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<taxonomicName id="2BD64D346D4FFFA2B8C7BE62F0D5281E" ID-CoL="7SZMD" authority="C. L. Koch, 1844" authorityName="C. L. Koch" authorityYear="1844" box="[151,738,1085,1111]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="onustus" subGenus="Adamaeus">
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<emphasis id="DEA2EAA56D4FFFA2B8C7BE62F33E281E" bold="true" box="[151,265,1085,1111]" italics="true" pageId="2" pageNumber="3">Damaeus</emphasis>
(
<emphasis id="DEA2EAA56D4FFFA2B94ABE62F3AE281E" bold="true" box="[282,409,1085,1111]" italics="true" pageId="2" pageNumber="3">Adamaeus</emphasis>
) 
<emphasis id="DEA2EAA56D4FFFA2B9FABE62F03F281E" bold="true" box="[426,520,1085,1111]" italics="true" pageId="2" pageNumber="3">onustus</emphasis>
<bibRefCitation id="88474B466D4FFFA2BA40BE62F0D5281E" author="Koch" box="[528,738,1085,1111]" pageId="2" pageNumber="3" refString="Koch, C. L. (1844) Deutschlands Crustaceen, Myriapoden und Arachniden. Regensburg, vol. 33 - 40." type="book chapter" year="1844">C. L. Koch, 1844</bibRefCitation>
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(
<figureCitation id="74ED2A326D4FFFA2B8CFBE00F33C283E" box="[159,267,1119,1143]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStartId-0="5.[151,250,1991,2014]" captionStartId-1="6.[151,250,1987,2010]" captionStartId-2="7.[151,250,2010,2033]" captionStartId-3="8.[151,250,1993,2016]" captionStartId-4="9.[151,250,1962,1985]" captionStartId-5="10.[151,250,2013,2036]" captionStartId-6="11.[151,250,1887,1910]" captionTargetBox-0="[249,1330,198,1955]" captionTargetBox-1="[206,1392,212,1928]" captionTargetBox-2="[197,1398,205,1972]" captionTargetBox-3="[319,1262,205,1961]" captionTargetBox-4="[306,1258,206,1913]" captionTargetBox-5="[217,1333,197,1974]" captionTargetBox-6="[165,1427,228,1848]" captionTargetId-0="figure@5.[243,1345,193,1970]" captionTargetId-1="figure@6.[177,1413,193,1959]" captionTargetId-2="figure@7.[175,1411,193,1983]" captionTargetId-3="figure@8.[315,1267,193,1968]" captionTargetId-4="figure@9.[295,1272,193,1934]" captionTargetId-5="figure@10.[217,1351,193,1984]" captionTargetId-6="figure@11.[151,1436,193,1856]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="10" captionTargetPageId-6="11" captionText-0="FIGURE 1. Damaeus onustus. (A) Adult, dorsal aspect, legs partially drawn, scale bar 100 μm; (B) central part of seta la." captionText-1="FIGURE 2. Damaeus onustus, adult, scale bars 100 μm. (A) Lateral aspect; (B) anogenital region. Legs partially drawn." captionText-2="FIGURE 3. Damaeus onustus, larva, dorsal aspect, scale bar 100 μm. Legs partially drawn." captionText-3="FIGURE 4. Damaeus onustus, scale bars 100 μm. (A) Anal region of larva; (B) anogenital region of protonymph." captionText-4="FIGURE 5. Damaeus onustus, anogenital region, scale bars 100 μm. (A) Deutonymph; (B) tritonymph." captionText-5="FIGURE 6. Damaeus onustus. (A) Tritonymph, dorsal aspect, legs partially drawn, scale bar 100 μm; (B) cornicle." captionText-6="FIGURE 7. Damaeus onustus, tritonymph, lateral aspect, legs partially drawn, scale bar 100 μm. (A) With humus protective camouflage; (B) without this camouflage." httpUri-0="https://zenodo.org/record/202940/files/figure.png" httpUri-1="https://zenodo.org/record/202941/files/figure.png" httpUri-2="https://zenodo.org/record/202942/files/figure.png" httpUri-3="https://zenodo.org/record/202944/files/figure.png" httpUri-4="https://zenodo.org/record/202945/files/figure.png" httpUri-5="https://zenodo.org/record/202946/files/figure.png" httpUri-6="https://zenodo.org/record/202947/files/figure.png" pageId="2" pageNumber="3">Figs. 1–7</figureCitation>
, 
<figureCitation id="74ED2A326D4FFFA2B946BE00F314283E" box="[278,291,1119,1143]" captionStart="FIGURE 8" captionStartId="12.[151,250,1633,1656]" captionTargetBox="[161,1435,202,1598]" captionTargetId="figure@12.[151,1436,193,1604]" captionTargetPageId="12" captionText="FIGURE 8. Tarsus I and part of tibia of tritonymph, paraxial aspect, scale bars 10 μm. (A) Damaeus onustus; (B) D. clavipes; (C) Kunstidamaeus tecticola." httpUri="https://zenodo.org/record/202948/files/figure.png" pageId="2" pageNumber="3">8</figureCitation>
A)
</paragraph>
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<taxonomicName id="2BD64D346D4FFFA2B8C7BEF6F3CA28F4" ID-CoL="7SZMD" authority="Grandjean 1943" authorityName="Grandjean" authorityYear="1943" box="[151,509,1192,1214]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="onustus">
<emphasis id="DEA2EAA56D4FFFA2B8C7BEF6F37D28F7" box="[151,330,1193,1214]" italics="true" pageId="2" pageNumber="3">Damaeus onustus</emphasis>
: 
<bibRefCitation id="88474B466D4FFFA2B907BEF7F3CA28F4" author="Grandjean" box="[343,509,1192,1214]" pageId="2" pageNumber="3" refString="Grandjean, F. (1943) Observations sur les Oribates. (16. serie). Bulletin du Museum D'Histoire naturelle, Paris, (2) 15, 410 - 417." type="journal article" year="1943">Grandjean 1943</bibRefCitation>
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; 
<bibRefCitation id="88474B466D4FFFA2BA57BEF6F09728F7" author="Hammen" box="[519,672,1192,1214]" pageId="2" pageNumber="3" refString="Hammen, L. van der (1952) The Oribatei (Acari) of the Netherlands. Zoologische Verhandelingen, Leiden, 17, 1 - 139." type="journal article" year="1952">Hammen 1952</bibRefCitation>
; 
<bibRefCitation id="88474B466D4FFFA2BAFABEF7F10E28F7" author="Sellnick" box="[682,825,1192,1214]" pageId="2" pageNumber="3" refString="Sellnick, M. (1960) Formenkreis: Hornmilben, Oribatei. In: Brohmers, P., Ehrmann, P., Ulmerm G. (Eds.): Die Tierwelt Mitteleuropas. Vol. 3, 4. Lief. (Erganzung). Quelle &amp; Meyer, Leipzig, pp. 45 - 134." type="journal article" year="1960">Sellnick 1960</bibRefCitation>
; 
<bibRefCitation id="88474B466D4FFFA2BB13BEF7F65128F7" author="Bulanova-Zachvatkina" box="[835,1126,1192,1214]" pageId="2" pageNumber="3" refString="Bulanova-Zachvatkina, E. M. (1967) Oribatid mites - Oribatida. Vysshaya Shkola Publishers, Moscow, pp. 1 - 254. (In Russian)." type="book chapter" year="1967">Bulanova-Zachvatkina 1967</bibRefCitation>
; 
<bibRefCitation id="88474B466D4FFFA2BC20BEF7F70828F4" author="Marshall" box="[1136,1343,1192,1214]" pageId="2" pageNumber="3" refString="Marshall, V. G., Reeves, R. M. &amp; Norton, R. A. (1987) Catalogue of the Oribatida (Acari) of Continental United States and Canada. Memoirs of the Entomological Society of Canada, 139, 1 - 418 + 6." type="journal article" year="1987">
<collectingCountry id="94C176276D4FFFA2BC20BEF7F6FD28F7" box="[1136,1226,1192,1214]" name="Marshall Islands" pageId="2" pageNumber="3">Marshall</collectingCountry>
<emphasis id="DEA2EAA56D4FFFA2BC81BEF6F6CB28F7" box="[1233,1276,1192,1214]" italics="true" pageId="2" pageNumber="3">et al</emphasis>
. 1987
</bibRefCitation>
. 
<taxonomicName id="2BD64D346D4FFFA2B8C7BE98F3EF2895" authority="Berlese, 1887" authorityName="Berlese" authorityYear="1887" box="[151,472,1222,1245]" class="Arachnida" family="Damaeidae" genus="Belba" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="geniculata">
<emphasis id="DEA2EAA56D4FFFA2B8C7BE98F3742894" box="[151,323,1223,1245]" italics="true" pageId="2" pageNumber="3">Belba geniculata</emphasis>
<bibRefCitation id="88474B466D4FFFA2B919BE99F3EF2895" author="Berlese" box="[329,472,1222,1244]" pageId="2" pageNumber="3" refString="Berlese, A. (1887) Acari, Myriapoda et Scorpiones hucusque in Italia reperta. Portici, Padova, vol. 36 - 44." type="book chapter" year="1887">Berlese, 1887</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="EC6936B76D4FFFA2B8C7BEB9F0252950" blockId="2.[151,1347,1192,1429]" pageId="2" pageNumber="3">
<taxonomicName id="2BD64D346D4FFFA2B8C7BEB9F03928B2" authority="Michael 1888" authorityName="Michael" authorityYear="1888" box="[151,526,1253,1275]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="geniculatus">
<emphasis id="DEA2EAA56D4FFFA2B8C7BEB9F34628B2" box="[151,369,1253,1275]" italics="true" pageId="2" pageNumber="3">Damaeus geniculatus</emphasis>
: 
<bibRefCitation id="88474B466D4FFFA2B92FBEBAF03928B2" author="Michael" box="[383,526,1253,1275]" pageId="2" pageNumber="3" refString="Michael, A. D. (1888) British Oribatidae. Vol 1 + 2. Ray Society, London, pp. 1 - 657." type="journal article" year="1888">Michael 1888</bibRefCitation>
</taxonomicName>
; 
<bibRefCitation id="88474B466D4FFFA2BA48BEBAF08728B2" author="Tragardh" box="[536,688,1253,1275]" pageId="2" pageNumber="3" refString="Tragardh, I. (1904) Monographie der arktischen Acariden. Fauna Arctica, 4 (1), 1 - 78." type="journal article" year="1904">Trägårdh 1904</bibRefCitation>
; 
<bibRefCitation id="88474B466D4FFFA2BAEABEBAF1DE28B2" author="Bulanova-Zachvatkina" box="[698,1001,1253,1275]" pageId="2" pageNumber="3" refString="Bulanova-Zachvatkina, E. M. (1957 b) Oribatid mites Damaeidae Berl. Genus Epidamaeus, gen. n. (Acariformes, Oribatei). Part II. Zoologicheskij Zhurnal, 36, 1792 - 1801. (In Russian)." type="journal article" year="1957" yearSuffix="b">Bulanova-Zachvatkina 1957b</bibRefCitation>
. 
<taxonomicName id="2BD64D346D4FFFA2B8C7BF5BF0392950" authority="Oudemans 1900" authorityName="Oudemans" authorityYear="1900" box="[151,526,1283,1306]" class="Arachnida" family="Damaeidae" genus="Oribata" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="geniculata">
<emphasis id="DEA2EAA56D4FFFA2B8C7BF5BF36F2953" box="[151,344,1284,1306]" italics="true" pageId="2" pageNumber="3">Oribata geniculata</emphasis>
: 
<bibRefCitation id="88474B466D4FFFA2B935BF5CF0392950" author="Oudemans" box="[357,526,1283,1305]" pageId="2" pageNumber="3" refString="Oudemans, A. C. (1900) New list of Dutch Acari. lst part. Tijdschrift voor Entomologie, 43, 150 - 171." type="journal article" year="1900">Oudemans 1900</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="EC6936B76D4FFFA2B8C7BF7CF0B22971" blockId="2.[151,1347,1192,1429]" box="[151,645,1314,1336]" pageId="2" pageNumber="3">
<taxonomicName id="2BD64D346D4FFFA2B8C7BF7CF3D02971" authority="Kulczynski 1902" authorityName="Kulczynski" authorityYear="1902" box="[151,487,1314,1336]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="geniculatus">
<collectingCountry id="94C176276D4FFFA2B8C7BF7CF29E2971" box="[151,169,1315,1336]" name="Iceland" pageId="2" pageNumber="3">
<emphasis id="DEA2EAA56D4FFFA2B8C7BF7CF29E2971" box="[151,169,1315,1336]" italics="true" pageId="2" pageNumber="3">O</emphasis>
</collectingCountry>
. 
<emphasis id="DEA2EAA56D4FFFA2B8E6BF7DF31D2971" box="[182,298,1314,1336]" italics="true" pageId="2" pageNumber="3">geniculatus</emphasis>
: 
<bibRefCitation id="88474B466D4FFFA2B968BF7DF3D02971" author="Kulczynski" box="[312,487,1314,1336]" pageId="2" pageNumber="3" refString="Kulczynski, M. (1902) Species oribatinarum (Oudms.) (Damaeinarum Michael) in Galicia collectae. Bulletin International de L'Academie des Sciences de Cracovie, Classe des sciences mathematiques et naturelles, 2, 89 - 96." type="journal article" year="1902">Kulczyński 1902</bibRefCitation>
</taxonomicName>
; 
<bibRefCitation id="88474B466D4FFFA2B9A1BF7DF0B62971" author="Sellnick" box="[497,641,1314,1336]" pageId="2" pageNumber="3" refString="Sellnick, M. (1928) Formenkreis: Hornmilben, Oribatei. In: Brohmers, P., Ehrmann, P., Ulmer, G. (Eds.): Die Tierwelt Mitteleuropas. Vol. 3, 4. Lief. (Teil 9). Quelle &amp; Meyer, Leipzig, pp. 1 - 41." type="journal article" year="1928">Sellnick 1928</bibRefCitation>
.
</paragraph>
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<taxonomicName id="2BD64D346D4FFFA2B8C7BF1EF3CB291E" authority="Oudemans 1929" authorityName="Oudemans" authorityYear="1929" box="[151,508,1345,1367]" class="Arachnida" family="Damaeidae" genus="Belba" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="geniculosa">
<emphasis id="DEA2EAA56D4FFFA2B8C7BF1EF371291E" box="[151,326,1345,1367]" italics="true" pageId="2" pageNumber="3">Belba geniculosa</emphasis>
: 
<bibRefCitation id="88474B466D4FFFA2B903BF1EF3CB291E" author="Oudemans" box="[339,508,1345,1367]" pageId="2" pageNumber="3" refString="Oudemans, A. C. (1929) Acarologische Aanteekeningen, XCVII. Entomologische Berichte, 7, 448 - 455." type="journal article" year="1929">Oudemans 1929</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="EC6936B76D4FFFA2B8C7BF3EF0D029DD" blockId="2.[151,1347,1192,1429]" pageId="2" pageNumber="3">
<taxonomicName id="2BD64D346D4FFFA2B8C7BF3EF06B293C" ID-CoL="7SZMD" authority="Norton 1978" authorityName="Norton" authorityYear="1978" box="[151,604,1376,1398]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="onustus" subGenus="Adamaeus">
<emphasis id="DEA2EAA56D4FFFA2B8C7BF3EF2C0293F" box="[151,247,1377,1398]" italics="true" pageId="2" pageNumber="3">Damaeus</emphasis>
(
<emphasis id="DEA2EAA56D4FFFA2B955BF3FF359293F" box="[261,366,1376,1398]" italics="true" pageId="2" pageNumber="3">Adamaeus</emphasis>
) 
<emphasis id="DEA2EAA56D4FFFA2B92DBF3EF3FD293F" box="[381,458,1377,1398]" italics="true" pageId="2" pageNumber="3">onustus</emphasis>
: 
<bibRefCitation id="88474B466D4FFFA2B988BF3EF06B293C" author="Norton" box="[472,604,1376,1398]" pageId="2" pageNumber="3" refString="Norton, R. A. (1978) Generic concepts in the Damaeidae (Acari: Oribatei). I. Three new taxa based on species of Nathan Banks. Acarologia, 20 (4), 603 - 621." type="journal article" year="1978">Norton 1978</bibRefCitation>
</taxonomicName>
; 
<bibRefCitation id="88474B466D4FFFA2BA36BF3FF162293C" author="Balogh" box="[614,853,1376,1398]" pageId="2" pageNumber="3" refString="Balogh, J. &amp; Balogh, P. (1992) The oribatid mites genera of the world. The Hungarian National Museum Press, Budapest, vol. 1, 1 - 263, vol. 2,1 - 375." type="journal article" year="1992">Balogh &amp; Balogh 1992</bibRefCitation>
; 
<bibRefCitation id="88474B466D4FFFA2BB0FBF3FF626293C" author="Perez-Inigo" box="[863,1041,1376,1398]" pageId="2" pageNumber="3" refString="Perez-Inigo, C. (1997) Acari. Oribatei. Gymnonota I. In: Ramos, A. et al. (Eds.): Fauna Iberica. Museo de Ciencias Naturales, Madrid, vol. 9, pp. 1 - 373." type="journal article" year="1997">Pérez-Íñigo 1997</bibRefCitation>
; 
<bibRefCitation id="88474B466D4FFFA2BC4BBF3FF6AC293C" author="Subias" box="[1051,1179,1376,1398]" pageId="2" pageNumber="3" refString="Subias, L. S. (2004) Listado sistematico, sinonimico y biogeografico de los Acaros Oribatidos (Acariformes, Oribatida) del mundo (1758 - 2002). Graellsia, 60, 3 - 305." type="journal article" year="2004">Subías 2004</bibRefCitation>
; 
<bibRefCitation id="88474B466D4FFFA2BCF5BF3FF72F293F" author="Miko" box="[1189,1304,1376,1398]" pageId="2" pageNumber="3" refString="Miko, L. (2006) Damaeidae. In: Weigmann, G.: Hornmilben (Oribatida), Die Tierwelt Deutschlands, Teil 76. Dahl, F. (Ed.), Goecke &amp; Evers, Keltern, pp. 179 - 207." type="journal article" year="2006">Miko 2006</bibRefCitation>
. 
<taxonomicName id="2BD64D346D4FFFA2B8C7BF20F3D629DD" ID-CoL="969df880-f097-4b35-af51-1db6763e2b20" authority="Schatz 1983" authorityName="Schatz" authorityYear="1983" box="[151,481,1406,1429]" class="Arachnida" family="Damaeidae" genus="Adamaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="onustus">
<emphasis id="DEA2EAA56D4FFFA2B8C7BF20F36429DC" box="[151,339,1407,1429]" italics="true" pageId="2" pageNumber="3">Adamaeus onustus</emphasis>
: 
<bibRefCitation id="88474B466D4FFFA2B931BF21F3D629DD" author="Schatz" box="[353,481,1406,1428]" pageId="2" pageNumber="3" refString="Schatz, H. (1983) U. - Ordn. Oribatei, Hornmilben. Catalogus faunae Austriae. Ein systematisches Verzeichnis aller auf osterreichischem Gebiet festgestellten Tierarten, 9, Osterreichische Akadademie der Wissenschaften, Wien, pp. 1 - 118." type="journal article" year="1983">Schatz 1983</bibRefCitation>
</taxonomicName>
; 
<bibRefCitation id="88474B466D4FFFA2B9BBBF21F0D429DD" author="Olszanowski" box="[491,739,1406,1429]" pageId="2" pageNumber="3" refString="Olszanowski, Z., Rajski, A. &amp; Niedbala, W. (1996) Catalogue of Polish fauna. Mites (Acari), moss mites (Oribatida). Sours Pozna n, 34 (9), 1 - 243. (In Polish)." type="journal article" year="1996">
Olszanowski 
<emphasis id="DEA2EAA56D4FFFA2BA24BFDFF09729DC" box="[628,672,1407,1429]" italics="true" pageId="2" pageNumber="3">et al</emphasis>
. 1996
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A4CC653C6D4FFFA4B8C7BF9EF34429DB" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3" type="diagnosis">
<paragraph id="EC6936B76D4FFFA2B8C7BF9EF0882B69" blockId="2.[151,1436,1473,2004]" pageId="2" pageNumber="3">
<emphasis id="DEA2EAA56D4FFFA2B8C7BF9EF3262993" bold="true" box="[151,273,1473,1498]" pageId="2" pageNumber="3">Diagnosis.</emphasis>
Adult of damaeid shape, large (1350–1600 μm), dark brown to almost black. Prodorsum subtriangular, notogaster strongly arched, almost spherical (
<figureCitation id="74ED2A326D4FFFA2BAF4BFBAF0DF29B4" box="[676,744,1509,1533]" captionStart="FIGURE 1" captionStartId="5.[151,250,1991,2014]" captionTargetBox="[249,1330,198,1955]" captionTargetId="figure@5.[243,1345,193,1970]" captionTargetPageId="5" captionText="FIGURE 1. Damaeus onustus. (A) Adult, dorsal aspect, legs partially drawn, scale bar 100 μm; (B) central part of seta la." httpUri="https://zenodo.org/record/202940/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
), cerotegument with short vermicular excrescences. Seta 
<emphasis id="DEA2EAA56D4FFFA2BDD6BFB9F7AB29B4" box="[1414,1436,1510,1533]" italics="true" pageId="2" pageNumber="3">in</emphasis>
shorter than 
<emphasis id="DEA2EAA56D4FFFA2B975BC55F3062A68" box="[293,305,1546,1569]" italics="true" pageId="2" pageNumber="3">c</emphasis>
1 (
<tableCitation id="A154030C6D4FFFA2B915BC56F3AD2A68" box="[325,410,1545,1569]" captionStart="TABLE 1" captionStartId="3.[151,235,290,311]" captionText="TABLE 1. Measurements of some morphological characters of juvenile stages of Damaeus onustus, D. clavipes, and Kunstidamaeus tecticola (mean measurements of 4 – 10 specimens in μm): larva (L), protonymph (PN), deutonymph (DN), tritonymph (TN) and adult (AD), nd – not developed." pageId="2" pageNumber="3">Table 1</tableCitation>
), seta 
<emphasis id="DEA2EAA56D4FFFA2B9B6BC55F3C82A68" box="[486,511,1546,1569]" italics="true" pageId="2" pageNumber="3">ex</emphasis>
short (
<figureCitation id="74ED2A326D4FFFA2BA00BC56F0A32A68" box="[592,660,1545,1569]" captionStart="FIGURE 2" captionStartId="6.[151,250,1987,2010]" captionTargetBox="[206,1392,212,1928]" captionTargetId="figure@6.[177,1413,193,1959]" captionTargetPageId="6" captionText="FIGURE 2. Damaeus onustus, adult, scale bars 100 μm. (A) Lateral aspect; (B) anogenital region. Legs partially drawn." httpUri="https://zenodo.org/record/202941/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
A). Sensillus approximately as long as seta 
<emphasis id="DEA2EAA56D4FFFA2BCD3BC55F6B82A68" box="[1155,1167,1546,1569]" italics="true" pageId="2" pageNumber="3">c</emphasis>
1. Apophyses 
<emphasis id="DEA2EAA56D4FFFA2BD7BBC55F70E2A68" box="[1323,1337,1546,1569]" italics="true" pageId="2" pageNumber="3">P</emphasis>
, 
<emphasis id="DEA2EAA56D4FFFA2BD18BC55F7A22A68" box="[1352,1429,1546,1569]" italics="true" pageId="2" pageNumber="3">Sp, Da</emphasis>
, 
<emphasis id="DEA2EAA56D4FFFA2B8C7BC6EF2812A01" box="[151,182,1585,1608]" italics="true" pageId="2" pageNumber="3">Ba</emphasis>
and 
<emphasis id="DEA2EAA56D4FFFA2B8BBBC6EF33D2A01" box="[235,266,1585,1608]" italics="true" pageId="2" pageNumber="3">Bp</emphasis>
well-developed, 
<emphasis id="DEA2EAA56D4FFFA2B99BBC6EF3EB2A01" box="[459,476,1585,1608]" italics="true" pageId="2" pageNumber="3">P</emphasis>
large, 
<emphasis id="DEA2EAA56D4FFFA2BA77BC6EF0742A01" box="[551,579,1585,1608]" italics="true" pageId="2" pageNumber="3">Sp</emphasis>
short, truncate, 
<emphasis id="DEA2EAA56D4FFFA2BAA7BC6EF12E2A01" box="[759,793,1585,1608]" italics="true" pageId="2" pageNumber="3">Da</emphasis>
, 
<emphasis id="DEA2EAA56D4FFFA2BB76BC6EF1722A01" box="[806,837,1585,1608]" italics="true" pageId="2" pageNumber="3">Ba</emphasis>
and 
<emphasis id="DEA2EAA56D4FFFA2BB2BBC6EF1AD2A01" box="[891,922,1585,1608]" italics="true" pageId="2" pageNumber="3">Bp</emphasis>
rounded, apophyses 
<emphasis id="DEA2EAA56D4FFFA2BCD4BC6EF6912A01" box="[1156,1190,1585,1608]" italics="true" pageId="2" pageNumber="3">Dp</emphasis>
and 
<emphasis id="DEA2EAA56D4FFFA2BC8CBC6EF6CF2A01" box="[1244,1272,1585,1608]" italics="true" pageId="2" pageNumber="3">Sa</emphasis>
weakly developed. Spinae adnatae (
<emphasis id="DEA2EAA56D4FFFA2B9CEBC0AF39E2A25" box="[414,425,1621,1644]" italics="true" pageId="2" pageNumber="3">s</emphasis>
. 
<emphasis id="DEA2EAA56D4FFFA2B9E0BC0AF3892A25" box="[432,446,1621,1644]" italics="true" pageId="2" pageNumber="3">a</emphasis>
.) relatively long, narrow, cone-shaped, and directed anteriorly, discidium (
<taxonomicName id="2BD64D346D4FFFA2BD70BC0AF7702A25" box="[1312,1351,1621,1644]" class="Insecta" family="Hesperiidae" genus="Dis" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="DEA2EAA56D4FFFA2BD70BC0AF7702A25" box="[1312,1351,1621,1644]" italics="true" pageId="2" pageNumber="3">Dis</emphasis>
</taxonomicName>
) rather short. Notogaster with 11 pairs of strong setae, nine in two longitudinal rows; anterior pair directed anteriorly, approximately as long as sensillus, other setae curved outside, length decreasing from anterior to posterior, setae of 
<emphasis id="DEA2EAA56D4FFFA2B8C7BC9EF2922A91" box="[151,165,1729,1752]" italics="true" pageId="2" pageNumber="3">p</emphasis>
-series rather short (
<figureCitation id="74ED2A326D4FFFA2B9DEBC9FF3E22A91" box="[398,469,1728,1752]" captionStart="FIGURE 2" captionStartId="6.[151,250,1987,2010]" captionTargetBox="[206,1392,212,1928]" captionTargetId="figure@6.[177,1413,193,1959]" captionTargetPageId="6" captionText="FIGURE 2. Damaeus onustus, adult, scale bars 100 μm. (A) Lateral aspect; (B) anogenital region. Legs partially drawn." httpUri="https://zenodo.org/record/202941/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
B). Coxisternal apophyses 
<emphasis id="DEA2EAA56D4FFFA2BB5ABC9FF1002A91" box="[778,823,1728,1752]" italics="true" pageId="2" pageNumber="3">E2a</emphasis>
, 
<emphasis id="DEA2EAA56D4FFFA2BB18BC9EF1562A91" box="[840,865,1729,1752]" italics="true" pageId="2" pageNumber="3">Va</emphasis>
and 
<emphasis id="DEA2EAA56D4FFFA2BBF1BC9EF1F72A91" box="[929,960,1729,1752]" italics="true" pageId="2" pageNumber="3">Vp</emphasis>
well-developed, rounded, apophysis 
<emphasis id="DEA2EAA56D4FFFA2BD3FBC9FF7AB2A91" box="[1391,1436,1728,1752]" italics="true" pageId="2" pageNumber="3">E2p</emphasis>
weakly developed. Formulae of leg setae and solenidia given in 
<tableCitation id="A154030C6D4FFFA2BBD2BCBBF1ED2AB5" box="[898,986,1764,1788]" captionStart="TABLE 2" captionStartId="14.[151,235,152,173]" captionText="TABLE 2. Formulae of leg setae (trochanter to tarsus, solenidia in parentheses) in developmental stages of Damaeus onustus," pageId="2" pageNumber="3">Table 2</tableCitation>
, dorsal coupled setae 
<emphasis id="DEA2EAA56D4FFFA2BC8BBCBBF6DE2AB5" box="[1243,1257,1764,1788]" italics="true" pageId="2" pageNumber="3">d</emphasis>
usually lost on genua I–III and tibiae I–IV. Legs monodactylous.
</paragraph>
<paragraph id="EC6936B76D4FFFA3B895BD73F07B2CBE" blockId="2.[151,1436,1473,2004]" lastBlockId="3.[151,1437,151,247]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
Juveniles slimmer than adult, elongated, unpigmented, legs slightly darker and approximately as long as body (
<tableCitation id="A154030C6D4FFFA2B8CEBD0FF2C62B21" box="[158,241,1872,1896]" captionStart="TABLE 1" captionStartId="3.[151,235,290,311]" captionText="TABLE 1. Measurements of some morphological characters of juvenile stages of Damaeus onustus, D. clavipes, and Kunstidamaeus tecticola (mean measurements of 4 – 10 specimens in μm): larva (L), protonymph (PN), deutonymph (DN), tritonymph (TN) and adult (AD), nd – not developed." pageId="2" pageNumber="3">Table 1</tableCitation>
). Prodorsal setae 
<emphasis id="DEA2EAA56D4FFFA2B9E9BD0EF3F82B21" box="[441,463,1873,1896]" italics="true" pageId="2" pageNumber="3">ro</emphasis>
and 
<emphasis id="DEA2EAA56D4FFFA2BA56BD0FF02D2B21" box="[518,538,1872,1896]" italics="true" pageId="2" pageNumber="3">le</emphasis>
relatively long, 
<emphasis id="DEA2EAA56D4FFFA2BA80BD0FF0D32B21" box="[720,740,1872,1896]" italics="true" pageId="2" pageNumber="3">le</emphasis>
more widely spaced than 
<emphasis id="DEA2EAA56D4FFFA2BC57BD0EF62A2B21" box="[1031,1053,1873,1896]" italics="true" pageId="2" pageNumber="3">ro</emphasis>
; setae 
<emphasis id="DEA2EAA56D4FFFA2BC3BBD0EF6B62B21" box="[1131,1153,1873,1896]" italics="true" pageId="2" pageNumber="3">in</emphasis>
and 
<emphasis id="DEA2EAA56D4FFFA2BCE6BD0EF6F82B21" box="[1206,1231,1873,1896]" italics="true" pageId="2" pageNumber="3">ex</emphasis>
short. Bothridium large, rounded, with funnel-like rim, sensillus setiform, finely barbed. Gastronotum of larva slightly elongated, truncate posteriorly; gastronotum of nymphs oval, rounded posteriorly. Most dorsal gastronotal setae long, curved, finely barbed, with flagellate ends; in larva setae raised, and inserted on large apophyses, in nymphs setae curved ventrally, and inserted on small apophyses. Prodorsum and gastronotum covered with granular cerotegument, covering also legs and setae. Cornicle thick, short, anteromedial to setae 
<emphasis id="DEA2EAA56D4EFFA3BB9CBAE3F1D52C9D" box="[972,994,188,212]" italics="true" pageId="3" pageNumber="4">la</emphasis>
. Nymphs carry compact humus mass attached to exuviae of previous instars.
</paragraph>
<caption id="B8A9663F6D4EFFA3B8C7BB7DF0092D3F" pageId="3" pageNumber="4">
<paragraph id="EC6936B76D4EFFA3B8C7BB7DF0092D3F" blockId="3.[151,1437,290,1678]" pageId="3" pageNumber="4">
<emphasis id="DEA2EAA56D4EFFA3B8C7BB7DF3322D71" bold="true" box="[151,261,290,312]" pageId="3" pageNumber="4">TABLE 1.</emphasis>
Measurements of some morphological characters of juvenile stages of 
<taxonomicName id="2BD64D346D4EFFA3BBB2BB7CF6A12D71" box="[994,1174,291,312]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="onustus">
<emphasis id="DEA2EAA56D4EFFA3BBB2BB7CF6A12D71" box="[994,1174,291,312]" italics="true" pageId="3" pageNumber="4">Damaeus onustus</emphasis>
</taxonomicName>
, 
<taxonomicName id="2BD64D346D4EFFA3BCF4BB7CF7202D71" box="[1188,1303,290,312]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="clavipes">
<emphasis id="DEA2EAA56D4EFFA3BCF4BB7CF6812D71" box="[1188,1206,291,312]" italics="true" pageId="3" pageNumber="4">D</emphasis>
. 
<emphasis id="DEA2EAA56D4EFFA3BC94BB7DF7202D71" box="[1220,1303,290,312]" italics="true" pageId="3" pageNumber="4">clavipes</emphasis>
</taxonomicName>
, and 
<taxonomicName id="2BD64D346D4EFFA3BD01BB7CF3792D1E" class="Arachnida" family="Damaeidae" genus="Kunstidamaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="tecticola">
<emphasis id="DEA2EAA56D4EFFA3BD01BB7CF3792D1E" italics="true" pageId="3" pageNumber="4">Kunstidamaeus tecticola</emphasis>
</taxonomicName>
(mean measurements of 4–10 specimens in μm): larva (L), protonymph (PN), deutonymph (DN), tritonymph (TN) and adult (AD), nd – not developed.
</paragraph>
</caption>
<paragraph id="EC6936B76D4EFFA3B8CFBBCEF77F2DEE" blockId="3.[151,1437,290,1678]" box="[159,1352,401,423]" pageId="3" pageNumber="4">Species Morphological character L PN DN TN AD</paragraph>
<paragraph id="EC6936B76D4EFFA3B8CFBBE4F77D2F00" blockId="3.[151,1437,290,1678]" pageId="3" pageNumber="4">
<taxonomicName id="2BD64D346D4EFFA3B8CFBBE4F3F92D99" authority="Body" authorityName="Body" box="[159,462,442,464]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="onustus">
<emphasis id="DEA2EAA56D4EFFA3B8CFBBE4F2862D99" box="[159,177,443,464]" italics="true" pageId="3" pageNumber="4">D</emphasis>
. 
<emphasis id="DEA2EAA56D4EFFA3B8EEBBE4F33C2D99" box="[190,267,443,464]" italics="true" pageId="3" pageNumber="4">onustus</emphasis>
Body
</taxonomicName>
length 611 829 994 1271 1434 Body width 304 423 553 701 929 Length of: seta 
<emphasis id="DEA2EAA56D4EFFA3BAD1B852F0A22E6A" box="[641,661,525,547]" italics="true" pageId="3" pageNumber="4">la</emphasis>
128 165 198 231 205 seta 
<emphasis id="DEA2EAA56D4EFFA3BAD1B867F0A22E04" box="[641,661,568,589]" italics="true" pageId="3" pageNumber="4">in</emphasis>
8 
<date id="986810776D4EFFA3BBCFB868F68C2E05" box="[927,1211,566,588]" pageId="3" pageNumber="4" value="1916-12-16">12 16 16</date>
80 seta 
<emphasis id="DEA2EAA56D4EFFA3BAD1B83EF0A02E3F" box="[641,663,609,630]" italics="true" pageId="3" pageNumber="4">ex</emphasis>
8 
<date id="986810776D4EFFA3BBCFB83FF68C2E3F" box="[927,1211,608,630]" pageId="3" pageNumber="4" value="1916-12-16">12 16 16</date>
12 seta 
<emphasis id="DEA2EAA56D4EFFA3BAD1B8D5F0BB2ED6" box="[641,652,650,671]" italics="true" pageId="3" pageNumber="4">c</emphasis>
1 295 289 290 292 239 seta 
<emphasis id="DEA2EAA56D4EFFA3BAD1B8E9F0BB2E82" box="[641,652,694,715]" italics="true" pageId="3" pageNumber="4">c</emphasis>
3 93 106 116 119 lost leg IV 722 962 1174 1434 1712 genital opening nd 96 139 205 244 anal opening 133 169 195 264 248
</paragraph>
<paragraph id="EC6936B76D4EFFA3B8CFB901F77D28A2" blockId="3.[151,1437,290,1678]" pageId="3" pageNumber="4">
<taxonomicName id="2BD64D346D4EFFA3B8CFB901F3F92F3A" authority="Body" authorityName="Body" box="[159,462,861,883]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="clavipes">
<emphasis id="DEA2EAA56D4EFFA3B8CFB901F2862F3A" box="[159,177,862,883]" italics="true" pageId="3" pageNumber="4">D</emphasis>
. 
<emphasis id="DEA2EAA56D4EFFA3B8EEB902F3262F3A" box="[190,273,861,883]" italics="true" pageId="3" pageNumber="4">clavipes</emphasis>
Body
</taxonomicName>
length 410 462 631 848 962 Body width 198 205 324 449 644 Length of: seta 
<emphasis id="DEA2EAA56D4EFFA3BAD1B9F0F0A42F8C" box="[641,659,943,965]" italics="true" pageId="3" pageNumber="4">le</emphasis>
96 112 129 172 191 seta 
<emphasis id="DEA2EAA56D4EFFA3BAD1B985F0A22FA6" box="[641,661,986,1007]" italics="true" pageId="3" pageNumber="4">in</emphasis>
46 42 40 40 86 seta 
<emphasis id="DEA2EAA56D4EFFA3BAD1BE5CF0A02851" box="[641,663,1027,1048]" italics="true" pageId="3" pageNumber="4">ex</emphasis>
16 16 19 23 31 seta 
<emphasis id="DEA2EAA56D4EFFA3BAD1BE73F0BB2808" box="[641,652,1068,1089]" italics="true" pageId="3" pageNumber="4">c</emphasis>
1 134 248 369 492 148 seta 
<emphasis id="DEA2EAA56D4EFFA3BAD1BE06F0BB2827" box="[641,652,1113,1134]" italics="true" pageId="3" pageNumber="4">c</emphasis>
3 48 98 101 102 lost leg IV 520 650 1141 1630 1597 genital opening nd 58 79 139 190 anal opening 85 92 112 168 193
</paragraph>
<paragraph id="EC6936B76D4EFFA3B8CFBF5FF0962B1E" blockId="3.[151,1437,290,1678]" lastBlockId="3.[151,1437,1746,2034]" pageId="3" pageNumber="4">
<taxonomicName id="2BD64D346D4EFFA3B8CFBF5FF3F9295C" authority="Body" authorityName="Body" box="[159,462,1279,1301]" class="Arachnida" family="Damaeidae" genus="Kunstidamaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="tecticola">
<emphasis id="DEA2EAA56D4EFFA3B8CFBF5FF287295C" box="[159,176,1280,1301]" italics="true" pageId="3" pageNumber="4">K</emphasis>
. 
<emphasis id="DEA2EAA56D4EFFA3B8ECBEA0F324295C" box="[188,275,1279,1301]" italics="true" pageId="3" pageNumber="4">tecticola</emphasis>
Body
</taxonomicName>
length 251 316 429 514 637 Body width 128 135 198 244 429 Length of: seta 
<emphasis id="DEA2EAA56D4EFFA3BAD1BF0DF0A42921" box="[641,659,1362,1384]" italics="true" pageId="3" pageNumber="4">le</emphasis>
42 62 72 77 120 seta 
<emphasis id="DEA2EAA56D4EFFA3BAD1BF23F0A229D8" box="[641,661,1404,1425]" italics="true" pageId="3" pageNumber="4">in</emphasis>
37 45 58 95 65 seta 
<emphasis id="DEA2EAA56D4EFFA3BAD1BFF9F0A029F2" box="[641,663,1446,1467]" italics="true" pageId="3" pageNumber="4">ex</emphasis>
35 62 40 88 48 seta 
<emphasis id="DEA2EAA56D4EFFA3BAD1BF90F0BB29AD" box="[641,652,1487,1508]" italics="true" pageId="3" pageNumber="4">c</emphasis>
1 166 212 286 343 81 seta 
<emphasis id="DEA2EAA56D4EFFA3BAD1BFA4F0BB2A59" box="[641,652,1531,1552]" italics="true" pageId="3" pageNumber="4">c</emphasis>
3 38 40 87 136 lost leg IV 248 304 436 715 880 genital opening nd 25 52 80 106 anal opening 48 52 71 114 108 
<emphasis id="DEA2EAA56D4EFFA3B897BC8DF0192AA2" bold="true" box="[199,558,1746,1771]" pageId="3" pageNumber="4">Description of juvenile stages.</emphasis>
Larva elongated (
<figureCitation id="74ED2A326D4EFFA3BAACBC8CF1762AA2" box="[764,833,1747,1771]" captionStart="FIGURE 3" captionStartId="7.[151,250,2010,2033]" captionTargetBox="[197,1398,205,1972]" captionTargetId="figure@7.[175,1411,193,1983]" captionTargetPageId="7" captionText="FIGURE 3. Damaeus onustus, larva, dorsal aspect, scale bar 100 μm. Legs partially drawn." httpUri="https://zenodo.org/record/202942/files/figure.png" pageId="3" pageNumber="4">Fig. 3</figureCitation>
), unpigmented. Prodorsum triangular, setae 
<emphasis id="DEA2EAA56D4EFFA3BD67BC8BF7792AA2" box="[1335,1358,1748,1771]" italics="true" pageId="3" pageNumber="4">ro</emphasis>
and 
<emphasis id="DEA2EAA56D4EFFA3BDD8BC8CF7AB2AA2" box="[1416,1436,1747,1771]" italics="true" pageId="3" pageNumber="4">le</emphasis>
rather long (
<tableCitation id="A154030C6D4EFFA3B973BCA8F34E2B46" box="[291,377,1783,1807]" captionStart="TABLE 1" captionStartId="3.[151,235,290,311]" captionText="TABLE 1. Measurements of some morphological characters of juvenile stages of Damaeus onustus, D. clavipes, and Kunstidamaeus tecticola (mean measurements of 4 – 10 specimens in μm): larva (L), protonymph (PN), deutonymph (DN), tritonymph (TN) and adult (AD), nd – not developed." pageId="3" pageNumber="4">Table 1</tableCitation>
), curved, finely barbed; 
<emphasis id="DEA2EAA56D4EFFA3BAC7BCA8F09C2B46" box="[663,683,1783,1807]" italics="true" pageId="3" pageNumber="4">le</emphasis>
slightly longer and more widely inserted than 
<emphasis id="DEA2EAA56D4EFFA3BC95BCA7F6EB2B46" box="[1221,1244,1784,1807]" italics="true" pageId="3" pageNumber="4">ro</emphasis>
. Setae 
<emphasis id="DEA2EAA56D4EFFA3BD63BCA7F77E2B46" box="[1331,1353,1784,1807]" italics="true" pageId="3" pageNumber="4">in</emphasis>
and 
<emphasis id="DEA2EAA56D4EFFA3BDD3BCA7F7AB2B46" box="[1411,1436,1784,1807]" italics="true" pageId="3" pageNumber="4">ex</emphasis>
short. Bothridium well developed, rounded, protruding from surface, with funnel-like rim; sensillus setiform, finely barbed. Lateral depression present above leg I.
</paragraph>
<paragraph id="EC6936B76D4EFFA4B895BD3CF72A2C9E" blockId="3.[151,1437,1746,2034]" lastBlockId="4.[151,1436,151,2002]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
Gastronotum of larva with 12 pairs of setae, including 
<emphasis id="DEA2EAA56D4EFFA3BB7FBD3CF10A2B32" box="[815,829,1891,1915]" italics="true" pageId="3" pageNumber="4">h</emphasis>
2 and 
<emphasis id="DEA2EAA56D4EFFA3BB29BD3CF1B02B32" box="[889,903,1891,1915]" italics="true" pageId="3" pageNumber="4">h</emphasis>
3 inserted lateral to posterior and middle part of anal opening, respectively (
<figureCitation id="74ED2A326D4EFFA3B983BDD5F02E2BEB" box="[467,537,1930,1954]" captionStart="FIGURE 4" captionStartId="8.[151,250,1993,2016]" captionTargetBox="[319,1262,205,1961]" captionTargetId="figure@8.[315,1267,193,1968]" captionTargetPageId="8" captionText="FIGURE 4. Damaeus onustus, scale bars 100 μm. (A) Anal region of larva; (B) anogenital region of protonymph." httpUri="https://zenodo.org/record/202944/files/figure.png" pageId="3" pageNumber="4">Fig. 4</figureCitation>
A). Gastronotal setae inserted on large apophyses; most setae long, except for shorter seta 
<emphasis id="DEA2EAA56D4EFFA3B973BDF0F3182B8F" box="[291,303,1967,1990]" italics="true" pageId="3" pageNumber="4">c</emphasis>
3 (
<tableCitation id="A154030C6D4EFFA3B913BDF1F3AE2B8F" box="[323,409,1966,1990]" captionStart="TABLE 1" captionStartId="3.[151,235,290,311]" captionText="TABLE 1. Measurements of some morphological characters of juvenile stages of Damaeus onustus, D. clavipes, and Kunstidamaeus tecticola (mean measurements of 4 – 10 specimens in μm): larva (L), protonymph (PN), deutonymph (DN), tritonymph (TN) and adult (AD), nd – not developed." pageId="3" pageNumber="4">Table 1</tableCitation>
), curved, finely barbed and distally pliable, often twisted. Seta 
<emphasis id="DEA2EAA56D4EFFA3BC27BDF1F6B22B8F" box="[1143,1157,1966,1990]" italics="true" pageId="3" pageNumber="4">h</emphasis>
2 rather short, 
<emphasis id="DEA2EAA56D4EFFA3BD78BDF1F7012B8F" box="[1320,1334,1966,1990]" italics="true" pageId="3" pageNumber="4">h</emphasis>
3 minute. Paraproctal valves (segment PS) with two pairs of vestigial setae. Cupule 
<emphasis id="DEA2EAA56D4EFFA3BB89BD89F1D82BA4" box="[985,1007,2006,2029]" italics="true" pageId="3" pageNumber="4">ia</emphasis>
located posterior to seta 
<emphasis id="DEA2EAA56D4EFFA3BD5BBD89F7202BA4" box="[1291,1303,2006,2029]" italics="true" pageId="3" pageNumber="4">c</emphasis>
3, cupule 
<emphasis id="DEA2EAA56D4EFFA3BDD0BD89F7AB2BA4" box="[1408,1436,2006,2029]" italics="true" pageId="3" pageNumber="4">im</emphasis>
posterior to seta 
<emphasis id="DEA2EAA56D49FFA4B900BAC7F35B2CF9" box="[336,364,152,176]" italics="true" pageId="4" pageNumber="5">lm</emphasis>
, cupule 
<emphasis id="DEA2EAA56D49FFA4B99BBAC6F3D62CF9" box="[459,481,153,176]" italics="true" pageId="4" pageNumber="5">ip</emphasis>
between setae 
<emphasis id="DEA2EAA56D49FFA4BADABAC7F0972CF9" box="[650,672,152,176]" italics="true" pageId="4" pageNumber="5">lp</emphasis>
and 
<emphasis id="DEA2EAA56D49FFA4BA85BAC7F0D42CF9" box="[725,739,152,176]" italics="true" pageId="4" pageNumber="5">h</emphasis>
2, and cupule 
<emphasis id="DEA2EAA56D49FFA4BB29BAC7F1B82CF9" box="[889,911,152,176]" italics="true" pageId="4" pageNumber="5">ih</emphasis>
lateral to posterior part of anal opening. Opisthosomal gland opening (
<emphasis id="DEA2EAA56D49FFA4B9FCBAE0F3E72C9E" box="[428,464,191,215]" italics="true" pageId="4" pageNumber="5">gla</emphasis>
) lateral to seta 
<emphasis id="DEA2EAA56D49FFA4BA2DBAE0F0BC2C9E" box="[637,651,191,215]" italics="true" pageId="4" pageNumber="5">h</emphasis>
2. Gastronotum with granular cerotegument, and few folds.
</paragraph>
<paragraph id="EC6936B76D49FFA4B895BABAF3302E05" blockId="4.[151,1436,151,2002]" pageId="4" pageNumber="5">
Nymphs more oval and stocky than larva, unpigmented. Gastronotum of protonymph with 12 pairs of setae, as three pairs of pseudanal setae (
<emphasis id="DEA2EAA56D49FFA4B9A5BB55F0342D68" box="[501,515,266,289]" italics="true" pageId="4" pageNumber="5">p</emphasis>
1– 
<emphasis id="DEA2EAA56D49FFA4BA48BB55F0112D68" box="[536,550,266,289]" italics="true" pageId="4" pageNumber="5">p</emphasis>
3) appear on segment PS (
<figureCitation id="74ED2A326D49FFA4BB1ABB56F1B82D68" box="[842,911,265,289]" captionStart="FIGURE 4" captionStartId="8.[151,250,1993,2016]" captionTargetBox="[319,1262,205,1961]" captionTargetId="figure@8.[315,1267,193,1968]" captionTargetPageId="8" captionText="FIGURE 4. Damaeus onustus, scale bars 100 μm. (A) Anal region of larva; (B) anogenital region of protonymph." httpUri="https://zenodo.org/record/202944/files/figure.png" pageId="4" pageNumber="5">Fig. 4</figureCitation>
B), and remain in successive instars (
<figureCitation id="74ED2A326D49FFA4BD62BB56F7B72D68" box="[1330,1408,265,289]" captionStart="FIGURE 5" captionStartId="9.[151,250,1962,1985]" captionTargetBox="[306,1258,206,1913]" captionTargetId="figure@9.[295,1272,193,1934]" captionTargetPageId="9" captionText="FIGURE 5. Damaeus onustus, anogenital region, scale bars 100 μm. (A) Deutonymph; (B) tritonymph." httpUri="https://zenodo.org/record/202945/files/figure.png" pageId="4" pageNumber="5">Figs. 5</figureCitation>
A, B), while setae of 
<emphasis id="DEA2EAA56D49FFA4B936BB6FF3432D01" box="[358,372,304,328]" italics="true" pageId="4" pageNumber="5">d</emphasis>
-series absent, leaving central part of gastronotum bare. Setae of 
<emphasis id="DEA2EAA56D49FFA4BC03BB6EF6562D01" box="[1107,1121,305,328]" italics="true" pageId="4" pageNumber="5">p</emphasis>
-series finely barbed, length decreasing from 
<emphasis id="DEA2EAA56D49FFA4B902BB0AF3572D25" box="[338,352,341,364]" italics="true" pageId="4" pageNumber="5">p</emphasis>
1 to 
<emphasis id="DEA2EAA56D49FFA4B9DABB0AF3AF2D25" box="[394,408,341,364]" italics="true" pageId="4" pageNumber="5">p</emphasis>
3. Genital valves of protonymph with one pair of short setae, two pairs added in deutonymph and two more pairs in tritonymph; all setae short. One pair of small aggenital setae appears in deutonymph, lateral to posterior part of genital valves, and remains in tritonymph. Genital papillae rather large. Paraproctal valves of protonymph (segment AD) and deutonymph (segment AN) bare, but those of tritonymph with two pairs of small anal setae (
<emphasis id="DEA2EAA56D49FFA4B944BBB7F3072DB6" box="[276,304,488,511]" italics="true" pageId="4" pageNumber="5">an</emphasis>
1 and 
<emphasis id="DEA2EAA56D49FFA4B93EBBB7F3BD2DB6" box="[366,394,488,511]" italics="true" pageId="4" pageNumber="5">an</emphasis>
2). In deutonymph three pairs of adanal setae (
<emphasis id="DEA2EAA56D49FFA4BBDDBBB8F19E2DB6" box="[909,937,487,511]" italics="true" pageId="4" pageNumber="5">ad</emphasis>
1– 
<emphasis id="DEA2EAA56D49FFA4BBEFBBB8F1EC2DB6" box="[959,987,487,511]" italics="true" pageId="4" pageNumber="5">ad</emphasis>
3) appear on segment AD, and remain in tritonymph; all setae slightly curved, and shorter than seta 
<emphasis id="DEA2EAA56D49FFA4BB63B850F1762E6F" box="[819,833,527,550]" italics="true" pageId="4" pageNumber="5">p</emphasis>
3. Anogenital region with granular cerotegument, and few folds.
</paragraph>
<paragraph id="EC6936B76D49FFA4B895B807F0182E94" blockId="4.[151,1436,151,2002]" pageId="4" pageNumber="5">
Prodorsum of nymphs relatively shorter than in larva. Prodorsal setae of tritonymph (
<figureCitation id="74ED2A326D49FFA4BCC5B807F6ED2E39" box="[1173,1242,600,624]" captionStart="FIGURE 6" captionStartId="10.[151,250,2013,2036]" captionTargetBox="[217,1333,197,1974]" captionTargetId="figure@10.[217,1351,193,1984]" captionTargetPageId="10" captionText="FIGURE 6. Damaeus onustus. (A) Tritonymph, dorsal aspect, legs partially drawn, scale bar 100 μm; (B) cornicle." httpUri="https://zenodo.org/record/202946/files/figure.png" pageId="4" pageNumber="5">Fig. 6</figureCitation>
) as in larva, two lateral depressions present, posterior to seta 
<emphasis id="DEA2EAA56D49FFA4BAD2B822F0A12EDC" box="[642,662,637,661]" italics="true" pageId="4" pageNumber="5">le</emphasis>
, and above leg I, and transverse ridge posterior to seta 
<emphasis id="DEA2EAA56D49FFA4BCA9B822F73A2EDC" box="[1273,1293,637,661]" italics="true" pageId="4" pageNumber="5">le</emphasis>
. Bothridium rather large, rounded, protruding from surface, with funnel-like rim; sensillus setiform, with short, rare barbs. Prodorsum with granular cerotegument.
</paragraph>
<paragraph id="EC6936B76D49FFA4B895B8B7F0C82FFC" blockId="4.[151,1436,151,2002]" pageId="4" pageNumber="5">
Gastronotal setae rather long, curved ventrally (
<figureCitation id="74ED2A326D49FFA4BABEB8B7F1022F49" box="[750,821,744,768]" captionStart="FIGURE 7" captionStartId="11.[151,250,1887,1910]" captionTargetBox="[165,1427,228,1848]" captionTargetId="figure@11.[151,1436,193,1856]" captionTargetPageId="11" captionText="FIGURE 7. Damaeus onustus, tritonymph, lateral aspect, legs partially drawn, scale bar 100 μm. (A) With humus protective camouflage; (B) without this camouflage." httpUri="https://zenodo.org/record/202947/files/figure.png" pageId="4" pageNumber="5">Fig. 7</figureCitation>
), inserted on marginal parts of gastronotum, covered with exuviae of previous instars and humus mass. After removal of humus camouflage, dorsocentral part bare, with short, thick cornicle (
<emphasis id="DEA2EAA56D49FFA4B9DAB96EF3A12F00" box="[394,406,817,841]" italics="true" pageId="4" pageNumber="5">k</emphasis>
), situated midway between cupules 
<emphasis id="DEA2EAA56D49FFA4BB63B96DF17E2F00" box="[819,841,818,841]" italics="true" pageId="4" pageNumber="5">ia</emphasis>
(anteromedial to setae 
<emphasis id="DEA2EAA56D49FFA4BC03B96EF65E2F00" box="[1107,1129,817,841]" italics="true" pageId="4" pageNumber="5">la</emphasis>
). All setae pliable in distal part, fragile and often broken. Cupules 
<emphasis id="DEA2EAA56D49FFA4BA02B909F05F2F24" box="[594,616,854,877]" italics="true" pageId="4" pageNumber="5">ia</emphasis>
and 
<emphasis id="DEA2EAA56D49FFA4BAF0B909F08B2F24" box="[672,700,854,877]" italics="true" pageId="4" pageNumber="5">im</emphasis>
located as in larva, cupule 
<emphasis id="DEA2EAA56D49FFA4BBA1B909F6302F24" box="[1009,1031,854,877]" italics="true" pageId="4" pageNumber="5">ip</emphasis>
ventral to seta 
<emphasis id="DEA2EAA56D49FFA4BCE4B90AF6FD2F24" box="[1204,1226,853,877]" italics="true" pageId="4" pageNumber="5">lp</emphasis>
, cupule 
<emphasis id="DEA2EAA56D49FFA4BD7BB90AF7782F24" box="[1323,1359,853,877]" italics="true" pageId="4" pageNumber="5">iad</emphasis>
lateral to anterior part of anal opening, cupules 
<emphasis id="DEA2EAA56D49FFA4BA31B925F0B52FD8" box="[609,642,890,913]" italics="true" pageId="4" pageNumber="5">ips</emphasis>
and 
<emphasis id="DEA2EAA56D49FFA4BAE9B926F0F82FD8" box="[697,719,889,913]" italics="true" pageId="4" pageNumber="5">ih</emphasis>
pushed laterally from cupule 
<emphasis id="DEA2EAA56D49FFA4BC71B926F6722FD8" box="[1057,1093,889,913]" italics="true" pageId="4" pageNumber="5">iad</emphasis>
, gland opening (
<emphasis id="DEA2EAA56D49FFA4BD54B926F71F2FD8" box="[1284,1320,889,913]" italics="true" pageId="4" pageNumber="5">gla</emphasis>
) posterior to cupule 
<emphasis id="DEA2EAA56D49FFA4B955B9C2F32C2FFC" box="[261,283,925,949]" italics="true" pageId="4" pageNumber="5">ih</emphasis>
. Gastronotum with granular cerotegument.
</paragraph>
<paragraph id="EC6936B76D49FFA4B895B99EF7B52FB6" blockId="4.[151,1436,151,2002]" pageId="4" pageNumber="5">
Tarsus I of tritonymph long and thin, with two solenidia, ω1 longer and thicker than ω2, seta 
<emphasis id="DEA2EAA56D49FFA4BCB8B99EF6C02F90" box="[1256,1271,961,985]" italics="true" pageId="4" pageNumber="5">ft</emphasis>
’ long, seta 
<emphasis id="DEA2EAA56D49FFA4BDD0B99EF7B82F90" box="[1408,1423,961,985]" italics="true" pageId="4" pageNumber="5">ft</emphasis>
” short, posterior to solenidion ω1 (
<figureCitation id="74ED2A326D49FFA4BA5BB9B8F07A2FB6" box="[523,589,999,1023]" captionStart="FIGURE 8" captionStartId="12.[151,250,1633,1656]" captionTargetBox="[161,1435,202,1598]" captionTargetId="figure@12.[151,1436,193,1604]" captionTargetPageId="12" captionText="FIGURE 8. Tarsus I and part of tibia of tritonymph, paraxial aspect, scale bars 10 μm. (A) Damaeus onustus; (B) D. clavipes; (C) Kunstidamaeus tecticola." httpUri="https://zenodo.org/record/202948/files/figure.png" pageId="4" pageNumber="5">Fig. 8</figureCitation>
A). Tibia I with two solenidia, φ1 long, with long coupled seta 
<emphasis id="DEA2EAA56D49FFA4BD58B9B8F7212FB6" box="[1288,1302,999,1023]" italics="true" pageId="4" pageNumber="5">d</emphasis>
, φ2 short.
</paragraph>
<paragraph id="EC6936B76D49FFA4B897BE52F12F28DB" blockId="4.[151,1436,151,2002]" pageId="4" pageNumber="5">
<emphasis id="DEA2EAA56D49FFA4B897BE52F08E286F" bold="true" box="[199,697,1037,1062]" pageId="4" pageNumber="5">Summary of ontogenetic transformations.</emphasis>
Setae 
<emphasis id="DEA2EAA56D49FFA4BB53BE50F12C286F" box="[771,795,1039,1062]" italics="true" pageId="4" pageNumber="5">ro</emphasis>
and 
<emphasis id="DEA2EAA56D49FFA4BB00BE51F153286F" box="[848,868,1038,1062]" italics="true" pageId="4" pageNumber="5">le</emphasis>
are rather long in all instars, and barbed, but in the juveniles setae 
<emphasis id="DEA2EAA56D49FFA4B913BE6CF36E2803" box="[323,345,1075,1098]" italics="true" pageId="4" pageNumber="5">ro</emphasis>
are inserted closer to the rostrum than in the adult. Seta 
<emphasis id="DEA2EAA56D49FFA4BB8DBE6CF1C42803" box="[989,1011,1075,1098]" italics="true" pageId="4" pageNumber="5">in</emphasis>
is short in the larva and nymphs, and rather long in the adult, while seta 
<emphasis id="DEA2EAA56D49FFA4BA75BE08F00A2827" box="[549,573,1111,1134]" italics="true" pageId="4" pageNumber="5">ex</emphasis>
is small in all instars. The bothridium is rather large, rounded in all instars, with a funnel-like rim, and the sensillus is long, setiform.
</paragraph>
<paragraph id="EC6936B76D49FFA4B895BEC1F34429DB" blockId="4.[151,1436,151,2002]" pageId="4" pageNumber="5">
The larva has 12 pairs of gastronotal setae, but in the protonymph some setae appear (
<emphasis id="DEA2EAA56D49FFA4BCCCBEC0F69D28FF" box="[1180,1194,1183,1206]" italics="true" pageId="4" pageNumber="5">p</emphasis>
-series) and some are lost (
<emphasis id="DEA2EAA56D49FFA4B882BE9DF2D72893" box="[210,224,1218,1242]" italics="true" pageId="4" pageNumber="5">d</emphasis>
-series), such that 12 pairs remain in all nymphs. The adult loses seta 
<emphasis id="DEA2EAA56D49FFA4BC40BE9CF62B2893" box="[1040,1052,1219,1242]" italics="true" pageId="4" pageNumber="5">c</emphasis>
3, so 11 pairs of notogastral setae remain. The formula of gastronotal setae in 
<taxonomicName id="2BD64D346D49FFA4BAD0BEB5F0C02948" box="[640,759,1258,1281]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="onustus">
<emphasis id="DEA2EAA56D49FFA4BAD0BEB5F0A32948" box="[640,660,1258,1281]" italics="true" pageId="4" pageNumber="5">D</emphasis>
. 
<emphasis id="DEA2EAA56D49FFA4BAF2BEB5F0C02948" box="[674,759,1258,1281]" italics="true" pageId="4" pageNumber="5">onustus</emphasis>
</taxonomicName>
is 
<date id="986810776D49FFA4BB48BEB6F1922948" box="[792,933,1257,1281]" pageId="4" pageNumber="5" value="1912-12-12" valueMax="1912-12-12" valueMin="1912-12-12">12-12-12-12</date>
-11 (larva to adult), those of coxisternal setae are: 3-1-2 (larva), 3-1-2-1 (protonymph), 3-1-2-2 (deutonymph) and 3-1-3-3 (tritonymph and adult). In the larva seta 
<emphasis id="DEA2EAA56D49FFA4B899BF6EF2D42900" box="[201,227,1329,1353]" italics="true" pageId="4" pageNumber="5">1c</emphasis>
is scaliform and covers the Claparède’s organ (see 
<bibRefCitation id="88474B466D49FFA4BB72BF6EF1EE2900" author="Grandjean" box="[802,985,1329,1353]" pageId="4" pageNumber="5" refString="Grandjean, F. (1955) L'organe de Claparede et son ecaille chez Damaeus onustus Koch. Bulletin du Museum D'Histoire naturelle, Paris, 27, 285 - 292." type="journal article" year="1955">Grandjean 1955</bibRefCitation>
). The formula of genital setae is 1-3-5-6 (protonymph to adult), and that of aggenital setae is 1-1-1 (deutonymph to adult). The formula of segments PS −AN is 03333-0333-022.
</paragraph>
</subSubSection>
<subSubSection id="A4CC653C6D49FFACB897BFC2F1A92BAD" lastPageId="12" lastPageNumber="13" pageId="4" pageNumber="5" type="distribution">
<paragraph id="EC6936B76D49FFA4B897BFC2F38829B7" blockId="4.[151,1436,151,2002]" pageId="4" pageNumber="5">
<emphasis id="DEA2EAA56D49FFA4B897BFC2F06029FF" bold="true" box="[199,599,1437,1462]" pageId="4" pageNumber="5">Distribution ecology and biology.</emphasis>
<taxonomicName id="2BD64D346D49FFA4BA30BFC0F11F29FF" box="[608,808,1439,1462]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="onustus">
<emphasis id="DEA2EAA56D49FFA4BA30BFC0F11F29FF" box="[608,808,1439,1462]" italics="true" pageId="4" pageNumber="5">Damaeus onustus</emphasis>
</taxonomicName>
is a widespread European species (
<bibRefCitation id="88474B466D49FFA4BC91BFC1F76629FF" author="Subias" box="[1217,1361,1438,1462]" pageId="4" pageNumber="5" refString="Subias, L. S. (2004) Listado sistematico, sinonimico y biogeografico de los Acaros Oribatidos (Acariformes, Oribatida) del mundo (1758 - 2002). Graellsia, 60, 3 - 305." type="journal article" year="2004">Subías 2004</bibRefCitation>
; 
<bibRefCitation id="88474B466D49FFA4BD0FBFC1F2E42993" author="Miko" pageId="4" pageNumber="5" refString="Miko, L. (2006) Damaeidae. In: Weigmann, G.: Hornmilben (Oribatida), Die Tierwelt Deutschlands, Teil 76. Dahl, F. (Ed.), Goecke &amp; Evers, Keltern, pp. 179 - 207." type="journal article" year="2006">Miko 2006</bibRefCitation>
), including 
<collectingCountry id="94C176276D49FFA4B938BF9DF3DB2993" box="[360,492,1474,1498]" name="Svalbard and Jan Mayen" pageId="4" pageNumber="5">Spitsbergen</collectingCountry>
(
<bibRefCitation id="88474B466D49FFA4BA53BF9DF1632993" author="Lebedeva" box="[515,852,1474,1498]" pageId="4" pageNumber="5" refString="Lebedeva, N. V. &amp; Lebedev, V. D. (2008) Transport of oribatid mites to the polar areas by birds. Integrative acarology. In: Proceedings of the 6 th European Congress, European Association of Acarologists, Montpellier, France, pp. 359 - 367" type="book chapter" year="2008">Lebedeva &amp; Lebedev 2008</bibRefCitation>
). It was recorded from several parts of 
<collectingCountry id="94C176276D49FFA4BD1EBF9DF7AB2993" box="[1358,1436,1474,1498]" name="Poland" pageId="4" pageNumber="5">Poland</collectingCountry>
(
<bibRefCitation id="88474B466D49FFA4B8CEBFB9F38429B7" author="Olszanowski" box="[158,435,1510,1534]" pageId="4" pageNumber="5" refString="Olszanowski, Z., Rajski, A. &amp; Niedbala, W. (1996) Catalogue of Polish fauna. Mites (Acari), moss mites (Oribatida). Sours Pozna n, 34 (9), 1 - 243. (In Polish)." type="journal article" year="1996">
Olszanowski 
<emphasis id="DEA2EAA56D49FFA4B968BFB8F35E29B7" box="[312,361,1510,1534]" italics="true" pageId="4" pageNumber="5">et al</emphasis>
. 1996
</bibRefCitation>
).
</paragraph>
<paragraph id="EC6936B76D49FFA4B895BC55F0D12AB3" blockId="4.[151,1436,151,2002]" pageId="4" pageNumber="5">
This species occurs in deciduous litter, mosses, bark and decaying wood (
<bibRefCitation id="88474B466D49FFA4BC7EBC55F6D22A6B" author="Willmann" box="[1070,1253,1546,1570]" pageId="4" pageNumber="5" refString="Willmann, C. (1931) Moosmilben oder Oribatiden (Oribatei). In: Dahl, F. (Ed.): Die Tierwelt Deutschland und der angrenzenden Meeresteile. Verlag Gustav Fisher, Jena, pp. 79 - 200." type="book chapter" year="1931">Willmann 1931</bibRefCitation>
; 
<bibRefCitation id="88474B466D49FFA4BCA5BC55F7A02A6B" author="Sellnick" box="[1269,1431,1546,1570]" pageId="4" pageNumber="5" refString="Sellnick, M. (1960) Formenkreis: Hornmilben, Oribatei. In: Brohmers, P., Ehrmann, P., Ulmerm G. (Eds.): Die Tierwelt Mitteleuropas. Vol. 3, 4. Lief. (Erganzung). Quelle &amp; Meyer, Leipzig, pp. 45 - 134." type="journal article" year="1960">Sellnick 1960</bibRefCitation>
; 
<bibRefCitation id="88474B466D49FFA4B8C7BC71F3A02A0F" author="Wood" box="[151,407,1582,1606]" pageId="4" pageNumber="5" refString="Wood, T. G. &amp; Lawton, J. H. (1973) Experimental Studies on the Respiratory Rates of Mites (Acari) from Beech-Woodland Leaf Litter. Oecologia, 12, 169 - 191." type="journal article" year="1973">Wood &amp; Lawton 1973</bibRefCitation>
; 
<bibRefCitation id="88474B466D49FFA4B9F4BC71F0632A0F" author="Wallwork" box="[420,596,1582,1606]" pageId="4" pageNumber="5" refString="Wallwork, J. A. (1976) The distribution and diversity of the Soil Fauna. Academic Press, London, pp. 1 - 331." type="book chapter" year="1976">Wallwork 1976</bibRefCitation>
; 
<bibRefCitation id="88474B466D49FFA4BA31BC70F10B2A0F" author="Maraun" box="[609,828,1582,1606]" pageId="4" pageNumber="5" refString="Maraun, M., Heethoff, M., Schneider, K., Scheu, S., Weigmann, G., Cianciolo, J., Thomas, R. H. &amp; Norton, R. A. (2004) Molecular phylogeny of oribatid mites (Oribatida, Acari): evidence for multiple radiations of parthenogenetic lineages. Experimental and Applied Acarology, 33, 183 - 201." type="journal article" year="2004">
Maraun 
<emphasis id="DEA2EAA56D49FFA4BAEFBC70F0C52A0F" box="[703,754,1582,1606]" italics="true" pageId="4" pageNumber="5">et al</emphasis>
. 2004
</bibRefCitation>
; 
<bibRefCitation id="88474B466D49FFA4BB18BC76F6BA2A0F" author="Zbikowska-Zdun" box="[840,1165,1577,1606]" pageId="4" pageNumber="5" refString="Zbikowska-Zdun, K., Piksa, K. &amp; Watrak, I. (2006) Diversity of mites (Acari: Oribatida) in selected microhabitats of the Bug River Protected Landscape Area. Biological Letters, 43 (2), 277 - 286." type="journal article" year="2006">
Żbikowska-Zdun 
<emphasis id="DEA2EAA56D49FFA4BC40BC70F6742A0F" box="[1040,1091,1582,1606]" italics="true" pageId="4" pageNumber="5">et al</emphasis>
. 2006
</bibRefCitation>
), but also in bushes and grasses, showing preference to loose accumulation of litter, with a great tolerance of humidity (
<bibRefCitation id="88474B466D49FFA4BCECBC0DF77F2A23" author="Rajski" box="[1212,1352,1618,1642]" pageId="4" pageNumber="5" refString="Rajski, A. (1967) Autecological-zoogeographical analysis of moss mites (Acari, Oribatei) on the basis of fauna in the Poznan Environs. Part I. Bulletin Entomologique de Pologne, 37 (1), 69 - 166." type="journal article" year="1967">Rajski 1967</bibRefCitation>
). In the Chiltern beech woods (South East 
<collectingCountry id="94C176276D49FFA4BA7DBC29F0BB2AC7" box="[557,652,1654,1678]" name="United Kingdom" pageId="4" pageNumber="5">England</collectingCountry>
) this species was abundant in beach litter accumulated in sheltered depressions, but was less abundant on steep slopes 
<bibRefCitation id="88474B466D49FFA4BAB7BCC5F19C2AFB" author="Wallwork" box="[743,939,1690,1714]" pageId="4" pageNumber="5" refString="Wallwork, J. A. (1976) The distribution and diversity of the Soil Fauna. Academic Press, London, pp. 1 - 331." type="book chapter" year="1976">Wallwork (1976)</bibRefCitation>
. In Botanical Garden on the campus of the University of Life Sciences in 
<collectingCountry id="94C176276D49FFA4B9BCBCE1F03C2A9F" box="[492,523,1726,1750]" name="American Samoa" pageId="4" pageNumber="5">Ås</collectingCountry>
(
<collectingCountry id="94C176276D49FFA4BA49BCE1F0402A9F" box="[537,631,1726,1750]" name="Norway" pageId="4" pageNumber="5">Norway</collectingCountry>
) it also occurs in Scots pine and shrub litter. 
<bibRefCitation id="88474B466D49FFA4BC3ABCE1F7322A9F" author="Schatz" box="[1130,1285,1726,1750]" pageId="4" pageNumber="5" refString="Schatz, H. (1983) U. - Ordn. Oribatei, Hornmilben. Catalogus faunae Austriae. Ein systematisches Verzeichnis aller auf osterreichischem Gebiet festgestellten Tierarten, 9, Osterreichische Akadademie der Wissenschaften, Wien, pp. 1 - 118." type="journal article" year="1983">Schatz (1983)</bibRefCitation>
considered it in forest, eurytopic, and microphytophagous species.
</paragraph>
<paragraph id="EC6936B76D49FFA4B895BD59F7762B2E" blockId="4.[151,1436,151,2002]" pageId="4" pageNumber="5">
<collectingCountry id="94C176276D49FFA4B895BD59F2D32B57" box="[197,228,1798,1822]" name="American Samoa" pageId="4" pageNumber="5">As</collectingCountry>
<taxonomicName id="2BD64D346D49FFA4B8BBBD58F3552B57" box="[235,354,1799,1822]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="onustus">
<emphasis id="DEA2EAA56D49FFA4B8BBBD58F2C82B57" box="[235,255,1799,1822]" italics="true" pageId="4" pageNumber="5">D</emphasis>
. 
<emphasis id="DEA2EAA56D49FFA4B95DBD58F3552B57" box="[269,354,1799,1822]" italics="true" pageId="4" pageNumber="5">onustus</emphasis>
</taxonomicName>
is relatively large, it was often cultured in the laboratory. 
<bibRefCitation id="88474B466D49FFA4BBBABD59F64B2B57" author="Pauly" box="[1002,1148,1798,1822]" pageId="4" pageNumber="5" refString="Pauly, F. (1956) Zur Biologie einiger Belbiden (Oribatei: Moosmilben) und zur Funktion ihrer pseudostigmatischen Organe. Zoologische Jahrbucher. Abteilung fur Systematik, Okologie und Geographie der Tiere, 84, 275 - 328." type="journal article" year="1956">Pauly (1956)</bibRefCitation>
investigated the development and biology of this species. When 
<taxonomicName id="2BD64D346D49FFA4BA31BD74F0ED2B0B" box="[609,730,1835,1858]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="onustus">
<emphasis id="DEA2EAA56D49FFA4BA31BD74F0422B0B" box="[609,629,1835,1858]" italics="true" pageId="4" pageNumber="5">D</emphasis>
. 
<emphasis id="DEA2EAA56D49FFA4BAD5BD74F0ED2B0B" box="[645,730,1835,1858]" italics="true" pageId="4" pageNumber="5">onustus</emphasis>
</taxonomicName>
fed on algae from tree bark, one female laid about 
<specimenCount id="FAD0FD3E6D49FFA4BD76BD75F7492B0A" box="[1318,1406,1834,1859]" pageId="4" pageNumber="5" type="egg">25 eggs</specimenCount>
in breeding season, whole development of species lasted about 140 days, and the adult lived about two years.
</paragraph>
<paragraph id="EC6936B76D49FFACB895BD2DF1A92BAD" blockId="4.[151,1436,151,2002]" lastBlockId="12.[151,1437,1730,2020]" lastPageId="12" lastPageNumber="13" pageId="4" pageNumber="5">
<bibRefCitation id="88474B466D49FFA4B895BD2DF3442BC3" author="Madge" box="[197,371,1906,1930]" pageId="4" pageNumber="5" refString="Madge, D. S. (1964 a) The water-relations of Belba geniculosa Oudms and other species of Oribatid mites. Acarologia, 6 (1), 199 - 223." type="journal article" year="1964" yearSuffix="a">Madge (1964 a</bibRefCitation>
, 
<bibRefCitation id="88474B466D49FFA4B9D0BD2DF3FE2BC3" author="Madge" box="[384,457,1906,1930]" pageId="4" pageNumber="5" refString="Madge, D. S. (1964 b) The humidity reactions of oribatid mites. Acarologia, 6 (3), 566 - 591." type="journal article" year="1964" yearSuffix="b">1964b</bibRefCitation>
, 
<bibRefCitation id="88474B466D49FFA4B985BD2DF02B2BC3" author="Madge" box="[469,540,1906,1930]" pageId="4" pageNumber="5" refString="Madge, D. S. (1964 c) The longevity of fasting oribatid mites. Acarologia, 6 (4), 718 - 729." type="journal article" year="1964" yearSuffix="c">1964c</bibRefCitation>
, 
<bibRefCitation id="88474B466D49FFA4BA79BD2DF0472BC3" author="Madge" box="[553,624,1906,1930]" pageId="4" pageNumber="5" refString="Madge, D. S. (1965 a) The effects of lethal temperatures on Oribatid mites. Acarologia, 7 (1), 121 - 130." type="journal article" year="1965" yearSuffix="a">1965a</bibRefCitation>
, 
<bibRefCitation id="88474B466D49FFA4BA2DBD2DF0F12BC3" author="Madge" box="[637,710,1906,1930]" pageId="4" pageNumber="5" refString="Madge, D. S. (1965 b) The behaviour of Belba geniculosa Oudms. and certain other species of Oribatid mites in controlled temperature gradients. Acarologia, 7 (2), 389 - 406." type="journal article" year="1965" yearSuffix="b">1965b</bibRefCitation>
, 
<bibRefCitation id="88474B466D49FFA4BA82BD2DF12F2BC3" author="Madge" box="[722,792,1906,1930]" pageId="4" pageNumber="5" refString="Madge, D. S. (1965 c) Further studies on the behaviour of Belba geniculosa Oudms. in relation to various environmental stimuli. Acarologia, 7 (4), 744 - 757." type="journal article" year="1965" yearSuffix="c">1965c</bibRefCitation>
) investigated the ecology of 
<taxonomicName id="2BD64D346D49FFA4BC33BD2CF6EB2BC3" box="[1123,1244,1907,1930]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="onustus">
<emphasis id="DEA2EAA56D49FFA4BC33BD2CF6402BC3" box="[1123,1143,1907,1930]" italics="true" pageId="4" pageNumber="5">D</emphasis>
. 
<emphasis id="DEA2EAA56D49FFA4BCD6BD2CF6EB2BC3" box="[1158,1244,1907,1930]" italics="true" pageId="4" pageNumber="5">onustus</emphasis>
</taxonomicName>
in very detailed, and inventive laboratory experiments, which considered water-relations, humidity reactions, longevity, lethal temperatures and behaviour of this species. From all interesting results the most interesting seem to be those connected with bearing the exuviae of previous instars by nymphs. When 
<taxonomicName id="2BD64D346D41FFACBB36BC99F1E82A94" box="[870,991,1734,1757]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="onustus">
<emphasis id="DEA2EAA56D41FFACBB36BC99F14D2A94" box="[870,890,1734,1757]" italics="true" pageId="12" pageNumber="13">D</emphasis>
. 
<emphasis id="DEA2EAA56D41FFACBBD9BC99F1E82A94" box="[905,991,1734,1757]" italics="true" pageId="12" pageNumber="13">onustus</emphasis>
</taxonomicName>
was exposed to temperatures 10–70o C, the critical temperature to waterproofing cuticular cover was lower for the juveniles (40o C) than for the adults (45o C), which may explain the ecological importance of bearing of exuviae and debris by nymphs. If debris was removed from the nymphs, evaporation of mites increased. Critical temperature for nymphs both with or without debris was then 40o C. This species controls well water-loss and temperature of body by well developed respiratory system. It also reacts well to air humidity by humidity receptors, which are probably located on forelegs, which is consistent with 
<bibRefCitation id="88474B466D41FFACB916BDFAF3ED2BF4" author="Pauly" box="[326,474,1957,1981]" pageId="12" pageNumber="13" refString="Pauly, F. (1956) Zur Biologie einiger Belbiden (Oribatei: Moosmilben) und zur Funktion ihrer pseudostigmatischen Organe. Zoologische Jahrbucher. Abteilung fur Systematik, Okologie und Geographie der Tiere, 84, 275 - 328." type="journal article" year="1956">Pauly (1956)</bibRefCitation>
. This species is more active in dry air than in moist air. 
<collectingCountry id="94C176276D41FFACBC14BDF9F6542BF7" box="[1092,1123,1958,1982]" name="American Samoa" pageId="12" pageNumber="13">As</collectingCountry>
a matter of radiant heat, the tritonymphs chose lower temperature (11.5o C) than the adults (13o C).
</paragraph>
</subSubSection>
<caption id="B8A9663F6D48FFA5B8C7BD98F75B2B94" box="[151,1388,1991,2014]" httpUri="https://zenodo.org/record/202940/files/figure.png" pageId="5" pageNumber="6" targetBox="[249,1330,198,1955]" targetPageId="5">
<paragraph id="EC6936B76D48FFA5B8C7BD98F75B2B94" blockId="5.[151,1388,1991,2014]" box="[151,1388,1991,2014]" pageId="5" pageNumber="6">
<emphasis id="DEA2EAA56D48FFA5B8C7BD98F3242B95" bold="true" box="[151,275,1991,2014]" pageId="5" pageNumber="6">FIGURE 1.</emphasis>
<taxonomicName id="2BD64D346D48FFA5B949BD97F3FB2B94" box="[281,460,1992,2013]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="onustus">
<emphasis id="DEA2EAA56D48FFA5B949BD97F3FB2B94" box="[281,460,1992,2013]" italics="true" pageId="5" pageNumber="6">Damaeus onustus</emphasis>
</taxonomicName>
. (A) Adult, dorsal aspect, legs partially drawn, scale bar 100 μm; (B) central part of seta 
<emphasis id="DEA2EAA56D48FFA5BD03BD98F7502B94" box="[1363,1383,1991,2013]" italics="true" pageId="5" pageNumber="6">la</emphasis>
.
</paragraph>
</caption>
<caption id="B8A9663F6D4BFFA6B8C7BD9CF7692B90" box="[151,1374,1987,2010]" httpUri="https://zenodo.org/record/202941/files/figure.png" pageId="6" pageNumber="7" targetBox="[206,1392,212,1928]" targetPageId="6">
<paragraph id="EC6936B76D4BFFA6B8C7BD9CF7692B90" blockId="6.[151,1374,1987,2010]" box="[151,1374,1987,2010]" pageId="6" pageNumber="7">
<emphasis id="DEA2EAA56D4BFFA6B8C7BD9CF3242B91" bold="true" box="[151,275,1987,2010]" pageId="6" pageNumber="7">FIGURE 2.</emphasis>
<taxonomicName id="2BD64D346D4BFFA6B949BD9BF3FB2B90" box="[281,460,1988,2009]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="onustus">
<emphasis id="DEA2EAA56D4BFFA6B949BD9BF3FB2B90" box="[281,460,1988,2009]" italics="true" pageId="6" pageNumber="7">Damaeus onustus</emphasis>
</taxonomicName>
, adult, scale bars 100 μm. (A) Lateral aspect; (B) anogenital region. Legs partially drawn.
</paragraph>
</caption>
<caption id="B8A9663F6D4AFFA7B8C7BD85F60C2BB8" box="[151,1083,2010,2033]" httpUri="https://zenodo.org/record/202942/files/figure.png" pageId="7" pageNumber="8" targetBox="[197,1398,205,1972]" targetPageId="7">
<paragraph id="EC6936B76D4AFFA7B8C7BD85F60C2BB8" blockId="7.[151,1083,2010,2033]" box="[151,1083,2010,2033]" pageId="7" pageNumber="8">
<emphasis id="DEA2EAA56D4AFFA7B8C7BD85F3252BB9" bold="true" box="[151,274,2010,2033]" pageId="7" pageNumber="8">FIGURE 3.</emphasis>
<taxonomicName id="2BD64D346D4AFFA7B949BD83F3FB2BB8" box="[281,460,2012,2033]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="onustus">
<emphasis id="DEA2EAA56D4AFFA7B949BD83F3FB2BB8" box="[281,460,2012,2033]" italics="true" pageId="7" pageNumber="8">Damaeus onustus</emphasis>
</taxonomicName>
, larva, dorsal aspect, scale bar 100 μm. Legs partially drawn.
</paragraph>
</caption>
<caption id="B8A9663F6D45FFA8B8C7BD96F72A2BA9" box="[151,1309,1993,2017]" httpUri="https://zenodo.org/record/202944/files/figure.png" pageId="8" pageNumber="9" targetBox="[319,1262,205,1961]" targetPageId="8">
<paragraph id="EC6936B76D45FFA8B8C7BD96F72A2BA9" blockId="8.[151,1309,1993,2017]" box="[151,1309,1993,2017]" pageId="8" pageNumber="9">
<emphasis id="DEA2EAA56D45FFA8B8C7BD96F3252B96" bold="true" box="[151,274,1993,2016]" pageId="8" pageNumber="9">FIGURE 4.</emphasis>
<taxonomicName id="2BD64D346D45FFA8B949BD94F3FB2BA9" box="[281,460,1995,2016]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="onustus">
<emphasis id="DEA2EAA56D45FFA8B949BD94F3FB2BA9" box="[281,460,1995,2016]" italics="true" pageId="8" pageNumber="9">Damaeus onustus</emphasis>
</taxonomicName>
, scale bars 100 μm. (A) Anal region of larva; (B) anogenital region of protonymph.
</paragraph>
</caption>
<caption id="B8A9663F6D44FFA9B8C7BDF5F6822B88" box="[151,1205,1962,1986]" httpUri="https://zenodo.org/record/202945/files/figure.png" pageId="9" pageNumber="10" targetBox="[306,1258,206,1913]" targetPageId="9">
<paragraph id="EC6936B76D44FFA9B8C7BDF5F6822B88" blockId="9.[151,1205,1962,1986]" box="[151,1205,1962,1986]" pageId="9" pageNumber="10">
<emphasis id="DEA2EAA56D44FFA9B8C7BDF5F3252B89" bold="true" box="[151,274,1962,1985]" pageId="9" pageNumber="10">FIGURE 5.</emphasis>
<taxonomicName id="2BD64D346D44FFA9B949BDF3F3FB2B88" box="[281,460,1964,1985]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="onustus">
<emphasis id="DEA2EAA56D44FFA9B949BDF3F3FB2B88" box="[281,460,1964,1985]" italics="true" pageId="9" pageNumber="10">Damaeus onustus</emphasis>
</taxonomicName>
, anogenital region, scale bars 100 μm. (A) Deutonymph; (B) tritonymph.
</paragraph>
</caption>
<caption id="B8A9663F6D47FFAAB8C7BD82F71F2BBA" box="[151,1320,2013,2036]" httpUri="https://zenodo.org/record/202946/files/figure.png" pageId="10" pageNumber="11" targetBox="[217,1333,197,1974]" targetPageId="10">
<paragraph id="EC6936B76D47FFAAB8C7BD82F71F2BBA" blockId="10.[151,1320,2013,2036]" box="[151,1320,2013,2036]" pageId="10" pageNumber="11">
<emphasis id="DEA2EAA56D47FFAAB8C7BD82F3252BBA" bold="true" box="[151,274,2013,2036]" pageId="10" pageNumber="11">FIGURE 6.</emphasis>
<taxonomicName id="2BD64D346D47FFAAB949BD80F3FB2BBD" box="[281,460,2015,2036]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="onustus">
<emphasis id="DEA2EAA56D47FFAAB949BD80F3FB2BBD" box="[281,460,2015,2036]" italics="true" pageId="10" pageNumber="11">Damaeus onustus</emphasis>
</taxonomicName>
. (A) Tritonymph, dorsal aspect, legs partially drawn, scale bar 100 μm; (B) cornicle.
</paragraph>
</caption>
<caption id="B8A9663F6D46FFABB8C7BD00F0092BDD" httpUri="https://zenodo.org/record/202947/files/figure.png" pageId="11" pageNumber="12" targetBox="[165,1427,228,1848]" targetPageId="11">
<paragraph id="EC6936B76D46FFABB8C7BD00F0092BDD" blockId="11.[151,1436,1887,1940]" pageId="11" pageNumber="12">
<emphasis id="DEA2EAA56D46FFABB8C7BD00F3232B3C" bold="true" box="[151,276,1887,1910]" pageId="11" pageNumber="12">FIGURE 7.</emphasis>
<taxonomicName id="2BD64D346D46FFABB94BBD3FF3E72B3C" box="[283,464,1888,1909]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="onustus">
<emphasis id="DEA2EAA56D46FFABB94BBD3FF3E72B3C" box="[283,464,1888,1909]" italics="true" pageId="11" pageNumber="12">Damaeus onustus</emphasis>
</taxonomicName>
, tritonymph, lateral aspect, legs partially drawn, scale bar 100 μm. (A) With humus protective camouflage; (B) without this camouflage.
</paragraph>
</caption>
<caption id="B8A9663F6D41FFACB8C7BC3EF3892ADF" httpUri="https://zenodo.org/record/202948/files/figure.png" pageId="12" pageNumber="13" targetBox="[161,1435,202,1598]" targetPageId="12">
<paragraph id="EC6936B76D41FFACB8C7BC3EF3892ADF" blockId="12.[151,1436,1633,1686]" pageId="12" pageNumber="13">
<emphasis id="DEA2EAA56D41FFACB8C7BC3EF3242A3F" bold="true" box="[151,275,1633,1656]" pageId="12" pageNumber="13">FIGURE 8.</emphasis>
Tarsus I and part of tibia of tritonymph, paraxial aspect, scale bars 10 μm. (A) 
<taxonomicName id="2BD64D346D41FFACBC69BC3DF6DB2A3E" box="[1081,1260,1634,1655]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="onustus">
<emphasis id="DEA2EAA56D41FFACBC69BC3DF6DB2A3E" box="[1081,1260,1634,1655]" italics="true" pageId="12" pageNumber="13">Damaeus onustus</emphasis>
</taxonomicName>
; (B) 
<taxonomicName id="2BD64D346D41FFACBD73BC3DF7A22A3E" box="[1315,1429,1633,1655]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="clavipes">
<emphasis id="DEA2EAA56D41FFACBD73BC3DF7022A3E" box="[1315,1333,1634,1655]" italics="true" pageId="12" pageNumber="13">D</emphasis>
. 
<emphasis id="DEA2EAA56D41FFACBD12BC3EF7A22A3E" box="[1346,1429,1633,1655]" italics="true" pageId="12" pageNumber="13">clavipes</emphasis>
</taxonomicName>
; (C) 
<taxonomicName id="2BD64D346D41FFACB8EFBCDFF38F2ADF" box="[191,440,1664,1686]" class="Arachnida" family="Damaeidae" genus="Kunstidamaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="tecticola">
<emphasis id="DEA2EAA56D41FFACB8EFBCDFF38F2ADF" box="[191,440,1664,1686]" italics="true" pageId="12" pageNumber="13">Kunstidamaeus tecticola</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="A4CC653C6D40FFADB895BAC6F2D72CBE" pageId="13" pageNumber="14" type="reference_group">
<paragraph id="EC6936B76D40FFADB895BAC6F2D72CBE" blockId="13.[151,1436,151,247]" pageId="13" pageNumber="14">
<taxonomicName id="2BD64D346D40FFADB895BAC6F3BB2CF9" ID-CoL="7SZMD" box="[197,396,153,176]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="onustus">
<emphasis id="DEA2EAA56D40FFADB895BAC6F3BB2CF9" box="[197,396,153,176]" italics="true" pageId="13" pageNumber="14">Damaeus onustus</emphasis>
</taxonomicName>
is an intermediate host of 
<taxonomicName id="2BD64D346D40FFADBAEEBAC7F65B2CE6" authority="Denegri 1993" authorityName="Denegri" authorityYear="1993" box="[702,1132,151,176]" class="Neoophora" family="Anoplocephalidae" genus="Cittotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="13" pageNumber="14" phylum="Platyhelminthes" rank="species" species="marmotae">
<emphasis id="DEA2EAA56D40FFADBAEEBAC7F1812CF9" box="[702,950,152,176]" italics="true" pageId="13" pageNumber="14">Cittotaenia marmotae</emphasis>
(
<bibRefCitation id="88474B466D40FFADBB96BAC8F6532CE6" author="Denegri" box="[966,1124,151,175]" pageId="13" pageNumber="14" refString="Denegri, G. M. (1993) Review of oribatid mites as intermediate hosts of the Anoplocephalidae. Experimental and Applied Acarology, 17 (8), 567 - 580." type="journal article" year="1993">Denegri 1993</bibRefCitation>
)
</taxonomicName>
and is a host of protozoan Apicomplexa— 
<taxonomicName id="2BD64D346D40FFADB91ABAE3F1BC2C9D" authority="Purrini &amp; Ormieres 1981" authorityName="Purrini &amp; Ormieres" authorityYear="1981" box="[330,907,188,212]" class="Sporozoa" family="Gregarinidae" genus="Erhardovina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eugregarinida" pageId="13" pageNumber="14" phylum="Protozoa" rank="species" species="bisphaera">
<emphasis id="DEA2EAA56D40FFADB91ABAE3F0672C9D" box="[330,592,188,212]" italics="true" pageId="13" pageNumber="14">Erhardovina bisphaera</emphasis>
(
<bibRefCitation id="88474B466D40FFADBA31BAE3F1B42C9D" author="Purrini" box="[609,899,188,212]" pageId="13" pageNumber="14" refString="Purrini, K. &amp; Ormieres, R. (1981) Uber vier neue Eugregarinen-Arten (Eugregrarinida, Sporozoa) der Hornmilben (Oribatei, Acarina). Zoologische Beitrage, 27, 123 - 132." type="journal article" year="1981">Purrini &amp; Ormieres 1981</bibRefCitation>
)
</taxonomicName>
and Microspora – 
<taxonomicName id="2BD64D346D40FFADBC38BAE2F2EB2CBE" authority="Geest et al. 2000" authorityName="Geest et al." authorityYear="2000" class="Microsporea" family="Nosematidae" genus="Nosema" kingdom="Plantae" order="Dissociodihaplophasida" pageId="13" pageNumber="14" phylum="Microsporidia" rank="species" species="acari">
<emphasis id="DEA2EAA56D40FFADBC38BAE2F7312C9D" box="[1128,1286,189,212]" italics="true" pageId="13" pageNumber="14">Nosema acari</emphasis>
(
<bibRefCitation id="88474B466D40FFADBD47BAE3F2E32CBE" author="Geest" pageId="13" pageNumber="14" refString="Geest, L. P. S., Elliot, S. L. van der, Breeuwer, J. A. J. &amp; Beerling E. A. M. (2000) Diseases of mites. Experimental and Applied Acarology, 24 (7), 497 - 560." type="journal article" year="2000">
Geest 
<emphasis id="DEA2EAA56D40FFADBD32BAE2F7A22C9D" box="[1378,1429,188,212]" italics="true" pageId="13" pageNumber="14">et al</emphasis>
. 2000
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>