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<document id="3BA8D510825EDD2788B29A57647398AE" ID-DOI="10.5281/zenodo.5333241" ID-ISSN="2345-7600" ID-Zenodo-Dep="5333241" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630173848801" checkinUser="diego" docAuthor="Tian, Ming-yi &amp; Deuve, Thierry" docDate="2007" docId="A0327D56FFD4FFC7FC1EFD89D9B8E089" docLanguage="en" docName="55rbz261-266.pdf" docOrigin="Raffles Bulletin of Zoology 55 (2)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Orthogonius cyclothorax Tian &amp; Deuve 2007, new species" docType="treatment" docVersion="3" lastPageNumber="264" masterDocId="5C0B052EFFD6FFC4FFB1FFC8DA46E210" masterDocTitle="A Taxonomic Treatment Of The Species Of Orthogonius Macleay (Coleoptera: Carabidae: Orthogoniini) Of Singapore" masterLastPageNumber="266" masterPageNumber="261" pageNumber="263" updateTime="1699174208107" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="06FC3FF3F7A5AECBB0875A0DD936A6E0">A Taxonomic Treatment Of The Species Of Orthogonius Macleay (Coleoptera: Carabidae: Orthogoniini) Of Singapore</mods:title>
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<mods:namePart id="BC71945BFB2EB4D847737426DB02BADA">Tian, Ming-yi</mods:namePart>
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<mods:namePart id="0EBF3E4CCCBF426615DC20530B969884">Deuve, Thierry</mods:namePart>
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<mods:title id="93F05742D8ED0B88CC7784B7EACF8D0E">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="264401E1811D5C1FD4D6B70F18EC6720">2007</mods:date>
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<mods:number id="A48B9E65CBCD0CADB69A52D12A2CF27F">2007-08-31</mods:number>
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<treatment id="A0327D56FFD4FFC7FC1EFD89D9B8E089" LSID="urn:lsid:plazi:treatment:A0327D56FFD4FFC7FC1EFD89D9B8E089" httpUri="http://treatment.plazi.org/id/A0327D56FFD4FFC7FC1EFD89D9B8E089" lastPageId="3" lastPageNumber="264" pageId="2" pageNumber="263">
<subSubSection id="60819FCBFFD4FFC6FC1EFD89DF1FE049" box="[943,1369,577,601]" pageId="2" pageNumber="263" type="nomenclature">
<paragraph id="2824CC40FFD4FFC6FC1EFD89DF1FE049" blockId="2.[943,1369,577,633]" box="[943,1369,577,601]" pageId="2" pageNumber="263">
<heading id="736C7B2CFFD4FFC6FC1EFD89DF1FE049" bold="true" box="[943,1369,577,601]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="263" reason="2">
<emphasis id="1AEF1052FFD4FFC6FC1EFD89DF1FE049" bold="true" box="[943,1369,577,601]" pageId="2" pageNumber="263">
<taxonomicName id="EF9BB7C3FFD4FFC6FC1EFD89DE82E049" ID-CoL="6SWY3" authority="Tian &amp; Deuve, 2007" authorityName="Tian &amp; Deuve" authorityYear="2007" box="[943,1220,577,601]" class="Insecta" family="Carabidae" genus="Orthogonius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="species" species="cyclothorax" status="sp. nov.">
<emphasis id="1AEF1052FFD4FFC6FC1EFD89DE82E049" bold="true" box="[943,1220,577,601]" italics="true" pageId="2" pageNumber="263">Orthogonius cyclothorax</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="01DCAD29FFD4FFC6FB62FD89DF1FE049" box="[1235,1369,577,601]" pageId="2" pageNumber="263" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="60819FCBFFD4FFC6FBF0FDAADE80E069" box="[1089,1222,610,633]" pageId="2" pageNumber="263" type="description">
<paragraph id="2824CC40FFD4FFC6FBF0FDAADE80E069" blockId="2.[943,1369,577,633]" box="[1089,1222,610,633]" pageId="2" pageNumber="263">
(
<figureCitation id="B0A0D0C5FFD4FFC6FBF8FDAADEF8E069" box="[1097,1214,610,633]" captionStart="Figs" captionStartId="3.[114,158,2036,2057]" captionTargetBox="[118,1453,1116,1998]" captionTargetId="figure-453@3.[85,1483,1085,2014]" captionTargetPageId="3" captionText="Figs. 711. Line drawings of structural features of Orthogonius cyclothorax, new species: 7, left fore tibia, dorsal view; 8, apex of right elytron; 9, abdominal ventrite VII of male; 10, aedeagal median lobe, right lateral view; 11, apical portion of aedeagal median lobe, dorsal view. Scale bars: 1 mm. A for Figs. 89; B for Figs. 7, 1011." figureDoi="http://doi.org/10.5281/zenodo.5333245" httpUri="https://zenodo.org/record/5333245/files/figure.png" pageId="2" pageNumber="263">Figs. 711</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="60819FCBFFD4FFC6FCF5FD68DE8AE11C" pageId="2" pageNumber="263" type="materials_examined">
<paragraph id="2824CC40FFD4FFC6FCF5FD68DE8AE11C" blockId="2.[836,1476,672,780]" pageId="2" pageNumber="263">
<emphasis id="1AEF1052FFD4FFC6FCF5FD68D9AEE0A5" bold="true" box="[836,1000,672,693]" pageId="2" pageNumber="263">
<emphasis id="1AEF1052FFD4FFC6FCF5FD68D988E0A5" bold="true" box="[836,974,672,693]" italics="true" pageId="2" pageNumber="263">Type material</emphasis>
.
</emphasis>
<materialsCitation id="98F3C61DFFD4FFC6FC41FD68DE8AE11C" collectingDate="2005-04-19" collectionCode="R, IRSNB" collectorName="P. Grootaert" country="Singapore" location="Swamp forest" pageId="2" pageNumber="263" specimenCount="1" typeStatus="holotype">
<typeStatus id="F72072E2FFD4FFC6FC41FD68DE0BE0A5" box="[1008,1101,672,693]" pageId="2" pageNumber="263" type="holotype">Holotype</typeStatus>
: male, labeled “Coll. I.
<collectionCode id="4E8A5485FFD4FFC6FA8AFD68DF08E0A5" box="[1339,1358,672,693]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="263">R</collectionCode>
. Sc. N. B. /
<collectingCountry id="508C8CD0FFD4FFC6FCF5FD75D9EEE0C2" box="[836,936,701,722]" name="Singapore" pageId="2" pageNumber="263">Singapore</collectingCountry>
, Nee Soon,
<collectingMethod id="F1DAB457FFD4FFC6FB94FD75DE3FE0C2" box="[1061,1145,701,722]" pageId="2" pageNumber="263">Mal trap</collectingMethod>
2,
<location id="2D449A9BFFD4FFC6FB2FFD75DF61E0C2" LSID="urn:lsid:plazi:treatment:A0327D56FFD4FFC7FC1EFD89D9B8E089:2D449A9BFFD4FFC6FB2FFD75DF61E0C2" box="[1182,1319,701,722]" country="Singapore" name="Swamp forest" pageId="2" pageNumber="263">Swamp forest</location>
, Station 25068.
<date id="5C25EA80FFD4FFC6FCF5FD12D982E0FF" box="[836,964,730,751]" pageId="2" pageNumber="263" value="2005-04-19">
<collectingDate id="4C611368FFD4FFC6FCF5FD12D982E0FF" box="[836,964,730,751]" pageId="2" pageNumber="263" value="2005-04-19">19 Apr. 2005</collectingDate>
</date>
.
<collectorName id="856EA996FFD4FFC6FC7CFD12DE79E0FF" box="[973,1087,730,751]" pageId="2" pageNumber="263">P. Grootaert</collectorName>
coll.”, collected at Nee Soon,
<collectingCountry id="508C8CD0FFD4FFC6FAD3FD12DF85E0FF" box="[1378,1475,730,751]" name="Singapore" pageId="2" pageNumber="263">Singapore</collectingCountry>
(type locality) and deposited in
<collectionCode id="4E8A5485FFD4FFC6FBCFFD3FDE8EE11C" box="[1150,1224,759,780]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="2" pageNumber="263">IRSNB</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="60819FCBFFD4FFC7FCF5FCFDD9FFE029" lastPageId="3" lastPageNumber="264" pageId="2" pageNumber="263" type="description">
<paragraph id="2824CC40FFD4FFC6FCF5FCFDDE01E17C" blockId="2.[836,1475,820,876]" pageId="2" pageNumber="263">
<emphasis id="1AEF1052FFD4FFC6FCF5FCFDD98DE15C" bold="true" box="[836,971,820,845]" pageId="2" pageNumber="263">
<emphasis id="1AEF1052FFD4FFC6FCF5FCFDD985E15D" bold="true" box="[836,963,821,845]" italics="true" pageId="2" pageNumber="263">Description</emphasis>
.
</emphasis>
Body: Length
<quantity id="EF6361A5FFD4FFC6FBC0FCFDDE93E15C" box="[1137,1237,821,844]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="2" pageNumber="263" unit="mm" value="10.5">10.5 mm</quantity>
; width
<quantity id="EF6361A5FFD4FFC6FA96FCFDDF38E15C" box="[1319,1406,821,844]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="2" pageNumber="263" unit="mm" value="4.5">4.5 mm</quantity>
. Form stout, moderately shiny.
</paragraph>
<paragraph id="2824CC40FFD4FFC6FCF5FC5DDF63E1FC" blockId="2.[836,1475,917,1004]" pageId="2" pageNumber="263">Color: Head dark brown, pronotum and elytra reddish brown; legs (except femora), ventral surface, palpi of mouthparts, antennae and labrum brown; femora yellow.</paragraph>
<paragraph id="2824CC40FFD4FFC6FCF5FBDDD9B0E67C" blockId="2.[836,1475,1045,1132]" pageId="2" pageNumber="263">Macrosculpture: Head and pronotum shallowly striate, impunctate, intervals 3, 5, 6 and 8 of elytra with sparse, small punctures.</paragraph>
<paragraph id="2824CC40FFD4FFC6FCF5FB5DDE38E6DC" blockId="2.[836,1475,1173,1228]" pageId="2" pageNumber="263">Microsculpture: Mesh pattern isodiametric on head, pronotum and elytra; microlines dense.</paragraph>
<paragraph id="2824CC40FFD4FFC6FCF5FB3DDE3AE7DC" blockId="2.[836,1476,1269,1484]" pageId="2" pageNumber="263">Head: Slightly narrower than long (HW/HL = 0.92), stout, evidently convex. Frontal impressions pit-like, small; clypeus bisetose, surface quite even. Eyes very large, prominent. Antennae rather long, extending to about one-fifths of elytra from base; pubescent from apical two-thirds of antennomere 4; antennomere 4 as long as antennomere 3, but less than two times of antennomere 2.</paragraph>
<paragraph id="2824CC40FFD4FFC6FCF5FA3DDFE1E4FC" blockId="2.[836,1476,1525,1772]" pageId="2" pageNumber="263">Mouthparts: Labrum six setose, anterior margin distinctly emarginate. Maxillae: palpi subcylindrical, palpomeres 3 and 4 subequal; palpomere 3 with several short setae at apex. Labium: submentum with one seta on either side; mentum without median tooth, asetose; glossal sclerite of ligula small, bisetose at apex; palpomeres with articles subcylindrical, palpomere 2 slightly longer than palpomere 3; palpomere 2 bisetose on inner margin with a few short setae at apex.</paragraph>
<paragraph id="2824CC40FFD4FFC6FCF5F8DDDE47EA1C" blockId="2.[836,1476,1813,2060]" pageId="2" pageNumber="263">Prothorax: Pronotum — form transverse (PW/PL = 1.82), widest at slightly posteriad middle; both anterior and posterior margins beaded; fore and hind angles round; sides evenly broadened; lateral expanded margins clearly defined, smooth and hardly reflexed; disc convex; anterior and posterior transverse impressions shallow, posteriolateral foveae small. Prosternum — intercoxal process distinctly bordered at apex.</paragraph>
<paragraph id="2824CC40FFD5FFC7FFC3FF0ADB9CE389" blockId="3.[114,754,194,409]" pageId="3" pageNumber="264">
Elytra: Form broadly ovate (EL/EW = 1.44), parallel at sides at middle portion, surface moderately convex; basal border complete; humeri square, apex broadly truncate, sutural angle nearly rectangular (
<figureCitation id="B0A0D0C5FFD5FFC7FE31FEEADB86E329" box="[384,448,290,313]" captionStart="Figs" captionStartId="3.[114,158,2036,2057]" captionTargetBox="[118,1453,1116,1998]" captionTargetId="figure-453@3.[85,1483,1085,2014]" captionTargetPageId="3" captionText="Figs. 711. Line drawings of structural features of Orthogonius cyclothorax, new species: 7, left fore tibia, dorsal view; 8, apex of right elytron; 9, abdominal ventrite VII of male; 10, aedeagal median lobe, right lateral view; 11, apical portion of aedeagal median lobe, dorsal view. Scale bars: 1 mm. A for Figs. 89; B for Figs. 7, 1011." figureDoi="http://doi.org/10.5281/zenodo.5333245" httpUri="https://zenodo.org/record/5333245/files/figure.png" pageId="3" pageNumber="264">Fig. 8</figureCitation>
); striae rather deep, intervals quite flat, subequal in width at middle portion; interval 7 flat, not carinate; interval 3 with middle and hind setiferous punctures, basal puncture absent.
</paragraph>
<paragraph id="2824CC40FFD5FFC7FFC3FE0ADB37E0A9" blockId="3.[114,753,450,697]" pageId="3" pageNumber="264">
Legs: Middle coxae glabrous. Femora — hind femur moderate, with two setae on posterior margin of ventral surface. Tibiae — fore tibia stout, gradually enlarged at apex, apical margin sinuate, outer angle rectangular (
<figureCitation id="B0A0D0C5FFD5FFC7FD13FDEAD8A5E029" box="[674,739,546,569]" captionStart="Figs" captionStartId="3.[114,158,2036,2057]" captionTargetBox="[118,1453,1116,1998]" captionTargetId="figure-453@3.[85,1483,1085,2014]" captionTargetPageId="3" captionText="Figs. 711. Line drawings of structural features of Orthogonius cyclothorax, new species: 7, left fore tibia, dorsal view; 8, apex of right elytron; 9, abdominal ventrite VII of male; 10, aedeagal median lobe, right lateral view; 11, apical portion of aedeagal median lobe, dorsal view. Scale bars: 1 mm. A for Figs. 89; B for Figs. 7, 1011." figureDoi="http://doi.org/10.5281/zenodo.5333245" httpUri="https://zenodo.org/record/5333245/files/figure.png" pageId="3" pageNumber="264">Fig. 7</figureCitation>
); middle tibia slender; hind tibia hardly dilated at apex, apical spurs long and sharp. Tarsi — hind tarsomere 4 shorter than tarsomere 3, shallowly emarginate at apex; all tarsal claws pectinate, pectens long.
</paragraph>
<paragraph id="2824CC40FFD5FFC7FFC3FD2AD85EE109" blockId="3.[114,753,738,793]" pageId="3" pageNumber="264">
Abdomen: Abdominal ventrite VII of male complete, without emargination at apical margin (
<figureCitation id="B0A0D0C5FFD5FFC7FE76FCCAD84DE109" box="[455,523,770,793]" captionStart="Figs" captionStartId="3.[114,158,2036,2057]" captionTargetBox="[118,1453,1116,1998]" captionTargetId="figure-453@3.[85,1483,1085,2014]" captionTargetPageId="3" captionText="Figs. 711. Line drawings of structural features of Orthogonius cyclothorax, new species: 7, left fore tibia, dorsal view; 8, apex of right elytron; 9, abdominal ventrite VII of male; 10, aedeagal median lobe, right lateral view; 11, apical portion of aedeagal median lobe, dorsal view. Scale bars: 1 mm. A for Figs. 89; B for Figs. 7, 1011." figureDoi="http://doi.org/10.5281/zenodo.5333245" httpUri="https://zenodo.org/record/5333245/files/figure.png" pageId="3" pageNumber="264">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="2824CC40FFD5FFC7FFC3FC8ADA83E1A9" blockId="3.[114,754,834,953]" pageId="3" pageNumber="264">
Male genitalia: Median lobe of aedeagus slightly expanded in middle portion, apex abruptly terminated and pointed; apical lamella small, longer than wide, obtuse at apex (
<figureCitation id="B0A0D0C5FFD5FFC7FD0DFC4ADAFFE1A9" captionStart="Figs" captionStartId="3.[114,158,2036,2057]" captionTargetBox="[118,1453,1116,1998]" captionTargetId="figure-453@3.[85,1483,1085,2014]" captionTargetPageId="3" captionText="Figs. 711. Line drawings of structural features of Orthogonius cyclothorax, new species: 7, left fore tibia, dorsal view; 8, apex of right elytron; 9, abdominal ventrite VII of male; 10, aedeagal median lobe, right lateral view; 11, apical portion of aedeagal median lobe, dorsal view. Scale bars: 1 mm. A for Figs. 89; B for Figs. 7, 1011." figureDoi="http://doi.org/10.5281/zenodo.5333245" httpUri="https://zenodo.org/record/5333245/files/figure.png" pageId="3" pageNumber="264">Figs. 1011</figureCitation>
).
</paragraph>
<paragraph id="2824CC40FFD5FFC7FFC3FC2ADEE6E329" blockId="3.[114,321,994,1017]" lastBlockId="3.[813,1454,193,313]" pageId="3" pageNumber="264">
Female: Unknown.
<emphasis id="1AEF1052FFD5FFC7FC9CFF09D986E2C9" bold="true" box="[813,960,193,217]" pageId="3" pageNumber="264">
<emphasis id="1AEF1052FFD5FFC7FC9CFF09D9E4E2C9" bold="true" box="[813,930,193,217]" italics="true" pageId="3" pageNumber="264">Etymology</emphasis>
.
</emphasis>
The specific epithet is a combination of the Greek adjectival prefix “cyclo-”, meaning round, and feminine noun “thorax”, referring to the form of the pronotum which has rounded fore and hind angles.
</paragraph>
<paragraph id="2824CC40FFD5FFC7FC9CFEA9D9FFE029" blockId="3.[813,1454,353,569]" pageId="3" pageNumber="264">
<emphasis id="1AEF1052FFD5FFC7FC9CFEA9D991E369" bold="true" box="[813,983,353,377]" pageId="3" pageNumber="264">
<emphasis id="1AEF1052FFD5FFC7FC9CFEA9D9F8E369" bold="true" box="[813,958,353,377]" italics="true" pageId="3" pageNumber="264">Comparisons</emphasis>
.
</emphasis>
This new species is similar to
<taxonomicName id="EF9BB7C3FFD5FFC7FAA2FEAADE4FE389" authority="Tian &amp; Deuve, 2006" authorityName="Tian &amp; Deuve" authorityYear="2006" class="Insecta" family="Carabidae" genus="Orthogonius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="medanensis">
<emphasis id="1AEF1052FFD5FFC7FAA2FEAADFE8E369" box="[1299,1454,354,377]" italics="true" pageId="3" pageNumber="264">O. medanensis</emphasis>
Tian &amp; Deuve, 2006
</taxonomicName>
of
<collectingRegion id="EA5F02A2FFD5FFC7FB9FFE4ADECCE389" box="[1070,1162,386,409]" country="Indonesia" name="Sumatera Utara" pageId="3" pageNumber="264">Sumatra</collectingRegion>
, but is markedly distinct in the following morphological features: its mentum is without seta (2 setae in
<taxonomicName id="EF9BB7C3FFD5FFC7FC5DFE0ADED3E3C9" authorityName="Tian &amp; Deuve" authorityYear="2006" box="[1004,1173,450,473]" class="Insecta" family="Carabidae" genus="Orthogonius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="medanensis">
<emphasis id="1AEF1052FFD5FFC7FC5DFE0ADED3E3C9" box="[1004,1173,450,473]" italics="true" pageId="3" pageNumber="264">O. medanensis</emphasis>
</taxonomicName>
), interval 3 of elytra is without basal setiferous punctures (evident in
<taxonomicName id="EF9BB7C3FFD5FFC7FABAFE2ADFE3E3E9" authorityName="Tian &amp; Deuve" authorityYear="2006" box="[1291,1445,482,505]" class="Insecta" family="Carabidae" genus="Orthogonius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="medanensis">
<emphasis id="1AEF1052FFD5FFC7FABAFE2ADFE3E3E9" box="[1291,1445,482,505]" italics="true" pageId="3" pageNumber="264">O. medanensis</emphasis>
</taxonomicName>
) and apical lamella of aedeagus is narrower (broader in
<taxonomicName id="EF9BB7C3FFD5FFC7FA25FDCAD9EFE029" authorityName="Tian &amp; Deuve" authorityYear="2006" class="Insecta" family="Carabidae" genus="Orthogonius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="medanensis">
<emphasis id="1AEF1052FFD5FFC7FA25FDCAD9EFE029" italics="true" pageId="3" pageNumber="264">O. medanensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="60819FCBFFD5FFC7FC9CFDA9D9B8E089" pageId="3" pageNumber="264" type="distribution">
<paragraph id="2824CC40FFD5FFC7FC9CFDA9D9B8E089" blockId="3.[813,1453,609,665]" pageId="3" pageNumber="264">
<emphasis id="1AEF1052FFD5FFC7FC9CFDA9D997E069" bold="true" box="[813,977,609,633]" pageId="3" pageNumber="264">
<emphasis id="1AEF1052FFD5FFC7FC9CFDA9D9F5E069" bold="true" box="[813,947,609,633]" italics="true" pageId="3" pageNumber="264">Distribution</emphasis>
.
</emphasis>
This species is known only from the
<typeStatus id="F72072E2FFD5FFC7FACEFDAADFEBE069" box="[1407,1453,610,633]" pageId="3" pageNumber="264">type</typeStatus>
locality,
<collectingCountry id="508C8CD0FFD5FFC7FC3AFD4AD9BDE089" box="[907,1019,642,665]" name="Singapore" pageId="3" pageNumber="264">Singapore</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>