193 lines
19 KiB
XML
193 lines
19 KiB
XML
<document id="D9C4FBD16CB6502890F1CFD9C8D36B29" ID-CLB-Dataset="26920" ID-DOI="10.11646/zootaxa.4599.1.1" ID-GBIF-Dataset="ea727a9b-324b-4c6b-a625-bc7cdd204473" ID-ISSN="1175-5326" ID-Zenodo-Dep="2668058" ID-ZooBank="7CDD24FF-5BA3-4D06-898D-A3E220515D4C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1557127209543" checkinUser="plazi" docAuthor="Longino, John T." docDate="2019" docId="5B56E872FFAF4A64FF328BF8FDA6EB98" docLanguage="en" docName="zootaxa.4599.1.1.pdf.imf" docOrigin="Zootaxa 4599 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Pheidole musacolor Longino 2019, new species" docType="treatment" docVersion="21" lastPageNumber="48" masterDocId="A76F900AFF804A4BFFA58B6FFFF4EF5B" masterDocTitle="Pheidole (Hymenoptera, Formicidae) of Middle American Wet Forest" masterLastPageNumber="126" masterPageNumber="1" pageNumber="48" updateTime="1698726215157" updateUser="ExternalLinkService">
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<mods:mods id="4C3A2BD98DE1FEBB5D85B0D2B58584C3" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="22FD54662C455EBA3EC2B5AEADA03AC5">
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<mods:title id="3B3595D2D2D0993770226B727CED1AFE">Pheidole (Hymenoptera, Formicidae) of Middle American Wet Forest</mods:title>
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<mods:name id="C0FB69F4E96890F9E037C3E504C010F7" type="personal">
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<mods:namePart id="D3342EE4AF73BB6ED710A367B24A726B">Longino, John T.</mods:namePart>
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<mods:typeOfResource id="BA3737A960F22092C4C897A745882F30">text</mods:typeOfResource>
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<mods:titleInfo id="A9A02BC69B613B0A416D69371D645CF5">
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<mods:title id="5493488EB9C432312197CE8310349C1A">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="136EB951A1556BAE281B6DF8EFEFC220">
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<mods:date id="822A655FBC57D872A1A581D0566D9816">2019</mods:date>
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<mods:detail id="F3D88AC021CD29FF4479D33926CA46F8" type="pubDate">
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<mods:number id="B1AFF9BCCE75BFCA9E23D44DA9B805C5">2019-05-06</mods:number>
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<mods:detail id="749433FAB0A8D974159BD58DA0F614F7" type="volume">
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<mods:number id="5EC6D49EBDD376F338187066FA3C91A3">4599</mods:number>
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<mods:detail id="069730F8A59F842EC61E2E6CBB36E2DF" type="issue">
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<mods:number id="9668B3B87BDB69BB5D98EB6C996F842E">1</mods:number>
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<mods:start id="0EAF0206BC5FF394D2D53C8D80173E3E">1</mods:start>
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<mods:classification id="038CAA4FE46582B1DB5E21EF519ACA31">journal article</mods:classification>
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<mods:identifier id="57565D3BC2159DDF018278AC735D0877" type="CLB-Dataset">26920</mods:identifier>
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<mods:identifier id="FD80BE7E75F9F4A3F89A44F219AB3ACA" type="DOI">10.11646/zootaxa.4599.1.1</mods:identifier>
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<mods:identifier id="E73DF4ED66209579726D2E90013CD0DC" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="CAB842C38A336AAF6642CB7BD835C7D5" type="Zenodo-Dep">2668058</mods:identifier>
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</mods:mods>
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<treatment id="5B56E872FFAF4A64FF328BF8FDA6EB98" ID-GBIF-Taxon="157062631" LSID="urn:lsid:plazi:treatment:5B56E872FFAF4A64FF328BF8FDA6EB98" httpUri="http://treatment.plazi.org/id/5B56E872FFAF4A64FF328BF8FDA6EB98" lastPageNumber="48" pageId="47" pageNumber="48">
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<subSubSection id="9BE50AEFFFAF4A64FF328BF8FEF0EF88" pageId="47" pageNumber="48" type="nomenclature">
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<paragraph id="D3405964FFAF4A64FF328BF8FDEAEFE9" blockId="47.[151,542,151,211]" box="[151,542,151,178]" pageId="47" pageNumber="48">
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<heading id="8808EE08FFAF4A64FF328BF8FDEAEFE9" bold="true" box="[151,542,151,178]" fontSize="11" level="1" pageId="47" pageNumber="48" reason="1">
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<emphasis id="E18B8576FFAF4A64FF328BF8FDEAEFE9" bold="true" box="[151,542,151,178]" pageId="47" pageNumber="48">
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<taxonomicName id="14FF22E7FFAF4A64FF328BF8FE73EFEA" authority="Longino, 2019" authorityName="Longino" authorityYear="2019" box="[151,391,151,177]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="musacolor" status="sp. nov.">
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<emphasis id="E18B8576FFAF4A64FF328BF8FE73EFEA" bold="true" box="[151,391,151,177]" italics="true" pageId="47" pageNumber="48">Pheidole musacolor</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="FAB8380DFFAF4A64FE2B8BF7FDEAEFE9" box="[398,542,152,178]" pageId="47" pageNumber="48" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="D3405964FFAF4A64FF328BD5FEF0EF88" blockId="47.[151,542,151,211]" box="[151,260,186,211]" pageId="47" pageNumber="48">
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(
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<figureCitation id="4BC445E1FFAF4A64FF3B8BD5FF08EF88" box="[158,252,186,211]" captionStart="PLATE 55" captionStartId="121.[151,238,1954,1977]" captionTargetId="figure@121.[202,1384,194,1933]" captionTargetPageId="121" captionText="PLATE 55." httpUri="https://zenodo.org/record/2668347/files/figure.png" pageId="47" pageNumber="48">Plate 55</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="9BE50AEFFFAF4A64FF328A6DFC6FEE40" box="[151,923,258,283]" pageId="47" pageNumber="48" type="description">
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<paragraph id="D3405964FFAF4A64FF328A6DFC6FEE40" blockId="47.[151,1437,258,1219]" box="[151,923,258,283]" pageId="47" pageNumber="48">
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<taxonomicName id="14FF22E7FFAF4A64FF328A6DFF0FEE40" authorityName="Biolleyi Forel" authorityYear="1908" box="[151,251,258,283]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
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<emphasis id="E18B8576FFAF4A64FF328A6DFF0FEE40" box="[151,251,258,283]" italics="true" pageId="47" pageNumber="48">Pheidole</emphasis>
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</taxonomicName>
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JTL-243: morphospecies code previously used on AntWeb.
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</paragraph>
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</subSubSection>
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<subSubSection id="9BE50AEFFFAF4A64FF628A48FDDAEE94" pageId="47" pageNumber="48" type="materials_examined">
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<paragraph id="D3405964FFAF4A64FF628A48FEA2EEF0" blockId="47.[151,1437,258,1219]" pageId="47" pageNumber="48">
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<materialsCitation id="63975339FFAF4A64FF628A48FE17EEDC" ID-GBIF-Occurrence="2237851541" collectingDate="2003-01-03" collectingDateMax="2015-06-19" collectingDateMin="2003-01-03" collectionCode="MUCR" country="Costa Rica" elevation="0" location="Cerro Platano" pageId="47" pageNumber="48" specimenCount="1" stateProvince="Limon" typeStatus="holotype">
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<typeStatus id="0C44E7C6FFAF4A64FF628A48FEABEE64" box="[199,351,295,319]" pageId="47" pageNumber="48" type="holotype">HOLOTYPE</typeStatus>
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: 1 major worker,
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<collectingCountry id="ABE819F4FFAF4A64FD8C8A48FD5CEE64" box="[553,680,294,319]" name="Costa Rica" pageId="47" pageNumber="48">Costa Rica</collectingCountry>
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,
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<collectingRegion id="113B9786FFAF4A64FD108A49FCF0EE64" box="[693,772,294,319]" country="Costa Rica" name="Limon" pageId="47" pageNumber="48">Limón</collectingRegion>
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:
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<location id="D6200FBFFFAF4A64FCB58A48FC5BEE64" LSID="urn:lsid:plazi:treatment:5B56E872FFAF4A64FF328BF8FDA6EB98:D6200FBFFFAF4A64FCB58A48FC5BEE64" box="[784,943,294,319]" country="Costa Rica" name="Cerro Platano" pageId="47" pageNumber="48" stateProvince="Limon">Cerro Plátano</location>
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, 9.86621 -83.24203 ±
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<elevation id="58D2BE57FFAF4A64FB0F8A49FACEEE64" box="[1194,1338,294,319]" isEnumeration="true" pageId="47" pageNumber="48">
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<quantity id="1407F481FFAF4A64FB0F8A49FB2AEE64" box="[1194,1246,294,319]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="47" pageNumber="48" unit="m" value="50.0">50m</quantity>
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,
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<quantity id="1407F481FFAF4A64FB4E8A48FACEEE64" box="[1259,1338,295,319]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="47" pageNumber="48" unit="m" value="1110.0">1110m</quantity>
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</elevation>
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,
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<date id="A7417FA4FFAF4A64FAE28A48FF26EE38" pageId="47" pageNumber="48" value="2015-06-19">
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<collectingDate id="B705864CFFAF4A64FAE28A48FF26EE38" pageId="47" pageNumber="48" value="2015-06-19">19-Jun- 2015</collectingDate>
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</date>
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, cloud forest, along steep ridge near peak, at bait (ADMAC, Ba-E-
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<date id="A7417FA4FFAF4A64FC408A24FBA9EE38" box="[997,1117,331,355]" pageId="47" pageNumber="48" value="2003-01-03" valueMax="2003-03-03" valueMin="2003-01-03">
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<collectingDate id="B705864CFFAF4A64FC408A24FBA9EE38" box="[997,1117,331,355]" pageId="47" pageNumber="48" value="2003-01-03" valueMax="2003-03-03" valueMin="2003-01-03">03-1-03-03</collectingDate>
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</date>
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) [
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<collectionCode id="B5EEC1A1FFAF4A64FBD38A24FB3AEE38" box="[1142,1230,331,355]" country="Costa Rica" name="Museo de Insectos" pageId="47" pageNumber="48">MUCR</collectionCode>
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, unique specimen identifier CASENT0636691]
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</materialsCitation>
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.
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<materialsCitation id="63975339FFAF4A64FE558A00FEA6EEF0" ID-GBIF-Occurrence="2237851453" collectingDate="2003-01-03" collectingDateMax="2015-06-19" collectingDateMin="2003-01-03" collectionCode="CAS, DZUP, JTLC, MCZC, USNM" country="Costa Rica" elevation="0" location="Cerro Platano" pageId="47" pageNumber="48" specimenCount="1" stateProvince="Limon" typeStatus="paratype">
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<typeStatus id="0C44E7C6FFAF4A64FE558A00FD7BEEDC" box="[496,655,367,391]" pageId="47" pageNumber="48" type="paratype">PARATYPES</typeStatus>
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: major, minor workers: same data as holotype [
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<collectionCode id="B5EEC1A1FFAF4A64FB108A00FB04EEDC" box="[1205,1264,367,391]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="47" pageNumber="48" type="Museum">CAS</collectionCode>
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,
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<collectionCode id="B5EEC1A1FFAF4A64FB5A8A00FABCEEDD" box="[1279,1352,367,390]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="47" pageNumber="48" type="University or college">DZUP</collectionCode>
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,
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<collectionCode id="B5EEC1A1FFAF4A64FAF08A00FA6CEEDC" box="[1365,1432,367,391]" pageId="47" pageNumber="48">JTLC</collectionCode>
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,
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<collectionCode id="B5EEC1A1FFAF4A64FF328AFCFF1DEEF0" box="[151,233,403,427]" pageId="47" pageNumber="48">MCZC</collectionCode>
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,
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<collectionCode id="B5EEC1A1FFAF4A64FF518AFCFEBFEEF0" box="[244,331,403,427]" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="47" pageNumber="48" type="Museum">USNM</collectionCode>
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]
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="D3405964FFAF4A64FF628AD9FDDAEE94" blockId="47.[151,1437,258,1219]" box="[199,558,438,463]" pageId="47" pageNumber="48">
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<emphasis id="E18B8576FFAF4A64FF628AD9FE51EE94" bold="true" box="[199,421,438,463]" pageId="47" pageNumber="48">Geographic range.</emphasis>
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<collectingCountry id="ABE819F4FFAF4A64FE098AD8FDDEEE94" box="[428,554,438,463]" name="Costa Rica" pageId="47" pageNumber="48">Costa Rica</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="9BE50AEFFFAF4A64FF628AB5FA93EDB4" pageId="47" pageNumber="48" type="diagnosis">
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<paragraph id="D3405964FFAF4A64FF628AB5FA93EDB4" blockId="47.[151,1437,258,1219]" pageId="47" pageNumber="48">
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<emphasis id="E18B8576FFAF4A64FF628AB5FE6FEEA8" bold="true" box="[199,411,474,499]" pageId="47" pageNumber="48">
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Diagnosis.
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<emphasis id="E18B8576FFAF4A64FEF78AB5FE6FEEA8" bold="true" box="[338,411,474,499]" italics="true" pageId="47" pageNumber="48">Minor</emphasis>
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</emphasis>
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: face smooth and shining; head evenly rounded posteriorly; occipital carina thin; promesonotal groove present, distinctly impressed; lateral pronotum smooth and shiny; katepisternum uniformly foveate; propodeal spines long, similar in length to posterior face of propodeum; gaster smooth and shining; abundant, flexuous, erect setae on mesosomal dorsum, tibiae, and gastral dorsum; color yellow.
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<emphasis id="E18B8576FFAF4A64FAA28929FABAED04" bold="true" box="[1287,1358,582,607]" italics="true" pageId="47" pageNumber="48">Major</emphasis>
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: inner hypostomal teeth widely spaced, acicular; scape base subterete, narrower than apical portion; face with irregular, rugulose foveolate sculpture anteriorly, to about level of eyes, remainder of face smooth and shiny; promesonotal groove impressed; propodeal spines about half length of posterior face of propodeum; gastral dorsum smooth and shining; abundant, erect, flexuous setae on face, sides of head, mesosomal dorsum, tibiae, and gastral dorsum.
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</paragraph>
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</subSubSection>
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<subSubSection id="9BE50AEFFFAF4A64FF628995FD89EC24" pageId="47" pageNumber="48" type="description">
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<paragraph id="D3405964FFAF4A64FF628995FD7FEC6C" blockId="47.[151,1437,258,1219]" pageId="47" pageNumber="48">
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<emphasis id="E18B8576FFAF4A64FF628995FDD9EC48" bold="true" box="[199,557,762,787]" pageId="47" pageNumber="48">Measurements, minor worker</emphasis>
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: HW 0.73, HL 0.87, SL 1.20, EL 0.19, WL 1.12, PSL 0.18, PTW 0.14, PPW 0.20, CI 85, SI 164, PSLI 25, PPI 140 (n=1).
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</paragraph>
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<paragraph id="D3405964FFAF4A64FF62882DFD89EC24" blockId="47.[151,1437,258,1219]" pageId="47" pageNumber="48">
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<emphasis id="E18B8576FFAF4A64FF62882DFDD9EC00" bold="true" box="[199,557,834,859]" pageId="47" pageNumber="48">Measurements, major worker</emphasis>
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: HW 1.52, HL 1.56, SL 1.18, EL 0.25, WL 1.42, PSL 0.18, PTW 0.26, PPW 0.35, CI 97, SI 77, PSLI 12, PPI 132 (n=1).
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</paragraph>
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</subSubSection>
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<subSubSection id="9BE50AEFFFAF4A64FF6288E5FB62ECF8" box="[199,1174,906,931]" pageId="47" pageNumber="48" type="biology_ecology">
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<paragraph id="D3405964FFAF4A64FF6288E5FB62ECF8" blockId="47.[151,1437,258,1219]" box="[199,1174,906,931]" pageId="47" pageNumber="48">
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<emphasis id="E18B8576FFAF4A64FF6288E5FED3ECF8" bold="true" box="[199,295,906,931]" pageId="47" pageNumber="48">Biology.</emphasis>
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This species occurs in montane wet forest. It is known from two ground baits.
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</paragraph>
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</subSubSection>
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<subSubSection id="9BE50AEFFFAF4A64FF6288C1FF1FEBC5" pageId="47" pageNumber="48" type="discussion">
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<paragraph id="D3405964FFAF4A64FF6288C1FF1FEBC5" blockId="47.[151,1437,258,1219]" pageId="47" pageNumber="48">
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<emphasis id="E18B8576FFAF4A64FF6288C1FEBCEC9C" bold="true" box="[199,328,942,967]" pageId="47" pageNumber="48">Comments</emphasis>
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. This species appears to be part of a complex of large, yellow species that occur in the mountains of southern
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<collectingCountry id="ABE819F4FFAF4A64FEB988BCFE6EECB0" box="[284,410,978,1003]" name="Costa Rica" pageId="47" pageNumber="48">Costa Rica</collectingCountry>
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. It is similar to the species
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<taxonomicName id="14FF22E7FFAF4A64FD6D88BBFC86ECB0" authorityName="Longino" authorityYear="2019" box="[712,882,978,1003]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="familiaparra">
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<emphasis id="E18B8576FFAF4A64FD6D88BBFC86ECB0" box="[712,882,978,1003]" italics="true" pageId="47" pageNumber="48">P. familiaparra</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="14FF22E7FFAF4A64FC2588BBFC1BECB0" authorityName="Longino" authorityYear="2019" box="[896,1007,980,1003]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="savegre">
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<emphasis id="E18B8576FFAF4A64FC2588BBFC1BECB0" box="[896,1007,980,1003]" italics="true" pageId="47" pageNumber="48">P. savegre</emphasis>
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</taxonomicName>
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, and
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<taxonomicName id="14FF22E7FFAF4A64FB8A88BBFB6CECB0" authorityName="Longino" authorityYear="2019" box="[1071,1176,980,1003]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="tinamu">
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<emphasis id="E18B8576FFAF4A64FB8A88BBFB6CECB0" box="[1071,1176,980,1003]" italics="true" pageId="47" pageNumber="48">P. tinamu</emphasis>
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</taxonomicName>
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from the Pacific slope. It is most similar to
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<taxonomicName id="14FF22E7FFAF4A64FED38898FE13EB55" authorityName="Longino" authorityYear="2019" box="[374,487,1015,1038]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="savegre">
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<emphasis id="E18B8576FFAF4A64FED38898FE13EB55" box="[374,487,1015,1038]" italics="true" pageId="47" pageNumber="48">P. savegre</emphasis>
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</taxonomicName>
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, differing in finer, lighter-colored pilosity, and the major worker has a smoother, less sculptured face. The minor worker of
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<taxonomicName id="14FF22E7FFAF4A64FDE98F73FD0DEB68" authorityName="Longino" authorityYear="2019" box="[588,761,1050,1075]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="familiaparra">
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<emphasis id="E18B8576FFAF4A64FDE98F73FD0DEB68" box="[588,761,1050,1075]" italics="true" pageId="47" pageNumber="48">P. familiaparra</emphasis>
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</taxonomicName>
|
|
has a more tapered posterior head and a more pronounced occipital carina. The minor worker of
|
|
<taxonomicName id="14FF22E7FFAF4A64FDE68F50FD59EB0D" authorityName="Longino" authorityYear="2019" box="[579,685,1087,1110]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="tinamu">
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|
<emphasis id="E18B8576FFAF4A64FDE68F50FD59EB0D" box="[579,685,1087,1110]" italics="true" pageId="47" pageNumber="48">P. tinamu</emphasis>
|
|
</taxonomicName>
|
|
is not readily distinguishable, but the major worker of
|
|
<taxonomicName id="14FF22E7FFAF4A64FABD8F50FA8AEB0D" authorityName="Longino" authorityYear="2019" box="[1304,1406,1087,1110]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="tinamu">
|
|
<emphasis id="E18B8576FFAF4A64FABD8F50FA8AEB0D" box="[1304,1406,1087,1110]" italics="true" pageId="47" pageNumber="48">P. tinamu</emphasis>
|
|
</taxonomicName>
|
|
is distinctive, with strongly heart-shaped head and strongly produced promesonotum. See additional notes under
|
|
<taxonomicName id="14FF22E7FFAF4A64FA228F0BFF11EBC5" authorityName="Longino" authorityYear="2019" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="tinamu">
|
|
<emphasis id="E18B8576FFAF4A64FA228F0BFF11EBC5" italics="true" pageId="47" pageNumber="48">P. tinamu</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="9BE50AEFFFAF4A64FF628FC5FDA6EB98" box="[199,594,1194,1219]" pageId="47" pageNumber="48" type="etymology">
|
|
<paragraph id="D3405964FFAF4A64FF628FC5FDA6EB98" blockId="47.[151,1437,258,1219]" box="[199,594,1194,1219]" pageId="47" pageNumber="48">
|
|
<emphasis id="E18B8576FFAF4A64FF628FC5FEBCEB98" bold="true" box="[199,328,1194,1219]" pageId="47" pageNumber="48">Etymology</emphasis>
|
|
: The color of a banana.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |