198 lines
25 KiB
XML
198 lines
25 KiB
XML
<document id="4ED104DDF79F1E7EBF4039297CF7708D" ID-DOI="10.11646/zootaxa.2462.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5312710" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629676798216" checkinUser="felipe" docAuthor="Hyman, Isabel T. & Ponder, Winston F." docDate="2010" docId="6413F378FFF96A65F28B72D4FD61FCD1" docLanguage="en" docName="zt02462p148.pdf" docOrigin="Zootaxa 2462 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2462.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Microcystinae Thiele 1931" docType="treatment" docVersion="4" lastPageNumber="112" masterDocId="982A8B00FF976A0AF21C766BFFEBFF8B" masterDocTitle="A morphological phylogenetic analysis and generic revision of Australian Helicarionidae (Gastropoda: Pulmonata: Stylommatophora), and an assessment of the relationships of the family 2462" masterLastPageNumber="148" masterPageNumber="1" pageNumber="111" updateTime="1699094437759" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="D6B684A9727F331D549C2F50A28CD904">
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<mods:title id="9AA78AAE005C63484571F6A312C14534">A morphological phylogenetic analysis and generic revision of Australian Helicarionidae (Gastropoda: Pulmonata: Stylommatophora), and an assessment of the relationships of the family 2462</mods:title>
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<mods:namePart id="C00AE1370CB1BF68867703366840A584">Hyman, Isabel T.</mods:namePart>
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<mods:roleTerm id="540B91A667CADF782F58B5CF463013FC">Author</mods:roleTerm>
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<mods:namePart id="8DF0A0018C486AE5CFEBCD6C33E41520">Ponder, Winston F.</mods:namePart>
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<mods:typeOfResource id="36E77F8D5AC8B3042E263D5EB099366B">text</mods:typeOfResource>
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<mods:title id="ACA6B361C40B9356513F6C2F7D331B84">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="7D7C00178BE16AE36A87C8DF5128508E">
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<mods:date id="80E0DDBF9B338917E523CAD33125BFDA">2010</mods:date>
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<mods:number id="D444EEC0EB114054E5F905004940B864">2010-05-14</mods:number>
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<mods:detail id="7B8122A3CE70C18FCD32DF7B56F042DB" type="volume">
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<mods:number id="8CC9BAE86B7CB10B4827C2232602AD17">2462</mods:number>
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<mods:detail id="194B65BDBDD183EB0FF842A35AF9294B" type="issue">
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<mods:location id="10D316E6E9CBE1D72F2EE9245AF211B1">
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<mods:url id="0D5557B7F53A787BEA8EEE6EB39617D9">https://biotaxa.org/Zootaxa/article/view/zootaxa.2462.1.1</mods:url>
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<mods:classification id="1C2B51D566CE8B280A1731CB3F29A522">journal article</mods:classification>
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<mods:identifier id="16AD38387F1966052621C549B4A3FCEC" type="DOI">10.11646/zootaxa.2462.1.1</mods:identifier>
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<mods:identifier id="674B8A17E5853944052657EB1E037060" type="ISSN">11755334</mods:identifier>
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<mods:identifier id="55B4841B4C6EEF517284710499AD667B" type="Zenodo-Dep">5312710</mods:identifier>
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<treatment id="6413F378FFF96A65F28B72D4FD61FCD1" ID-DOI="http://doi.org/10.5281/zenodo.5321095" ID-Zenodo-Dep="5321095" LSID="urn:lsid:plazi:treatment:6413F378FFF96A65F28B72D4FD61FCD1" httpUri="http://treatment.plazi.org/id/6413F378FFF96A65F28B72D4FD61FCD1" lastPageId="111" lastPageNumber="112" pageId="110" pageNumber="111">
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<subSubSection id="A4A011E5FFF96A64F28B72D4FE3BFB52" box="[151,464,1215,1241]" pageId="110" pageNumber="111" type="nomenclature">
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<paragraph id="EC05426EFFF96A64F28B72D4FE3BFB52" blockId="110.[151,464,1215,1241]" box="[151,464,1215,1241]" pageId="110" pageNumber="111">
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<heading id="B74DF502FFF96A64F28B72D4FE3BFB52" bold="true" box="[151,464,1215,1241]" fontSize="11" level="1" pageId="110" pageNumber="111" reason="1">
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<emphasis id="DECE9E7CFFF96A64F28B72D4FE3BFB52" bold="true" box="[151,464,1215,1241]" pageId="110" pageNumber="111">
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Subfamily
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<taxonomicName id="2BBA39EDFFF96A64F30372D4FE3BFB52" ID-CoL="7NXTF" authorityName="Thiele" authorityYear="1931" box="[287,464,1215,1241]" class="Gastropoda" family="Euconulidae" kingdom="Animalia" order="Stylommatophora" pageId="110" pageNumber="111" phylum="Mollusca" rank="subFamily" subFamily="Microcystinae">Microcystinae</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="A4A011E5FFF96A64F28B7361FAAFF87F" pageId="110" pageNumber="111" type="discussion">
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<paragraph id="EC05426EFFF96A64F28B7361FB90F99F" blockId="110.[151,1436,1290,2036]" pageId="110" pageNumber="111">
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<emphasis id="DECE9E7CFFF96A64F28B7361FEFBFAAF" bold="true" box="[151,272,1290,1316]" pageId="110" pageNumber="111">Remarks.</emphasis>
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In our analyses
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<taxonomicName id="2BBA39EDFFF96A64F3CB7361FD95FAAF" authorityName="Thiele" authorityYear="1931" box="[471,638,1290,1316]" class="Gastropoda" family="Euconulidae" kingdom="Animalia" order="Stylommatophora" pageId="110" pageNumber="111" phylum="Mollusca" rank="subFamily" subFamily="Microcystinae">Microcystinae</taxonomicName>
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is monophyletic only if
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<taxonomicName id="2BBA39EDFFF96A64F1BB7360FB6FFAAF" box="[935,1156,1291,1316]" class="Gastropoda" family="Euconulidae" genus="Liardetia" kingdom="Animalia" order="Stylommatophora" pageId="110" pageNumber="111" phylum="Mollusca" rank="species" species="doliolum">
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<emphasis id="DECE9E7CFFF96A64F1BB7360FB6FFAAF" box="[935,1156,1291,1316]" italics="true" pageId="110" pageNumber="111">Liardetia doliolum</emphasis>
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</taxonomicName>
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is excluded (
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<figureCitation id="74815EEBFFF96A64F7397361FA61FAAF" box="[1317,1418,1290,1316]" captionStart="FIGURE 1" captionStartId="13.[151,255,1878,1902]" captionTargetBox="[174,1417,215,1826]" captionTargetId="figure-53@13.[154,1439,197,1848]" captionTargetPageId="13" captionText="FIGURE 1. Majority-rule consensus tree (1,743 trees, length 117 steps) from the morphological analysis containing representatives of all groups. Families and subfamilies are shaded and are labelled in bold if the group is not monophyletic." figureDoi="http://doi.org/10.5281/zenodo.5312714" httpUri="https://zenodo.org/record/5312714/files/figure.png" pageId="110" pageNumber="111">Figure 1</figureCitation>
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).
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<bibRefCitation id="882B3F9FFFF96A64F28B7359FED9FAC7" author="Baker, H. B." box="[151,306,1330,1356]" pageId="110" pageNumber="111" pagination="1 - 102" refId="ref72753" refString="Baker, H. B. (1938) Zonitid snails from Pacific Islands Part 1: Southern genera of Microcystinae. Bernice P. Bishop Museum Bulletin, 158, 1 - 102." type="journal article" year="1938">Baker (1938)</bibRefCitation>
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noted similarities between the shell of
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<taxonomicName id="2BBA39EDFFF96A64F0E47358FC8FFAC7" authorityName="Gude" authorityYear="1913" box="[760,868,1331,1356]" class="Gastropoda" family="Euconulidae" genus="Liardetia" kingdom="Animalia" order="Stylommatophora" pageId="110" pageNumber="111" phylum="Mollusca" rank="genus">
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<emphasis id="DECE9E7CFFF96A64F0E47358FC8FFAC7" box="[760,868,1331,1356]" italics="true" pageId="110" pageNumber="111">Liardetia</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="2BBA39EDFFF96A64F1807359FBC5FAC7" authorityName="H. B. Baker" authorityYear="1928" box="[924,1070,1330,1356]" class="Gastropoda" family="Euconulidae" kingdom="Animalia" order="Stylommatophora" pageId="110" pageNumber="106" phylum="Mollusca" rank="subFamily" subFamily="Euconulinae">Euconulinae</taxonomicName>
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and
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<taxonomicName id="2BBA39EDFFF96A64F67A7359FB32FAC7" box="[1126,1241,1330,1356]" genus="The" pageId="110" pageNumber="83" rank="species" species="radula">the radula</taxonomicName>
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of
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<taxonomicName id="2BBA39EDFFF96A64F6E37358FA80FAC7" authorityName="Gude" authorityYear="1913" box="[1279,1387,1331,1356]" class="Gastropoda" family="Euconulidae" genus="Liardetia" kingdom="Animalia" order="Stylommatophora" pageId="110" pageNumber="111" phylum="Mollusca" rank="genus">
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<emphasis id="DECE9E7CFFF96A64F6E37358FA80FAC7" box="[1279,1387,1331,1356]" italics="true" pageId="110" pageNumber="111">Liardetia</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="2BBA39EDFFF96A64F28B7330FF1DFAFF" authorityName="W. T. Blanford" authorityYear="1863" box="[151,246,1371,1396]" class="Gastropoda" family="Chronidae" genus="Kaliella" kingdom="Animalia" order="Stylommatophora" pageId="110" pageNumber="111" phylum="Mollusca" rank="genus">
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<emphasis id="DECE9E7CFFF96A64F28B7330FF1DFAFF" box="[151,246,1371,1396]" italics="true" pageId="110" pageNumber="111">Kaliella</emphasis>
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</taxonomicName>
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(currently placed in
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<taxonomicName id="2BBA39EDFFF96A64F3F67331FD8CFAFF" box="[490,615,1370,1396]" class="Gastropoda" family="Chronidae" kingdom="Animalia" order="Stylommatophora" pageId="110" pageNumber="111" phylum="Mollusca" rank="family">Chronidae</taxonomicName>
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;
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<bibRefCitation id="882B3F9FFFF96A64F0687331FCC3FAFF" author="Hausdorf, B." box="[628,808,1370,1396]" pageId="110" pageNumber="111" pagination="35 - 66" refId="ref74906" refString="Hausdorf, B. (1998) Phylogeny of the Limacoidea sensu lato (Gastropoda: Stylommatophora). Journal of Molluscan Studies, 64, 35 - 66." type="journal article" year="1998">Hausdorf 1998</bibRefCitation>
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). However, while there are some differences between
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<taxonomicName id="2BBA39EDFFF96A64F28B73E8FEF5FA17" box="[151,286,1411,1436]" class="Gastropoda" family="Euconulidae" genus="Liardetia" kingdom="Animalia" order="Stylommatophora" pageId="110" pageNumber="111" phylum="Mollusca" rank="species" species="doliolum">
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<emphasis id="DECE9E7CFFF96A64F28B73E8FEF5FA17" box="[151,286,1411,1436]" italics="true" pageId="110" pageNumber="111">L. doliolum</emphasis>
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</taxonomicName>
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and the other
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<taxonomicName id="2BBA39EDFFF96A64F3DA73E9FD85FA17" authorityName="Thiele" authorityYear="1931" box="[454,622,1410,1436]" class="Gastropoda" family="Euconulidae" kingdom="Animalia" order="Stylommatophora" pageId="110" pageNumber="111" phylum="Mollusca" rank="subFamily" subFamily="Microcystinae">Microcystinae</taxonomicName>
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included in the current study (e.g., bursa copulatrix at the junction of the penis and vagina rather than the base of the penis; presence of an epiphallic caecum), the majority of the characters place this species in
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<taxonomicName id="2BBA39EDFFF96A64F03473B9FD3FFA67" authorityName="Thiele" authorityYear="1931" box="[552,724,1490,1516]" class="Gastropoda" family="Euconulidae" kingdom="Animalia" order="Stylommatophora" pageId="110" pageNumber="111" phylum="Mollusca" rank="subFamily" subFamily="Microcystinae">Microcystinae</taxonomicName>
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. When
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<taxonomicName id="2BBA39EDFFF96A64F12673B8FBB9FA67" box="[826,1106,1490,1516]" class="Gastropoda" family="Helicarionidae" genus="Durgella" kingdom="Animalia" order="Stylommatophora" pageId="110" pageNumber="111" phylum="Mollusca" rank="species" species="hatchongi">
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<emphasis id="DECE9E7CFFF96A64F12673B8FC48FA67" box="[826,931,1491,1516]" italics="true" pageId="110" pageNumber="111">Durgella</emphasis>
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cf.
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<emphasis id="DECE9E7CFFF96A64F1C173B8FBB9FA67" box="[989,1106,1491,1516]" italics="true" pageId="110" pageNumber="111">hatchongi</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="2BBA39EDFFF96A64F68573B8FAFDFA67" authority="Hyman & Ponder, 2010" authorityName="Hyman & Ponder" authorityYear="2010" box="[1177,1302,1491,1516]" class="Gastropoda" family="Cystopeltidae" genus="Cystopelta" kingdom="Animalia" order="Stylommatophora" pageId="110" pageNumber="111" phylum="Mollusca" rank="genus" status="sp. nov.">
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<emphasis id="DECE9E7CFFF96A64F68573B8FAFDFA67" box="[1177,1302,1491,1516]" italics="true" pageId="110" pageNumber="111">Cystopelta</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="C5FD2307FFF96A64F73873B9FA81FA67" box="[1316,1386,1490,1516]" pageId="110" pageNumber="111" rank="species">n. sp.</taxonomicNameLabel>
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are excluded,
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<taxonomicName id="2BBA39EDFFF96A64F3137390FE7DF99F" box="[271,406,1531,1556]" class="Gastropoda" family="Euconulidae" genus="Liardetia" kingdom="Animalia" order="Stylommatophora" pageId="110" pageNumber="111" phylum="Mollusca" rank="species" species="doliolum">
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<emphasis id="DECE9E7CFFF96A64F3137390FE7DF99F" box="[271,406,1531,1556]" italics="true" pageId="110" pageNumber="111">L. doliolum</emphasis>
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</taxonomicName>
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is included in a monophyletic
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<taxonomicName id="2BBA39EDFFF96A64F11C7391FC43F99F" authorityName="Thiele" authorityYear="1931" box="[768,936,1530,1556]" class="Gastropoda" family="Euconulidae" kingdom="Animalia" order="Stylommatophora" pageId="110" pageNumber="111" phylum="Mollusca" rank="subFamily" subFamily="Microcystinae">Microcystinae</taxonomicName>
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(data not shown).
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</paragraph>
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<paragraph id="EC05426EFFF96A64F2DF7049FDF1F907" blockId="110.[151,1436,1290,2036]" pageId="110" pageNumber="111">
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<bibRefCitation id="882B3F9FFFF96A64F2DF7049FE62F9B7" author="Hausdorf, B." box="[195,393,1570,1596]" pageId="110" pageNumber="111" pagination="35 - 66" refId="ref74906" refString="Hausdorf, B. (1998) Phylogeny of the Limacoidea sensu lato (Gastropoda: Stylommatophora). Journal of Molluscan Studies, 64, 35 - 66." type="journal article" year="1998">Hausdorf (1998)</bibRefCitation>
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gave the following synapomorphies for this group: bursa copulatrix male side, partial detachment of the prostate, ovoviviparous reproduction. These characters are also included in
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<bibRefCitation id="882B3F9FFFF96A64F70F7021FF19F907" author="Schileyko, A. A." pageId="110" pageNumber="111" refId="ref76865" refString="Schileyko, A. A. (2002 a) Treatise on Recent Terrestrial Pulmonate Molluscs, Part 8: Punctidae, Helicodiscidae, Discidae, Cystopeltidae, Euconulidae, Trochomorphidae. Ruthenica, Supplement 2, Moscow." type="book" year="2002">Schileyko’s (2002a)</bibRefCitation>
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description of the group.
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</paragraph>
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<paragraph id="EC05426EFFF96A64F2DF70F1FDCEF8DF" blockId="110.[151,1436,1290,2036]" pageId="110" pageNumber="111">
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Several of the species included in
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<taxonomicName id="2BBA39EDFFF96A64F04670F1FCE9F93F" authorityName="Thiele" authorityYear="1931" box="[602,770,1690,1716]" class="Gastropoda" family="Euconulidae" kingdom="Animalia" order="Stylommatophora" pageId="110" pageNumber="111" phylum="Mollusca" rank="subFamily" subFamily="Microcystinae">Microcystinae</taxonomicName>
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in the current study were placed in subfamilies of
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<taxonomicName id="2BBA39EDFFF96A64F77C70F1FEFAF957" authorityName="Bourguignat" authorityYear="1877" class="Gastropoda" family="Helicarionidae" kingdom="Animalia" order="Stylommatophora" pageId="110" pageNumber="111" phylum="Mollusca" rank="family">Helicarionidae</taxonomicName>
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by
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<bibRefCitation id="882B3F9FFFF96A64F34A70A9FDD0F957" author="Schileyko, A. A." box="[342,571,1730,1756]" pageId="110" pageNumber="111" refId="ref76904" refString="Schileyko, A. A. (2002 b) Treatise on Recent Terrestrial Pulmonate Molluscs, Part 9: Helicarionidae, Gymnarionidae, Rhysotinidae, Ariophantidae. Ruthenica, Supplement 2, Moscow." type="book" year="2002">Schileyko (2002b)</bibRefCitation>
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(e.g.,
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<taxonomicName id="2BBA39EDFFF96A64F08370A8FCE4F957" authorityName="Preston" authorityYear="1913" box="[671,783,1731,1756]" class="Gastropoda" family="Euconulidae" genus="Roybellia" kingdom="Animalia" order="Stylommatophora" pageId="110" pageNumber="111" phylum="Mollusca" rank="genus">
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<emphasis id="DECE9E7CFFF96A64F08370A8FCE4F957" box="[671,783,1731,1756]" italics="true" pageId="110" pageNumber="111">Roybellia</emphasis>
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</taxonomicName>
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in Geotrochinae,
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<taxonomicName id="2BBA39EDFFF96A64F61B70A8FB4BF957" authorityName="Iredale" authorityYear="1944" box="[1031,1184,1731,1756]" class="Gastropoda" family="Euconulidae" genus="Melloconcha" kingdom="Animalia" order="Stylommatophora" pageId="110" pageNumber="111" phylum="Mollusca" rank="genus">
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<emphasis id="DECE9E7CFFF96A64F61B70A8FB4BF957" box="[1031,1184,1731,1756]" italics="true" pageId="110" pageNumber="111">Melloconcha</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="2BBA39EDFFF96A64F6A770A8FAB5F957" authorityName="Iredale" authorityYear="1944" box="[1211,1374,1731,1756]" class="Gastropoda" family="Euconulidae" genus="Innesoconcha" kingdom="Animalia" order="Stylommatophora" pageId="110" pageNumber="111" phylum="Mollusca" rank="genus">
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<emphasis id="DECE9E7CFFF96A64F6A770A8FAB5F957" box="[1211,1374,1731,1756]" italics="true" pageId="110" pageNumber="111">Innesoconcha</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="2BBA39EDFFF96A64F28B7080FE8FF88F" authorityName="Preston" authorityYear="1913" box="[151,356,1771,1796]" class="Gastropoda" family="Euconulidae" genus="Mathewsoconcha" kingdom="Animalia" order="Stylommatophora" pageId="110" pageNumber="111" phylum="Mollusca" rank="genus">
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<emphasis id="DECE9E7CFFF96A64F28B7080FE8FF88F" box="[151,356,1771,1796]" italics="true" pageId="110" pageNumber="111">Mathewsoconcha</emphasis>
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</taxonomicName>
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in Helicarioninae) based on shell data. This will be discussed further in future papers [Hyman & Ponder, unpublished data based on
|
||
<bibRefCitation id="882B3F9FFFF96A64F0D47179FC90F8A7" author="Hyman, I. T." box="[712,891,1810,1836]" pageId="110" pageNumber="111" refId="ref75213" refString="Hyman, I. T. (2005) Taxonomy, systematics and evolutionary trends in Helicarionidae (Mollusca, Pulmonata), PhD thesis, University of Sydney, 583 pp." type="book" year="2005">Hyman (2005)</bibRefCitation>
|
||
], where the
|
||
<taxonomicName id="2BBA39EDFFF96A64F6127179FB5EF8A7" authorityName="Thiele" authorityYear="1931" box="[1038,1205,1810,1836]" class="Gastropoda" family="Euconulidae" kingdom="Animalia" order="Stylommatophora" pageId="110" pageNumber="111" phylum="Mollusca" rank="subFamily" subFamily="Microcystinae">Microcystinae</taxonomicName>
|
||
of Lord Howe and
|
||
<collectingRegion id="2E7E8C8CFFF96A64F28B7151FF1FF8DF" box="[151,244,1850,1876]" country="United Kingdom" name="Norfolk" pageId="110" pageNumber="111">Norfolk</collectingRegion>
|
||
Islands will be described.
|
||
</paragraph>
|
||
<paragraph id="EC05426EFFF96A64F2DF7109FC21F8F7" blockId="110.[151,1436,1290,2036]" box="[195,970,1890,1916]" pageId="110" pageNumber="111">
|
||
Based on our analyses, key characters defining
|
||
<taxonomicName id="2BBA39EDFFF96A64F0F37109FC7CF8F7" authorityName="Thiele" authorityYear="1931" box="[751,919,1890,1916]" class="Gastropoda" family="Euconulidae" kingdom="Animalia" order="Stylommatophora" pageId="110" pageNumber="111" phylum="Mollusca" rank="subFamily" subFamily="Microcystinae">Microcystinae</taxonomicName>
|
||
are:
|
||
</paragraph>
|
||
<paragraph id="EC05426EFFF96A64F2DF71E1FAAFF87F" blockId="110.[151,1436,1290,2036]" pageId="110" pageNumber="111">Shell rarely reduced. Mantle with no visible minor blood vessels. Genital system ovoviviparous; bursa copulatrix usually very short, inserted on penis; stimulator absent; epiphallic retractor caecum and flagellum absent in most taxa. Spermatophore small, peanut-shaped, soft-walled; attached to inner wall of penis.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A4A011E5FFF86A65F2DA76FCFD61FCD1" pageId="111" pageNumber="112" type="description">
|
||
<paragraph id="EC05426EFFF86A65F2DA76FCFD61FCD1" blockId="111.[151,1437,151,858]" pageId="111" pageNumber="112">
|
||
<emphasis id="DECE9E7CFFF86A65F2DA76FCFEB5FF3A" bold="true" box="[198,350,151,177]" pageId="111" pageNumber="112">Description.</emphasis>
|
||
Shell present, rarely reduced, ranging in diameter from ~
|
||
<quantity id="2B42EF8BFFF86A65F60F76F3FB9BFF39" box="[1043,1136,152,178]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="111" pageNumber="112" unit="mm" value="2.5">2.5 mm</quantity>
|
||
in
|
||
<taxonomicName id="2BBA39EDFFF86A65F68576F3FEDCFF51" authority="Preston, 1913" authorityName="Preston" authorityYear="1913" class="Gastropoda" family="Helicarionidae" genus="Pittoconcha" kingdom="Animalia" order="Stylommatophora" pageId="111" pageNumber="112" phylum="Mollusca" rank="species" species="concinna">
|
||
<emphasis id="DECE9E7CFFF86A65F68576F3FA70FF3A" box="[1177,1435,152,177]" italics="true" pageId="111" pageNumber="112">Pittoconcha concinna</emphasis>
|
||
Preston, 1913
|
||
</taxonomicName>
|
||
to
|
||
<quantity id="2B42EF8BFFF86A65F34076ABFE46FF51" box="[348,429,192,218]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="111" pageNumber="112" unit="mm" value="20.0">20 mm</quantity>
|
||
in
|
||
<taxonomicName id="2BBA39EDFFF86A65F3CE76ABFC49FF51" authority="(Gray, 1834)" baseAuthorityName="Gray" baseAuthorityYear="1834" box="[466,930,192,218]" class="Gastropoda" family="Euconulidae" genus="Mathewsoconcha" kingdom="Animalia" order="Stylommatophora" pageId="111" pageNumber="112" phylum="Mollusca" rank="species" species="phillipii">
|
||
<emphasis id="DECE9E7CFFF86A65F3CE76ABFCE8FF52" box="[466,771,192,217]" italics="true" pageId="111" pageNumber="112">Mathewsoconcha phillipii</emphasis>
|
||
(
|
||
<bibRefCitation id="882B3F9FFFF86A65F10D76ABFC72FF51" author="Gray, J. E." box="[785,921,192,218]" pageId="111" pageNumber="112" pagination="63 - 68" refId="ref74735" refString="Gray, J. E. (1834) Characters of new species of shells. Proceedings of the Zoological Society of London, 1834, 63 - 68." type="journal article" year="1834">Gray, 1834</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Shell shape and sculpture variable. Mantle laps absent, or only right lap present, or both laps present; mantle laps long and narrow in shape; not fused. Mantle lobes small. Sole of foot tripartite; caudal apparatus formed from curled up sole (
|
||
<bibRefCitation id="882B3F9FFFF86A65F6BF777BFAB4FEA1" author="Muratov, I. V." box="[1187,1375,272,298]" pageId="111" pageNumber="112" pagination="5 - 26" refId="ref75844" refString="Muratov, I. V. (1999) Analysis of the phylogenetic relationships and their systematic implications in the Limacoinei (= Zonitinia) infraorder (Gastropoda, Pulmonata, Geophila). Ruthenica, 9, 5 - 26." type="journal article" year="1999">Muratov’s 1999</bibRefCitation>
|
||
helicarionid
|
||
<typeStatus id="3301FCCCFFF86A65F2E27753FEDEFED9" box="[254,309,312,338]" pageId="111" pageNumber="112">type</typeStatus>
|
||
); caudal apparatus very small to moderately large; caudal foss horizontal slit in tail. Kidney unilobed; mantle with no visible minor blood vessels, pigmentation absent or present, mantle gland absent or present. Genital system ovoviviparous; oviduct non-glandular. Neither carrefour nor talon embedded in albumen gland. Post-uterine oviduct very long; capsular gland absent. Bursa copulatrix very short in most taxa; inserted on penis; duct of bursa copulatrix wide; not distinguishable from bursa copulatrix (in
|
||
<taxonomicName id="2BBA39EDFFF86A65F6FF77B3FF28FD92" class="Gastropoda" family="Euconulidae" genus="Liardetia" kingdom="Animalia" order="Stylommatophora" pageId="111" pageNumber="112" phylum="Mollusca" rank="species" species="doliolum">
|
||
<emphasis id="DECE9E7CFFF86A65F6FF77B3FF28FD92" italics="true" pageId="111" pageNumber="112">Liardetia doliolum</emphasis>
|
||
</taxonomicName>
|
||
bursa copulatrix is short, inserted at junction of penis and post-uterine oviduct and duct of bursa copulatrix is narrow, distinguishable from bursa copulatrix). Stimulator absent. Epiphallus enters penis through simple pore, rarely two fleshy lips or verge; penis internally with no sculpture, or with longitudinal ridges or longitudinal, circular or transverse pilasters. Penial sheath present, enclosing coiled penis and epiphallus; closed at distal end; penis retractor muscle attached to penis or to epiphallus near penis junction. Epiphallic retractor caecum absent (most taxa) or present (
|
||
<taxonomicName id="2BBA39EDFFF86A65F0CA74A3FC58FD6A" box="[726,947,712,737]" class="Gastropoda" family="Euconulidae" genus="Liardetia" kingdom="Animalia" order="Stylommatophora" pageId="111" pageNumber="112" phylum="Mollusca" rank="species" species="doliolum">
|
||
<emphasis id="DECE9E7CFFF86A65F0CA74A3FC58FD6A" box="[726,947,712,737]" italics="true" pageId="111" pageNumber="112">Liardetia doliolum</emphasis>
|
||
</taxonomicName>
|
||
); epiphallic flagellum absent. Spermatophore small, peanut-shaped, soft-walled; attached to inner wall of penis. Radula relatively long and narrow. Central tooth with small to moderately small ectocones; mesocone lanceolate. Lateral and marginal tooth fields distinguishable. Tooth rows straight.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |