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<document id="0A872564EA31BB6F91069E983FA31E67" ID-CLB-Dataset="36828" ID-DOI="10.11646/zootaxa.3824.1.1" ID-GBIF-Dataset="df227efc-13d9-421e-92ed-f9aa417b0cf7" ID-ISSN="1175-5326" ID-Zenodo-Dep="286404" ID-ZooBank="023D78D0-4182-48D2-BAEB-CDA6473CF585" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460443577100" checkinUser="plazi" docAuthor="Webster, Mark &amp; Bohach, Lisa L." docDate="2014" docId="B10C8793FFDDFFA961B5F8AEFED08094" docLanguage="en" docName="zt03824p066.pdf" docOrigin="Zootaxa 3824 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Lochmanolenellus primus" docType="treatment" docVersion="6" lastPageNumber="34" masterDocId="4D35FFEBFFFDFF886122FFE2FFE18379" masterDocTitle="Systematic revision of the trilobite genera Laudonia and Lochmanolenellus (Olenelloidea) from the lower Dyeran (Cambrian Series 2) of western Laurentia" masterLastPageNumber="66" masterPageNumber="1" pageNumber="33" updateTime="1698333190807" updateUser="ExternalLinkService">
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<mods:title id="C389B263E83A3D20A0894E1B18436E53">Systematic revision of the trilobite genera Laudonia and Lochmanolenellus (Olenelloidea) from the lower Dyeran (Cambrian Series 2) of western Laurentia</mods:title>
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<mods:name id="ED052EC40D3AB87C504A72ECE37E2BCB" type="personal">
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<mods:namePart id="6050898F078D3CB0FECE067C9B2C7A4A">Webster, Mark</mods:namePart>
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<mods:namePart id="31AA4EAF0CB106A3837B3964E202B2D7">Bohach, Lisa L.</mods:namePart>
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<mods:typeOfResource id="E963AFF071E1BBB6DE5438B363047561">text</mods:typeOfResource>
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<mods:title id="E3E19A05949DCAB79D148E55C210BFAA">Zootaxa</mods:title>
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<mods:part id="C80A5C1C4D9839071E6913516F19A2EB">
<mods:date id="5555F0CD574D5E8DD30C15257991165D">2014</mods:date>
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<mods:number id="198F1F67BF7C4620F01DC013C9A9260B">3824</mods:number>
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<mods:identifier id="1B18B8CFD52144201CCD3EFB402CD258" type="CLB-Dataset">36828</mods:identifier>
<mods:identifier id="414F5D6ACA2A56CFA07683DE9D241535" type="DOI">10.11646/zootaxa.3824.1.1</mods:identifier>
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<treatment id="B10C8793FFDDFFA961B5F8AEFED08094" ID-DOI="http://doi.org/10.5281/zenodo.6129728" ID-GBIF-Taxon="127668557" ID-Zenodo-Dep="6129728" LSID="urn:lsid:plazi:treatment:B10C8793FFDDFFA961B5F8AEFED08094" httpUri="http://treatment.plazi.org/id/B10C8793FFDDFFA961B5F8AEFED08094" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
<subSubSection id="71BF650EFFDDFFA861B5F8AEFEDB84FE" pageId="32" pageNumber="33" type="nomenclature">
<paragraph id="391A3685FFDDFFA861B5F8AEFDEE841F" blockId="32.[151,527,1868,1927]" box="[151,527,1868,1894]" pageId="32" pageNumber="33">
<heading id="625281E9FFDDFFA861B5F8AEFDEE841F" bold="true" box="[151,527,1868,1894]" fontSize="11" level="1" pageId="32" pageNumber="33" reason="1">
<emphasis id="0BD1EA97FFDDFFA861B5F8AEFDEE841F" bold="true" box="[151,527,1868,1894]" pageId="32" pageNumber="33">
<taxonomicName id="FEA54D06FFDDFFA861B5F8AEFE18841F" box="[151,505,1868,1894]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="primus">
<emphasis id="0BD1EA97FFDDFFA861B5F8AEFE99841F" bold="true" box="[151,376,1868,1894]" italics="true" pageId="32" pageNumber="33">Lochmanolenellus</emphasis>
cf.
<emphasis id="0BD1EA97FFDDFFA86086F8AEFE18841F" bold="true" box="[420,505,1868,1894]" italics="true" pageId="32" pageNumber="33">primus</emphasis>
</taxonomicName>
2
</emphasis>
</heading>
</paragraph>
<paragraph id="391A3685FFDDFFA861B5F88CFEDB84FE" blockId="32.[151,527,1868,1927]" box="[151,314,1902,1927]" pageId="32" pageNumber="33">Fig. 10.410.7</paragraph>
</subSubSection>
<subSubSection id="71BF650EFFDDFFA961B5F854FAA78265" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="materials_examined">
<paragraph id="391A3685FFDDFFA861B5F854FE19848A" blockId="32.[151,1436,1974,2035]" pageId="32" pageNumber="33">
<emphasis id="0BD1EA97FFDDFFA861B5F854FEE784B6" bold="true" box="[151,262,1974,1999]" pageId="32" pageNumber="33">Material.</emphasis>
One cephalon from
<collectionCode id="5FB4AE40FFDDFFA860DBF855FDCE84B6" box="[505,559,1975,1999]" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="32" pageNumber="33">GSC</collectionCode>
locality 95189 (
<collectionCode id="5FB4AE40FFDDFFA863D3F855FCC884B6" box="[753,809,1975,1999]" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="32" pageNumber="33">GSC</collectionCode>
137491), and three cephala from
<collectionCode id="5FB4AE40FFDDFFA86595F855FB0F84B6" box="[1207,1262,1975,1999]" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="32" pageNumber="33">GSC</collectionCode>
locality 95190 (
<collectionCode id="5FB4AE40FFDDFFA861BCF839FF37848A" box="[158,214,2011,2035]" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="32" pageNumber="33">GSC</collectionCode>
137488 to
<collectionCode id="5FB4AE40FFDDFFA86077F839FE6A848A" box="[341,395,2011,2035]" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="32" pageNumber="33">GSC</collectionCode>
137490).
</paragraph>
<paragraph id="391A3685FFDCFFA961E5FF75FAA78265" blockId="33.[151,1437,151,1005]" pageId="33" pageNumber="34">
<emphasis id="0BD1EA97FFDCFFA961E5FF75FEB883C9" bold="true" box="[199,345,151,176]" pageId="33" pageNumber="34">Occurrence.</emphasis>
<collectingCountry id="41B27615FFDCFFA96051FF7AFE0E83C9" box="[371,495,152,176]" name="Canada" pageId="33" pageNumber="34">CANADA</collectingCountry>
:
<emphasis id="0BD1EA97FFDCFFA9632EFF7AFC1383C9" box="[524,1010,152,176]" italics="true" pageId="33" pageNumber="34">
Cassiar Mountains, British
<collectingCountry id="41B27615FFDCFFA96258FF7AFC0D83C9" box="[890,1004,152,176]" name="United States of America" pageId="33" pageNumber="34">Columbia</collectingCountry>
:
</emphasis>
<collectionCode id="5FB4AE40FFDCFFA96529FF7AFBA083C9" box="[1035,1089,152,176]" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="33" pageNumber="34">GSC</collectionCode>
locality 95189, limestone approximately10.6 metres above the base of Unit 4 of the Rosella Formation; and
<collectionCode id="5FB4AE40FFDCFFA9651DFF5EFB9483AD" box="[1087,1141,188,212]" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="33" pageNumber="34">GSC</collectionCode>
locality 95190, limestone approximately12.5 metres above the base of Unit 4 of the Rosella Formation; both in Section 2 of
<bibRefCitation id="5D344B74FFDCFFA965FCFF3DFF1E8265" author="Fritz" pageId="33" pageNumber="34" refString="Fritz, W. H. (1978) Upper (carbonate) part of Atan Group, Lower Cambrian, north-central British Columbia. Geological Survey of Canada, Paper, 78 - 1 A, 7 - 16." type="book chapter" year="1978" yearSuffix="a">Fritz (1978, textfig. 3.1a)</bibRefCitation>
, on the north bank of the Dease River, approximately 2.7 kilometres (
<quantity id="FE5D9B60FFDCFFA9652FFEE6FB978264" box="[1037,1142,260,285]" metricMagnitude="3" metricUnit="m" metricValue="2.7358848" pageId="33" pageNumber="34" unit="mi" value="1.7">1.7 miles</quantity>
) east of McDame.
</paragraph>
</subSubSection>
<subSubSection id="71BF650EFFDCFFA961E5FEC5FED08094" pageId="33" pageNumber="34" type="discussion">
<paragraph id="391A3685FFDCFFA961E5FEC5FED08094" blockId="33.[151,1437,151,1005]" pageId="33" pageNumber="34">
<emphasis id="0BD1EA97FFDCFFA961E5FEC5FEAD8239" bold="true" box="[199,332,295,320]" pageId="33" pageNumber="34">Discussion.</emphasis>
Four poorly preserved cephala from the Cassiar Mountains are possibly conspecific with
<taxonomicName id="FEA54D06FFDCFFA961B5FEAEFE59821D" box="[151,440,332,356]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="primus">
<emphasis id="0BD1EA97FFDCFFA961B5FEAEFE59821D" box="[151,440,332,356]" italics="true" pageId="33" pageNumber="34">Lochmanolenellus primus</emphasis>
</taxonomicName>
. The smallest cephalon is in phase 3 of cephalic development (Fig. 10.4; sagittal cephalic length estimated to be approximately
<quantity id="FE5D9B60FFDCFFA9636BFE92FD5582F1" box="[585,692,368,392]" metricMagnitude="-3" metricUnit="m" metricValue="1.77" pageId="33" pageNumber="34" unit="mm" value="1.77">1.77 mm</quantity>
), and the other three are earliest to mid-phase 5 (Fig. 10.510.7; sagittal cephalic lengths estimated to be approximately
<quantity id="FE5D9B60FFDCFFA9620AFE76FC6482D5" box="[808,901,404,429]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="33" pageNumber="34" unit="mm" value="2.6">2.6 mm</quantity>
,
<quantity id="FE5D9B60FFDCFFA962B4FE76FC1282D5" box="[918,1011,404,429]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="33" pageNumber="34" unit="mm" value="5.7">5.7 mm</quantity>
, and
<quantity id="FE5D9B60FFDCFFA9651BFE76FB7782D5" box="[1081,1174,404,429]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="33" pageNumber="34" unit="mm" value="6.8">6.8 mm</quantity>
, respectively). All are therefore smaller than the
<typeStatus id="E61E8827FFDCFFA960E3FE55FDC282A9" box="[449,547,439,464]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
of the species. Most of their differences from the
<typeStatus id="E61E8827FFDCFFA9657CFE55FB2182A9" box="[1118,1216,439,464]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
are consistent with ontogenetic trends known to occur within other species of the genus: when arranged by size, these specimens document (1) a progressive anterior migration of the genal spine base from a position transversely opposite the lateral margin of L2 to a position transversely opposite the lateral margin of L3; (2) a progressive rotation of the distal portion of the posterior cephalic margin into a more directly anteriorly oriented direction; (3) a proportional shortening of the ocular lobe such that the posterior tip migrates from a position transversely opposite the posterior half of the lateral margin of L1 to a position transversely opposite the anterior half of the lateral margin of L1; and (4) a proportional widening (tr.) of the cephalon, especially in the posterior portion of the extraocular area and the proximal portion of the posterior cephalic margin. The specimens potentially differ from
<taxonomicName id="FEA54D06FFDCFFA96554FD3BFB0E8196" box="[1142,1263,728,752]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="primus">
<emphasis id="0BD1EA97FFDCFFA96554FD3BFB0E8196" box="[1142,1263,728,752]" italics="true" pageId="33" pageNumber="34">Lo. primus</emphasis>
</taxonomicName>
in possessing a more arcuate (convex posteriorly) proximal portion of the posterior cephalic margin, such that the intergenal spine base is slightly more anteriorly located (Fig. 10.6), and in possessing a distal portion of the posterior cephalic margin that is slightly more strongly anteriorly oriented than might be expected for their size (Fig. 10.6, 10.7). Until such differences can be unambiguously resolved as representing either intraspecific variation or interspecific disparity, the Canadian specimens are conservatively left in open nomenclature and treated as
<taxonomicName id="FEA54D06FFDCFFA965D7FC6FFA7D80DD" box="[1269,1436,908,933]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="primus">
<emphasis id="0BD1EA97FFDCFFA965D7FC6FFAFB80DD" box="[1269,1306,909,932]" italics="true" pageId="33" pageNumber="34">Lo.</emphasis>
cf.
<emphasis id="0BD1EA97FFDCFFA9646CFC6FFA7D80DD" box="[1358,1436,909,932]" italics="true" pageId="33" pageNumber="34">primus</emphasis>
</taxonomicName>
2—this name also emphasizes that the Canadian material might not represent the same species as
<taxonomicName id="FEA54D06FFDCFFA965CAFC53FA6780BE" box="[1256,1414,943,968]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="primus">
<emphasis id="0BD1EA97FFDCFFA965CAFC53FAEC80B1" box="[1256,1293,945,968]" italics="true" pageId="33" pageNumber="34">Lo.</emphasis>
cf.
<emphasis id="0BD1EA97FFDCFFA9641AFC52FA6780BE" box="[1336,1414,944,967]" italics="true" pageId="33" pageNumber="34">primus</emphasis>
</taxonomicName>
1 from
<collectingCountry id="41B27615FFDCFFA961F6FC36FECC8094" box="[212,301,980,1005]" name="Mexico" pageId="33" pageNumber="34">Mexico</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>