283 lines
42 KiB
XML
283 lines
42 KiB
XML
<document id="1C374CFBC8D4D4F48C6543530BA5C80D" ID-CLB-Dataset="28990" ID-DOI="10.11646/zootaxa.4461.1.7" ID-GBIF-Dataset="1fbcced0-ec5a-4afe-bcb1-496990e850c9" ID-ISSN="1175-5326" ID-Zenodo-Dep="1459872" ID-ZooBank="3FACE488-20DC-46A4-A511-026B7F2DBEA2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539336504258" checkinUser="plazi" docAuthor="Evsyukov, Aleksandr, Golovatch, Sergei & Reip, Hans S." docDate="2018" docId="610E87FB9301FF83FF0DF15BFE451C33" docLanguage="en" docName="zootaxa.4461.1.7.pdf" docOrigin="Zootaxa 4461 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Julus khostensis Evsyukov & Golovatch & Reip 2018, sp. n." docType="treatment" docVersion="6" lastPageNumber="114" masterDocId="9D37FF839316FF9AFF9AF75AFFFE1E2B" masterDocTitle="The millipede genus Julus Linnaeus, 1758 in the Caucasus (Diplopoda: Julida: Julidae)" masterLastPageNumber="117" masterPageNumber="89" pageNumber="112" updateTime="1698579428612" updateUser="ExternalLinkService">
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<mods:title id="7B44E2B87CBC417F2549D1C6F1C4D528">The millipede genus Julus Linnaeus, 1758 in the Caucasus (Diplopoda: Julida: Julidae)</mods:title>
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<mods:name id="62EA132B994D1B2C81DB6FC0D0B2552E" type="personal">
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<mods:namePart id="FFCC7ED55E052B9C2A1C1C40F16E747A">Evsyukov, Aleksandr</mods:namePart>
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<mods:namePart id="365F5F23BF2A30A3A8723A611128CD38">Golovatch, Sergei</mods:namePart>
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<mods:namePart id="925CCF526069BF9EAE35E00ABE7412A5">Reip, Hans S.</mods:namePart>
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<mods:title id="7DF07D2AC033EA7FC33A0AFF255B3D06">Zootaxa</mods:title>
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<mods:part id="8F6C62A63F23C29809950DC6DA6F3E63">
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<mods:date id="EF6D7C5D3371F008CE123CCEA2641103">2018</mods:date>
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<mods:number id="ECC1A6EBE6E4404EBC8D756BBDBF14EB">2018-08-20</mods:number>
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<mods:number id="63CC22055843B351EA0BDF2FFE4433A3">4461</mods:number>
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<treatment id="610E87FB9301FF83FF0DF15BFE451C33" ID-DOI="http://doi.org/10.5281/zenodo.5978679" ID-GBIF-Taxon="148701768" ID-Zenodo-Dep="5978679" LSID="urn:lsid:plazi:treatment:610E87FB9301FF83FF0DF15BFE451C33" httpUri="http://treatment.plazi.org/id/610E87FB9301FF83FF0DF15BFE451C33" lastPageId="25" lastPageNumber="114" pageId="23" pageNumber="112">
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<subSubSection id="A1BD65669301FF8DFF0DF15BFD681817" pageId="23" pageNumber="112" type="nomenclature">
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<paragraph id="E91836ED9301FF8DFF0DF15BFE561830" blockId="23.[151,662,1537,1597]" box="[151,424,1537,1563]" pageId="23" pageNumber="112">
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<heading id="B25081819301FF8DFF0DF15BFE561830" bold="true" box="[151,424,1537,1563]" fontSize="11" level="1" pageId="23" pageNumber="112" reason="1">
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<emphasis id="DBD3EAFF9301FF8DFF0DF15BFE561830" bold="true" box="[151,424,1537,1563]" pageId="23" pageNumber="112">
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<taxonomicName id="2EA74D6E9301FF8DFF0DF15BFEA01830" ID-CoL="849L6" authority="Evsyukov & Golovatch & Reip, 2018" authorityName="Evsyukov & Golovatch & Reip" authorityYear="2018" box="[151,350,1537,1563]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="23" pageNumber="112" phylum="Arthropoda" rank="species" species="khostensis" status="sp. nov.">
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<emphasis id="DBD3EAFF9301FF8DFF0DF15BFEA01830" bold="true" box="[151,350,1537,1563]" italics="true" pageId="23" pageNumber="112">Julus khostensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="C0E057849301FF8DFEFFF15BFE561830" box="[357,424,1537,1563]" pageId="23" pageNumber="112" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="E91836ED9301FF8DFF0DF17EFD681817" blockId="23.[151,662,1537,1597]" box="[151,662,1572,1597]" pageId="23" pageNumber="112">
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<figureCitation id="719C2A689301FF8DFF0DF17EFF0D1817" box="[151,243,1572,1597]" captionStart="FIGURE 1" captionStartId="4.[151,249,1820,1842]" captionTargetBox="[344,1236,854,1775]" captionTargetId="figure@4.[340,1248,841,1798]" captionTargetPageId="4" captionText="FIGURE 1. Antennae in Caucasian species of Julus, lateral views. A: J. alexandrae Evsyukov, 2016. B: J. colchicus Lohmander, 1936. C: J. jedryczkowskii Golovatch, 1981. D: J. kubanus Verhoeff, 1921. E: J. lignaui Verhoeff, 1910. F: J. lindholmi Lohmander, 1936. G: J. dagestanus sp. n. H: J. khostensis sp. n. Scale bar: 0.3 mm." httpUri="https://zenodo.org/record/1459874/files/figure.png" pageId="23" pageNumber="112">Figs 1H</figureCitation>
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,
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<figureCitation id="719C2A689301FF8DFF64F17EFEE91817" box="[254,279,1572,1596]" captionStart="FIGURE 2" captionStartId="5.[151,249,1601,1623]" captionTargetBox="[340,1248,985,1580]" captionTargetId="figure@5.[340,1248,985,1580]" captionTargetPageId="5" captionText="FIGURE 2. Gnathochilaria in Caucasian species of Julus, ventral views. A: J. alexandrae Evsyukov, 2016. B: J. colchicus Lohmander, 1936. C: J. jedryczkowskii Golovatch, 1981. D: J. kubanus Verhoeff, 1921. E: J. lignaui Verhoeff, 1910. F: J. lindholmi Lohmander, 1936. G: J. subalpinus Lohmander, 1936. H: J. dagestanus sp. n. I: J. khostensis sp. n. Scale bars: 0.3 mm." httpUri="https://zenodo.org/record/1459876/files/figure.png" pageId="23" pageNumber="112">
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<date id="9D19102D9301FF8DFF64F17EFEE91817" box="[254,279,1572,1596]" pageId="23" pageNumber="112">2I</date>
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</figureCitation>
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,
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<figureCitation id="719C2A689301FF8DFEBEF17EFEBD1817" box="[292,323,1572,1596]" captionStart="FIGURE 3" captionStartId="6.[151,249,1499,1521]" captionTargetBox="[340,1248,481,1476]" captionTargetId="figure@6.[340,1248,481,1478]" captionTargetPageId="6" captionText="FIGURE 3. Male legs 1 in Julus alexandrae Evsyukov, 2016 (A), J. colchicus Lohmander, 1936 (B–D), J. jedryczkowskii Golovatch, 1981 (E–F), J. kubanus Verhoeff, 1921 (G–I), J. lignaui Verhoeff, 1910 (J–M), J. lindholmi Lohmander, 1936 (N– P), J. subalpinus Lohmander, 1936 (Q–R), J. dagestanus sp. n. (S) and J. khostensis sp. n. (T). A: from the Rostov-on-Don Region. B: from Krasnaya Polyana, Krasnodar Province. C: from Sakhrai, Adygea Republic. D: syntype from Gagry, Abkhazia, after Lohmander (1936). E: from North Ossetia. F: paratype from North Ossetia, after Golovatch (1981). G: from Maikop, Adygea Republic. H: from Pskhu, Abkhazia. I: from Aksaut, Karachaevo-Cherkessia, after Lohmander (1936). J: from Krasnaya Polyana, Krasnodar Province. K: from Teberda, Karachaevo-Cherkessia. L: from Mestia, Svanetia, Georgia. M: from Teberda, Karachaevo-Cherkessia. N: from Svanetia, Georgia. O: from near Teberda, Karachaevo-Cherkessia. P: holotype from Olginskoe, Abkhazia, after Lohmander (1936). Q: from North Ossetia. R: syntype from Mount Djuga, Krasnodar Province, after Lohmander (1936). S: holotype from Dagestan. T: holotype from Khosta, Krasnodar Province. Oral views B–O; caudal views A, P. Abbreviations: cx, coxa; te, telopodite remnant. Scale bars: 0.3 mm (A–C, E, G, H, J–O, Q, S, T), 0.2 mm (F), not to scale (D, I, P, R)." httpUri="https://zenodo.org/record/1459878/files/figure.png" pageId="23" pageNumber="112">3T</figureCitation>
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,
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<figureCitation id="719C2A689301FF8DFED5F17FFE8D1817" box="[335,371,1573,1596]" captionStart="FIGURE 7" captionStartId="11.[151,250,767,789]" captionTargetBox="[349,1245,180,761]" captionTargetId="figure@11.[340,1248,180,763]" captionTargetPageId="11" captionText="FIGURE 7. Male legs 2 in Julus subalpinus Lohmander, 1936 (A–B), J. dagestanus sp. n. (C) and J. khostensis sp. n. (D). A: from North Ossetia. B: syntype from Mount Djuga, Krasnodar Province, after Lohmander (1936). C: holotype from Dagestan. D: holotype from Khosta, Krasnodar Province. Oral views A, C, D; lateral view B. Abbreviations: alp, anterolateral process of coxa 2; mp, main process of coxa 2; p, penes. Scale bars: 0.3 mm." httpUri="https://zenodo.org/record/1459886/files/figure.png" pageId="23" pageNumber="112">7D</figureCitation>
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,
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<figureCitation id="719C2A689301FF8DFEE5F17EFE2F1817" box="[383,465,1572,1596]" captionStart="FIGURE 13" captionStartId="18.[151,250,1689,1711]" captionTargetBox="[340,1248,482,1662]" captionTargetId="figure@18.[340,1248,481,1668]" captionTargetPageId="18" captionText="FIGURE 13. Male gonopods in Julus dagestanus sp. n. (A–C holotype from Dagestan) and J. khostensis sp. n. (D–E holotype from Khosta, Krasnodar Province). A: promere with flagellum, mesal view. B: promere, caudal view. C: left gonopods, lateral view. D: promere with flagella, caudal view. E: right gonopods, lateral view. Scale bars: 0.3 mm. Abbreviations: fl, flagellum; la, lamella on promere; la1 and la2, lamella 1 and lamella 2 on promere; lo, lateral outgrowth of opisthomere; ms, mesomere; op, opisthomere; pr, promere." httpUri="https://zenodo.org/record/1459898/files/figure.png" pageId="23" pageNumber="112">13D–E</figureCitation>
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,
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<figureCitation id="719C2A689301FF8DFE46F17EFDF11816" box="[476,527,1572,1597]" captionStart="FIGURE 15" captionStartId="20.[151,249,1687,1709]" captionTargetBox="[340,1248,1178,1666]" captionTargetId="figure@20.[340,1248,1178,1666]" captionTargetPageId="20" captionText="FIGURE 15. Ventral edge of male segments 7 in Caucasian species of Julus. A: J. alexandrae Evsyukov, 2016. B: J. colchicus Lohmander, 1936 C: J. jedryczkowskii Golovatch, 1981. D: J. kubanus Verhoeff, 1921. E: J. lignaui Verhoeff, 1910. F: J. lindholmi Lohmander, 1936. G: J. subalpinus Lohmander, 1936. H: J. khostensis sp. n. Scale bars: 0.3 mm." httpUri="https://zenodo.org/record/1459902/files/figure.png" pageId="23" pageNumber="112">15H</figureCitation>
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,
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<figureCitation id="719C2A689301FF8DFD80F17EFDBC1816" box="[538,578,1572,1597]" captionStart="FIGURE 16" captionStartId="21.[151,249,1286,1308]" captionTargetBox="[340,1247,583,1264]" captionTargetId="figure@21.[340,1247,583,1264]" captionTargetPageId="21" captionText="FIGURE 16. Striation on metazonites in Caucasian species of Julus. A: J. alexandrae Evsyukov, 2016. B: J. colchicus Lohmander, 1936. C: J. jedryczkowskii Golovatch, 1981. D: J. kubanus Verhoeff, 1921. E: J. lignaui Verhoeff, 1910. F: J. lindholmi Lohmander, 1936. G: J. subalpinus Lohmander, 1936. H: J. dagestanus sp. n. I: J. khostensis sp. n. Pictures taken not to scale." httpUri="https://zenodo.org/record/1459904/files/figure.png" pageId="23" pageNumber="112">
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<date id="9D19102D9301FF8DFD80F17EFDBC1816" box="[538,578,1572,1597]" pageId="23" pageNumber="112" value="2016-01">16I</date>
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</figureCitation>
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, Map 2
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</paragraph>
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</subSubSection>
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<subSubSection id="A1BD65669301FF8DFF0DF136FDAC1882" pageId="23" pageNumber="112" type="materials_examined">
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<paragraph id="E91836ED9301FF8DFF0DF136FDAC1882" blockId="23.[151,1436,1644,2029]" pageId="23" pageNumber="112">
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<emphasis id="DBD3EAFF9301FF8DFF0DF136FE7A18AE" bold="true" box="[151,388,1644,1669]" pageId="23" pageNumber="112">Material examined.</emphasis>
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Holotype male (ZMUM), Russia, Krasnodar Prov., Khosta,
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<taxonomicName id="2EA74D6E9301FF8DFBDEF137FB7C18AF" box="[1092,1154,1645,1668]" class="Pinopsida" family="Taxaceae" genus="Taxus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="23" pageNumber="112" phylum="Tracheophyta" rank="genus">
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<emphasis id="DBD3EAFF9301FF8DFBDEF137FB7C18AF" box="[1092,1154,1645,1668]" italics="true" pageId="23" pageNumber="112">Taxus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="2EA74D6E9301FF8DFB58F137FAF818AF" box="[1218,1286,1645,1668]" class="Magnoliopsida" family="Buxaceae" genus="Buxus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Buxales" pageId="23" pageNumber="112" phylum="Tracheophyta" rank="genus">
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<emphasis id="DBD3EAFF9301FF8DFB58F137FAF818AF" box="[1218,1286,1645,1668]" italics="true" pageId="23" pageNumber="112">Buxus</emphasis>
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</taxonomicName>
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grove, Cave Tisovaya I,
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<date id="9D19102D9301FF8DFE81F1CAFE631882" box="[283,413,1680,1705]" pageId="23" pageNumber="112" value="1985-09-27">27.IX.1985</date>
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, leg. N. Myuge.
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</paragraph>
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</subSubSection>
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<subSubSection id="A1BD65669301FF8DFF5DF1EEFB6E19AA" pageId="23" pageNumber="112" type="diagnosis">
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<paragraph id="E91836ED9301FF8DFF5DF1EEFB6E19AA" blockId="23.[151,1436,1644,2029]" pageId="23" pageNumber="112">
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<emphasis id="DBD3EAFF9301FF8DFF5DF1EEFEBF18E6" bold="true" box="[199,321,1716,1741]" pageId="23" pageNumber="112">Diagnosis.</emphasis>
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The promere is similar to that in
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<taxonomicName id="2EA74D6E9301FF8DFD2DF1EFFCD118E6" authorityName="Verhoeff" authorityYear="1921" box="[695,815,1716,1741]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="23" pageNumber="112" phylum="Arthropoda" rank="species" species="kubanus">
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<emphasis id="DBD3EAFF9301FF8DFD2DF1EFFCD118E6" box="[695,815,1716,1741]" italics="true" pageId="23" pageNumber="112">J. kubanus</emphasis>
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</taxonomicName>
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, but the processes on male leg-pair 2 resemble those of
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<taxonomicName id="2EA74D6E9301FF8DFF0DF183FEDE18DA" authorityName="Lohmander" authorityYear="1936" box="[151,288,1752,1777]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="23" pageNumber="112" phylum="Arthropoda" rank="species" species="lindholmi">
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<emphasis id="DBD3EAFF9301FF8DFF0DF183FEDE18DA" box="[151,288,1752,1777]" italics="true" pageId="23" pageNumber="112">J. lindholmi</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="2EA74D6E9301FF8DFEC5F183FDDC18DA" authorityName="Golovatch" authorityYear="1981" box="[351,546,1752,1777]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="23" pageNumber="112" phylum="Arthropoda" rank="species" species="jedryczkowskii">
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<emphasis id="DBD3EAFF9301FF8DFEC5F183FDDC18DA" box="[351,546,1752,1777]" italics="true" pageId="23" pageNumber="112">J. jedryczkowskii</emphasis>
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</taxonomicName>
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. Differs clearly from all congeners by the structure of the flagellum on the promere and the conformation of the opisthomere. Coxa (
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<emphasis id="DBD3EAFF9301FF8DFC86F1A6FCC8193E" bold="true" box="[796,822,1788,1813]" pageId="23" pageNumber="112">cx</emphasis>
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) of male leg-pair 1 with a large lateral outgrowth (
|
||
<figureCitation id="719C2A689301FF8DFAF5F1A6FF471913" captionStart="FIGURE 3" captionStartId="6.[151,249,1499,1521]" captionTargetBox="[340,1248,481,1476]" captionTargetId="figure@6.[340,1248,481,1478]" captionTargetPageId="6" captionText="FIGURE 3. Male legs 1 in Julus alexandrae Evsyukov, 2016 (A), J. colchicus Lohmander, 1936 (B–D), J. jedryczkowskii Golovatch, 1981 (E–F), J. kubanus Verhoeff, 1921 (G–I), J. lignaui Verhoeff, 1910 (J–M), J. lindholmi Lohmander, 1936 (N– P), J. subalpinus Lohmander, 1936 (Q–R), J. dagestanus sp. n. (S) and J. khostensis sp. n. (T). A: from the Rostov-on-Don Region. B: from Krasnaya Polyana, Krasnodar Province. C: from Sakhrai, Adygea Republic. D: syntype from Gagry, Abkhazia, after Lohmander (1936). E: from North Ossetia. F: paratype from North Ossetia, after Golovatch (1981). G: from Maikop, Adygea Republic. H: from Pskhu, Abkhazia. I: from Aksaut, Karachaevo-Cherkessia, after Lohmander (1936). J: from Krasnaya Polyana, Krasnodar Province. K: from Teberda, Karachaevo-Cherkessia. L: from Mestia, Svanetia, Georgia. M: from Teberda, Karachaevo-Cherkessia. N: from Svanetia, Georgia. O: from near Teberda, Karachaevo-Cherkessia. P: holotype from Olginskoe, Abkhazia, after Lohmander (1936). Q: from North Ossetia. R: syntype from Mount Djuga, Krasnodar Province, after Lohmander (1936). S: holotype from Dagestan. T: holotype from Khosta, Krasnodar Province. Oral views B–O; caudal views A, P. Abbreviations: cx, coxa; te, telopodite remnant. Scale bars: 0.3 mm (A–C, E, G, H, J–O, Q, S, T), 0.2 mm (F), not to scale (D, I, P, R)." httpUri="https://zenodo.org/record/1459878/files/figure.png" pageId="23" pageNumber="112">Fig. 3T</figureCitation>
|
||
). Male leg-pair 2 (
|
||
<figureCitation id="719C2A689301FF8DFE0CF07AFE0F1913" box="[406,497,1824,1849]" captionStart="FIGURE 7" captionStartId="11.[151,250,767,789]" captionTargetBox="[349,1245,180,761]" captionTargetId="figure@11.[340,1248,180,763]" captionTargetPageId="11" captionText="FIGURE 7. Male legs 2 in Julus subalpinus Lohmander, 1936 (A–B), J. dagestanus sp. n. (C) and J. khostensis sp. n. (D). A: from North Ossetia. B: syntype from Mount Djuga, Krasnodar Province, after Lohmander (1936). C: holotype from Dagestan. D: holotype from Khosta, Krasnodar Province. Oral views A, C, D; lateral view B. Abbreviations: alp, anterolateral process of coxa 2; mp, main process of coxa 2; p, penes. Scale bars: 0.3 mm." httpUri="https://zenodo.org/record/1459886/files/figure.png" pageId="23" pageNumber="112">Fig. 7D</figureCitation>
|
||
) with two pairs of coxal processes. Gonopodal promere (
|
||
<emphasis id="DBD3EAFF9301FF8DFB0AF07AFB521912" bold="true" box="[1168,1196,1824,1849]" pageId="23" pageNumber="112">pr</emphasis>
|
||
) narrow, with a thin lamella (
|
||
<emphasis id="DBD3EAFF9301FF8DFF63F01EFEF11976" bold="true" box="[249,271,1860,1885]" pageId="23" pageNumber="112">la</emphasis>
|
||
) (
|
||
<figureCitation id="719C2A689301FF8DFEBDF01EFE701977" box="[295,398,1860,1885]" captionStart="FIGURE 13" captionStartId="18.[151,250,1689,1711]" captionTargetBox="[340,1248,482,1662]" captionTargetId="figure@18.[340,1248,481,1668]" captionTargetPageId="18" captionText="FIGURE 13. Male gonopods in Julus dagestanus sp. n. (A–C holotype from Dagestan) and J. khostensis sp. n. (D–E holotype from Khosta, Krasnodar Province). A: promere with flagellum, mesal view. B: promere, caudal view. C: left gonopods, lateral view. D: promere with flagella, caudal view. E: right gonopods, lateral view. Scale bars: 0.3 mm. Abbreviations: fl, flagellum; la, lamella on promere; la1 and la2, lamella 1 and lamella 2 on promere; lo, lateral outgrowth of opisthomere; ms, mesomere; op, opisthomere; pr, promere." httpUri="https://zenodo.org/record/1459898/files/figure.png" pageId="23" pageNumber="112">Fig. 13D</figureCitation>
|
||
). Each flagellum (
|
||
<emphasis id="DBD3EAFF9301FF8DFDC6F01EFD931976" bold="true" box="[604,621,1860,1885]" pageId="23" pageNumber="112">fl</emphasis>
|
||
) with a lamina in apical part. Mesomere (
|
||
<emphasis id="DBD3EAFF9301FF8DFBA5F01EFB9F1976" bold="true" box="[1087,1121,1860,1885]" pageId="23" pageNumber="112">ms</emphasis>
|
||
) rounded, opisthomere (
|
||
<emphasis id="DBD3EAFF9301FF8DFAEFF01EFA6C1976" bold="true" box="[1397,1426,1860,1885]" pageId="23" pageNumber="112">op</emphasis>
|
||
) bifurcate (
|
||
<figureCitation id="719C2A689301FF8DFE90F032FE9119AB" box="[266,367,1896,1921]" captionStart="FIGURE 13" captionStartId="18.[151,250,1689,1711]" captionTargetBox="[340,1248,482,1662]" captionTargetId="figure@18.[340,1248,481,1668]" captionTargetPageId="18" captionText="FIGURE 13. Male gonopods in Julus dagestanus sp. n. (A–C holotype from Dagestan) and J. khostensis sp. n. (D–E holotype from Khosta, Krasnodar Province). A: promere with flagellum, mesal view. B: promere, caudal view. C: left gonopods, lateral view. D: promere with flagella, caudal view. E: right gonopods, lateral view. Scale bars: 0.3 mm. Abbreviations: fl, flagellum; la, lamella on promere; la1 and la2, lamella 1 and lamella 2 on promere; lo, lateral outgrowth of opisthomere; ms, mesomere; op, opisthomere; pr, promere." httpUri="https://zenodo.org/record/1459898/files/figure.png" pageId="23" pageNumber="112">Fig. 13E</figureCitation>
|
||
). Striations on metazonae regular and parallel to one another (
|
||
<figureCitation id="719C2A689301FF8DFBBCF032FB7D19AA" box="[1062,1155,1896,1921]" captionStart="FIGURE 16" captionStartId="21.[151,249,1286,1308]" captionTargetBox="[340,1247,583,1264]" captionTargetId="figure@21.[340,1247,583,1264]" captionTargetPageId="21" captionText="FIGURE 16. Striation on metazonites in Caucasian species of Julus. A: J. alexandrae Evsyukov, 2016. B: J. colchicus Lohmander, 1936. C: J. jedryczkowskii Golovatch, 1981. D: J. kubanus Verhoeff, 1921. E: J. lignaui Verhoeff, 1910. F: J. lindholmi Lohmander, 1936. G: J. subalpinus Lohmander, 1936. H: J. dagestanus sp. n. I: J. khostensis sp. n. Pictures taken not to scale." httpUri="https://zenodo.org/record/1459904/files/figure.png" pageId="23" pageNumber="112">
|
||
Fig.
|
||
<date id="9D19102D9301FF8DFBC1F032FB7D19AA" box="[1115,1155,1896,1921]" pageId="23" pageNumber="112" value="2016-01">16I</date>
|
||
</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A1BD65669301FF8DFF5DF0D6FB08198E" box="[199,1270,1932,1957]" pageId="23" pageNumber="112" type="etymology">
|
||
<paragraph id="E91836ED9301FF8DFF5DF0D6FB08198E" blockId="23.[151,1436,1644,2029]" box="[199,1270,1932,1957]" pageId="23" pageNumber="112">
|
||
<emphasis id="DBD3EAFF9301FF8DFF5DF0D6FE5A198E" bold="true" box="[199,420,1932,1957]" pageId="23" pageNumber="112">Derivatio nominis.</emphasis>
|
||
To emphasize Khosta, Greater Sochi, a city near the
|
||
<emphasis id="DBD3EAFF9301FF8DFC6DF0D7FB84198F" box="[1015,1146,1933,1956]" italics="true" pageId="23" pageNumber="112">terra typica</emphasis>
|
||
; adjective.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A1BD65669301FF83FF5DF0EAFE451C33" lastPageId="25" lastPageNumber="114" pageId="23" pageNumber="112" type="description">
|
||
<paragraph id="E91836ED9301FF8DFF5DF0EAFC3719C6" blockId="23.[151,1436,1644,2029]" pageId="23" pageNumber="112">
|
||
<emphasis id="DBD3EAFF9301FF8DFF5DF0EAFEA719E2" bold="true" box="[199,345,1968,1993]" pageId="23" pageNumber="112">Description.</emphasis>
|
||
Length
|
||
<quantity id="2E5F9B089301FF8DFE21F0EAFDF019E3" box="[443,526,1968,1992]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" pageId="23" pageNumber="112" unit="mm" value="29.0">29 mm</quantity>
|
||
, width
|
||
<quantity id="2E5F9B089301FF8DFDFCF0EAFD3F19E3" box="[614,705,1968,1992]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="23" pageNumber="112" unit="mm" value="1.8">1.8 mm</quantity>
|
||
. Body segment formula 45+1+T. Body marbled greyish, venter lighter; antennae and anal valves greyish brown; legs yellow; eyes black.
|
||
</paragraph>
|
||
<paragraph id="E91836ED930EFF82FF0DF422FCCA1D86" blockId="24.[151,1437,888,941]" pageId="24" pageNumber="113">
|
||
<emphasis id="DBD3EAFF930EFF82FF0DF422FF171DA5" bold="true" box="[151,233,888,910]" pageId="24" pageNumber="113">MAP 2.</emphasis>
|
||
Distribution of
|
||
<taxonomicName id="2EA74D6E930EFF82FE11F422FE3E1DA5" authorityName="Evsyukov & Golovatch & Reip" authorityYear="2018" box="[395,448,888,910]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="24" pageNumber="113" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DBD3EAFF930EFF82FE11F422FE3E1DA5" box="[395,448,888,910]" italics="true" pageId="24" pageNumber="113">Julus</emphasis>
|
||
</taxonomicName>
|
||
species:
|
||
<taxonomicName id="2EA74D6E930EFF82FD87F423FCD91DA5" authority="Verhoeff, 1921" authorityName="Verhoeff" authorityYear="1921" box="[541,807,888,910]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="24" pageNumber="113" phylum="Arthropoda" rank="species" species="kubanus">
|
||
<emphasis id="DBD3EAFF930EFF82FD87F423FD771DA5" box="[541,649,888,910]" italics="true" pageId="24" pageNumber="113">J. kubanus</emphasis>
|
||
<bibRefCitation id="8D364B1C930EFF82FD15F422FCD91DA5" box="[655,807,888,910]" pageId="24" pageNumber="113" refId="ref20337" refString="Verhoeff, K. W. (1921) Chilognathen-Studien (91. Diplopoden-Aufsatz). Archiv fur Naturgeschichte, 85 A (12), 23 - 80." type="journal article">Verhoeff, 1921</bibRefCitation>
|
||
</taxonomicName>
|
||
(green circle),
|
||
<taxonomicName id="2EA74D6E930EFF82FC5AF423FB011DA5" authority="Lohmander, 1936" authorityName="Lohmander" authorityYear="1936" box="[960,1279,888,910]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="24" pageNumber="113" phylum="Arthropoda" rank="species" species="subalpinus">
|
||
<emphasis id="DBD3EAFF930EFF82FC5AF423FBB81DA5" box="[960,1094,888,910]" italics="true" pageId="24" pageNumber="113">J. subalpinus</emphasis>
|
||
<bibRefCitation id="8D364B1C930EFF82FBD6F422FB011DA5" box="[1100,1279,888,910]" pageId="24" pageNumber="113" refId="ref19572" refString="Lohmander, H. (1936) Uber die Diplopoden des Kaukasusgebietes. Goteborgs Kungliga Vetenskaps- och Vitterhets-Samhalles Handlingar, 5, 5 B (1), 1 - 196." type="journal article">Lohmander, 1936</bibRefCitation>
|
||
</taxonomicName>
|
||
(red square),
|
||
<taxonomicName id="2EA74D6E930EFF82FA10F423FEF71D86" authority="Evsyukov & Golovatch & Reip, 2018" authorityName="Evsyukov & Golovatch & Reip" authorityYear="2018" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="24" pageNumber="113" phylum="Arthropoda" rank="species" species="dagestanus" status="sp. nov.">
|
||
<emphasis id="DBD3EAFF930EFF82FA10F423FEF71D86" italics="true" pageId="24" pageNumber="113">J. dagestanus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C0E05784930EFF82FE95F4CDFEB61D86" box="[271,328,919,941]" pageId="24" pageNumber="113" rank="species">
|
||
<emphasis id="DBD3EAFF930EFF82FE95F4CDFEB61D86" bold="true" box="[271,328,919,941]" pageId="24" pageNumber="113">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(blue triangle),
|
||
<taxonomicName id="2EA74D6E930EFF82FE71F4C2FD951D86" authority="Evsyukov & Golovatch & Reip, 2018" authorityName="Evsyukov & Golovatch & Reip" authorityYear="2018" box="[491,619,919,941]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="24" pageNumber="113" phylum="Arthropoda" rank="species" species="khostensis" status="sp. nov.">
|
||
<emphasis id="DBD3EAFF930EFF82FE71F4C2FD951D86" box="[491,619,919,941]" italics="true" pageId="24" pageNumber="113">J. khostensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C0E05784930EFF82FDEBF4CDFD541D86" box="[625,682,919,941]" pageId="24" pageNumber="113" rank="species">
|
||
<emphasis id="DBD3EAFF930EFF82FDEBF4CDFD541D86" bold="true" box="[625,682,919,941]" pageId="24" pageNumber="113">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(purple star).
|
||
</paragraph>
|
||
<paragraph id="E91836ED930EFF82FF5DF480FB7C1A11" blockId="24.[151,1436,985,1154]" pageId="24" pageNumber="113">
|
||
Eye patches subtriangular, each composed of
|
||
<emphasis id="DBD3EAFF930EFF82FD55F481FD141DD9" box="[719,746,987,1010]" italics="true" pageId="24" pageNumber="113">ca</emphasis>
|
||
40 ocelli. Epicranial setae 1+1, supralabral setae 2+2, labral setae 10+10. Genae unmodified. Antennae short and slender,
|
||
<emphasis id="DBD3EAFF930EFF82FC0EF4A5FC1A1A3D" box="[916,996,1023,1046]" italics="true" pageId="24" pageNumber="113">in situ</emphasis>
|
||
reaching back to body segment 3. Antennomeres 5 and 6 each with a distal corolla of sensory bacilliform sensilla (
|
||
<figureCitation id="719C2A68930EFF82FB81F37BFB8A1A11" box="[1051,1140,1057,1082]" captionStart="FIGURE 1" captionStartId="4.[151,249,1820,1842]" captionTargetBox="[344,1236,854,1775]" captionTargetId="figure@4.[340,1248,841,1798]" captionTargetPageId="4" captionText="FIGURE 1. Antennae in Caucasian species of Julus, lateral views. A: J. alexandrae Evsyukov, 2016. B: J. colchicus Lohmander, 1936. C: J. jedryczkowskii Golovatch, 1981. D: J. kubanus Verhoeff, 1921. E: J. lignaui Verhoeff, 1910. F: J. lindholmi Lohmander, 1936. G: J. dagestanus sp. n. H: J. khostensis sp. n. Scale bar: 0.3 mm." httpUri="https://zenodo.org/record/1459874/files/figure.png" pageId="24" pageNumber="113">Fig. 1H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="E91836ED930EFF82FF5DF31FFCB71AA9" blockId="24.[151,1436,985,1154]" pageId="24" pageNumber="113">
|
||
Lamellae linguales bare (artifact or abnormality?). A paramedian pair of pronounced swellings with a row of
|
||
<emphasis id="DBD3EAFF930EFF82FF0DF331FF4C1AA9" box="[151,178,1131,1154]" italics="true" pageId="24" pageNumber="113">ca</emphasis>
|
||
12 setae per swelling parabasally, below mentum (
|
||
<figureCitation id="719C2A68930EFF82FD77F333FCC21AA9" box="[749,828,1129,1154]" captionStart="FIGURE 2" captionStartId="5.[151,249,1601,1623]" captionTargetBox="[340,1248,985,1580]" captionTargetId="figure@5.[340,1248,985,1580]" captionTargetPageId="5" captionText="FIGURE 2. Gnathochilaria in Caucasian species of Julus, ventral views. A: J. alexandrae Evsyukov, 2016. B: J. colchicus Lohmander, 1936. C: J. jedryczkowskii Golovatch, 1981. D: J. kubanus Verhoeff, 1921. E: J. lignaui Verhoeff, 1910. F: J. lindholmi Lohmander, 1936. G: J. subalpinus Lohmander, 1936. H: J. dagestanus sp. n. I: J. khostensis sp. n. Scale bars: 0.3 mm." httpUri="https://zenodo.org/record/1459876/files/figure.png" pageId="24" pageNumber="113">
|
||
Fig.
|
||
<date id="9D19102D930EFF82FCBBF330FCC21AA9" box="[801,828,1130,1154]" pageId="24" pageNumber="113">2I</date>
|
||
</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="BDD86665930EFF82FF0DF1ABFF00190D" httpUri="https://zenodo.org/record/1459914/files/figure.png" pageId="24" pageNumber="113" startId="24.[151,249,1777,1799]" targetBox="[341,1242,1249,1720]" targetPageId="24">
|
||
<paragraph id="E91836ED930EFF82FF0DF1ABFF00190D" blockId="24.[151,1436,1777,1830]" pageId="24" pageNumber="113">
|
||
<emphasis id="DBD3EAFF930EFF82FF0DF1ABFEDD192C" bold="true" box="[151,291,1777,1799]" pageId="24" pageNumber="113">FIGURE 20.</emphasis>
|
||
Relatioship between the number of podous body rings and body diameter in males of
|
||
<taxonomicName id="2EA74D6E930EFF82FB51F1ABFAFE192C" authorityName="Evsyukov & Golovatch & Reip" authorityYear="2018" box="[1227,1280,1777,1799]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="24" pageNumber="113" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DBD3EAFF930EFF82FB51F1ABFAFE192C" box="[1227,1280,1777,1799]" italics="true" pageId="24" pageNumber="113">Julus</emphasis>
|
||
</taxonomicName>
|
||
species in the Caucasus.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E91836ED930EFF83FF5DF008FC481EFF" blockId="24.[151,1436,1874,2007]" lastBlockId="25.[151,1437,151,536]" lastPageId="25" lastPageNumber="114" pageId="24" pageNumber="113">
|
||
Body subcylindrical and slender. Suture dividing pro- and metazonae distinctly constricting both. Prozonae smooth, without striae. Metazonae with striations reaching the hind margin,
|
||
<emphasis id="DBD3EAFF930EFF82FC76F022FBF919A4" box="[1004,1031,1912,1935]" italics="true" pageId="24" pageNumber="113">ca</emphasis>
|
||
10 striae per about dorsal quarter of metazonital surface, i.e. between axial line and ozopore. Striae on metazonae regular and parallel to one another (
|
||
<figureCitation id="719C2A68930EFF82FF05F0E4FF0119FC" box="[159,255,1982,2007]" captionStart="FIGURE 16" captionStartId="21.[151,249,1286,1308]" captionTargetBox="[340,1247,583,1264]" captionTargetId="figure@21.[340,1247,583,1264]" captionTargetPageId="21" captionText="FIGURE 16. Striation on metazonites in Caucasian species of Julus. A: J. alexandrae Evsyukov, 2016. B: J. colchicus Lohmander, 1936. C: J. jedryczkowskii Golovatch, 1981. D: J. kubanus Verhoeff, 1921. E: J. lignaui Verhoeff, 1910. F: J. lindholmi Lohmander, 1936. G: J. subalpinus Lohmander, 1936. H: J. dagestanus sp. n. I: J. khostensis sp. n. Pictures taken not to scale." httpUri="https://zenodo.org/record/1459904/files/figure.png" pageId="24" pageNumber="113">
|
||
Fig.
|
||
<date id="9D19102D930EFF82FF4DF0E4FF0119FC" box="[215,255,1982,2007]" pageId="24" pageNumber="113" value="2016-01">16I</date>
|
||
</figureCitation>
|
||
). A whorl of sparse, thin, short setae at hind edge of metazonae, setae gradually growing denser and somewhat longer towards telson. Telson with sparse long setae on epiproct, the latter carrying a small apical clawshaped projection directed caudad. Anal valves convex, densely setose.
|
||
</paragraph>
|
||
<paragraph id="E91836ED930FFF83FF5DF7BAFCBB1FA3" blockId="25.[151,1437,151,536]" pageId="25" pageNumber="114">
|
||
MALE. Pregonopodal legs more incrassate than postgonopodal ones. Segments 6 and 7 wider than others. Ventral edge of segment 7 with small curved lamellae bordering the gonopodal aperture (
|
||
<figureCitation id="719C2A68930FFF83FBEFF65EFB221F36" box="[1141,1244,260,285]" captionStart="FIGURE 15" captionStartId="20.[151,249,1687,1709]" captionTargetBox="[340,1248,1178,1666]" captionTargetId="figure@20.[340,1248,1178,1666]" captionTargetPageId="20" captionText="FIGURE 15. Ventral edge of male segments 7 in Caucasian species of Julus. A: J. alexandrae Evsyukov, 2016. B: J. colchicus Lohmander, 1936 C: J. jedryczkowskii Golovatch, 1981. D: J. kubanus Verhoeff, 1921. E: J. lignaui Verhoeff, 1910. F: J. lindholmi Lohmander, 1936. G: J. subalpinus Lohmander, 1936. H: J. khostensis sp. n. Scale bars: 0.3 mm." httpUri="https://zenodo.org/record/1459902/files/figure.png" pageId="25" pageNumber="114">Fig. 15H</figureCitation>
|
||
). Suture between coxa (
|
||
<emphasis id="DBD3EAFF930FFF83FF47F67DFF061F6B" bold="true" box="[221,248,295,320]" pageId="25" pageNumber="114">cx</emphasis>
|
||
) and a 1-segmented telopodite remnant (
|
||
<emphasis id="DBD3EAFF930FFF83FD54F67DFD1A1F6B" bold="true" box="[718,740,295,320]" pageId="25" pageNumber="114">te</emphasis>
|
||
) well-developed;
|
||
<emphasis id="DBD3EAFF930FFF83FC28F67DFC331F6B" bold="true" box="[946,973,295,320]" pageId="25" pageNumber="114">cx</emphasis>
|
||
with a large lateral outgrowth;
|
||
<emphasis id="DBD3EAFF930FFF83FAADF67DFAB31F6B" bold="true" box="[1335,1357,295,320]" pageId="25" pageNumber="114">te</emphasis>
|
||
with a group of thin setae (
|
||
<figureCitation id="719C2A68930FFF83FEE4F616FE291F4F" box="[382,471,332,357]" captionStart="FIGURE 3" captionStartId="6.[151,249,1499,1521]" captionTargetBox="[340,1248,481,1476]" captionTargetId="figure@6.[340,1248,481,1478]" captionTargetPageId="6" captionText="FIGURE 3. Male legs 1 in Julus alexandrae Evsyukov, 2016 (A), J. colchicus Lohmander, 1936 (B–D), J. jedryczkowskii Golovatch, 1981 (E–F), J. kubanus Verhoeff, 1921 (G–I), J. lignaui Verhoeff, 1910 (J–M), J. lindholmi Lohmander, 1936 (N– P), J. subalpinus Lohmander, 1936 (Q–R), J. dagestanus sp. n. (S) and J. khostensis sp. n. (T). A: from the Rostov-on-Don Region. B: from Krasnaya Polyana, Krasnodar Province. C: from Sakhrai, Adygea Republic. D: syntype from Gagry, Abkhazia, after Lohmander (1936). E: from North Ossetia. F: paratype from North Ossetia, after Golovatch (1981). G: from Maikop, Adygea Republic. H: from Pskhu, Abkhazia. I: from Aksaut, Karachaevo-Cherkessia, after Lohmander (1936). J: from Krasnaya Polyana, Krasnodar Province. K: from Teberda, Karachaevo-Cherkessia. L: from Mestia, Svanetia, Georgia. M: from Teberda, Karachaevo-Cherkessia. N: from Svanetia, Georgia. O: from near Teberda, Karachaevo-Cherkessia. P: holotype from Olginskoe, Abkhazia, after Lohmander (1936). Q: from North Ossetia. R: syntype from Mount Djuga, Krasnodar Province, after Lohmander (1936). S: holotype from Dagestan. T: holotype from Khosta, Krasnodar Province. Oral views B–O; caudal views A, P. Abbreviations: cx, coxa; te, telopodite remnant. Scale bars: 0.3 mm (A–C, E, G, H, J–O, Q, S, T), 0.2 mm (F), not to scale (D, I, P, R)." httpUri="https://zenodo.org/record/1459878/files/figure.png" pageId="25" pageNumber="114">Fig. 3T</figureCitation>
|
||
). Leg-pair 2 (
|
||
<figureCitation id="719C2A68930FFF83FDEEF616FD301F4F" box="[628,718,332,357]" captionStart="FIGURE 7" captionStartId="11.[151,250,767,789]" captionTargetBox="[349,1245,180,761]" captionTargetId="figure@11.[340,1248,180,763]" captionTargetPageId="11" captionText="FIGURE 7. Male legs 2 in Julus subalpinus Lohmander, 1936 (A–B), J. dagestanus sp. n. (C) and J. khostensis sp. n. (D). A: from North Ossetia. B: syntype from Mount Djuga, Krasnodar Province, after Lohmander (1936). C: holotype from Dagestan. D: holotype from Khosta, Krasnodar Province. Oral views A, C, D; lateral view B. Abbreviations: alp, anterolateral process of coxa 2; mp, main process of coxa 2; p, penes. Scale bars: 0.3 mm." httpUri="https://zenodo.org/record/1459886/files/figure.png" pageId="25" pageNumber="114">Fig. 7D</figureCitation>
|
||
) with two pairs of coxal processes. Anterolateral process (
|
||
<emphasis id="DBD3EAFF930FFF83FAF7F616FA6C1F4E" bold="true" box="[1389,1426,332,357]" pageId="25" pageNumber="114">alp</emphasis>
|
||
) relatively long and slender. Main one (
|
||
<emphasis id="DBD3EAFF930FFF83FDD2F635FD911FA3" bold="true" box="[584,623,367,392]" pageId="25" pageNumber="114">mp</emphasis>
|
||
) short and slender.
|
||
</paragraph>
|
||
<paragraph id="E91836ED930FFF83FF5DF6CEFBD21FDF" blockId="25.[151,1437,151,536]" pageId="25" pageNumber="114">
|
||
All three parts of each gonopod (
|
||
<figureCitation id="719C2A68930FFF83FDA7F6CEFD5E1F87" box="[573,672,404,429]" captionStart="FIGURE 13" captionStartId="18.[151,250,1689,1711]" captionTargetBox="[340,1248,482,1662]" captionTargetId="figure@18.[340,1248,481,1668]" captionTargetPageId="18" captionText="FIGURE 13. Male gonopods in Julus dagestanus sp. n. (A–C holotype from Dagestan) and J. khostensis sp. n. (D–E holotype from Khosta, Krasnodar Province). A: promere with flagellum, mesal view. B: promere, caudal view. C: left gonopods, lateral view. D: promere with flagella, caudal view. E: right gonopods, lateral view. Scale bars: 0.3 mm. Abbreviations: fl, flagellum; la, lamella on promere; la1 and la2, lamella 1 and lamella 2 on promere; lo, lateral outgrowth of opisthomere; ms, mesomere; op, opisthomere; pr, promere." httpUri="https://zenodo.org/record/1459898/files/figure.png" pageId="25" pageNumber="114">Fig. 13E</figureCitation>
|
||
) forming a tight block. Promere (
|
||
<emphasis id="DBD3EAFF930FFF83FB84F6CEFBC41F86" bold="true" box="[1054,1082,404,429]" pageId="25" pageNumber="114">pr</emphasis>
|
||
) longer than opisthomere (
|
||
<emphasis id="DBD3EAFF930FFF83FAF4F6CEFA751F86" bold="true" box="[1390,1419,404,429]" pageId="25" pageNumber="114">op</emphasis>
|
||
), flagella (
|
||
<emphasis id="DBD3EAFF930FFF83FE9BF6EDFEEC1FFB" bold="true" box="[257,274,439,464]" pageId="25" pageNumber="114">fl</emphasis>
|
||
) well-developed;
|
||
<emphasis id="DBD3EAFF930FFF83FE73F6EDFE041FFB" bold="true" box="[489,506,439,464]" pageId="25" pageNumber="114">fl</emphasis>
|
||
with a lamina in apical part;
|
||
<emphasis id="DBD3EAFF930FFF83FCF0F6EDFC781FFB" bold="true" box="[874,902,439,464]" pageId="25" pageNumber="114">pr</emphasis>
|
||
narrow, with a thin lamella (
|
||
<emphasis id="DBD3EAFF930FFF83FB76F6EDFAFC1FFB" bold="true" box="[1260,1282,439,464]" pageId="25" pageNumber="114">la</emphasis>
|
||
) (
|
||
<figureCitation id="719C2A68930FFF83FABAF6EDFA701FFB" box="[1312,1422,439,464]" captionStart="FIGURE 13" captionStartId="18.[151,250,1689,1711]" captionTargetBox="[340,1248,482,1662]" captionTargetId="figure@18.[340,1248,481,1668]" captionTargetPageId="18" captionText="FIGURE 13. Male gonopods in Julus dagestanus sp. n. (A–C holotype from Dagestan) and J. khostensis sp. n. (D–E holotype from Khosta, Krasnodar Province). A: promere with flagellum, mesal view. B: promere, caudal view. C: left gonopods, lateral view. D: promere with flagella, caudal view. E: right gonopods, lateral view. Scale bars: 0.3 mm. Abbreviations: fl, flagellum; la, lamella on promere; la1 and la2, lamella 1 and lamella 2 on promere; lo, lateral outgrowth of opisthomere; ms, mesomere; op, opisthomere; pr, promere." httpUri="https://zenodo.org/record/1459898/files/figure.png" pageId="25" pageNumber="114">Fig. 13D</figureCitation>
|
||
); mesomere (
|
||
<emphasis id="DBD3EAFF930FFF83FE80F686FEC21FDE" bold="true" box="[282,316,476,501]" pageId="25" pageNumber="114">ms</emphasis>
|
||
) rounded; op and lateral outgrowth (
|
||
<emphasis id="DBD3EAFF930FFF83FD40F686FD0E1FDE" bold="true" box="[730,752,476,501]" pageId="25" pageNumber="114">lo</emphasis>
|
||
) pointed at apex (
|
||
<figureCitation id="719C2A68930FFF83FC21F686FBE11FDF" box="[955,1055,476,501]" captionStart="FIGURE 13" captionStartId="18.[151,250,1689,1711]" captionTargetBox="[340,1248,482,1662]" captionTargetId="figure@18.[340,1248,481,1668]" captionTargetPageId="18" captionText="FIGURE 13. Male gonopods in Julus dagestanus sp. n. (A–C holotype from Dagestan) and J. khostensis sp. n. (D–E holotype from Khosta, Krasnodar Province). A: promere with flagellum, mesal view. B: promere, caudal view. C: left gonopods, lateral view. D: promere with flagella, caudal view. E: right gonopods, lateral view. Scale bars: 0.3 mm. Abbreviations: fl, flagellum; la, lamella on promere; la1 and la2, lamella 1 and lamella 2 on promere; lo, lateral outgrowth of opisthomere; ms, mesomere; op, opisthomere; pr, promere." httpUri="https://zenodo.org/record/1459898/files/figure.png" pageId="25" pageNumber="114">Fig. 13E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="E91836ED930FFF83FF5DF55AFE451C33" blockId="25.[151,1437,151,536]" box="[199,443,511,536]" pageId="25" pageNumber="114">FEMALE. Unknown.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |