185 lines
31 KiB
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185 lines
31 KiB
XML
<document id="1DF5936A766BB08D97DDEE7AEB6E885B" ID-CLB-Dataset="8118" ID-DOI="10.5252/z2010n1a2" ID-GBIF-Dataset="4cecd899-5900-4411-8446-0d9552b1b098" ID-ISSN="1638-9387" ID-Zenodo-Dep="4520753" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1612808309234" checkinUser="felipe" docAuthor="Randriamanantsoa, Richard, Aberlenc, Henri-Pierre, Ralisoa, Olga Bakoly, Ratnadass, Alain & Vercambre, Bernard" docDate="2010" docId="901F8794FF9F746DFD4FFB20FC1FFBE3" docLanguage="fr" docName="Zoosystema.32.1.19-72.pdf" docOrigin="Zoosystema 32 (1)" docSource="http://www.bioone.org/doi/abs/10.5252/z2010n1a2" docStyle="DocumentStyle:E638A3C99FA728B529662B46F84D75FD.3:Zoosystema.2009-2010.journal_article" docStyleId="E638A3C99FA728B529662B46F84D75FD" docStyleName="Zoosystema.2009-2010.journal_article" docStyleVersion="3" docTitle="Hexodon unicolor" docType="treatment" docVersion="12" lastPageNumber="31" masterDocId="6C26FFECFF947461FFB5FFEDFF94FFE8" masterDocTitle="Les larves des Scarabaeoidea (Insecta, Coleoptera) en riziculture pluviale des régions de haute et moyenne altitudes du Centre de Madagascar" masterLastPageNumber="72" masterPageNumber="19" pageNumber="30" updateTime="1698928337295" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="29EF762766FCC15AA67F9938A47FB5D9">Les larves des Scarabaeoidea (Insecta, Coleoptera) en riziculture pluviale des régions de haute et moyenne altitudes du Centre de Madagascar</mods:title>
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<mods:namePart id="C305A7910BAD71DD24DBF49B69F51504">Randriamanantsoa, Richard</mods:namePart>
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<mods:affiliation id="74DD5648EBFA7F6D4BA8318AE9856DF4">Fofifa - URP Scrid, 110 Antsirabe (Madagascar) r _ randriamanantsoa @ yahoo. fr</mods:affiliation>
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<mods:namePart id="A9EEA5F04F923D53A3C0CE4B3736028D">Aberlenc, Henri-Pierre</mods:namePart>
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<mods:affiliation id="92384C6EBA9DC7A563E0E0FD2596F87F">Cirad Bios, UMR CBGP (INRA / IRD / Cirad / Montpellier SupAgro), Campus international de Baillarguet, Ta A- 55 / L, F- 34398 Montpellier cedex 5 (France) henri-pierre. aberlenc @ cirad. fr</mods:affiliation>
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<mods:nameIdentifier id="F6FC90A6469BD3BE2019FB0F65BD27E4" type="email">aberlenc@cirad.fr</mods:nameIdentifier>
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<mods:namePart id="FD62ADD2C3BA505B4F4B44BE557B3D81">Ralisoa, Olga Bakoly</mods:namePart>
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<mods:affiliation id="E6EB4924E8BD85ADDFE47B415CB18F53">Département d’Entomologie, Faculté des Sciences, Université d’Antananarivo, Antananarivo 101 (Madagascar)</mods:affiliation>
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<mods:namePart id="46920843C8BF23FF0789F21918FFC4EF">Ratnadass, Alain</mods:namePart>
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<mods:affiliation id="AA1825DD05DCF2617432097856CD1FAB">Cirad, UPR HortSys - ICRISAT BP 12404, Niamey (Niger) a. ratnadass @ cgiar. org</mods:affiliation>
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<mods:nameIdentifier id="5CD203DC4F4C12531ADDFF49B45BEBC2" type="email">ratnadass@cgiar.org</mods:nameIdentifier>
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<mods:namePart id="3BD5E184575D53F88127D3A8EE241C0E">Vercambre, Bernard</mods:namePart>
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<mods:affiliation id="73B3BFCBE8CB3CD4C7D1A4EA7019E1F1">Cirad, UPR Systèmes de Cultures annuels, avenue Agropolis, F- 34398 Montpellier cedex 5 (France) bernard. vercambre @ cirad. fr</mods:affiliation>
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<mods:nameIdentifier id="26B654FCA36C0E26C4E01888B8FBD487" type="email">vercambre@cirad.fr</mods:nameIdentifier>
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<mods:title id="0C3391ED8D735690D92C4A84E9E327E0">Zoosystema</mods:title>
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<mods:date id="5ED998338B5197B726F8EE4C0101B51D">2010</mods:date>
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<mods:number id="A87012B72D650AA4B7DA7E44C1CE073F">2010-03-31</mods:number>
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<treatment id="901F8794FF9F746DFD4FFB20FC1FFBE3" ID-DOI="http://doi.org/10.5281/zenodo.4549536" ID-GBIF-Taxon="177425664" ID-Zenodo-Dep="4549536" LSID="urn:lsid:plazi:treatment:901F8794FF9F746DFD4FFB20FC1FFBE3" httpUri="http://treatment.plazi.org/id/901F8794FF9F746DFD4FFB20FC1FFBE3" lastPageId="12" lastPageNumber="31" pageId="11" pageNumber="30">
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<subSubSection id="50AC6509FF9F746AFD4FFB20FCE1FA8D" pageId="11" pageNumber="30" type="nomenclature">
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<paragraph id="18093682FF9F746AFD4FFB20FBF0FB01" blockId="11.[735,1151,1229,1320]" box="[762,1124,1229,1257]" pageId="11" pageNumber="30">
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<taxonomicName id="DFB64D01FF9F746AFD4FFB20FBF0FB01" authority="(Olivier, 1789)" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[762,1124,1229,1257]" class="Insecta" family="Dynastidae" genus="Hexodon" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="30" phylum="Arthropoda" rank="species" species="unicolor">
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<emphasis id="2AC2EA90FF9F746AFD4FFB20FC28FB00" bold="true" box="[762,956,1229,1256]" italics="true" pageId="11" pageNumber="30">Hexodon unicolor</emphasis>
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(Olivier, 1789)
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</taxonomicName>
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</paragraph>
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<paragraph id="18093682FF9F746AFD5EFB00FBEBFAC0" blockId="11.[735,1151,1229,1320]" pageId="11" pageNumber="30">
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(
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<figureCitation id="808D2A07FF9F746AFD47FB00FCDFFAE0" box="[754,843,1261,1288]" captionStart="FIG" captionStartId="29.[113,124,1567,1584]" captionTargetBox="[152,1172,222,1545]" captionTargetId="figure-18@29.[148,709,692,1169]" captionTargetPageId="29" captionText="FIG. 2. — Larves de Scarabaeoidea de Madagascar, habitus: A, Bricoptis variolosa (Gory & Percheron, 1833); B, Celidota parvula (Janson, 1881); C, Anochilia bifida (Olivier, 1789); D, Euryomia argentea (Olivier, 1789); E, Heteronychus arator rugifrons (Fairmaire, 1871); F, Heteronychus bituberculatus (Kolbe, 1900); G, Heteronychus plebeius (Klug, 1833); H, Hexodon unicolor (Olivier, 1789); I, Heteroconus paradoxus (Endrödi,1968); J, Paramorphochelus cornutus (Nonfried,1892); K, Encya sikorai (Brenske,1891); L, Apicencya waterloti (Dewailly, 1950); M, Empecta scutata (Fairmaire,1901); N, Hoplochelus betanimena (Künckel,1887); O, Triodontus nitidulus (Guérin, 1844). Échelles: A-I, K, L, N, 5 mm; J, O, 2 mm; M, 3 mm." figureDoi="http://doi.org/10.5281/zenodo.4520757" httpUri="https://zenodo.org/record/4520757/files/figure.png" pageId="11" pageNumber="30">Figs 2H</figureCitation>
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;
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<figureCitation id="808D2A07FF9F746AFCE9FB00FCEFFAE0" box="[860,891,1261,1288]" captionStart="FIG" captionStartId="31.[113,124,1635,1652]" captionTargetBox="[124,1194,218,1601]" captionTargetId="figure-14@31.[106,1195,214,1601]" captionTargetPageId="31" captionText="FIG. 4. — Larves de Scarabaeoidea de Madagascar,raster: A, sp. D1; B, Hexodon unicolor (Olivier, 1789); C, sp. D2; D, Heteroconus paradoxus (Endrödi, 1968); E, Paranodon coquerelii (Fairmaire, 1871); F, Paramorphochelus cornutus (Nonfried, 1892); G, sp. M1; H, Encya sikorai (Brenske, 1891). Échelles: A-D, H, 2 mm; E-G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520763" httpUri="https://zenodo.org/record/4520763/files/figure.png" pageId="11" pageNumber="30">4B</figureCitation>
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;
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<figureCitation id="808D2A07FF9F746AFC38FB00FC38FAE0" box="[909,940,1261,1288]" captionStart="FIG" captionStartId="33.[113,124,1635,1652]" captionTargetBox="[117,1206,214,1443]" captionTargetId="figure-14@33.[117,1206,214,1443]" captionTargetPageId="33" captionText="FIG. 6. — Larves de Scarabaeoidea de Madagascar: A-F, tête; A, Anochilia bifida (Olivier, 1789); B, Hexodon unicolor (Olivier, 1789); C, sp.D2; D, Paramorphochelus cornutus (Nonfried,1892); E, Empecta scutata (Fairmaire,1901); F, sp.S2; G, H, épipharynx; G, Bricoptis variolosa (Gory & Percheron, 1833); H, Celidota parvula (Janson, 1881). Échelles: A-C, 2 mm; D-F, 1 mm; G, H, 0,2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520767" httpUri="https://zenodo.org/record/4520767/files/figure.png" pageId="11" pageNumber="30">6B</figureCitation>
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;
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<figureCitation id="808D2A07FF9F746AFC08FB00FC74FAE0" box="[957,992,1261,1288]" captionStart="FIG" captionStartId="34.[113,124,1635,1652]" captionTargetBox="[155,1168,220,1591]" captionTargetId="figure-21@34.[155,1168,220,1591]" captionTargetPageId="34" captionText="FIG. 7. — Larves de Scarabaeoidea de Madagascar,épipharynx: A, Anochilia bifida (Olivier,1789); B, Euryomia argentea (Olivier,1789); C, Heteronychus arator rugifrons (Fairmaire, 1871); D, Heteronychus bituberculatus (Kolbe, 1900); E, F, sp. D1; G, Hexodon unicolor (Olivier, 1789); H, sp. D2. Échelles: A, B, D, E, G, 0,5 mm; C, 0,2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520771" httpUri="https://zenodo.org/record/4520771/files/figure.png" pageId="11" pageNumber="30">7G</figureCitation>
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;
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<figureCitation id="808D2A07FF9F746AFC47FB00FBB0FAE0" box="[1010,1060,1261,1288]" captionStart="FIG" captionStartId="37.[113,124,1589,1606]" captionTargetBox="[146,1173,219,1547]" captionTargetId="figure-14@37.[146,1173,218,1547]" captionTargetPageId="37" captionText="FIG. 10. — Larves de Scarabaeoidea de Madagascar,hypopharynx: A, Bricoptis variolosa (Gory & Percheron,1833); B, Celidota parvula (Janson, 1881); C, Anochilia bifida (Olivier, 1789); D, Euryomia argentea (Olivier, 1789); E, Heteronychus arator rugifrons (Fairmaire, 1871); F, Heteronychus bituberculatus (Kolbe, 1900); G, Heteronychus plebeius (Klug, 1833); H, Hexodon unicolor (Olivier, 1789); I, Heteroconus paradoxus (Endrödi,1968); J, Paranodon coquerelii (Fairmaire,1871); K, Paramorphochelus cornutus (Nonfried,1892); L, Encya sikorai (Brenske, 1891). Échelles: A, J, 0,2 mm; C, D, F, G, I, 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.4520783" httpUri="https://zenodo.org/record/4520783/files/figure.png" pageId="11" pageNumber="30">10H</figureCitation>
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;
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<figureCitation id="808D2A07FF9F746AFB83FB00FBFCFAE0" box="[1078,1128,1261,1288]" captionStart="FIG" captionStartId="39.[113,124,1567,1584]" captionTargetBox="[118,1207,216,1486]" captionTargetId="figure-14@39.[118,1207,216,1486]" captionTargetPageId="39" captionText="FIG. 12. — Larves de Scarabaeoidea de Madagascar: A-E, hypopharynx; A, Enaria melanictera (Klug, 1833); B, Apicencya waterloti (Dewailly, 1950); C, Empecta scutata (Fairmaire, 1901); D, Hoplochelus betanimena (Künckel, 1887); E, Triodontus nitidulus (Guérin, 1844); F-O, antenne; F, Anochilia bifida (Olivier, 1789); G, Heteronychus plebeius (Klug, 1833); H, Hexodon unicolor (Olivier, 1789); I, Heteroconus paradoxus (Endrödi,1968); J, Paranodon coquerelii (Fairmaire,1871); K-M, Encya sikorai (Brenske,1891); L, M, plages sensorielles de l’article apical; N, Apicencya waterloti (Dewailly, 1950); O, Empecta scutata (Fairmaire, 1901). Échelles: A, F, G, I, J, O, 0,5 mm; E, L, M, 0,2 mm; H, K, N, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520787" httpUri="https://zenodo.org/record/4520787/files/figure.png" pageId="11" pageNumber="30">12H</figureCitation>
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;
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<figureCitation id="808D2A07FF9F746AFD6AFAE0FD6CFAC0" box="[735,760,1293,1320]" captionStart="FIG" captionStartId="41.[113,124,1589,1606]" captionTargetBox="[161,1157,219,1557]" captionTargetId="figure-14@41.[157,1165,214,1560]" captionTargetPageId="41" captionText="FIG. 14. — Larves de Scarabaeoidea de Madagascar,mandibule: A, D, H, L, P, T, X, gauche,face dorsale; B, E, I, M, Q, U, Y, gauche, face ventrale; C, F, J, N, R, V, Z1, droite, face dorsale; G, K, O, S, W, Z2, droite, face ventrale; A-C, Bricoptis variolosa (Gory & Percheron, 1833); D-G, Anochilia bifida (Olivier, 1789); H-K, Euryomia argentea (Olivier, 1789); L-O, Heteronychus arator rugifrons (Fairmaire,1871); P-S, Heteronychus bituberculatus (Kolbe,1900); T-W, Hexodon unicolor (Olivier,1789); X-Z2, Heteroconus paradoxus (Endrödi, 1968). Échelles: A-C, H-O, 0,5 mm; D-G, P-Z2, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520795" httpUri="https://zenodo.org/record/4520795/files/figure.png" pageId="11" pageNumber="30">14</figureCitation>
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T-W; 16N; 19D; 21R-T; 24N, O)
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</paragraph>
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<paragraph id="18093682FF9F746AFD10FAA1FCE1FA8D" blockId="11.[674,1211,1356,1605]" box="[677,885,1356,1383]" pageId="11" pageNumber="30">MATÉRIEL EXAMINÉ</paragraph>
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</subSubSection>
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<subSubSection id="50AC6509FF9F746DFD10FA81FC1FFBE3" lastPageId="12" lastPageNumber="31" pageId="11" pageNumber="30" type="description">
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<paragraph id="18093682FF9F746AFD10FA81FBDFF9AD" blockId="11.[674,1211,1356,1605]" pageId="11" pageNumber="30">
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Larves provenant d’Andranomanelatra, prélevées dans une parcelle menée en système de culture à base de Riz pluvial en haut de pente. La larve est indifférente à la culture et améliore la structure du sol (
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<bibRefCitation id="7C274B73FF9F746AFD79FA06FC3EF9EE" author="RATNADASS A. & MICHELLON R. & RANDRIAMANANTSOA R. & SEGUY L." box="[716,938,1515,1542]" pageId="11" pageNumber="30" pagination="589 - 602" refId="ref24459" refString="RATNADASS A., MICHELLON R., RANDRIAMANANTSOA R. & SEGUY L. 2006. - Effects of soil and plant management on crop pests and diseases, in UPHOFF N., BALL A., FERNANDES E., HERREN H., HUSSON O., LAING M., PALM C., PRETTY J., SANCHEZ P., SANGINGA N. & THIES J. (eds), Biological Approaches to Sustainable Soil Systems. CRC Press-Taylor & Francis, Boca Raton, Florida: 589 - 602." type="book chapter" year="2006">
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Ratnadass
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<emphasis id="2AC2EA90FF9F746AFC8EFA06FCFFF9ED" box="[827,875,1515,1541]" italics="true" pageId="11" pageNumber="30">et al.</emphasis>
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2006
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</bibRefCitation>
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).
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<typeStatus id="C70D8820FF9F746AFC0DFA06FC78F9EE" box="[952,1004,1515,1542]" pageId="11" pageNumber="30">Type</typeStatus>
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de sol: ferrallitique. Spécimens conservés dans les collections du Fofifa ( larves), et du Cirad-CBGP (
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<specimenCount id="0EB0FD0BFF9F746AFC40F9C7FBD4F9AD" box="[1013,1088,1578,1605]" pageId="11" pageNumber="30" type="larva">1 larve</specimenCount>
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).
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</paragraph>
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<paragraph id="18093682FF9F746AFD10F984FCA1F96A" blockId="11.[677,1014,1641,1700]" box="[677,821,1641,1668]" pageId="11" pageNumber="30">DESCRIPTION</paragraph>
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<paragraph id="18093682FF9F746AFD10F964FC62F94C" blockId="11.[677,1014,1641,1700]" box="[677,1014,1673,1700]" pageId="11" pageNumber="30">
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Couleur blanc ivoire (
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<figureCitation id="808D2A07FF9F746AFC25F964FC7DF94C" box="[912,1001,1673,1700]" captionStart="FIG" captionStartId="29.[113,124,1567,1584]" captionTargetBox="[152,1172,222,1545]" captionTargetId="figure-18@29.[148,709,692,1169]" captionTargetPageId="29" captionText="FIG. 2. — Larves de Scarabaeoidea de Madagascar, habitus: A, Bricoptis variolosa (Gory & Percheron, 1833); B, Celidota parvula (Janson, 1881); C, Anochilia bifida (Olivier, 1789); D, Euryomia argentea (Olivier, 1789); E, Heteronychus arator rugifrons (Fairmaire, 1871); F, Heteronychus bituberculatus (Kolbe, 1900); G, Heteronychus plebeius (Klug, 1833); H, Hexodon unicolor (Olivier, 1789); I, Heteroconus paradoxus (Endrödi,1968); J, Paramorphochelus cornutus (Nonfried,1892); K, Encya sikorai (Brenske,1891); L, Apicencya waterloti (Dewailly, 1950); M, Empecta scutata (Fairmaire,1901); N, Hoplochelus betanimena (Künckel,1887); O, Triodontus nitidulus (Guérin, 1844). Échelles: A-I, K, L, N, 5 mm; J, O, 2 mm; M, 3 mm." figureDoi="http://doi.org/10.5281/zenodo.4520757" httpUri="https://zenodo.org/record/4520757/files/figure.png" pageId="11" pageNumber="30">Fig. 2H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="18093682FF98746DFFC4FF35FE96FF1A" blockId="12.[111,648,216,401]" box="[113,258,216,242]" pageId="12" pageNumber="31">
|
||
<emphasis id="2AC2EA90FF98746DFFC4FF35FE96FF1A" box="[113,258,216,242]" italics="true" pageId="12" pageNumber="31">
|
||
Tête (
|
||
<figureCitation id="808D2A07FF98746DFF1FFF35FF6DFF1A" box="[170,249,216,242]" captionStart="FIG" captionStartId="33.[113,124,1635,1652]" captionTargetBox="[117,1206,214,1443]" captionTargetId="figure-14@33.[117,1206,214,1443]" captionTargetPageId="33" captionText="FIG. 6. — Larves de Scarabaeoidea de Madagascar: A-F, tête; A, Anochilia bifida (Olivier, 1789); B, Hexodon unicolor (Olivier, 1789); C, sp.D2; D, Paramorphochelus cornutus (Nonfried,1892); E, Empecta scutata (Fairmaire,1901); F, sp.S2; G, H, épipharynx; G, Bricoptis variolosa (Gory & Percheron, 1833); H, Celidota parvula (Janson, 1881). Échelles: A-C, 2 mm; D-F, 1 mm; G, H, 0,2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520767" httpUri="https://zenodo.org/record/4520767/files/figure.png" pageId="12" pageNumber="31">Fig. 6B</figureCitation>
|
||
)
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="18093682FF98746DFFC4FF15FE45FE79" blockId="12.[111,648,216,401]" pageId="12" pageNumber="31">Couleur brunâtre. Épicrâne avec 2 soies dorsoépicraniales et 2 soies épicraniales latéro-externes disposées symétriquement. Une soie sur chaque côté de l’angle frontal. Clypéus transverse avec une soie clypéale externe sur chaque côté.</paragraph>
|
||
<paragraph id="18093682FF98746DFFC4FE58FDD0FD67" blockId="12.[111,648,437,655]" pageId="12" pageNumber="31">
|
||
<emphasis id="2AC2EA90FF98746DFFC4FE58FF43FE27" bold="true" box="[113,215,437,463]" pageId="12" pageNumber="31">Antennes</emphasis>
|
||
(
|
||
<figureCitation id="808D2A07FF98746DFF56FE5BFED2FE39" box="[227,326,438,465]" captionStart="FIG" captionStartId="39.[113,124,1567,1584]" captionTargetBox="[118,1207,216,1486]" captionTargetId="figure-14@39.[118,1207,216,1486]" captionTargetPageId="39" captionText="FIG. 12. — Larves de Scarabaeoidea de Madagascar: A-E, hypopharynx; A, Enaria melanictera (Klug, 1833); B, Apicencya waterloti (Dewailly, 1950); C, Empecta scutata (Fairmaire, 1901); D, Hoplochelus betanimena (Künckel, 1887); E, Triodontus nitidulus (Guérin, 1844); F-O, antenne; F, Anochilia bifida (Olivier, 1789); G, Heteronychus plebeius (Klug, 1833); H, Hexodon unicolor (Olivier, 1789); I, Heteroconus paradoxus (Endrödi,1968); J, Paranodon coquerelii (Fairmaire,1871); K-M, Encya sikorai (Brenske,1891); L, M, plages sensorielles de l’article apical; N, Apicencya waterloti (Dewailly, 1950); O, Empecta scutata (Fairmaire, 1901). Échelles: A, F, G, I, J, O, 0,5 mm; E, L, M, 0,2 mm; H, K, N, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520787" httpUri="https://zenodo.org/record/4520787/files/figure.png" pageId="12" pageNumber="31">Fig. 12H</figureCitation>
|
||
). Article II approximativement deux fois plus long que les autres (les longueurs respectives des articles sont: I = 1,14; II = 2,14; III = 1,28; IV = 1). Une aire sensorielle apico-latérale sur l’avant-dernier article. Dernier article antennaire avec 5 plages sensorielles: 1 petite plage apicale ronde, 2 ventrales écartées et 2 dorsales adjacentes.
|
||
</paragraph>
|
||
<paragraph id="18093682FF98746DFFC4FD5EFEF7FC23" blockId="12.[111,648,691,971]" pageId="12" pageNumber="31">
|
||
<emphasis id="2AC2EA90FF98746DFFC4FD5EFF23FD25" bold="true" box="[113,183,691,717]" pageId="12" pageNumber="31">Labre.</emphasis>
|
||
Transverse, asymétrique. Zone médiane du bord antérieur un peu saillante, formant presque un lobe médian, légèrement différencié du côté droit, avec 4 soies antérieures externes et 2 soies antérieures internes écartées. De part et d’autre, 1 soie latéro-antérieure externe, 2 soies latéro-antérieures internes, puis 1 longue soie latérale. 2 longues soies labro-internes. 4 soies postérieures (en 2 groupes transverses de 2 soies).
|
||
</paragraph>
|
||
<paragraph id="18093682FF98746DFFC4FC1DFDE8FAE0" blockId="12.[111,648,1008,1288]" pageId="12" pageNumber="31">
|
||
<emphasis id="2AC2EA90FF98746DFFC4FC1DFF60FBE3" bold="true" box="[113,244,1008,1035]" pageId="12" pageNumber="31">Épipharynx</emphasis>
|
||
(
|
||
<figureCitation id="808D2A07FF98746DFEB0FC1DFEF4FBE3" box="[261,352,1008,1035]" captionStart="FIG" captionStartId="34.[113,124,1635,1652]" captionTargetBox="[155,1168,220,1591]" captionTargetId="figure-21@34.[155,1168,220,1591]" captionTargetPageId="34" captionText="FIG. 7. — Larves de Scarabaeoidea de Madagascar,épipharynx: A, Anochilia bifida (Olivier,1789); B, Euryomia argentea (Olivier,1789); C, Heteronychus arator rugifrons (Fairmaire, 1871); D, Heteronychus bituberculatus (Kolbe, 1900); E, F, sp. D1; G, Hexodon unicolor (Olivier, 1789); H, sp. D2. Échelles: A, B, D, E, G, 0,5 mm; C, 0,2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520771" httpUri="https://zenodo.org/record/4520771/files/figure.png" pageId="12" pageNumber="31">Fig. 7G</figureCitation>
|
||
). Haptomère formant une saillie redressée bifide, sclérifiée et émoussée. Acroparia avec de fortes soies. Aire pédiale glabre, avec un sillon anguleux, en arc de cercle vers la base. Chaetoparia dissymétriques à rangs concentriques de soies convergeant vers l’aire pédiale, plus fortes près de l’haptomère et sur la périphérie de l’aire pédiale, plus fines vers l’extérieur. Acanthoparia avec 6 à 9 courtes soies lamelliformes incurvées vers l’avant.
|
||
</paragraph>
|
||
<paragraph id="18093682FF98746DFFC4FAC1FE6BF9AD" blockId="12.[111,648,1324,1605]" pageId="12" pageNumber="31">
|
||
<emphasis id="2AC2EA90FF98746DFFC4FAC1FF63FAAE" bold="true" box="[113,247,1324,1350]" pageId="12" pageNumber="31">Mandibules</emphasis>
|
||
(
|
||
<figureCitation id="808D2A07FF98746DFEBDFAC0FEC3FAA0" box="[264,343,1325,1352]" captionStart="FIG" captionStartId="41.[113,124,1589,1606]" captionTargetBox="[161,1157,219,1557]" captionTargetId="figure-14@41.[157,1165,214,1560]" captionTargetPageId="41" captionText="FIG. 14. — Larves de Scarabaeoidea de Madagascar,mandibule: A, D, H, L, P, T, X, gauche,face dorsale; B, E, I, M, Q, U, Y, gauche, face ventrale; C, F, J, N, R, V, Z1, droite, face dorsale; G, K, O, S, W, Z2, droite, face ventrale; A-C, Bricoptis variolosa (Gory & Percheron, 1833); D-G, Anochilia bifida (Olivier, 1789); H-K, Euryomia argentea (Olivier, 1789); L-O, Heteronychus arator rugifrons (Fairmaire,1871); P-S, Heteronychus bituberculatus (Kolbe,1900); T-W, Hexodon unicolor (Olivier,1789); X-Z2, Heteroconus paradoxus (Endrödi, 1968). Échelles: A-C, H-O, 0,5 mm; D-G, P-Z2, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520795" httpUri="https://zenodo.org/record/4520795/files/figure.png" pageId="12" pageNumber="31">Fig. 14</figureCitation>
|
||
T-W). Aire scissoriale de la mandibule gauche avec une forte dent apicale, suivie d’une dent arrondie et d’une petite dent pointue. Aire scissoriale de la mandibule droite avec une forte dent apicale, suivie d’une forte dent arrondie. Mandibules gauche et droite avec chacune 6 à 8 soies baso-latérales et 2 soies dorso-externes. Sur la face ventrale des 2 mandibules, présence d’une aire stridulatoire formée de 35 bourrelets parallèles transverses.
|
||
</paragraph>
|
||
<paragraph id="18093682FF98746DFFC4F984FB08FEFB" blockId="12.[111,647,1641,1700]" lastBlockId="12.[677,1209,216,275]" pageId="12" pageNumber="31">
|
||
<emphasis id="2AC2EA90FF98746DFFC4F984FF5DF96B" bold="true" box="[113,201,1641,1667]" pageId="12" pageNumber="31">Maxilles</emphasis>
|
||
(
|
||
<figureCitation id="808D2A07FF98746DFF60F984FEA1F96C" box="[213,309,1641,1668]" captionStart="FIG" captionStartId="43.[113,124,1567,1584]" captionTargetBox="[152,1180,217,1546]" captionTargetId="figure-14@43.[151,1175,219,1547]" captionTargetPageId="43" captionText="FIG. 16. — Larves de Scarabaeoidea de Madagascar: A-D, mandibule, Triodontus nitidulus (Guérin, 1844); A, gauche, face dorsale; B, gauche, face ventrale; C, droite, face dorsale; D, droite, face ventrale; E-O, maxille, face dorsale; E, Bricoptis variolosa (Gory & Percheron, 1833); F, Celidota parvula (Janson, 1881); G, Anochilia bifida (Olivier, 1789); H, Euryomia argentea (Olivier, 1789); I, J, Heteronychus arator rugifrons (Fairmaire, 1871); J, dents stridulatoires; K, L, Heteronychus bituberculatus (Kolbe,1900); L, dents stridulatoires; M, Heteronychus plebeius (Klug,1833); N, Hexodon unicolor (Olivier,1789); O, Heteroconus paradoxus (Endrödi,1968). Échelles: A-D, N, O, 0,5 mm; E, F, H, I, K, 0,4 mm; G, 1 mm; J, 0,01 mm; L, 0,02 mm." figureDoi="http://doi.org/10.5281/zenodo.4520805" httpUri="https://zenodo.org/record/4520805/files/figure.png" pageId="12" pageNumber="31">Fig. 16N</figureCitation>
|
||
). Galea et lacinia soudées. Lacinia à uncus avec 2 dents. Galea à uncus sclérifié. Sur la face dorsale, aire stridulatoire composée de 9 à 10 dents irrégulières aux bords arrondis émoussés.
|
||
</paragraph>
|
||
<paragraph id="18093682FF98746DFD10FEDAFBCAFE18" blockId="12.[674,1210,311,496]" pageId="12" pageNumber="31">
|
||
<emphasis id="2AC2EA90FF98746DFD10FEDAFCA3FEB9" bold="true" box="[677,823,311,337]" pageId="12" pageNumber="31">Hypopharynx</emphasis>
|
||
(
|
||
<figureCitation id="808D2A07FF98746DFCF7FEDAFC36FEBA" box="[834,930,311,338]" captionStart="FIG" captionStartId="37.[113,124,1589,1606]" captionTargetBox="[146,1173,219,1547]" captionTargetId="figure-14@37.[146,1173,218,1547]" captionTargetPageId="37" captionText="FIG. 10. — Larves de Scarabaeoidea de Madagascar,hypopharynx: A, Bricoptis variolosa (Gory & Percheron,1833); B, Celidota parvula (Janson, 1881); C, Anochilia bifida (Olivier, 1789); D, Euryomia argentea (Olivier, 1789); E, Heteronychus arator rugifrons (Fairmaire, 1871); F, Heteronychus bituberculatus (Kolbe, 1900); G, Heteronychus plebeius (Klug, 1833); H, Hexodon unicolor (Olivier, 1789); I, Heteroconus paradoxus (Endrödi,1968); J, Paranodon coquerelii (Fairmaire,1871); K, Paramorphochelus cornutus (Nonfried,1892); L, Encya sikorai (Brenske, 1891). Échelles: A, J, 0,2 mm; C, D, F, G, I, 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.4520783" httpUri="https://zenodo.org/record/4520783/files/figure.png" pageId="12" pageNumber="31">Fig.10H</figureCitation>
|
||
). Glosses avec une vingtaine à une trentaine de courtes et fortes soies au milieu et à la base, ainsi qu’avec des touffes de longues soies vers l’apex et sur les côtés. Oncyli transverses, avec une dent sclérifiée postérieure subtriangulaire et une forte dent antérieure, sclérifiée et dressée.
|
||
</paragraph>
|
||
<paragraph id="18093682FF98746DFD10FDF8FD79FDC7" blockId="12.[675,1211,533,718]" box="[677,749,533,559]" pageId="12" pageNumber="31">
|
||
<emphasis id="2AC2EA90FF98746DFD10FDF8FD79FDC7" box="[677,749,533,559]" italics="true" pageId="12" pageNumber="31">Thorax</emphasis>
|
||
</paragraph>
|
||
<paragraph id="18093682FF98746DFD10FDD9FD7EFD26" blockId="12.[675,1211,533,718]" pageId="12" pageNumber="31">
|
||
<emphasis id="2AC2EA90FF98746DFD10FDD9FD73FDA6" bold="true" box="[677,743,564,590]" pageId="12" pageNumber="31">Pattes</emphasis>
|
||
(
|
||
<figureCitation id="808D2A07FF98746DFD46FDD9FCC2FDA7" box="[755,854,564,591]" captionStart="FIG" captionStartId="46.[113,124,1635,1652]" captionTargetBox="[121,1206,215,1527]" captionTargetId="figure-21@46.[117,1206,214,1542]" captionTargetPageId="46" captionText="FIG. 19. — Larves de Scarabaeoidea de Madagascar, pattes: A, Heteronychus arator rugifrons (Fairmaire, 1871); B, Heteronychus bituberculatus (Kolbe, 1900); C, Heteronychus plebeius (Klug, 1833); D, Hexodon unicolor (Olivier, 1789); E, Heteroconus paradoxus (Endrödi, 1968); F, Paranodon coquerelii (Fairmaire, 1871). Échelles: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520815" httpUri="https://zenodo.org/record/4520815/files/figure.png" pageId="12" pageNumber="31">Fig. 19D</figureCitation>
|
||
). Longues, de longueur croissante de P1 à P3, toutes de structure analogue: coxa, trochanter, fémur, tibiotarse, griffe forte, longue, aiguë, anguleuse à la base, incurvée et sclérifiée à l’apex.
|
||
</paragraph>
|
||
<paragraph id="18093682FF98746DFD10FD1FFC93FCE4" blockId="12.[674,1211,754,1035]" box="[677,775,754,780]" pageId="12" pageNumber="31">
|
||
<emphasis id="2AC2EA90FF98746DFD10FD1FFC93FCE4" box="[677,775,754,780]" italics="true" pageId="12" pageNumber="31">Abdomen</emphasis>
|
||
</paragraph>
|
||
<paragraph id="18093682FF98746DFD10FCFFFC1FFBE3" blockId="12.[674,1211,754,1035]" pageId="12" pageNumber="31">
|
||
<emphasis id="2AC2EA90FF98746DFD10FCFFFD7FFCC4" bold="true" box="[677,747,786,812]" pageId="12" pageNumber="31">Raster</emphasis>
|
||
(
|
||
<figureCitation id="808D2A07FF98746DFD4FFCFFFCDAFCC5" box="[762,846,786,813]" captionStart="FIG" captionStartId="31.[113,124,1635,1652]" captionTargetBox="[124,1194,218,1601]" captionTargetId="figure-14@31.[106,1195,214,1601]" captionTargetPageId="31" captionText="FIG. 4. — Larves de Scarabaeoidea de Madagascar,raster: A, sp. D1; B, Hexodon unicolor (Olivier, 1789); C, sp. D2; D, Heteroconus paradoxus (Endrödi, 1968); E, Paranodon coquerelii (Fairmaire, 1871); F, Paramorphochelus cornutus (Nonfried, 1892); G, sp. M1; H, Encya sikorai (Brenske, 1891). Échelles: A-D, H, 2 mm; E-G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520763" httpUri="https://zenodo.org/record/4520763/files/figure.png" pageId="12" pageNumber="31">Fig. 4B</figureCitation>
|
||
). Du côté dorsal de la fente anale transverse, nombreuses soies courtes et épaisses, orientées vers l’avant, mêlées de quelques longues soies plus fines. Une étroite bande glabre le long de la fente anale du côté dorsal.Teges avec de nombreuses et très épaisses soies aplaties et recourbées à l’extrémité du côté ventral de la fente anale. Barbula avec de rares et longues soies.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |