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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.121.30254" ID-GBIF-Dataset="b12b6eed-6d17-4e2b-b290-f5c01bf79271" ID-PMC="PMC6509104" ID-Pensoft-Pub="1314-2003-121-53" ID-Pensoft-UUID="A90CEB1FFFFDE325FFC1FFD53E14FF93" ID-PubMed="31118872" ID-Zenodo-Dep="2702029" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1314-2003-121-53" ModsDocOrigin="PhytoKeys 121" ModsDocTitle="Typification of eight current and seven related names and a new section in the genus Bromus (Bromeae, Pooideae, Poaceae)" checkinTime="1556793052295" checkinUser="pensoft" docAuthor="Llamas, Felix &amp; Acedo, Carmen" docDate="2019" docId="636E9E2DEF005C463D43BFD3AE44EB56" docLanguage="en" docName="PhytoKeys 121: 53-72" docOrigin="PhytoKeys 121" docSource="http://dx.doi.org/10.3897/phytokeys.121.30254" docTitle="Bromus hordeaceus subsp. divaricatus Kerguelen, Soc. Echange Pl. Vasc. Eur. Bassin Medit., Bull. 18: 27. 1981" docType="treatment" docVersion="6" id="A90CEB1FFFFDE325FFC1FFD53E14FF93" lastPageNumber="59" masterDocId="A90CEB1FFFFDE325FFC1FFD53E14FF93" masterDocTitle="Typification of eight current and seven related names and a new section in the genus Bromus (Bromeae, Pooideae, Poaceae)" masterLastPageNumber="72" masterPageNumber="53" pageNumber="58" updateTime="1668139280325" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Typification of eight current and seven related names and a new section in the genus Bromus (Bromeae, Pooideae, Poaceae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Llamas, Felix</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Acedo, Carmen</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>121</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>53</mods:start>
<mods:end>72</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.121.30254</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.121.30254</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-121-53</mods:identifier>
<mods:identifier type="Pensoft-UUID">A90CEB1FFFFDE325FFC1FFD53E14FF93</mods:identifier>
<mods:identifier type="Zenodo-Dep">2702029</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="156890807" LSID="urn:lsid:plazi:treatment:636E9E2DEF005C463D43BFD3AE44EB56" httpUri="http://treatment.plazi.org/id/636E9E2DEF005C463D43BFD3AE44EB56" lastPageId="6" lastPageNumber="59" pageId="5" pageNumber="58">
<subSubSection pageId="5" pageNumber="58" type="nomenclature">
<paragraph pageId="5" pageNumber="58">
<taxonomicName LSID="5C5A266B-129E-563A-A0B6-5F70E8023DCC" authority="L. subsp. divaricatus (Bonnier &amp; Layens) Kerguelen, Soc. Echange Pl. Vasc. Eur. Bassin Medit., Bull. 18: 27. (1981)" class="Liliopsida" family="Poaceae" genus="Bromus" higherTaxonomySource="CoL" infraspecific-rank="subsp." kingdom="Plantae" lsidName="Bromus hordeaceus" order="Poales" pageId="5" pageNumber="58" phylum="Tracheophyta" rank="species" species="hordeaceus">
<pageBreakToken pageId="5" pageNumber="58" start="start">Bromus</pageBreakToken>
hordeaceus L. subsp. divaricatus (Bonnier &amp; Layens)
<normalizedToken originalValue="Kerguélen">Kerguelen</normalizedToken>
, Soc. Echange Pl. Vasc. Eur. Bassin
<normalizedToken originalValue="Médit">Medit</normalizedToken>
., Bull. 18: 27. (1981)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="58" type="reference_group">
<paragraph pageId="5" pageNumber="58">
<taxonomicName authority="Guss. subsp. divaricatus" class="Liliopsida" family="Poaceae" genus="Bromus" higherTaxonomySource="CoL" infraspecific-rank="subsp." kingdom="Plantae" lsidName="Bromus intermedius" order="Poales" pageId="5" pageNumber="58" phylum="Tracheophyta" rank="species" species="intermedius" subspecies="divaricatus">Bromus intermedius Guss. subsp. divaricatus</taxonomicName>
Bonnier &amp; Layens, Tabl. Syn. Pl. Vasc. Fl. Fr. 369. (1894), non
<taxonomicName family="Poaceae" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="B. divaricatus" order="Poales" pageId="5" pageNumber="58" rank="species" species="divaricatus">
<emphasis italics="true" pageId="5" pageNumber="58">B. divaricatus</emphasis>
</taxonomicName>
sensu Rhode ex DC. Type Protologue: [France] &quot;Endroits incultes, sables&quot;. Type: [France], (lectotype, designated here: [figure] in Bonnier and Layens, Tabl. Syn. Pl. Vasc. Fl. Fr. 369. (1894); epitype, designated here: [France]: Pornichet, Loire
<normalizedToken originalValue="Inférieure">Inferieure</normalizedToken>
[Loire-Atlantique], sea shore, June 25 1856, Lloyd, BM [BM001067302 image! as
<taxonomicName class="Liliopsida" family="Poaceae" genus="Bromus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bromus molliformis" order="Poales" pageId="5" pageNumber="58" phylum="Tracheophyta" rank="species" species="molliformis">
<emphasis italics="true" pageId="5" pageNumber="58">Bromus molliformis</emphasis>
</taxonomicName>
Lloyd]). (Figure 5)
</paragraph>
<paragraph pageId="5" pageNumber="58">
<taxonomicName class="Liliopsida" family="Poaceae" genus="Bromus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bromus molliformis" order="Poales" pageId="5" pageNumber="58" phylum="Tracheophyta" rank="species" species="molliformis">Bromus molliformis</taxonomicName>
J.Lloyd ex Billot, Fl. Gall. &amp; Germ. Exsicc. (Haguenau) 1: 297-298. (Feb 1854). Type Protologue: &quot;Loire
<normalizedToken originalValue="Inférieure”">Inferieure&quot;</normalizedToken>
[Loire-Atlantique]. Type: [France]. Le Roc-Saint-Luc, commune de Pissotte,
<normalizedToken originalValue="près">pres</normalizedToken>
de Fontenay-le-Comte (
<normalizedToken originalValue="Vendée">Vendee</normalizedToken>
); June 3, 1853, (lectotype, designated here: P [P02381530 image!]; isolectotypes: P [P03354928, P03354936, P03486702, P03364411]). (Figure 6)
</paragraph>
<paragraph pageId="5" pageNumber="58">
<taxonomicName class="Liliopsida" family="Poaceae" genus="Serrafalcus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Serrafalcus lloydianus" order="Poales" pageId="5" pageNumber="58" phylum="Tracheophyta" rank="species" species="lloydianus">Serrafalcus lloydianus</taxonomicName>
Godr. &amp; Gren. in Grenier and Godron, Fl. France 3: 591 (1855). Type Protologue: &quot;Hab. Sables maritimes; Cannes,
<normalizedToken originalValue="Hyères">Hyeres</normalizedToken>
, Montpellier, etc.; littoral de
<normalizedToken originalValue="l’Océan">l'Ocean</normalizedToken>
depuis Bayonne
<normalizedToken originalValue="jusqu’à">jusqu'a</normalizedToken>
<normalizedToken originalValue="l’embouchure">l'embouchure</normalizedToken>
de la Loire&quot;.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="58" type="materials_examined">
<paragraph pageId="5" pageNumber="58">Type.</paragraph>
<paragraph pageId="5" pageNumber="58">
Based on
<taxonomicName authority="Guss. subsp. divaricatus" class="Liliopsida" family="Gramineae" genus="Bromus" higherTaxonomySource="CoL" infraspecific-rank="subsp." kingdom="Plantae" lsidName="Bromus intermedius" order="Poales" pageId="5" pageNumber="58" phylum="Tracheophyta" rank="species" species="intermedius" subspecies="divaricatus">Bromus intermedius Guss. subsp. divaricatus</taxonomicName>
Bonnier &amp; Layens.
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="59" pageId="5" pageNumber="58" type="remarks">
<paragraph pageId="5" pageNumber="58">Remarks.</paragraph>
<paragraph pageId="5" pageNumber="58">
<taxonomicName class="Liliopsida" family="Gramineae" genus="Bromus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bromus hordeaceus" order="Poales" pageId="5" pageNumber="58" phylum="Tracheophyta" rank="species" species="hordeaceus">
<emphasis italics="true" pageId="5" pageNumber="58">Bromus hordeaceus</emphasis>
</taxonomicName>
L. is a very variable species with a complex nomenclatural history at subspecies level. The origin of many problems in the group is the name
<taxonomicName family="Poaceae" kingdom="Plantae" lsidName="B. molliformis" order="Poales" pageId="5" pageNumber="58" rank="species" species="molliformis">
<emphasis italics="true" pageId="5" pageNumber="58">B. molliformis</emphasis>
</taxonomicName>
J.Lloyd, Fl. Loire-Inf. (
<bibRefCitation author="Lloyd, J" journalOrPublisher="Repertorium Specierum Novarum Regni Vegetabilis" pageId="10" pageNumber="63" refId="B21" refString="Lloyd, J, 1844. Flore de la Loire-Inferieure. Prosper Sevire, Nantes. https://gallica.bnf.fr/ark:/12148/bpt6k984852/f4.image" title="Flore de la Loire-Inferieure. Prosper Sevire, Nantes. https: // gallica. bnf. fr / ark: / 12148 / bpt 6 k 984852 / f 4. image" year="1844">Lloyd 1844</bibRefCitation>
: 314-315), which was invalidly published as a provisional name &quot;je proposerais, si
<normalizedToken originalValue="c’était">c'etait</normalizedToken>
une
<normalizedToken originalValue="espèce">espece</normalizedToken>
nouvelle, de
<normalizedToken originalValue="l’appeler">l'appeler</normalizedToken>
<taxonomicName lsidName="Br. molliformis" pageId="5" pageNumber="58" rank="species" species="molliformis">
<emphasis italics="true" pageId="5" pageNumber="58">Br. molliformis</emphasis>
</taxonomicName>
&quot;. The name was later validated as
<taxonomicName class="Liliopsida" family="Gramineae" genus="Bromus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bromus molliformis" order="Poales" pageId="5" pageNumber="58" phylum="Tracheophyta" rank="species" species="molliformis">
<emphasis italics="true" pageId="5" pageNumber="58">Bromus molliformis</emphasis>
</taxonomicName>
J.Lloyd ex Billot, Fl. Gall. &amp; Germ. Exsicc. (Haguenau) 2: Cent. 14. 1854. Billot supplies a brief description for &quot;cette
<normalizedToken originalValue="espèce">espece</normalizedToken>
<normalizedToken originalValue="donnée">donnee</normalizedToken>
par M.Lloyd dans sa
<emphasis italics="true" pageId="5" pageNumber="58">
Flore de la
<normalizedToken originalValue="Loire-Inférieure">Loire-Inferieure</normalizedToken>
</emphasis>
sous le nom de
<taxonomicName family="Poaceae" kingdom="Plantae" lsidName="B. divaricatitus" order="Poales" pageId="5" pageNumber="58" rank="species" species="divaricatitus">
<emphasis italics="true" pageId="5" pageNumber="58">B. divaricatitus</emphasis>
</taxonomicName>
Rohde?&quot; Billot (Feb. 1854: 297-298) neglects to mention several details of
<normalizedToken originalValue="Lloyd’s">Lloyd's</normalizedToken>
description of
<taxonomicName family="Poaceae" kingdom="Plantae" lsidName="B. molliformis" order="Poales" pageId="5" pageNumber="58" rank="species" species="molliformis">
<emphasis italics="true" pageId="5" pageNumber="58">B. molliformis</emphasis>
</taxonomicName>
and describes the species as follows:
</paragraph>
<paragraph pageId="5" pageNumber="58">
&quot;Racine fibreuse. Chaume de 2-4
<normalizedToken originalValue="décimètres">decimetres</normalizedToken>
. Feuilles et graines
<normalizedToken originalValue="inférieures">inferieures</normalizedToken>
mollement poilues. Panicule oblongue, droite,
<normalizedToken originalValue="étalée">etalee</normalizedToken>
,
<emphasis italics="true" pageId="5" pageNumber="58">
<normalizedToken originalValue="resserrée">resserree</normalizedToken>
<normalizedToken originalValue="après">apres</normalizedToken>
la floraison
</emphasis>
;
<normalizedToken originalValue="pédoncules">pedoncules</normalizedToken>
courts, simples. Epillets oblongs,
<normalizedToken originalValue="étalés">etales</normalizedToken>
,
<emphasis italics="true" pageId="5" pageNumber="58">velus</emphasis>
;
<normalizedToken originalValue="arête">arete</normalizedToken>
<normalizedToken originalValue="égalant">egalant</normalizedToken>
la glumelle,
<normalizedToken originalValue="d’abord">d'abord</normalizedToken>
droite,
<normalizedToken originalValue="à">a</normalizedToken>
la fin
<emphasis italics="true" pageId="5" pageNumber="58">
<normalizedToken originalValue="tortillée">tortillee</normalizedToken>
<normalizedToken originalValue="divariquée">divariquee</normalizedToken>
</emphasis>
,
<normalizedToken originalValue="insérée">inseree</normalizedToken>
<normalizedToken originalValue="à">a</normalizedToken>
1 1/2
<normalizedToken originalValue="millimètre">millimetre</normalizedToken>
du sommet obtus,
<normalizedToken originalValue="échancré”">echancre&quot;</normalizedToken>
.
</paragraph>
<paragraph pageId="5" pageNumber="58">
<bibRefCitation author="Billot, C" journalOrPublisher="Archives de la Flore de France et d'Allemagne" pageId="9" pageNumber="62" pagination="287 - 298" refId="B5" refString="Billot, C, 1854. Flora Galliae et Germaniae Exsiccata. Archives de la Flore de France et d'Allemagne 1: 287 - 298" title="Flora Galliae et Germaniae Exsiccata." volume="1" year="1854">Billot (1854)</bibRefCitation>
publishes the first validating description for
<taxonomicName class="Liliopsida" family="Gramineae" genus="Bromus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bromus molliformis" order="Poales" pageId="5" pageNumber="58" phylum="Tracheophyta" rank="species" species="molliformis">
<emphasis italics="true" pageId="5" pageNumber="58">Bromus molliformis</emphasis>
</taxonomicName>
. He includes the reference of the features to differentiate the new species from the closely related
<taxonomicName family="Poaceae" kingdom="Plantae" lsidName="B. hordeaceus" order="Poales" pageId="5" pageNumber="58" rank="species" species="hordeaceus">
<emphasis italics="true" pageId="5" pageNumber="58">B. hordeaceus</emphasis>
</taxonomicName>
L. and from
<taxonomicName family="Poaceae" kingdom="Plantae" lsidName="B. divaricatus" order="Poales" pageId="5" pageNumber="58" rank="species" species="divaricatus">
<emphasis italics="true" pageId="5" pageNumber="58">B. divaricatus</emphasis>
</taxonomicName>
Rhode. He sold his exsiccata to several herbaria. There is not a register of those herbaria. Currently, some of them are included in P where some original specimens with its species number (1386) are preserved. Additionally, there are several names and combinations at several ranks, both validly and invalidly published and various nomenclatural changes. Nevertheless,
<taxonomicName family="Poaceae" kingdom="Plantae" lsidName="B. molliformis" order="Poales" pageId="5" pageNumber="58" rank="species" species="molliformis">
<emphasis italics="true" pageId="5" pageNumber="58">B. molliformis</emphasis>
</taxonomicName>
J.Lloyd ex Billot remained untypified and still lacked a stable nomenclature.
</paragraph>
<paragraph lastPageId="6" lastPageNumber="59" pageId="5" pageNumber="58">
<taxonomicName class="Liliopsida" family="Poaceae" genus="Serrafalcus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Serrafalcus lloydianus" order="Poales" pageId="5" pageNumber="58" phylum="Tracheophyta" rank="species" species="lloydianus">
<emphasis italics="true" pageId="5" pageNumber="58">Serrafalcus lloydianus</emphasis>
</taxonomicName>
Godr. &amp; Gren. (1855: 591) is a superfluous name for
<taxonomicName family="Poaceae" kingdom="Plantae" lsidName="B. molliformis" order="Poales" pageId="5" pageNumber="58" rank="species" species="molliformis">
<emphasis italics="true" pageId="5" pageNumber="58">B. molliformis</emphasis>
</taxonomicName>
J.Lloyd ex Billot.
<bibRefCitation author="Grenier, M" journalOrPublisher="Bulletin de la Societe Botanique de France" pageId="9" pageNumber="62" refId="B10" refString="Grenier, M, Godron, M, 1855. Flore de France. J.-B Bailliere, Paris. https://biodiversitylibrary.org/page/6122574" title="Flore de France. J. - B Bailliere, Paris. https: // biodiversitylibrary. org / page / 6122574" year="1855">Grenier and Godron (1855)</bibRefCitation>
listed several synonyms, including &quot;
<taxonomicName family="Poaceae" kingdom="Plantae" lsidName="B. divaricatus" order="Poales" pageId="5" pageNumber="58" rank="species" species="divaricatus">
<emphasis italics="true" pageId="5" pageNumber="58">B. divaricatus</emphasis>
</taxonomicName>
Lloyd non Rhode&quot; and
<taxonomicName family="Poaceae" kingdom="Plantae" lsidName="B. molliformis" order="Poales" pageId="5" pageNumber="58" rank="species" species="molliformis">
<emphasis italics="true" pageId="5" pageNumber="58">B. molliformis</emphasis>
</taxonomicName>
J.Lloyd ex Billot
<pageBreakToken pageId="6" pageNumber="59" start="start">[</pageBreakToken>
sub.
<taxonomicName family="Poaceae" kingdom="Plantae" lsidName="B. molliformis" order="Poales" pageId="6" pageNumber="59" rank="species" species="molliformis">
<emphasis italics="true" pageId="6" pageNumber="59">B. molliformis</emphasis>
</taxonomicName>
Lloyd]. They also mention
<normalizedToken originalValue="Billot’s">Billot's</normalizedToken>
exsiccate n. 1586; and include his new species in the complex of species having
<normalizedToken originalValue="“arête">&quot;arete</normalizedToken>
<normalizedToken originalValue="…">...</normalizedToken>
tordue sur
<normalizedToken originalValue="elle-même">elle-meme</normalizedToken>
et
<normalizedToken originalValue="divariquée”">divariquee&quot;</normalizedToken>
. Therefore, this name does not need a Lectoype as it is a superfluous and illegitimate renaming (Art. 52.1) of
<taxonomicName class="Liliopsida" family="Gramineae" genus="Bromus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bromus molliformis" order="Poales" pageId="6" pageNumber="59" phylum="Tracheophyta" rank="species" species="molliformis">
<emphasis italics="true" pageId="6" pageNumber="59">Bromus molliformis</emphasis>
</taxonomicName>
J.Lloyd ex Billot and has the same type we select here for that name.
</paragraph>
<paragraph pageId="6" pageNumber="59">
<bibRefCitation author="Bonnier, G" journalOrPublisher="Archives de la Flore de France et d'Allemagne" pageId="9" pageNumber="62" refId="B6" refString="Bonnier, G, Layens, G, 1894. Tableaux synoptiques des plantes vasculaires de la Flore de la France. Paul Dupont, Paris. https://doi.org/10.5962/bhl.title.323" title="Tableaux synoptiques des plantes vasculaires de la Flore de la France. Paul Dupont, Paris. https: // doi. org / 10.5962 / bhl. title. 323" year="1894">Bonnier and Layens (1894</bibRefCitation>
: 369) describe
<taxonomicName authority="Guss. subsp. divaricatus" class="Liliopsida" family="Gramineae" genus="Bromus" higherTaxonomySource="CoL" infraspecific-rank="subsp." kingdom="Plantae" lsidName="Bromus intermedius" order="Poales" pageId="6" pageNumber="59" phylum="Tracheophyta" rank="species" species="intermedius" subspecies="divaricatus">Bromus intermedius Guss. subsp. divaricatus</taxonomicName>
Bonnier &amp; Layens, based on
<taxonomicName family="Poaceae" kingdom="Plantae" lsidName="B. divaricatus" order="Poales" pageId="6" pageNumber="59" rank="species" species="divaricatus">
<emphasis italics="true" pageId="6" pageNumber="59">B. divaricatus</emphasis>
</taxonomicName>
[sensu] Lloyd, non Rhode, with the characteristics of
<normalizedToken originalValue="Lloyd’s">Lloyd's</normalizedToken>
plant.
<bibRefCitation author="Kerguelen, M" journalOrPublisher="Lejeunia" pageId="10" pageNumber="63" refId="B18" refString="Kerguelen, M, 1981. N 8913 [17] Bromusmolliformis. Societe pour l'Echange des Plantes Vasculaires de l'Europe et du Bassin Mediterraneen 18: 27." title="N 8913 [17] Bromusmolliformis. Societe pour l'Echange des Plantes Vasculaires de l'Europe et du Bassin Mediterraneen 18: 27." year="1981">
<normalizedToken originalValue="Kerguélen">Kerguelen</normalizedToken>
(1981
</bibRefCitation>
: 27) combines it to
<taxonomicName authority="subsp divaricatus" class="Liliopsida" family="Gramineae" genus="Bromus" higherTaxonomySource="CoL" infraspecific-rank="subsp" kingdom="Plantae" lsidName="Bromus hordeaceus" order="Poales" pageId="6" pageNumber="59" phylum="Tracheophyta" rank="species" species="hordeaceus" subspecies="divaricatus">Bromus hordeaceus subsp divaricatus</taxonomicName>
(Bonnier &amp; Layens)
<normalizedToken originalValue="Kerguélen">Kerguelen</normalizedToken>
. Bonnier and Layens type material is unknown (
<bibRefCitation author="Stafleu, FA" journalOrPublisher="Feddes Repertorium" pageId="10" pageNumber="63" refId="B28" refString="Stafleu, FA, Cowan, RS, 1976. Taxonomic literature (2nd ed.) Utrecht: Bohn, Scheltema &amp; Holkema, v.1: A-G. http://www.sil.si.edu/DigitalCollections/tl-2/browse.cfm?vol=1page/11" title="Taxonomic literature (2 nd ed.) Utrecht: Bohn, Scheltema &amp; Holkema, v. 1: A-G. http: // www. sil. si. edu / DigitalCollections / tl- 2 / browse. cfm? vol = 1 page / 11" year="1976">Stafleu and Cowan 1976</bibRefCitation>
).
<bibRefCitation author="Kerguelen, M" journalOrPublisher="Lejeunia" pageId="9" pageNumber="62" pagination="1 - 343" refId="B17" refString="Kerguelen, M, 1975. Les Gramineae (Poaceae) de la Flore Francaise. Essai de mise au point taxonomique et nomenclaturale. Lejeunia 75: 1 - 343" title="Les Gramineae (Poaceae) de la Flore Francaise. Essai de mise au point taxonomique et nomenclaturale." volume="75" year="1975">
<normalizedToken originalValue="Kerguélen">Kerguelen</normalizedToken>
(1975
</bibRefCitation>
: 104) mentions
<normalizedToken originalValue="Lloyd’s">Lloyd's</normalizedToken>
type material is in herbarium NTM, pointing out two localities &quot;Pornic, Saint Brevin&quot; indicated by Lloyd. Nevertheless, no original material is preserved in NTM (Mary Laury Guerin, Com. pers. 2018).
</paragraph>
<paragraph pageId="6" pageNumber="59">
<taxonomicName authority="L." class="Liliopsida" family="Gramineae" genus="Bromus" higherTaxonomySource="CoL" infraspecific-rank="var." kingdom="Plantae" lsidName="Bromus hordeaceus var. molliformis" order="Poales" pageId="6" pageNumber="59" phylum="Tracheophyta" rank="variety" species="hordeaceus" variety="molliformis">Bromus hordeaceus L. var. molliformis</taxonomicName>
(J.Lloyd ex Billot)
<bibRefCitation author="Halacsy, E" journalOrPublisher="Bulletin de la Societe Botanique de France" pageId="9" pageNumber="62" refId="B13" refString="Halacsy, E, 1904. Conspectus Florae Graecae III. Guilelmi Engelmann, Lipsiae. http://bibdigital.rjb.csic.es/spa/Libro.php?Libro=2294&amp;Pagina=1" title="Conspectus Florae Graecae III. Guilelmi Engelmann, Lipsiae. http: // bibdigital. rjb. csic. es / spa / Libro. php? Libro = 2294 &amp; Pagina = 1" year="1904">
<normalizedToken originalValue="Halácsy">Halacsy</normalizedToken>
(1904
</bibRefCitation>
: 396). This is another combination of
<taxonomicName family="Poaceae" kingdom="Plantae" lsidName="B. molliformis" order="Poales" pageId="6" pageNumber="59" rank="species" species="molliformis">
<emphasis italics="true" pageId="6" pageNumber="59">B. molliformis</emphasis>
</taxonomicName>
J.Lloyd and
<normalizedToken originalValue="Halácsy">Halacsy</normalizedToken>
pointed out its main traits &quot;Panicula magis conferta, ramis brevissime, spiculis densius et longius velutinu-pilosis, aristis demum extrorso subcurvatis&quot;.
</paragraph>
<paragraph pageId="6" pageNumber="59">
<taxonomicName authority="L. subsp. molliformis" class="Liliopsida" family="Gramineae" genus="Bromus" higherTaxonomySource="CoL" infraspecific-rank="subsp." kingdom="Plantae" lsidName="Bromus hordeaceus" order="Poales" pageId="6" pageNumber="59" phylum="Tracheophyta" rank="species" species="hordeaceus" subspecies="molliformis">Bromus hordeaceus L. subsp. molliformis</taxonomicName>
(J.Lloyd) Maire &amp; Weiller (1955: 255). This post-1953 combination, without direct citation to the basionym, nor to a potentially validating Latin description, but only to &quot;
<taxonomicName class="Liliopsida" family="Gramineae" genus="Bromus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bromus molliformis" order="Poales" pageId="6" pageNumber="59" phylum="Tracheophyta" rank="species" species="molliformis">
<emphasis italics="true" pageId="6" pageNumber="59">Bromus molliformis</emphasis>
</taxonomicName>
Lloyd, Fl. Loire-Inf. 315 (1844)&quot;, is another combination not validly published.
</paragraph>
<paragraph pageId="6" pageNumber="59">
As
<taxonomicName authority="subsp. divaricatus" class="Liliopsida" family="Gramineae" genus="Bromus" higherTaxonomySource="CoL" infraspecific-rank="subsp." kingdom="Plantae" lsidName="Bromus intermedius" order="Poales" pageId="6" pageNumber="59" phylum="Tracheophyta" rank="species" species="intermedius" subspecies="divaricatus">Bromus intermedius subsp. divaricatus</taxonomicName>
Bonnier &amp; Layens is an untypified name, we choose, as lectotype, the figure in
<bibRefCitation author="Bonnier, G" journalOrPublisher="Archives de la Flore de France et d'Allemagne" pageId="9" pageNumber="62" refId="B6" refString="Bonnier, G, Layens, G, 1894. Tableaux synoptiques des plantes vasculaires de la Flore de la France. Paul Dupont, Paris. https://doi.org/10.5962/bhl.title.323" title="Tableaux synoptiques des plantes vasculaires de la Flore de la France. Paul Dupont, Paris. https: // doi. org / 10.5962 / bhl. title. 323" year="1894">Bonnier and Layens (1894</bibRefCitation>
: 369) that is definitely original material. However, as is common with figures, it is difficult to observe some diagnostic characteristics. Thus, we select, as epitype, a specimen collected by Lloyd, conserved in BM, consistent with the protologue. It is a sheet bearing two collections. We choose the specimen on the left from Pornichet (barcode BM001067302).
</paragraph>
<caption ID-Zenodo-Dep="2702123" doi="10.3897/phytokeys.119.30254.figure5" httpUri="https://binary.pensoft.net/fig/297552" pageId="6" pageNumber="59" start="Figure 5" startId="F5">
<paragraph pageId="6" pageNumber="59">
<emphasis bold="true" pageId="6" pageNumber="59">Figure 5.</emphasis>
The epitype of
<taxonomicName authority="L. subsp divaricatus" class="Liliopsida" family="Gramineae" genus="Bromus" higherTaxonomySource="CoL" infraspecific-rank="subsp" kingdom="Plantae" lsidName="Bromus hordeaceus" order="Poales" pageId="6" pageNumber="59" phylum="Tracheophyta" rank="species" species="hordeaceus" subspecies="divaricatus">Bromus hordeaceus L. subsp divaricatus</taxonomicName>
(Bonnier &amp; Layens)
<normalizedToken originalValue="Kerguélen">Kerguelen</normalizedToken>
is the specimen on the left with barcode BM001067302 (collected and identified by Lloyd as
<taxonomicName family="Poaceae" kingdom="Plantae" lsidName="B. molliformis" order="Poales" pageId="6" pageNumber="59" rank="species" species="molliformis">
<emphasis italics="true" pageId="6" pageNumber="59">B. molliformis</emphasis>
</taxonomicName>
; preserved at BM (Permanent URL: http://data.nhm.ac.uk/object/9a6e0684-9e9b-4951-ac7e-ba13ad34ed31). Reproduced with permission.
</paragraph>
</caption>
<caption ID-Zenodo-Dep="2702125" doi="10.3897/phytokeys.119.30254.figure6" httpUri="https://binary.pensoft.net/fig/297553" pageId="6" pageNumber="59" start="Figure 6" startId="F6">
<paragraph pageId="6" pageNumber="59">
<emphasis bold="true" pageId="6" pageNumber="59">Figure 6.</emphasis>
Lectotype for
<taxonomicName class="Liliopsida" family="Gramineae" genus="Bromus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bromus molliformis" order="Poales" pageId="6" pageNumber="59" phylum="Tracheophyta" rank="species" species="molliformis">
<emphasis italics="true" pageId="6" pageNumber="59">Bromus molliformis</emphasis>
</taxonomicName>
J.Lloyd ex Billot at P. It is a specimen preserved at the MNHN Collection Vascular plants. Specimen with barcode P02381530. (Available at http://coldb.mnhn.fr/catalognumber/mnhn/p/p02381530) Reproduced with permission.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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