylum="Arthropoda" rank="subSpecies" species="mexicana" subSpecies="prima">
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<emphasis id="0BD1EA97FFE7FF9261F7F909FE0D8479" box="[213,492,1771,1792]" italics="true" pageId="26" pageNumber="27">Wanneria mexicana prima</emphasis>
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Lochman
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</taxonomicName>
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in Cooper
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<emphasis id="0BD1EA97FFE7FF9263F9F909FCEC8479" box="[731,781,1770,1792]" italics="true" pageId="26" pageNumber="27">et al</emphasis>
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. (part), pp. 96–98, pl. 18, fig. 3 only [not pl. 18, fig. 1, =
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<taxonomicName id="FEA54D06FFE7FF9261E5F8EBFE0C8466" box="[199,493,1801,1823]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="primus">
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<emphasis id="0BD1EA97FFE7FF9261E5F8EBFE608466" box="[199,385,1801,1823]" italics="true" pageId="26" pageNumber="27">Lochmanolenellus</emphasis>
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cf.
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<emphasis id="0BD1EA97FFE7FF926085F8E8FE0C8466" box="[423,493,1802,1823]" italics="true" pageId="26" pageNumber="27">primus</emphasis>
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</taxonomicName>
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1; not pl. 18, fig. 2, = genus and species indet.].
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</paragraph>
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<paragraph id="391A3685FFE7FF9261B5F8CAFC1A8447" blockId="26.[151,1436,1770,2038]" box="[151,1019,1832,1854]" pageId="26" pageNumber="27">
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1952
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<taxonomicName id="FEA54D06FFE7FF9261F2F8CBFDA48447" authority="Lochman" authorityName="Lochman" box="[208,581,1832,1854]" class="Trilobita" family="Holmiidae" genus="Wanneria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="26" pageNumber="27" phylum="Arthropoda" rank="subSpecies" species="mexicana" subSpecies="prima">
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<emphasis id="0BD1EA97FFE7FF9261F2F8CBFE3A8447" box="[208,475,1833,1854]" italics="true" pageId="26" pageNumber="27">Wanneria mexicana prima</emphasis>
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Lochman
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</taxonomicName>
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; Cooper & Arellano in Cooper
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<emphasis id="0BD1EA97FFE7FF9262A1F8CBFC518447" box="[899,944,1832,1854]" italics="true" pageId="26" pageNumber="27">et al</emphasis>
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., p. 19.
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</paragraph>
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<paragraph id="391A3685FFE7FF9261B5F8A4FCE48425" blockId="26.[151,1436,1770,2038]" box="[151,773,1862,1885]" pageId="26" pageNumber="27">
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1956
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<emphasis id="0BD1EA97FFE7FF9261F2F8A4FE958424" box="[208,372,1862,1885]" italics="true" pageId="26" pageNumber="27">Laudonia prima</emphasis>
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(Lochman); Harrington, pp. 56, 60, 61.
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</paragraph>
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<paragraph id="391A3685FFE7FF9261B5F887FCA98402" blockId="26.[151,1436,1770,2038]" box="[151,840,1893,1915]" pageId="26" pageNumber="27">
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1971
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<emphasis id="0BD1EA97FFE7FF9261F2F884FD918402" box="[208,624,1893,1915]" italics="true" pageId="26" pageNumber="27">
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<taxonomicName id="FEA54D06FFE7FF9261F2F884FED38402" box="[208,306,1894,1915]" class="Trilobita" family="Holmiidae" genus="Wanneria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">Wanneria</taxonomicName>
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(“Laudonia”)
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<taxonomicName id="FEA54D06FFE7FF9260EEF884FD918402" authority="Lochman" authorityName="Lochman" box="[460,624,1894,1915]" class="Trilobita" family="Holmiidae" genus="Wanneria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="26" pageNumber="27" phylum="Arthropoda" rank="subSpecies" species="mexicana" subSpecies="prima">mexicana prima</taxonomicName>
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</emphasis>
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Lochman; Hu, p. 79.
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</paragraph>
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<paragraph id="391A3685FFE7FF9261B5F866FD8F84E3" blockId="26.[151,1436,1770,2038]" box="[151,622,1924,1946]" pageId="26" pageNumber="27">
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1972
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<emphasis id="0BD1EA97FFE7FF9261F2F866FE9584E3" box="[208,372,1924,1946]" italics="true" pageId="26" pageNumber="27">Laudonia prima</emphasis>
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(Lochman); Fritz, p. 27.
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</paragraph>
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<paragraph id="391A3685FFE7FF9261B5F840FCCB84C1" blockId="26.[151,1436,1770,2038]" box="[151,810,1954,1976]" pageId="26" pageNumber="27">
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1984
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<taxonomicName id="FEA54D06FFE7FF9261F2F841FDA484C1" authority="Lochman" authorityName="Lochman" box="[208,581,1954,1976]" class="Trilobita" family="Holmiidae" genus="Wanneria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="26" pageNumber="27" phylum="Arthropoda" rank="subSpecies" species="mexicana" subSpecies="prima">
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<emphasis id="0BD1EA97FFE7FF9261F2F841FE3A84C1" box="[208,475,1955,1976]" italics="true" pageId="26" pageNumber="27">Wanneria mexicana prima</emphasis>
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Lochman
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</taxonomicName>
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; Stewart
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<emphasis id="0BD1EA97FFE7FF926380F841FD2E84C1" box="[674,719,1954,1976]" italics="true" pageId="26" pageNumber="27">et al</emphasis>
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., fig. 19.
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</paragraph>
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<paragraph id="391A3685FFE7FF9261B5F823FD0B84AE" blockId="26.[151,1436,1770,2038]" box="[151,746,1985,2007]" pageId="26" pageNumber="27">
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1992
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<emphasis id="0BD1EA97FFE7FF9261F2F823FED084AE" box="[208,305,1985,2007]" italics="true" pageId="26" pageNumber="27">Laudonia</emphasis>
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?
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<taxonomicName id="FEA54D06FFE7FF926060F820FDFC84AE" authority="Lochman" authorityName="Lochman" box="[322,541,1985,2007]" class="Trilobita" family="Holmiidae" genus="Wanneria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="mexicana">
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<emphasis id="0BD1EA97FFE7FF926060F820FE4284AE" box="[322,419,1986,2007]" italics="true" pageId="26" pageNumber="27">mexicana</emphasis>
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(Lochman)
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</taxonomicName>
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; Fritz, pp. 5, 12, 26.
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</paragraph>
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<paragraph id="391A3685FFE7FF9261B5F802FC37848F" blockId="26.[151,1436,1770,2038]" box="[151,982,2016,2038]" pageId="26" pageNumber="27">
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1998
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<taxonomicName id="FEA54D06FFE7FF9261F2F802FD8A848F" authority="Lochman" authorityName="Lochman" box="[208,619,2016,2038]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="mexicana">
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<emphasis id="0BD1EA97FFE7FF9261F2F802FE10848F" box="[208,497,2016,2038]" italics="true" pageId="26" pageNumber="27">Lochmanolenellus mexicana</emphasis>
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(Lochman)
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</taxonomicName>
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; Lieberman, pp. 61, 62, 74, fig. 4.4.
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</paragraph>
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<paragraph id="391A3685FFE6FF9361B5FF7AFCD48392" blockId="27.[151,914,152,296]" pageId="27" pageNumber="28">
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||
1999
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<taxonomicName id="FEA54D06FFE6FF9361F2FF7AFD8A83D7" authority="Lochman" authorityName="Lochman" box="[208,619,152,174]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="mexicana">
|
||
<emphasis id="0BD1EA97FFE6FF9361F2FF7AFE1083D7" box="[208,497,152,174]" italics="true" pageId="27" pageNumber="28">Lochmanolenellus mexicana</emphasis>
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(Lochman)
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</taxonomicName>
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; Lieberman, pp. 116–118. 1999
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<taxonomicName id="FEA54D06FFE6FF9361F2FF54FD8A83B5" authority="Lochman" authorityName="Lochman" box="[208,619,182,205]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="mexicana">
|
||
<emphasis id="0BD1EA97FFE6FF9361F2FF54FE1083B4" box="[208,497,182,205]" italics="true" pageId="27" pageNumber="28">Lochmanolenellus mexicana</emphasis>
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(Lochman)
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</taxonomicName>
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; Smith & Lieberman, p. 462. 2002
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<taxonomicName id="FEA54D06FFE6FF9361F2FF37FD8A8392" authority="Lochman" authorityName="Lochman" box="[208,619,213,235]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="mexicana">
|
||
<emphasis id="0BD1EA97FFE6FF9361F2FF37FE108392" box="[208,497,213,235]" italics="true" pageId="27" pageNumber="28">Lochmanolenellus mexicana</emphasis>
|
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(Lochman)
|
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</taxonomicName>
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; Lieberman, p. 699.
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</paragraph>
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<paragraph id="391A3685FFE6FF9361B5FF16FCC98273" blockId="27.[151,914,152,296]" box="[151,808,244,266]" pageId="27" pageNumber="28">
|
||
2003
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<taxonomicName id="FEA54D06FFE6FF9361F2FF16FD8A8273" authority="Lochman" authorityName="Lochman" box="[208,619,244,266]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="mexicana">
|
||
<emphasis id="0BD1EA97FFE6FF9361F2FF16FE108273" box="[208,497,244,266]" italics="true" pageId="27" pageNumber="28">Lochmanolenellus mexicana</emphasis>
|
||
(Lochman)
|
||
</taxonomicName>
|
||
; Lieberman, p. 63.
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||
</paragraph>
|
||
<paragraph id="391A3685FFE6FF9361B5FEF0FCB38251" blockId="27.[151,914,152,296]" box="[151,850,274,296]" pageId="27" pageNumber="28">
|
||
2003
|
||
<taxonomicName id="FEA54D06FFE6FF9361F2FEF0FD8A8251" authority="Lochman" authorityName="Lochman" box="[208,619,274,296]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="mexicana">
|
||
<emphasis id="0BD1EA97FFE6FF9361F2FEF0FE108251" box="[208,497,274,296]" italics="true" pageId="27" pageNumber="28">Lochmanolenellus mexicana</emphasis>
|
||
(Lochman)
|
||
</taxonomicName>
|
||
; Jell & Adrain, p. 399.
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</paragraph>
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</subSubSection>
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<subSubSection id="71BF650EFFE6FF9361B5FEB7FF5D813F" pageId="27" pageNumber="28" type="diagnosis">
|
||
<paragraph id="391A3685FFE6FF9361B5FEB7FF5D813F" blockId="27.[151,1437,341,2022]" pageId="27" pageNumber="28">
|
||
<emphasis id="0BD1EA97FFE6FF9361B5FEB7FEF08217" bold="true" box="[151,273,341,366]" pageId="27" pageNumber="28">Diagnosis.</emphasis>
|
||
Cephalon pentagonal in outline; genal spine base transversely opposite posterior portion of lateral margins of L3; distal portion of posterior cephalic margin oriented anterolaterally at slightly less than 15° relative to an exsagittal line when traced toward base of genal spine. Distance (tr.) between genal spine bases approximately 152% of cephalic length (sag.). Angle between distal portion of posterior cephalic margin and outer margin of proximal portion of intergenal spine approximately 144°. Length (sag.) of LA approximately 65% of maximum width of LA. Posterior tip of ocular lobe transversely opposite area of contact between axial furrow and S1.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="71BF650EFFE6FF9461E5FDB3FEE68674" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" type="description">
|
||
<paragraph id="391A3685FFE6FF9461E5FDB3FEE68674" blockId="27.[151,1437,341,2022]" lastBlockId="28.[151,1437,151,1941]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
|
||
<emphasis id="0BD1EA97FFE6FF9361E5FDB3FEB88113" bold="true" box="[199,345,593,618]" pageId="27" pageNumber="28">Description.</emphasis>
|
||
Cephalic length of only known specimen approximately
|
||
<quantity id="FE5D9B60FFE6FF9362C1FDB3FBDB8110" box="[995,1082,593,617]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" pageId="27" pageNumber="28" unit="mm" value="9.7">9.7 mm</quantity>
|
||
(sag.). Cephalon pentagonal in outline, widest (tr.) at base of genal spines. Proximal portion of posterior cephalic margin oriented weakly posteriorly by approximately 10° relative to a transverse line when traced abaxially, flexing anteriorly at a rounded adgenal angle located slightly less than half way along cephalic margin from axial furrow to base of genal spine, distal portion of posterior cephalic margin oriented anterolaterally at slightly less than 15° relative to an exsagittal line when traced abaxially. Genal spine stout, broad-based; angle between distal portion of posterior cephalic margin and inner margin of proximal portion of spine approximately 75°, spines gently curving into slightly more posterior orientation along their length; tips not preserved but spine length estimated to be approximately one-half cephalic length (sag.); spine base at contact with distal portion of posterior cephalic margin transversely opposite posterior portion of lateral margins of L3. Distance (tr.) between genal spine bases approximately 152% of cephalic length (sag.). Intergenal spine slender, broad-based, angle between distal portion of posterior cephalic margin and outer margin of proximal portion of spine approximately 144°; tips not preserved but spine length estimated to be between one-third and one-half cephalic length (sag.); spine base slightly distal to adgenal angle. Cephalic border defined by broad, trough-like border furrow interrupted only by intergenal ridge as it merges with border at base of intergenal spine; border furrow somewhat shallower and narrower along proximal portion of posterior cephalic border. Anterior cephalic border narrowest at sagittal axis, broadens distally, width opposite junction of ocular lobes with LA approximately 15% length (sag.) of glabella; broadest portion of border located at confluence with genal spines; posterior border tapers to point adaxially, barely touches axial furrow at posterior-most portion of LO. Border well rounded dorsally anterior to LA, somewhat more broadly dorsally arched at and between bases of genal and intergenal spines. Preglabellar field absent; LA slopes anteriorly almost vertically into trough-like cephalic border furrow that is slightly wider than anterior border at sagittal axis. Plectrum not developed. Glabella club-shaped in outline, 83% of cephalic length (sag.), moderately dorsally convex (tr.), strongly dorsally arched (sag.), summit formed by posterior portion of LA and L3. Maximum width of LA 153% basal glabellar width (tr.). Posterior margin of glabella convex posteriorly. SO deep only abaxially, abaxial end slightly anterior to adaxial end. LO slightly subtrapezoidal, narrows slightly anteriorly (transverse width across SO approximately 95% basal glabellar width), length (exsag.) approximately 11% of glabellar length (sag.), very strongly dorsally convex (tr.). S1 deepest abaxially, approximately parallel to SO. L1 subquadrate, transverse width across S1 approximately 95% basal glabellar width; length (exsag.) approximately 15% of glabellar length (sag.). S2 deepest abaxially, connected to axial furrow, strongly arcuate (convex anteriorly) on either side of sagittal axis, adaxial portion slightly posterior to distal portion. L2 broadly V-shaped, lateral margins strongly diverging anteriorly, transverse width across S2 approximately 121% basal glabellar width, distance (exsag.) between contact of S1 with axial furrow and contact of S2 with axial furrow approximately 15% of glabellar length (sag.). S3 transglabellar, oriented anterolaterally away from axis until contact with ocular lobes where it contacts anteriorly converging axial furrow, the two together thus forming a strongly caret-shaped furrow on either side of sagittal axis, deepest at apex of each caret. L3 broadly M-shaped, lateral margins diverging anteriorly until contact with inner margin of ocular lobes, then converging anteriorly until contact with S3; transverse width of glabella at point of contact between axial furrow and inner margin of ocular lobes approximately 136% of basal glabellar width, distance (exsag.) between contact of S2 with axial furrow and contact of axial furrow with inner margin of ocular lobes approximately 4% of glabellar length (sag.). LA transversely oblate in outline, anterior margin bluntly rounded and rather bull-nosed; maximum width (tr.) at contact with outer margin of ocular lobes, length (sag.) approximately 65% of maximum width and 40% of glabellar length (sag.), maximum width (tr.) approximately 153% of basal glabellar width; moderately dorsally convex (tr.), strongly dorsally convex (sag.); prominently inflated relative to extraocular area, lateral margins clearly defined from anterior portion of extraocular area by axial furrow. Shallow, weakly arcuate (convex posteriorly) preocular furrow runs inwards and anteriorly from contact of LA with outer margin of ocular lobes; not incised over axis. Very shallow transocular furrow runs anterolaterally across ocular lobes from point of contact between S3 and axial furrow to preocular furrow, isolating ocular lobes from LA. Axis of LO chipped, but preserves base of axial node or spine. Prominent ovoid lateral swellings on L1, somewhat more subdued lateral swellings on L2, weak lateral swellings on anterior portion of LO; lateral swellings on LO merge into interocular area, interrupting axial furrow. Axial furrow broad, deepest at lateral margins of L2 and L3. Ocular lobes divergent from exsagittal axis by approximately 23°, crescentic; posterior tip transversely opposite area of contact between axial furrow and S1. Deep ocular furrow runs along length of ocular lobe; anteriorly, ocular furrow curves slightly outwards to merge into a “triple junction” of furrows at point where preocular furrow meets the axial furrow at the margin of LA, thus isolating outer band of ocular lobe from LA; posteriorly, ocular furrow curves slightly inwards to meet inner margin of ocular lobe so that posterior tip of ocular lobe is located on outer band. Inner band of ocular lobe convex dorsally (tr.). All but innermost lip of outer band of ocular lobe not preserved (worn or broken off); width (tr.) and dorsal convexity of outer band unknown. Circumocular suture defines large visual surface (not preserved) that occupied much of length of outer wall of ocular lobe; portion of outer wall of ocular lobe below circumocular suture forms prominent, steep eye socle approximately one-quarter of estimated vertical height of visual surface. Interocular area slopes down from inner margin of ocular lobe to axial furrow; surface convex upward, giving arched appearance; approximately half width (tr.) of inner band of ocular lobe opposite S2. Extraocular area climbs moderately steeply out of broad border furrow then arches to produce a moderately vaulted transverse profile. In vicinity of ocular lobe, extraocular area abruptly changes slope to form a steep-walled, horizontally topped (tr. and exsag.) extraocular platform upon which ocular lobe sits; extraocular platform subcrescentic in outline when viewed from above, width (tr.) of flat-topped portion of platform greatest at midlength of ocular lobes; anteriorly, extraocular platform narrows to a thin, subdued ridge that extends beyond anterior limit of ocular lobe, descending in vertical height and running parallel to lateral margin of LA then fading into anterior border furrow; posteriorly, extraocular platform progressively narrows (tr.) and wedges out against vaulted posterolateral portion of extraocular area just posterior to posterior tip of ocular lobe. Prominent intergenal ridge extends from interocular area opposite lateral margin of L1 to posterior cephalic border at base of intergenal spine; ridge broadens distally and merges smoothly into border, interrupting border furrow. Only known specimen exfoliated, so details of prosopon (if developed) unknown. Hypostome, rostral plate, thorax, and pygidium unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="71BF650EFFE1FF9461E5FAF5FAB884EC" pageId="28" pageNumber="29" type="materials_examined">
|
||
<paragraph id="391A3685FFE1FF9461E5FAF5FE24862D" blockId="28.[151,1437,151,1941]" pageId="28" pageNumber="29">
|
||
<emphasis id="0BD1EA97FFE1FF9461E5FAF5FEDA8649" bold="true" box="[199,315,1303,1328]" pageId="28" pageNumber="29">
|
||
<typeStatus id="E61E8827FFE1FF9461E5FAF5FED78649" box="[199,310,1303,1328]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<collectionCode id="5FB4AE40FFE1FF946064FAFAFE768649" LSID="urn:lsid:biocol.org:col:34871" box="[326,407,1304,1328]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="28" pageNumber="29">USNM</collectionCode>
|
||
115681 (
|
||
<figureCitation id="A19E2A00FFE1FF94632BFAF5FDB08649" box="[521,593,1303,1328]" captionStart="FIGURE 8" captionStartId="29.[151,250,890,912]" captionTargetBox="[164,1432,197,860]" captionTargetId="figure@29.[151,1436,181,869]" captionTargetPageId="29" captionText="FIGURE 8. Holotype cephalon of Lochmanolenellus primus (Lochman in Cooper et al., 1952); USNM 115681. 1, Dorsal view. 2, Anterior view. 3, Left lateral view. 4, Oblique anterolateral view. All x 4. From upper portion of Puerto Blanco Formation at USNM locality 801 c, Cerros de la Proveedora, Caborca area, Sonora, Mexico." httpUri="https://zenodo.org/record/286412/files/figure.png" pageId="28" pageNumber="29">Fig. 8</figureCitation>
|
||
; also figured by Lochman in
|
||
<bibRefCitation id="5D344B74FFE1FF946297FAFAFB7D8649" author="Cooper" box="[949,1180,1303,1328]" pageId="28" pageNumber="29" refString="Cooper, G. A., Arellano, A. R. V., Johnson, J. H., Okulitch, V. J., Stoyanow, A. & Lochman, C. (1952) Cambrian stratigraphy and paleontology near Caborca, northwestern Sonora, Mexico. Smithsonian Miscellaneous Collections, 119 (1), 1 - 184." type="journal article" year="1952">
|
||
Cooper
|
||
<emphasis id="0BD1EA97FFE1FF946530FAFBFBA98649" box="[1042,1096,1304,1328]" italics="true" pageId="28" pageNumber="29">et al</emphasis>
|
||
., 1952
|
||
</bibRefCitation>
|
||
, pl. 18, fig. 3 and by
|
||
<bibRefCitation id="5D344B74FFE1FF9461B5FADEFEB8862D" author="Lieberman" box="[151,345,1340,1365]" pageId="28" pageNumber="29" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman, 1998</bibRefCitation>
|
||
, fig. 4.4).
|
||
</paragraph>
|
||
<paragraph id="391A3685FFE1FF9461E5FABDFC5A869C" blockId="28.[151,1437,151,1941]" pageId="28" pageNumber="29">
|
||
<emphasis id="0BD1EA97FFE1FF9461E5FABDFE628601" bold="true" box="[199,387,1375,1400]" pageId="28" pageNumber="29">Other material.</emphasis>
|
||
Lochman (in
|
||
<bibRefCitation id="5D344B74FFE1FF946307FA82FCE48601" author="Cooper" box="[549,773,1375,1400]" pageId="28" pageNumber="29" refString="Cooper, G. A., Arellano, A. R. V., Johnson, J. H., Okulitch, V. J., Stoyanow, A. & Lochman, C. (1952) Cambrian stratigraphy and paleontology near Caborca, northwestern Sonora, Mexico. Smithsonian Miscellaneous Collections, 119 (1), 1 - 184." type="journal article" year="1952">
|
||
Cooper
|
||
<emphasis id="0BD1EA97FFE1FF9463A3FA83FD538601" box="[641,690,1376,1400]" italics="true" pageId="28" pageNumber="29">et al</emphasis>
|
||
., 1952
|
||
</bibRefCitation>
|
||
, p. 97) listed
|
||
<collectionCode id="5FB4AE40FFE1FF946280FA82FC128601" LSID="urn:lsid:biocol.org:col:34871" box="[930,1011,1376,1400]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="28" pageNumber="29">USNM</collectionCode>
|
||
115682,
|
||
<collectionCode id="5FB4AE40FFE1FF94657CFA82FB4E8601" LSID="urn:lsid:biocol.org:col:34871" box="[1118,1199,1376,1400]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="28" pageNumber="29">USNM</collectionCode>
|
||
115683, and
|
||
<collectionCode id="5FB4AE40FFE1FF946469FA82FA7D8601" LSID="urn:lsid:biocol.org:col:34871" box="[1355,1436,1376,1400]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="28" pageNumber="29">USNM</collectionCode>
|
||
115684 as
|
||
<typeStatus id="E61E8827FFE1FF94602CFA67FE9886E4" box="[270,377,1413,1437]" pageId="28" pageNumber="29" type="paratype">paratypes</typeStatus>
|
||
of “
|
||
<taxonomicName id="FEA54D06FFE1FF946089FA67FD3386E5" box="[427,722,1413,1436]" class="Trilobita" family="Holmiidae" genus="Wanneria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="28" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="mexicana" subSpecies="prima">
|
||
<emphasis id="0BD1EA97FFE1FF946089FA67FD3386E5" box="[427,722,1413,1436]" italics="true" pageId="28" pageNumber="29">Wanneria mexicana prima</emphasis>
|
||
</taxonomicName>
|
||
”. All these specimens are poorly preserved in highly weathered limestone, and none can be unambiguously identified to the species level. They are herein treated as
|
||
<taxonomicName id="FEA54D06FFE1FF9461B5FA2EFE3C869D" box="[151,477,1484,1509]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="primus">
|
||
<emphasis id="0BD1EA97FFE1FF9461B5FA2EFE85869D" box="[151,356,1484,1508]" italics="true" pageId="28" pageNumber="29">Lochmanolenellus</emphasis>
|
||
cf.
|
||
<emphasis id="0BD1EA97FFE1FF9460ADFA2FFE3C869D" box="[399,477,1485,1508]" italics="true" pageId="28" pageNumber="29">primus</emphasis>
|
||
</taxonomicName>
|
||
1 and an indeterminate genus and species.
|
||
</paragraph>
|
||
<paragraph id="391A3685FFE1FF9461E5FA0DFC23847C" blockId="28.[151,1437,151,1941]" pageId="28" pageNumber="29">
|
||
Lochman (in
|
||
<bibRefCitation id="5D344B74FFE1FF946049FA12FDBB8571" author="Cooper" box="[363,602,1519,1544]" pageId="28" pageNumber="29" refString="Cooper, G. A., Arellano, A. R. V., Johnson, J. H., Okulitch, V. J., Stoyanow, A. & Lochman, C. (1952) Cambrian stratigraphy and paleontology near Caborca, northwestern Sonora, Mexico. Smithsonian Miscellaneous Collections, 119 (1), 1 - 184." type="journal article" year="1952">
|
||
Cooper
|
||
<emphasis id="0BD1EA97FFE1FF9460EEFA13FDE28571" box="[460,515,1520,1544]" italics="true" pageId="28" pageNumber="29">et al</emphasis>
|
||
., 1952
|
||
</bibRefCitation>
|
||
, p. 97) noted similarity between the Mexican material and Canadian specimens from
|
||
<collectionCode id="5FB4AE40FFE1FF946073F9F6FE438555" LSID="urn:lsid:biocol.org:col:34871" box="[337,418,1556,1580]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="28" pageNumber="29">USNM</collectionCode>
|
||
locality 61k at Mumm Peak, and suggested that the Canadian specimens were conspecific with
|
||
<taxonomicName id="FEA54D06FFE1FF9461F2F9DAFE1B8536" box="[208,506,1592,1615]" class="Trilobita" family="Holmiidae" genus="Wanneria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="28" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="mexicana" subSpecies="prima">
|
||
<emphasis id="0BD1EA97FFE1FF9461F2F9DAFE1B8536" box="[208,506,1592,1615]" italics="true" pageId="28" pageNumber="29">Wanneria mexicana prima</emphasis>
|
||
</taxonomicName>
|
||
. However, those Canadian specimens represent
|
||
<emphasis id="0BD1EA97FFE1FF94653FF9DAFB198529" box="[1053,1272,1592,1616]" italics="true" pageId="28" pageNumber="29">
|
||
Laudonia
|
||
<taxonomicName id="FEA54D06FFE1FF9465B2F9DAFB198529" box="[1168,1272,1592,1616]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="bispinata">bispinata</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="0BD1EA97FFE1FF946413F9DAFEE0850D" italics="true" pageId="28" pageNumber="29">
|
||
Laudonia
|
||
<taxonomicName id="FEA54D06FFE1FF9461B5F9BFFEE0850D" box="[151,257,1629,1652]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="amputata">amputata</taxonomicName>
|
||
</emphasis>
|
||
(
|
||
<bibRefCitation id="5D344B74FFE1FF946033F9BEFE39850C" author="Harrington" box="[273,472,1628,1653]" pageId="28" pageNumber="29" refString="Harrington, H. J. (1956) Olenellidae with advanced cephalic spines. Journal of Paleontology, 30, 56 - 61." type="journal article" year="1956">Harrington, 1956</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="5D344B74FFE1FF9460C7F9BEFD87850D" author="Fritz" box="[485,614,1628,1653]" pageId="28" pageNumber="29" refString="Fritz, W. H. (1992) Walcott's Lower Cambrian olenellid trilobites collection 61 k, Mount Robson area, Canadian Rocky Mountains. Geological Survey of Canada Bulletin, 432, 1 - 65." type="journal article" year="1992">Fritz, 1992</bibRefCitation>
|
||
). Specimens originally identified as “
|
||
<emphasis id="0BD1EA97FFE1FF946530F9BEFB9F850D" box="[1042,1150,1628,1652]" italics="true" pageId="28" pageNumber="29">Laudonia</emphasis>
|
||
” by
|
||
<bibRefCitation id="5D344B74FFE1FF946594F9BEFA7D850C" author="Nelson" box="[1206,1436,1628,1653]" pageId="28" pageNumber="29" refString="Nelson, C. A. (1976) Late Precambrian-Early Cambrian stratigraphic and faunal succession of eastern California and the Precambrian-Cambrian boundary. In: Moore, J. N. & Fritsche, A. E. (Eds.), Depositional environments of Lower Paleozoic rocks in the White-Inyo Mountains, Inyo County, California. Pacific Coast Paleogeography Field Guide, 1, pp. 31 - 42." type="journal article" year="1976">Nelson (1976, pl. 5)</bibRefCitation>
|
||
from the upper part of the Poleta Formation in the White-Inyo Mountains and Death Valley regions of Inyo County, California, were assigned to
|
||
<taxonomicName id="FEA54D06FFE1FF9460FEF946FD1E85C5" box="[476,767,1700,1724]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="primus">
|
||
<emphasis id="0BD1EA97FFE1FF9460FEF946FD1E85C5" box="[476,767,1700,1724]" italics="true" pageId="28" pageNumber="29">Lochmanolenellus primus</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="5D344B74FFE1FF946208F946FC0E85C5" author="Lieberman" box="[810,1007,1700,1725]" pageId="28" pageNumber="29" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman (1998</bibRefCitation>
|
||
, p. 74; 1999, pp. 116, 118). However, these specimens are herein reassigned to
|
||
<taxonomicName id="FEA54D06FFE1FF9463A4F92BFCA88599" box="[646,841,1736,1760]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="trapezoidalis">
|
||
<emphasis id="0BD1EA97FFE1FF9463A4F92BFCA88599" box="[646,841,1736,1760]" italics="true" pageId="28" pageNumber="29">Lo. trapezoidalis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="FEA54D06FFE1FF9462AEF92BFB618599" box="[908,1152,1735,1760]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="subquadratus">
|
||
<emphasis id="0BD1EA97FFE1FF9462AEF92BFC508599" box="[908,945,1737,1760]" italics="true" pageId="28" pageNumber="29">Lo.</emphasis>
|
||
cf.
|
||
<emphasis id="0BD1EA97FFE1FF9462C5F92AFB618599" box="[999,1152,1736,1760]" italics="true" pageId="28" pageNumber="29">subquadratus</emphasis>
|
||
</taxonomicName>
|
||
1.
|
||
<collectingCountry id="41B27615FFE1FF94658CF92AFB2C8599" box="[1198,1229,1736,1760]" name="American Samoa" pageId="28" pageNumber="29">As</collectingCountry>
|
||
a result of these reassignments,
|
||
<taxonomicName id="FEA54D06FFE1FF946061F90FFE5C847D" box="[323,445,1773,1796]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="primus">
|
||
<emphasis id="0BD1EA97FFE1FF946061F90FFE5C847D" box="[323,445,1773,1796]" italics="true" pageId="28" pageNumber="29">Lo. primus</emphasis>
|
||
</taxonomicName>
|
||
is definitively known only from the
|
||
<typeStatus id="E61E8827FFE1FF94627AF90EFC5F847C" box="[856,958,1772,1797]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="391A3685FFE1FF9461E5F8EDFAB884EC" blockId="28.[151,1437,151,1941]" pageId="28" pageNumber="29">
|
||
<emphasis id="0BD1EA97FFE1FF9461E5F8EDFEB88451" bold="true" box="[199,345,1807,1832]" pageId="28" pageNumber="29">Occurrence.</emphasis>
|
||
<collectingCountry id="41B27615FFE1FF946046F8F2FE368451" box="[356,471,1808,1832]" name="Mexico" pageId="28" pageNumber="29">MEXICO</collectingCountry>
|
||
:
|
||
<emphasis id="0BD1EA97FFE1FF9460C4F8F2FD62845E" box="[486,643,1808,1831]" italics="true" pageId="28" pageNumber="29">Sonora State:</emphasis>
|
||
<collectionCode id="5FB4AE40FFE1FF9463ACF8F2FD3E8451" LSID="urn:lsid:biocol.org:col:34871" box="[654,735,1808,1832]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="28" pageNumber="29">USNM</collectionCode>
|
||
locality 801c, “
|
||
<taxonomicName id="FEA54D06FFE1FF9462BDF8F2FBB18451" authority="Beds" authorityName="Beds" box="[927,1104,1807,1832]" class="Trilobita" family="Holmiidae" genus="Wanneria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="0BD1EA97FFE1FF9462BDF8F2FBEA845E" box="[927,1035,1808,1831]" italics="true" pageId="28" pageNumber="29">Wanneria</emphasis>
|
||
Beds
|
||
</taxonomicName>
|
||
” of upper portion of Puerto Blanco Formation, 180 metres (
|
||
<quantity id="FE5D9B60FFE1FF946321F8D6FD828434" box="[515,611,1844,1869]" metricMagnitude="2" metricUnit="m" metricValue="1.79832" pageId="28" pageNumber="29" unit="ft" value="590.0">590 feet</quantity>
|
||
) above base of measured section and 113 metres (
|
||
<quantity id="FE5D9B60FFE1FF946585F8D6FAE98434" box="[1191,1288,1844,1869]" metricMagnitude="2" metricUnit="m" metricValue="1.130808" pageId="28" pageNumber="29" unit="ft" value="371.0">371 feet</quantity>
|
||
) below local base of Proveedora Quartzite, western flank of Cerros de la Proveedora, 11 kilometres (six to seven miles) west of Caborca (Cooper & Arellano in
|
||
<bibRefCitation id="5D344B74FFE1FF946322F89EFD3B84EC" author="Cooper" box="[512,730,1916,1941]" pageId="28" pageNumber="29" refString="Cooper, G. A., Arellano, A. R. V., Johnson, J. H., Okulitch, V. J., Stoyanow, A. & Lochman, C. (1952) Cambrian stratigraphy and paleontology near Caborca, northwestern Sonora, Mexico. Smithsonian Miscellaneous Collections, 119 (1), 1 - 184." type="journal article" year="1952">
|
||
Cooper
|
||
<emphasis id="0BD1EA97FFE1FF946378F89FFD6A84ED" box="[602,651,1916,1940]" italics="true" pageId="28" pageNumber="29">et al</emphasis>
|
||
., 1952
|
||
</bibRefCitation>
|
||
; Lochman in
|
||
<bibRefCitation id="5D344B74FFE1FF946256F89EFBAE84EC" author="Cooper" box="[884,1103,1916,1941]" pageId="28" pageNumber="29" refString="Cooper, G. A., Arellano, A. R. V., Johnson, J. H., Okulitch, V. J., Stoyanow, A. & Lochman, C. (1952) Cambrian stratigraphy and paleontology near Caborca, northwestern Sonora, Mexico. Smithsonian Miscellaneous Collections, 119 (1), 1 - 184." type="journal article" year="1952">
|
||
Cooper
|
||
<emphasis id="0BD1EA97FFE1FF9462EFF89FFC1E84ED" box="[973,1023,1916,1940]" italics="true" pageId="28" pageNumber="29">et al</emphasis>
|
||
., 1952
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="5D344B74FFE1FF946578F89EFAAC84EC" author="Lochman-Balk" box="[1114,1357,1916,1941]" pageId="28" pageNumber="29" refString="Lochman-Balk, C. (1956) The Cambrian of the Rocky Mountains and southwest deserts of the United States and adjoining Sonora Province, Mexico. XX Congreso Geologico Internacional XX Sesion, Mexico. El Sistema Cambrico, su Paleogeografia y el problema de su base. Part II: Australia, America, pp. 529 - 662." type="book chapter" year="1956">Lochman-Balk, 1956</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="6DDA660DFFE0FF9561B5FC98FBDB80B4" httpUri="https://zenodo.org/record/286412/files/figure.png" pageId="29" pageNumber="30" targetBox="[164,1432,197,860]" targetPageId="29">
|
||
<paragraph id="391A3685FFE0FF9561B5FC98FBDB80B4" blockId="29.[151,1436,890,973]" pageId="29" pageNumber="30">
|
||
<emphasis id="0BD1EA97FFE0FF9561B5FC98FEF480F6" bold="true" box="[151,277,890,912]" pageId="29" pageNumber="30">FIGURE 8.</emphasis>
|
||
Holotype cephalon of
|
||
<taxonomicName id="FEA54D06FFE0FF956328FC98FCF380E9" box="[522,786,890,912]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="primus">
|
||
<emphasis id="0BD1EA97FFE0FF956328FC98FCF380E9" box="[522,786,890,912]" italics="true" pageId="29" pageNumber="30">Lochmanolenellus primus</emphasis>
|
||
</taxonomicName>
|
||
(Lochman in Cooper
|
||
<emphasis id="0BD1EA97FFE0FF9562DCFC99FBCF80E9" box="[1022,1070,890,912]" italics="true" pageId="29" pageNumber="30">et al</emphasis>
|
||
., 1952); USNM 115681. 1, Dorsal view. 2, Anterior view. 3, Left lateral view. 4, Oblique anterolateral view. All x4. From upper portion of Puerto Blanco Formation at USNM locality 801c, Cerros de la Proveedora, Caborca area, Sonora, Mexico.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="71BF650EFFE0FF9561E5FC16FA8C851C" pageId="29" pageNumber="30" type="discussion">
|
||
<paragraph id="391A3685FFE0FF9561E5FC16FA8C851C" blockId="29.[151,1436,1012,2031]" pageId="29" pageNumber="30">
|
||
<emphasis id="0BD1EA97FFE0FF9561E5FC16FEAD8774" bold="true" box="[199,332,1012,1037]" pageId="29" pageNumber="30">Discussion.</emphasis>
|
||
<collectingCountry id="41B27615FFE0FF956077FC17FE958774" box="[341,372,1013,1037]" name="American Samoa" pageId="29" pageNumber="30">As</collectingCountry>
|
||
can be seen from the synonymy list (see previous), there is some inconsistency in previous literature regarding the specific name of this taxon—it has variably been referred to as
|
||
<taxonomicName id="FEA54D06FFE0FF956544FBF5FAFD8757" box="[1126,1308,1047,1070]" class="Trilobita" family="Holmiidae" genus="Wanneria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="subSpecies" species="mexicana" subSpecies="prima">
|
||
<emphasis id="0BD1EA97FFE0FF956544FBF5FAFD8757" box="[1126,1308,1047,1070]" italics="true" pageId="29" pageNumber="30">mexicana prima</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="FEA54D06FFE0FF956408FBF5FA748757" box="[1322,1429,1047,1070]" class="Trilobita" family="Holmiidae" genus="Wanneria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="mexicana">
|
||
<emphasis id="0BD1EA97FFE0FF956408FBF5FA748757" box="[1322,1429,1047,1070]" italics="true" pageId="29" pageNumber="30">mexicana</emphasis>
|
||
</taxonomicName>
|
||
, or
|
||
<emphasis id="0BD1EA97FFE0FF956197FBDAFF198736" box="[181,248,1080,1103]" italics="true" pageId="29" pageNumber="30">prima</emphasis>
|
||
. In her initial description of
|
||
<taxonomicName id="FEA54D06FFE0FF95631EFBDAFC728729" authority="Lochman" authorityName="Lochman" box="[572,915,1079,1104]" class="Trilobita" family="Holmiidae" genus="Wanneria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="mexicana">
|
||
<emphasis id="0BD1EA97FFE0FF95631EFBDAFCFB8736" box="[572,794,1080,1103]" italics="true" pageId="29" pageNumber="30">Wanneria mexicana</emphasis>
|
||
, Lochman
|
||
</taxonomicName>
|
||
(in
|
||
<bibRefCitation id="5D344B74FFE0FF9562E2FBDAFB7D8729" author="Cooper" box="[960,1180,1079,1104]" pageId="29" pageNumber="30" refString="Cooper, G. A., Arellano, A. R. V., Johnson, J. H., Okulitch, V. J., Stoyanow, A. & Lochman, C. (1952) Cambrian stratigraphy and paleontology near Caborca, northwestern Sonora, Mexico. Smithsonian Miscellaneous Collections, 119 (1), 1 - 184." type="journal article" year="1952">
|
||
Cooper
|
||
<emphasis id="0BD1EA97FFE0FF956538FBDAFBAA8736" box="[1050,1099,1079,1103]" italics="true" pageId="29" pageNumber="30">et al</emphasis>
|
||
., 1952
|
||
</bibRefCitation>
|
||
, p. 96) noted that “[i]n this species it is presumed that the long, medium-sized genal spines would lie at the posterolateral angles, and the intergenal spines would be small and in a short distance on the posterior margin”. She then proceeded to describe
|
||
<taxonomicName id="FEA54D06FFE0FF9561B5FB7EFE8F87CA" box="[151,366,1180,1203]" class="Trilobita" family="Holmiidae" genus="Wanneria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="subSpecies" species="mexicana" subSpecies="prima">
|
||
<emphasis id="0BD1EA97FFE0FF9561B5FB7EFE8F87CA" box="[151,366,1180,1203]" italics="true" pageId="29" pageNumber="30">W. mexicana prima</emphasis>
|
||
</taxonomicName>
|
||
as a variety (subspecies) of that species. She noted that “(t)he cephalon of the variety is like that of the species in all proportions and details except for the position and size of the genal and intergenal spines which are placed as follows: At the posterolateral angle at the end of the strong posterior facial suture ridge lie mediumsized intergenal spines, projecting somewhat laterally; the long, heavy genal spines lie on a line with the first glabellar furrow, and the spine projects nearly straight outward at the side of the cephalon, with a slight upward flexure at the base” (Lochman in
|
||
<bibRefCitation id="5D344B74FFE0FF95632DFAA1FD0A8622" author="Cooper" box="[527,747,1346,1371]" pageId="29" pageNumber="30" refString="Cooper, G. A., Arellano, A. R. V., Johnson, J. H., Okulitch, V. J., Stoyanow, A. & Lochman, C. (1952) Cambrian stratigraphy and paleontology near Caborca, northwestern Sonora, Mexico. Smithsonian Miscellaneous Collections, 119 (1), 1 - 184." type="journal article" year="1952">
|
||
Cooper
|
||
<emphasis id="0BD1EA97FFE0FF95634BFAA1FD7A8623" box="[617,667,1346,1370]" italics="true" pageId="29" pageNumber="30">et al</emphasis>
|
||
., 1952
|
||
</bibRefCitation>
|
||
, pp. 96–97). In a subsequent paragraph, Lochman (in
|
||
<bibRefCitation id="5D344B74FFE0FF956468FAA1FEF68605" author="Cooper" pageId="29" pageNumber="30" refString="Cooper, G. A., Arellano, A. R. V., Johnson, J. H., Okulitch, V. J., Stoyanow, A. & Lochman, C. (1952) Cambrian stratigraphy and paleontology near Caborca, northwestern Sonora, Mexico. Smithsonian Miscellaneous Collections, 119 (1), 1 - 184." type="journal article" year="1952">
|
||
Cooper
|
||
<emphasis id="0BD1EA97FFE0FF9561B5FA86FF268605" box="[151,199,1380,1404]" italics="true" pageId="29" pageNumber="30">et al</emphasis>
|
||
., 1952
|
||
</bibRefCitation>
|
||
, p. 97) reiterated this distinction—she noted that several deeply weathered cephala co-occurred with
|
||
<taxonomicName id="FEA54D06FFE0FF9564A2FA86FEB186E4" class="Trilobita" family="Holmiidae" genus="Wanneria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="subSpecies" species="mexicana" subSpecies="prima">
|
||
<emphasis id="0BD1EA97FFE0FF9564A2FA86FEB186E4" italics="true" pageId="29" pageNumber="30">W. mexicana prima</emphasis>
|
||
</taxonomicName>
|
||
at Locality 801c and that, although the glabella and ocular lobes of these specimens were too poorly preserved for definitive generic identification, their genal spines were located at the “posterolateral angle of the cephalon” and that they “may represent the normal form of the species,
|
||
<taxonomicName id="FEA54D06FFE0FF956532FA2AFB1386A6" box="[1040,1266,1480,1503]" class="Trilobita" family="Holmiidae" genus="Wanneria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="mexicana">
|
||
<emphasis id="0BD1EA97FFE0FF956532FA2AFB1386A6" box="[1040,1266,1480,1503]" italics="true" pageId="29" pageNumber="30">Wanneria mexicana</emphasis>
|
||
</taxonomicName>
|
||
” (Lochman in
|
||
<bibRefCitation id="5D344B74FFE0FF9561B5FA0BFE90857B" author="Cooper" box="[151,369,1513,1538]" pageId="29" pageNumber="30" refString="Cooper, G. A., Arellano, A. R. V., Johnson, J. H., Okulitch, V. J., Stoyanow, A. & Lochman, C. (1952) Cambrian stratigraphy and paleontology near Caborca, northwestern Sonora, Mexico. Smithsonian Miscellaneous Collections, 119 (1), 1 - 184." type="journal article" year="1952">
|
||
Cooper
|
||
<emphasis id="0BD1EA97FFE0FF9561D2FA08FEC38578" box="[240,290,1513,1537]" italics="true" pageId="29" pageNumber="30">et al</emphasis>
|
||
., 1952
|
||
</bibRefCitation>
|
||
, p. 97). No specimen of this “normal form” was figured, and
|
||
<taxonomicName id="FEA54D06FFE0FF956503FA08FB4E8578" box="[1057,1199,1514,1537]" class="Trilobita" family="Holmiidae" genus="Wanneria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="mexicana">
|
||
<emphasis id="0BD1EA97FFE0FF956503FA08FB4E8578" box="[1057,1199,1514,1537]" italics="true" pageId="29" pageNumber="30">W. mexicana</emphasis>
|
||
</taxonomicName>
|
||
is therefore a nomen nudum. However, it is clear that Lochman (in
|
||
<bibRefCitation id="5D344B74FFE0FF9563BDF9E9FC98855A" author="Cooper" box="[671,889,1546,1571]" pageId="29" pageNumber="30" refString="Cooper, G. A., Arellano, A. R. V., Johnson, J. H., Okulitch, V. J., Stoyanow, A. & Lochman, C. (1952) Cambrian stratigraphy and paleontology near Caborca, northwestern Sonora, Mexico. Smithsonian Miscellaneous Collections, 119 (1), 1 - 184." type="journal article" year="1952">
|
||
Cooper
|
||
<emphasis id="0BD1EA97FFE0FF9563DBF9E9FCCA855B" box="[761,811,1546,1570]" italics="true" pageId="29" pageNumber="30">et al</emphasis>
|
||
., 1952
|
||
</bibRefCitation>
|
||
) intended the (sub)specific name
|
||
<emphasis id="0BD1EA97FFE0FF9565D4F9E9FAD8855B" box="[1270,1337,1547,1570]" italics="true" pageId="29" pageNumber="30">prima</emphasis>
|
||
to apply to the morphotype with the advanced genal spines. The
|
||
<typeStatus id="E61E8827FFE0FF956224F9CEFCD7853D" box="[774,822,1580,1604]" pageId="29" pageNumber="30">type</typeStatus>
|
||
species of
|
||
<taxonomicName id="FEA54D06FFE0FF956291F9CEFB60853D" box="[947,1153,1580,1604]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="0BD1EA97FFE0FF956291F9CEFB60853D" box="[947,1153,1580,1604]" italics="true" pageId="29" pageNumber="30">Lochmanolenellus</emphasis>
|
||
</taxonomicName>
|
||
is therefore here referred to as
|
||
<taxonomicName id="FEA54D06FFE0FF9561F0F9ACFEAA851C" box="[210,331,1614,1637]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="primus">
|
||
<emphasis id="0BD1EA97FFE0FF9561F0F9ACFEAA851C" box="[210,331,1614,1637]" italics="true" pageId="29" pageNumber="30">Lo. primus</emphasis>
|
||
</taxonomicName>
|
||
(Lochman in
|
||
<bibRefCitation id="5D344B74FFE0FF9560CBF9AFFD23851C" author="Cooper" box="[489,706,1612,1637]" pageId="29" pageNumber="30" refString="Cooper, G. A., Arellano, A. R. V., Johnson, J. H., Okulitch, V. J., Stoyanow, A. & Lochman, C. (1952) Cambrian stratigraphy and paleontology near Caborca, northwestern Sonora, Mexico. Smithsonian Miscellaneous Collections, 119 (1), 1 - 184." type="journal article" year="1952">
|
||
Cooper
|
||
<emphasis id="0BD1EA97FFE0FF956361F9ACFD95851C" box="[579,628,1613,1637]" italics="true" pageId="29" pageNumber="30">et al</emphasis>
|
||
., 1952
|
||
</bibRefCitation>
|
||
) rather than
|
||
<taxonomicName id="FEA54D06FFE0FF956272F9ACFC09851C" box="[848,1000,1614,1637]" class="Trilobita" family="Holmiidae" genus="Wanneria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="mexicana">
|
||
<emphasis id="0BD1EA97FFE0FF956272F9ACFC09851C" box="[848,1000,1614,1637]" italics="true" pageId="29" pageNumber="30">Lo. mexicana</emphasis>
|
||
</taxonomicName>
|
||
(Lochman in
|
||
<bibRefCitation id="5D344B74FFE0FF9565A7F9AFFA81851C" author="Cooper" box="[1157,1376,1612,1637]" pageId="29" pageNumber="30" refString="Cooper, G. A., Arellano, A. R. V., Johnson, J. H., Okulitch, V. J., Stoyanow, A. & Lochman, C. (1952) Cambrian stratigraphy and paleontology near Caborca, northwestern Sonora, Mexico. Smithsonian Miscellaneous Collections, 119 (1), 1 - 184." type="journal article" year="1952">
|
||
Cooper
|
||
<emphasis id="0BD1EA97FFE0FF9565FCF9ACFAF1851C" box="[1246,1296,1613,1637]" italics="true" pageId="29" pageNumber="30">et al</emphasis>
|
||
., 1952
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="71BF650EFFE0FF9761E5F98CFBCB87C2" lastPageId="31" lastPageNumber="32" pageId="29" pageNumber="30" type="reference_group">
|
||
<paragraph id="391A3685FFE0FF9761E5F98CFD2A8119" blockId="29.[151,1436,1012,2031]" lastBlockId="31.[151,1436,151,1211]" lastPageId="31" lastPageNumber="32" pageId="29" pageNumber="30">
|
||
<taxonomicName id="FEA54D06FFE0FF9561E5F98CFE0A85FF" box="[199,491,1646,1670]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="primus">
|
||
<emphasis id="0BD1EA97FFE0FF9561E5F98CFE0A85FF" box="[199,491,1646,1670]" italics="true" pageId="29" pageNumber="30">Lochmanolenellus primus</emphasis>
|
||
</taxonomicName>
|
||
was included in cladistic analyses of the Olenelloidea by
|
||
<bibRefCitation id="5D344B74FFE0FF95655DF98CFAAA85FE" author="Lieberman" box="[1151,1355,1646,1671]" pageId="29" pageNumber="30" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman (1998)</bibRefCitation>
|
||
and of the
|
||
<taxonomicName id="FEA54D06FFE0FF9561E2F970FEA285D2" box="[192,323,1682,1707]" class="Trilobita" family="Biceratopsidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="subFamily" subFamily="Bristoliinae">Bristoliinae</taxonomicName>
|
||
by Lieberman (1999). The present study reveals several coding inaccuracies for this species in both analyses. A plectrum is not present (
|
||
<bibRefCitation id="5D344B74FFE0FF95636AF954FCF185B6" author="Lieberman" box="[584,784,1718,1743]" pageId="29" pageNumber="30" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman, 1998</bibRefCitation>
|
||
, character 4, state 1; this had originally been coded by
|
||
<bibRefCitation id="5D344B74FFE0FF9561B5F938FE84858A" author="Lieberman" box="[151,357,1754,1779]" pageId="29" pageNumber="30" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman [1998]</bibRefCitation>
|
||
as present [state 0]). LA is clearly separated from the anterior extraocular area by a furrow (
|
||
<bibRefCitation id="5D344B74FFE0FF9561BCF91CFDB3846E" author="Lieberman" box="[158,594,1790,1815]" pageId="29" pageNumber="30" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman, 1998, character 6, state 0</bibRefCitation>
|
||
) as was stated by
|
||
<bibRefCitation id="5D344B74FFE0FF95620CF91CFC1D846E" author="Lieberman" box="[814,1020,1790,1815]" pageId="29" pageNumber="30" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman (1998)</bibRefCitation>
|
||
in his discussion for this character, although in his character-taxon matrix this is coded as not prominently separated (state 1). The sagittal length of LA (
|
||
<bibRefCitation id="5D344B74FFE0FF9561EFF8A4FDC18426" author="Lieberman" box="[205,544,1862,1887]" pageId="29" pageNumber="30" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman, 1998, character 9</bibRefCitation>
|
||
) is approximately 135% of the sagittal length of LO and L1, falling between states 0 (equal to the length of LO and L1) and 1 (150% of the length of LO and L1) as defined by
|
||
<bibRefCitation id="5D344B74FFE0FF956401F888FD6484DE" author="Lieberman" pageId="29" pageNumber="30" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman (1998; this was originally coded as state 0)</bibRefCitation>
|
||
. A prominent shelf (the eye socle) separates the ocular lobe from the extraocular area (
|
||
<bibRefCitation id="5D344B74FFE0FF956042F850FDA58496" author="Lieberman" pageId="29" pageNumber="30" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman, 1998, character 16, state 0; this had originally been coded as state 1 [ocular lobe merges smoothly into extraocular area]</bibRefCitation>
|
||
). “Lateral lobes” (= lateral swellings herein) are developed on LO (
|
||
<bibRefCitation id="5D344B74FFE0FF97643FF834FEE383C9" author="Lieberman" lastPageId="31" lastPageNumber="32" pageId="29" pageNumber="30" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman, 1998</bibRefCitation>
|
||
, character 42, state 1; these had been coded as absent [state 0] by
|
||
<bibRefCitation id="5D344B74FFE2FF976513FF75FB1F83C9" author="Lieberman" box="[1073,1278,151,176]" pageId="31" pageNumber="32" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman [1998]</bibRefCitation>
|
||
)—the lateral swellings on LO are subtle in comparison to those on L2 and especially on L1, but are comparable in development to those present on LO of
|
||
<taxonomicName id="FEA54D06FFE2FF9760EFFF02FCA78381" authority="Fritz, 1972" authorityName="Fritz" authorityYear="1972" box="[461,838,223,248]" class="Dothideomycetes" family="Patellariaceae" genus="Holmiella" kingdom="Fungi" order="Patellariales" pageId="31" pageNumber="32" phylum="Ascomycota" rank="species" species="preancora">
|
||
<emphasis id="0BD1EA97FFE2FF9760EFFF02FD5D8381" box="[461,700,224,248]" italics="true" pageId="31" pageNumber="32">Holmiella preancora</emphasis>
|
||
<bibRefCitation id="5D344B74FFE2FF9763E7FF3DFCA78381" author="Fritz" box="[709,838,223,248]" pageId="31" pageNumber="32" refString="Fritz, W. H. (1972) Lower Cambrian trilobites from the Sekwi Formation type section, Mackenzie Mountains, northwest Canada. Geological Survey of Canada Bulletin, 212, 1 - 90." type="journal article" year="1972">Fritz, 1972</bibRefCitation>
|
||
</taxonomicName>
|
||
, for example. An anterior ocular line is not present (
|
||
<bibRefCitation id="5D344B74FFE2FF9761BCFEE6FE848265" author="Lieberman" box="[158,357,260,285]" pageId="31" pageNumber="32" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman, 1998</bibRefCitation>
|
||
, character 46, state 1; this had been coded as visible [state 0] by
|
||
<bibRefCitation id="5D344B74FFE2FF976579FEE6FACA8265" author="Lieberman" box="[1115,1323,260,285]" pageId="31" pageNumber="32" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman [1998]</bibRefCitation>
|
||
). A genal ridge is not present (
|
||
<bibRefCitation id="5D344B74FFE2FF976058FEC5FDDF8239" author="Lieberman" box="[378,574,295,320]" pageId="31" pageNumber="32" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman, 1998</bibRefCitation>
|
||
, character 47, state 1; this had been coded as prominently developed [state 0] by
|
||
<bibRefCitation id="5D344B74FFE2FF976198FEAEFE69821D" author="Lieberman" box="[186,392,332,357]" pageId="31" pageNumber="32" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman [1998]</bibRefCitation>
|
||
). The posterior tips of the ocular lobes (
|
||
<bibRefCitation id="5D344B74FFE2FF97626AFEAEFB43821D" author="Lieberman" box="[840,1186,332,357]" pageId="31" pageNumber="32" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman, 1998, character 23</bibRefCitation>
|
||
) were correctly coded as opposite S1 (state 5) by
|
||
<bibRefCitation id="5D344B74FFE2FF9760E9FE8DFD7882F1" author="Lieberman" box="[459,665,367,392]" pageId="31" pageNumber="32" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman (1998)</bibRefCitation>
|
||
, but incorrectly as opposite L1 (equivalent to state 2) by Lieberman (1999,
|
||
<taxonomicName id="FEA54D06FFE2FF9761CAFE76FE8A82D4" box="[232,363,404,429]" class="Trilobita" family="Biceratopsidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="subFamily" subFamily="Bristoliinae">Bristoliinae</taxonomicName>
|
||
analysis, character 3)—the 1999
|
||
<taxonomicName id="FEA54D06FFE2FF9763C9FE76FC8F82D4" box="[747,878,404,429]" class="Trilobita" family="Biceratopsidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="subFamily" subFamily="Bristoliinae">Bristoliinae</taxonomicName>
|
||
analysis included no equivalent state to 23(5) of the 1998 analysis. The genal spine angle (
|
||
<bibRefCitation id="5D344B74FFE2FF97634BFE55FC2082A9" author="Lieberman" box="[617,961,439,464]" pageId="31" pageNumber="32" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman, 1998, character 52</bibRefCitation>
|
||
) was correctly coded as opposite L3 (state 3) by
|
||
<bibRefCitation id="5D344B74FFE2FF9761FAFE3EFE44828D" author="Lieberman" box="[216,421,476,501]" pageId="31" pageNumber="32" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman (1998)</bibRefCitation>
|
||
, but as opposite the distal tip of S3 by Lieberman (1999,
|
||
<taxonomicName id="FEA54D06FFE2FF97650AFE3EFB4A828C" box="[1064,1195,476,501]" class="Trilobita" family="Biceratopsidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="subFamily" subFamily="Bristoliinae">Bristoliinae</taxonomicName>
|
||
analysis, character 1, state 3)—the 1999
|
||
<taxonomicName id="FEA54D06FFE2FF97604EFE1DFE0E8161" box="[364,495,511,536]" class="Trilobita" family="Biceratopsidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="subFamily" subFamily="Bristoliinae">Bristoliinae</taxonomicName>
|
||
analysis included no equivalent state to 52(3) of the 1998 analysis. A faint furrow is developed across the ocular lobe at the contact with LA (Lieberman, 1999,
|
||
<taxonomicName id="FEA54D06FFE2FF9762C9FDC6FB8F8144" box="[1003,1134,548,573]" class="Trilobita" family="Biceratopsidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="subFamily" subFamily="Bristoliinae">Bristoliinae</taxonomicName>
|
||
analysis, character 7, state 1; this was coded as absent by Lieberman [1999]).
|
||
</paragraph>
|
||
<caption id="6DDA660DFFE3FF9661B5F924FE418489" httpUri="https://zenodo.org/record/286413/files/figure.png" pageId="30" pageNumber="31" targetBox="[358,1227,208,1696]" targetPageId="30">
|
||
<paragraph id="391A3685FFE3FF9661B5F924FE418489" blockId="30.[151,1436,1734,2032]" pageId="30" pageNumber="31">
|
||
<emphasis id="0BD1EA97FFE3FF9661B5F924FEF485A2" bold="true" box="[151,277,1734,1756]" pageId="30" pageNumber="31">FIGURE 9.</emphasis>
|
||
Bivariate scatterplots depicting intraspecific variation and interspecific disparity in various aspects of cephalic morphology among morphologically mature
|
||
<taxonomicName id="FEA54D06FFE3FF966349F906FCC48583" box="[619,805,1764,1786]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="0BD1EA97FFE3FF966349F906FCC48583" box="[619,805,1764,1786]" italics="true" pageId="30" pageNumber="31">Lochmanolenellus</emphasis>
|
||
</taxonomicName>
|
||
species. 1, Orientation of the distal portion of the posterior cephalic margin (distal PCM). 2, Proportional cephalic width, measured as the ratio of the transverse distance between the genal spine bases to glabella length. 3, Proportional anterior advancement of the genal spine bases. 4, Relative location of the adgenal angle along the posterior cephalic margin (PCM), quantified as the ratio of the [linear distance from the innermost tip of the PCM to the adgenal angle] to the [linear distance from the innermost tip of the PCM to the adgenal angle + linear distance from the adgenal angle to the base of the genal spine where it contacts the distal PCM]. 5, Degree of lateral flaring of the intergenal spine, quantified as the angle between the outer margin of the proximal portion of the intergenal spine and the distal portion of the posterior cephalic margin (distal PCM). 6, Proportional width of LA. 7, Proportional length of LA. 8, Proportional length of L2.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="391A3685FFE2FF9761E5FD8EFBCB87C2" blockId="31.[151,1436,151,1211]" pageId="31" pageNumber="32">
|
||
The
|
||
<typeStatus id="E61E8827FFE2FF9761DBFD8EFEBA81FC" box="[249,347,620,645]" pageId="31" pageNumber="32" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="FEA54D06FFE2FF9760A2FD8EFD4281FD" box="[384,675,620,644]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="primus">
|
||
<emphasis id="0BD1EA97FFE2FF9760A2FD8EFD4281FD" box="[384,675,620,644]" italics="true" pageId="31" pageNumber="32">Lochmanolenellus primus</emphasis>
|
||
</taxonomicName>
|
||
is easily distinguished from
|
||
<taxonomicName id="FEA54D06FFE2FF9762C6FD8FFB4381FD" box="[996,1186,620,644]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="trapezoidalis">
|
||
<emphasis id="0BD1EA97FFE2FF9762C6FD8FFB4381FD" box="[996,1186,620,644]" italics="true" pageId="31" pageNumber="32">Lo. trapezoidalis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="FEA54D06FFE2FF9765FAFD8FFA7D81FD" box="[1240,1436,620,644]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="subquadratus">
|
||
<emphasis id="0BD1EA97FFE2FF9765FAFD8FFA7D81FD" box="[1240,1436,620,644]" italics="true" pageId="31" pageNumber="32">Lo. subquadratus</emphasis>
|
||
</taxonomicName>
|
||
by its pentagonal rather than subquadrate or trapezoidal cephalic outline (Figs 9.1, 9.2, 18.8) and by its less strongly anteriorly advanced genal spine bases (Fig. 9.3). However,
|
||
<taxonomicName id="FEA54D06FFE2FF976285FD4DFBC281BF" box="[935,1059,687,710]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="primus">
|
||
<emphasis id="0BD1EA97FFE2FF976285FD4DFBC281BF" box="[935,1059,687,710]" italics="true" pageId="31" pageNumber="32">Lo. primus</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="FEA54D06FFE2FF97659CFD4DFA9C81BF" box="[1214,1405,686,710]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="pentagonalis">
|
||
<emphasis id="0BD1EA97FFE2FF97659CFD4DFA9C81BF" box="[1214,1405,686,710]" italics="true" pageId="31" pageNumber="32">Lo. pentagonalis</emphasis>
|
||
</taxonomicName>
|
||
in those traits (Figs 9.1–9.3, 18.8), and the two can be most readily distinguished by differences in (1) the relative location of the adgenal angle along the posterior cephalic margin, (2) the degree of lateral flaring of the intergenal spines, and (3) the proportional width of LA. The adgenal angle of
|
||
<taxonomicName id="FEA54D06FFE2FF976294FCF1FBD28053" box="[950,1075,787,810]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="primus">
|
||
<emphasis id="0BD1EA97FFE2FF976294FCF1FBD28053" box="[950,1075,787,810]" italics="true" pageId="31" pageNumber="32">Lo. primus</emphasis>
|
||
</taxonomicName>
|
||
is located proportionally more distally along the posterior cephalic margin than that of
|
||
<taxonomicName id="FEA54D06FFE2FF976233FCD7FC2E8035" box="[785,975,820,844]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="trapezoidalis">
|
||
<emphasis id="0BD1EA97FFE2FF976233FCD7FC2E8035" box="[785,975,820,844]" italics="true" pageId="31" pageNumber="32">Lo. trapezoidalis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="FEA54D06FFE2FF976525FCD7FB2D8035" box="[1031,1228,820,844]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="subquadratus">
|
||
<emphasis id="0BD1EA97FFE2FF976525FCD7FB2D8035" box="[1031,1228,820,844]" italics="true" pageId="31" pageNumber="32">Lo. subquadratus</emphasis>
|
||
</taxonomicName>
|
||
, but slightly more proximally than that of
|
||
<taxonomicName id="FEA54D06FFE2FF976080FCB4FDBE8014" box="[418,607,853,877]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="pentagonalis">
|
||
<emphasis id="0BD1EA97FFE2FF976080FCB4FDBE8014" box="[418,607,853,877]" italics="true" pageId="31" pageNumber="32">Lo. pentagonalis</emphasis>
|
||
</taxonomicName>
|
||
(Fig. 9.4). The intergenal spines of
|
||
<taxonomicName id="FEA54D06FFE2FF9762D4FCB4FB918014" box="[1014,1136,854,877]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="primus">
|
||
<emphasis id="0BD1EA97FFE2FF9762D4FCB4FB918014" box="[1014,1136,854,877]" italics="true" pageId="31" pageNumber="32">Lo. primus</emphasis>
|
||
</taxonomicName>
|
||
flare outwards (relative to the orientation of the distal portion of the posterior cephalic margin) more strongly than do those of
|
||
<taxonomicName id="FEA54D06FFE2FF976455FC95FEC880C9" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="trapezoidalis">
|
||
<emphasis id="0BD1EA97FFE2FF976455FC95FEC880C9" italics="true" pageId="31" pageNumber="32">Lo. trapezoidalis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="FEA54D06FFE2FF976047FC7BFDCD80C9" box="[357,556,920,944]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="subquadratus">
|
||
<emphasis id="0BD1EA97FFE2FF976047FC7BFDCD80C9" box="[357,556,920,944]" italics="true" pageId="31" pageNumber="32">Lo. subquadratus</emphasis>
|
||
</taxonomicName>
|
||
, but slightly less strongly than do those of
|
||
<taxonomicName id="FEA54D06FFE2FF976508FC7BFB0B80C9" box="[1066,1258,920,944]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="pentagonalis">
|
||
<emphasis id="0BD1EA97FFE2FF976508FC7BFB0B80C9" box="[1066,1258,920,944]" italics="true" pageId="31" pageNumber="32">Lo. pentagonalis</emphasis>
|
||
</taxonomicName>
|
||
(Fig. 9.5). The glabella of
|
||
<taxonomicName id="FEA54D06FFE2FF976002FC58FE4180A8" box="[288,416,954,977]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="primus">
|
||
<emphasis id="0BD1EA97FFE2FF976002FC58FE4180A8" box="[288,416,954,977]" italics="true" pageId="31" pageNumber="32">Lo. primus</emphasis>
|
||
</taxonomicName>
|
||
is proportionally wider across LA (tr.) relative to basal glabellar width (tr.) than
|
||
<taxonomicName id="FEA54D06FFE2FF976455FC58FEC9808A" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="pentagonalis">
|
||
<emphasis id="0BD1EA97FFE2FF976455FC58FEC9808A" italics="true" pageId="31" pageNumber="32">Lo. pentagonalis</emphasis>
|
||
</taxonomicName>
|
||
(Fig. 9.6). Another diagnostic feature of
|
||
<taxonomicName id="FEA54D06FFE2FF976210FC39FC54808B" box="[818,949,987,1010]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="primus">
|
||
<emphasis id="0BD1EA97FFE2FF976210FC39FC54808B" box="[818,949,987,1010]" italics="true" pageId="31" pageNumber="32">Lo. primus</emphasis>
|
||
</taxonomicName>
|
||
is the length of LA, which is slightly proportionally shorter (relative to glabella length, for specimens of that size) than in other species of the genus (Fig. 9.7).
|
||
<collectingCountry id="41B27615FFE2FF9761F4FBFCFF14874F" box="[214,245,1054,1078]" name="American Samoa" pageId="31" pageNumber="32">As</collectingCountry>
|
||
a result, both L1 and L2 are typically slightly proportionally longer (relative to glabella length, for specimens of that size) in
|
||
<taxonomicName id="FEA54D06FFE2FF97609EFBDDFDD7872F" box="[444,566,1087,1110]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="primus">
|
||
<emphasis id="0BD1EA97FFE2FF97609EFBDDFDD7872F" box="[444,566,1087,1110]" italics="true" pageId="31" pageNumber="32">Lo. primus</emphasis>
|
||
</taxonomicName>
|
||
than in other species (Fig. 9.8; although some specimens of
|
||
<taxonomicName id="FEA54D06FFE2FF9765FDFBDDFA7D872F" box="[1247,1436,1086,1110]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="pentagonalis">
|
||
<emphasis id="0BD1EA97FFE2FF9765FDFBDDFA7D872F" box="[1247,1436,1086,1110]" italics="true" pageId="31" pageNumber="32">Lo. pentagonalis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="FEA54D06FFE2FF9761E5FB83FE678701" box="[199,390,1120,1144]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="trapezoidalis">
|
||
<emphasis id="0BD1EA97FFE2FF9761E5FB83FE678701" box="[199,390,1120,1144]" italics="true" pageId="31" pageNumber="32">Lo. trapezoidalis</emphasis>
|
||
</taxonomicName>
|
||
approach or match these proportions). Together these features serve to differentiate the only definitively known specimen of
|
||
<taxonomicName id="FEA54D06FFE2FF97632EFB60FD6887E0" box="[524,649,1154,1177]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="primus">
|
||
<emphasis id="0BD1EA97FFE2FF97632EFB60FD6887E0" box="[524,649,1154,1177]" italics="true" pageId="31" pageNumber="32">Lo. primus</emphasis>
|
||
</taxonomicName>
|
||
from all other congeneric species. The validity of these distinctions should be re-checked if and when additional material of
|
||
<taxonomicName id="FEA54D06FFE2FF97622DFB41FC6887C3" box="[783,905,1187,1210]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="primus">
|
||
<emphasis id="0BD1EA97FFE2FF97622DFB41FC6887C3" box="[783,905,1187,1210]" italics="true" pageId="31" pageNumber="32">Lo. primus</emphasis>
|
||
</taxonomicName>
|
||
is discovered.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |