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<document id="39054A7F03C359270C4A13DA4409D90D" ID-DOI="10.5281/zenodo.5172560" ID-ISSN="1942-1354" ID-Zenodo-Dep="5172560" ID-ZooBank="06B12AE7-A3D7-4D79-96BA-2BDD12B2250C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628270342172" checkinUser="felipe" docAuthor="Hawks, David C." docDate="2017" docId="545787BBD41AFF9E1AF0F8B5FD7ED9A2" docLanguage="en" docName="InsectaMundi.0544.1-9.pdf" docOrigin="Insecta Mundi 2017 (544)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Chrysina falcifera Hawks 2017, new species" docType="treatment" docVersion="3" lastPageNumber="2" masterDocId="A86EFFC3D418FF9D1A30FFB9FFE1DF78" masterDocTitle="Five new species of Chrysina Kirby (Coleoptera: Scarabaeidae: Rutelinae)" masterLastPageNumber="9" masterPageNumber="1" pageNumber="1" updateTime="1699038469858" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="D627988DD448D6BD5A3EB525773C081F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AFECA58DB80CD3025ABC18CD9D1C71FD">
<mods:title id="B89FC2E82E84B491FE9874C1FAC288D7">Five new species of Chrysina Kirby (Coleoptera: Scarabaeidae: Rutelinae)</mods:title>
</mods:titleInfo>
<mods:name id="9596D43B773C07717479F9606C999398" type="personal">
<mods:role id="81175644B1D3716F336DE8FD41C6E83F">
<mods:roleTerm id="C3AE2F513EB20D25605435991B7DE1AC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="049B5310D426C05EFC5A48A3A3779CA2">Hawks, David C.</mods:namePart>
</mods:name>
<mods:typeOfResource id="5814F9B0D4B9629125E572EC5A118BE9">text</mods:typeOfResource>
<mods:relatedItem id="4165689F74DF9094B89D652C91BE3FC1" type="host">
<mods:titleInfo id="D63E115C2752DAD4C47FFBB7EAD45125">
<mods:title id="CFB5192D98B593A4A7229A4C47DD3D77">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="9CC632AA3C5CE3700D954F1B3FFD23A7">
<mods:date id="795E10750D57F0F6C886F6618A11BFE8">2017</mods:date>
<mods:detail id="24349E8AB007DBBF2796A243B88EB77A" type="pubDate">
<mods:number id="0D069DF991F4FF81840B9CC5E3564234">2017-04-12</mods:number>
</mods:detail>
<mods:detail id="BBF63503FEA0772CBE3E740289FB83BD" type="volume">
<mods:number id="C21B83E4618C1EC1B481A3040E3472AC">2017</mods:number>
</mods:detail>
<mods:detail id="BA8F54C416B06333255593E1A369FFFD" type="issue">
<mods:number id="C27A8B963C1807FD0224E9FC75A94D8C">544</mods:number>
</mods:detail>
<mods:extent id="55F810E5F06CAA24F2EB3D8C1777268E" unit="page">
<mods:start id="AECC90D14B8B30CCAE70B3D6FB209936">1</mods:start>
<mods:end id="993B8DC327848BA8D60229A1D9AA84EF">9</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="3297F48D819E17D936DEF7EC6E802CF6">journal article</mods:classification>
<mods:identifier id="2C389609D432C40DAEAAB4E4DC606328" type="DOI">10.5281/zenodo.5172560</mods:identifier>
<mods:identifier id="FD01A86B4B7A94F473A9B592654BAB49" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="E2A5B8D220B47B542C9B7EDCDC5DCB35" type="Zenodo-Dep">5172560</mods:identifier>
<mods:identifier id="B34943D1B82B855F1770C6BC5DDA98A3" type="ZooBank">06B12AE7-A3D7-4D79-96BA-2BDD12B2250C</mods:identifier>
</mods:mods>
<treatment id="545787BBD41AFF9E1AF0F8B5FD7ED9A2" LSID="urn:lsid:plazi:treatment:545787BBD41AFF9E1AF0F8B5FD7ED9A2" httpUri="http://treatment.plazi.org/id/545787BBD41AFF9E1AF0F8B5FD7ED9A2" lastPageId="3" lastPageNumber="2" pageId="2" pageNumber="1">
<subSubSection id="94E46526D41AFF9F1AF0F8B5FD37D85C" box="[192,726,1804,1829]" pageId="2" pageNumber="1" type="nomenclature">
<paragraph id="DC4136ADD41AFF9F1AF0F8B5FD37D85C" blockId="2.[192,726,1804,1861]" box="[192,726,1804,1829]" pageId="2" pageNumber="1">
<heading id="870981C1D41AFF9F1AF0F8B5FD37D85C" bold="true" box="[192,726,1804,1829]" fontSize="10" level="2" pageId="2" pageNumber="1" reason="3">
<emphasis id="EE8AEABFD41AFF9F1AF0F8B5FD37D85C" bold="true" box="[192,726,1804,1829]" pageId="2" pageNumber="1">
<taxonomicName id="1BFE4D2ED41AFF9F1AF0F8B5FDC5D85D" authority="Hawks" authorityName="Hawks" authorityYear="2017" box="[192,548,1804,1829]" class="Insecta" family="Rutelidae" genus="Chrysina" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1" phylum="Arthropoda" rank="species" species="falcifera" status="sp. nov.">
<emphasis id="EE8AEABFD41AFF9F1AF0F8B5FE5DD85C" bold="true" box="[192,444,1804,1828]" italics="true" pageId="2" pageNumber="1">Chrysina falcifera</emphasis>
Hawks
</taxonomicName>
,
<taxonomicNameLabel id="F5B957C4D41AFF9F1801F8B4FD37D85C" box="[561,726,1804,1829]" pageId="2" pageNumber="1" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="94E46526D41AFF9F1AF0F895FE65D83D" box="[192,388,1836,1861]" pageId="2" pageNumber="1" type="description">
<paragraph id="DC4136ADD41AFF9F1AF0F895FE65D83D" blockId="2.[192,726,1804,1861]" box="[192,388,1836,1861]" pageId="2" pageNumber="1">
<heading id="870981C1D41AFF9F1AF0F895FE65D83D" bold="true" box="[192,388,1836,1861]" fontSize="10" level="2" pageId="2" pageNumber="1" reason="3">
<emphasis id="EE8AEABFD41AFF9F1AF0F895FE65D83D" bold="true" box="[192,388,1836,1861]" pageId="2" pageNumber="1">
<figureCitation id="44C52A28D41AFF9F1AF0F895FEA0D83D" box="[192,321,1836,1861]" captionStart="Figures 1–5" captionStartId="9.[192,288,1527,1549]" captionTargetBox="[195,1435,197,1510]" captionTargetId="figure-53@9.[192,1440,192,1518]" captionTargetPageId="9" captionText="Figures 1–5. Chrysina adult dorsal habitus photographs (not to scale). 1) Chrysina falcifer Hawks, new species. 2) Chrysina galbina Hawks, new species. 3) Chrysina juxtaprasina Hawks, new species. 4) Chrysina paulseni Hawks, new species. 5) Chrysina sagacita Hawks, new species." figureDoi="http://doi.org/10.5281/zenodo.5172562" httpUri="https://zenodo.org/record/5172562/files/figure.png" pageId="2" pageNumber="1">Figures 1</figureCitation>
,
<figureCitation id="44C52A28D41AFF9F1B60F894FE9FD83D" box="[336,382,1837,1861]" captionStart="Figures 6–20" captionStartId="10.[192,288,1444,1466]" captionTargetBox="[274,1350,220,1414]" captionTargetId="figure-131@10.[256,1379,192,1440]" captionTargetPageId="10" captionText="Figures 6–20. Chrysina genitalia photographs (not to scale). 6–8) Chrysina falcifer Hawks, new species, male genital capsule. 6) Dorsal view. 7) Ventral view. 8) Apical view. 9–11) Chrysina galbina Hawks, new species, male genital capsule. 9) Dorsal view. 10) Ventral view. 11) Apical view. 12–13) Chrysina juxtaprasina Hawks, new species, male genital capsule. 12) Dorsal view. 13) Ventral view. 14–16) Chrysina paulseni Hawks, new species, male genital capsule. 14) Dorsal view. 15) Ventral view. 16) Apical view. 17, 18) Chrysina sagacita Hawks, new species, male genital capsule. 17) Dorsal view. 18) Ventral view. 19) Chrysina juxtaprasina Hawks, new species, female inferior plates. 20) Chrysina paulseni Hawks, new species, female inferior plates." figureDoi="http://doi.org/10.5281/zenodo.5172566" httpUri="https://zenodo.org/record/5172566/files/figure.png" pageId="2" pageNumber="1">6–8</figureCitation>
.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="94E46526D41AFF9E1AF0F8D4FCDDDE22" lastPageId="3" lastPageNumber="2" pageId="2" pageNumber="1" type="materials_examined">
<paragraph id="DC4136ADD41AFF9F1AF0F8D4FA41D8FC" blockId="2.[192,1440,1901,1925]" box="[192,1440,1901,1925]" pageId="2" pageNumber="1">
<emphasis id="EE8AEABFD41AFF9F1AF0F8D4FEB2D8FD" bold="true" box="[192,339,1901,1925]" pageId="2" pageNumber="1">Type data.</emphasis>
<materialsCitation id="6C963CF0D41AFF9F1B6BF8D4FA41D8FC" box="[347,1440,1901,1925]" collectionCode="BMNH" country="Costa Rica" location="Pirris" municipality="June" pageId="2" pageNumber="1" specimenCount="1" typeStatus="holotype">
<typeStatus id="0345880FD41AFF9F1B6BF8D4FE2BD8FD" box="[347,458,1901,1925]" pageId="2" pageNumber="1" type="holotype">Holotype</typeStatus>
male (deposited at
<collectionCode id="BAEFAE68D41AFF9F18F0F8D4FCFDD8FD" box="[704,796,1901,1925]" pageId="2" pageNumber="1">BMNH</collectionCode>
), labeled: a) handwritten “
<location id="D9216076D41AFF9F1E57F8D4FB4CD8FD" LSID="urn:lsid:plazi:treatment:545787BBD41AFF9E1AF0F8B5FD7ED9A2:D9216076D41AFF9F1E57F8D4FB4CD8FD" box="[1127,1197,1901,1925]" country="Costa Rica" municipality="June" name="Pirris" pageId="2" pageNumber="1">Pirris</location>
/
<collectingCountry id="A4E9763DD41AFF9F1EF6F8D4FAABD8FD" box="[1222,1354,1901,1925]" name="Costa Rica" pageId="2" pageNumber="1">Costa Rica</collectingCountry>
/
<collectingMunicipality id="3C25ACD7D41AFF9F1F53F8D4FA41D8FC" box="[1379,1440,1901,1924]" pageId="2" pageNumber="1">June</collectingMunicipality>
</materialsCitation>
</paragraph>
<paragraph id="DC4136ADD41BFF9E1AF0FF7BFCDDDE22" blockId="3.[192,1440,194,346]" pageId="3" pageNumber="2">
“
<taxonomicName id="1BFE4D2ED41BFF9E1AFAFF7BFEDADFA2" authorityName="Kirby" authorityYear="1828" box="[202,315,194,218]" class="Insecta" family="Rutelidae" genus="Chrysina" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="EE8AEABFD41BFF9E1AFAFF7BFEDADFA2" box="[202,315,194,218]" italics="true" pageId="3" pageNumber="2">Chrysina</emphasis>
</taxonomicName>
/
<taxonomicName id="1BFE4D2ED41BFF9E1B60FF7BFE57DFA2" authorityName="Hawks" authorityYear="2017" box="[336,438,194,218]" class="Insecta" family="Rutelidae" genus="Chrysina" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="falcifera">
<emphasis id="EE8AEABFD41BFF9E1B60FF7BFE57DFA2" box="[336,438,194,218]" italics="true" pageId="3" pageNumber="2">falcifera</emphasis>
</taxonomicName>
<materialsCitation id="6C963CF0D41BFF9E1B8CFF7BFD3BDF81" collectorName="C. Rica" location="Hawks" pageId="3" pageNumber="2" specimenCount="2" typeStatus="holotype">
male symbol /
<location id="D9216076D41BFF9E185AFF7BFD22DFA2" LSID="urn:lsid:plazi:treatment:545787BBD41AFF9E1AF0F8B5FD7ED9A2:D9216076D41BFF9E185AFF7BFD22DFA2" box="[618,707,194,218]" name="Hawks" pageId="3" pageNumber="2">Hawks</location>
, 2017 /
<typeStatus id="0345880FD41BFF9E192DFF7BFC5DDFA2" box="[797,956,194,218]" pageId="3" pageNumber="2" type="holotype">HOLOTYPE</typeStatus>
”.
<typeStatus id="0345880FD41BFF9E19E1FF7BFBA3DFA2" box="[977,1090,194,218]" pageId="3" pageNumber="2" type="paratype">Paratype</typeStatus>
female labeled: a) handwritten “Savanillas / de Pirris /
<collectorName id="710B537BD41BFF9E1829FF5BFD93DF82" box="[537,626,226,250]" pageId="3" pageNumber="2">C. Rica</collectorName>
5.1900”
</materialsCitation>
;
<materialsCitation id="6C963CF0D41BFF9E18D7FF5BFBACDF82" box="[743,1101,226,250]" location="Nevinson Coll." pageId="3" pageNumber="2" specimenCount="1" typeStatus="paratype">
b) “
<location id="D9216076D41BFF9E1921FF5BFC26DF82" LSID="urn:lsid:plazi:treatment:545787BBD41AFF9E1AF0F8B5FD7ED9A2:D9216076D41BFF9E1921FF5BFC26DF82" box="[785,967,226,250]" name="Nevinson Coll." pageId="3" pageNumber="2">Nevinson Coll.</location>
/ 1918-14”
</materialsCitation>
; c) handwritten “
<taxonomicName id="1BFE4D2ED41BFF9E1F15FF5BFA71DF82" authorityName="Burmeister" authorityYear="1844" box="[1317,1424,226,250]" class="Insecta" family="Scarabaeidae" genus="Plusiotis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="EE8AEABFD41BFF9E1F15FF5BFA71DF82" box="[1317,1424,226,250]" italics="true" pageId="3" pageNumber="2">Plusiotis</emphasis>
</taxonomicName>
/
<emphasis id="EE8AEABFD41BFF9E1AF0FEBBFEAEDE61" box="[192,335,258,281]" italics="true" pageId="3" pageNumber="2">marginatus</emphasis>
/Waterh.”; d) on yellow paper, “
<taxonomicName id="1BFE4D2ED41BFF9E18E7FEBBFCA9DE62" authorityName="Kirby" authorityYear="1828" box="[727,840,258,282]" class="Insecta" family="Rutelidae" genus="Chrysina" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="EE8AEABFD41BFF9E18E7FEBBFCA9DE62" box="[727,840,258,282]" italics="true" pageId="3" pageNumber="2">Chrysina</emphasis>
</taxonomicName>
/
<taxonomicName id="1BFE4D2ED41BFF9E196FFEBBFC24DE62" authorityName="Hawks" authorityYear="2017" box="[863,965,258,282]" class="Insecta" family="Rutelidae" genus="Chrysina" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="falcifera">
<emphasis id="EE8AEABFD41BFF9E196FFEBBFC24DE62" box="[863,965,258,282]" italics="true" pageId="3" pageNumber="2">falcifera</emphasis>
</taxonomicName>
female symbol / Hawks, 2017 /
<collectionCode id="BAEFAE68D41BFF9E1F7CFEBBFEECDE42" pageId="3" pageNumber="2">PARA- TYPE</collectionCode>
”.
<materialsCitation id="6C963CF0D41BFF9E1B14FE9BFBF0DE42" box="[292,1041,290,314]" country="Costa Rica" location="Costa Rica" pageId="3" pageNumber="2" specimenCount="1" typeStatus="paratype">
<typeStatus id="0345880FD41BFF9E1B14FE9BFE74DE42" box="[292,405,290,314]" pageId="3" pageNumber="2" type="paratype">Paratype</typeStatus>
male (
<figureCitation id="44C52A28D41BFF9E1BDCFE9BFDD0DE42" box="[492,561,290,314]" captionStart="Figures 1–5" captionStartId="9.[192,288,1527,1549]" captionTargetBox="[195,1435,197,1510]" captionTargetId="figure-53@9.[192,1440,192,1518]" captionTargetPageId="9" captionText="Figures 1–5. Chrysina adult dorsal habitus photographs (not to scale). 1) Chrysina falcifer Hawks, new species. 2) Chrysina galbina Hawks, new species. 3) Chrysina juxtaprasina Hawks, new species. 4) Chrysina paulseni Hawks, new species. 5) Chrysina sagacita Hawks, new species." figureDoi="http://doi.org/10.5281/zenodo.5172562" httpUri="https://zenodo.org/record/5172562/files/figure.png" pageId="3" pageNumber="2">Fig. 1</figureCitation>
) labeled: a) handwritten “
<collectingCountry id="A4E9763DD41BFF9E194EFE9BFBE7DE42" box="[894,1030,290,314]" name="Costa Rica" pageId="3" pageNumber="2">Costa Rica</collectingCountry>
”
</materialsCitation>
; b) on yellow paper, “
<taxonomicName id="1BFE4D2ED41BFF9E1F2EFE9BFA6EDE42" authorityName="Kirby" authorityYear="1828" box="[1310,1423,290,314]" class="Insecta" family="Rutelidae" genus="Chrysina" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="EE8AEABFD41BFF9E1F2EFE9BFA6EDE42" box="[1310,1423,290,314]" italics="true" pageId="3" pageNumber="2">Chrysina</emphasis>
</taxonomicName>
/
<taxonomicName id="1BFE4D2ED41BFF9E1AF0FEFBFEC7DE22" authorityName="Hawks" authorityYear="2017" box="[192,294,322,346]" class="Insecta" family="Rutelidae" genus="Chrysina" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="falcifera">
<emphasis id="EE8AEABFD41BFF9E1AF0FEFBFEC7DE22" box="[192,294,322,346]" italics="true" pageId="3" pageNumber="2">falcifera</emphasis>
</taxonomicName>
male symbol / Hawks, 2017 /
<typeStatus id="0345880FD41BFF9E18A4FEFBFCCFDE22" box="[660,814,322,346]" pageId="3" pageNumber="2" type="paratype">PARATYPE</typeStatus>
”.
</paragraph>
</subSubSection>
<subSubSection id="94E46526D41BFF9E1AF0FE38FE0ADA42" pageId="3" pageNumber="2" type="description">
<paragraph id="DC4136ADD41BFF9E1AF0FE38FA7ADD62" blockId="3.[192,1441,385,954]" pageId="3" pageNumber="2">
<emphasis id="EE8AEABFD41BFF9E1AF0FE38FDDBDEE2" bold="true" box="[192,570,385,410]" pageId="3" pageNumber="2">
Description,
<typeStatus id="0345880FD41BFF9E1B40FE3BFE08DEE2" box="[368,489,386,410]" pageId="3" pageNumber="2" type="holotype">holotype</typeStatus>
male.
</emphasis>
Length
<quantity id="1B069B48D41BFF9E18AEFE38FCE8DEE1" box="[670,777,385,409]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" pageId="3" pageNumber="2" unit="mm" value="22.5">22.5 mm</quantity>
, width
<quantity id="1B069B48D41BFF9E1951FE38FC2FDEE1" box="[865,974,385,409]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="3" pageNumber="2" unit="mm" value="11.5">11.5 mm</quantity>
. Color of dorsal surface of head, pronotum, elytra, and pygidium mostly iridescent yellowish green; clypeus and pronotum edged with golden green; elytral margin with well-defined metallic golden band (includes apical calli); ventral surface of body iridescent green; femora and tibiae iridescent yellow-green; tarsi iridescent cupreous; antennal scape iridescent yellowish green dorsally; remainder of antennomeres present brown (clubs missing).
</paragraph>
<paragraph id="DC4136ADD41BFF9E1AC0FD9BFD75DDE2" blockId="3.[192,1441,385,954]" pageId="3" pageNumber="2">
<emphasis id="EE8AEABFD41BFF9E1AC0FD9BFEA0DD41" bold="true" box="[240,321,546,570]" pageId="3" pageNumber="2">Head.</emphasis>
Form broadly rounded. Dorsal surface of head with small, weakly impressed punctures, mixed with fine punctures. Clypeus convex in lateral view. Anterior border of labrum bisinuate, with 2 emarginations. Mandibles hidden in dorsal view. Eye size moderate with ratio of interocular distance to width of pronotum at base = 1: 2.1.
</paragraph>
<paragraph id="DC4136ADD41BFF9E1AC0FD1BFA73DC42" blockId="3.[192,1441,385,954]" pageId="3" pageNumber="2">
<emphasis id="EE8AEABFD41BFF9E1AC0FD1BFEBDDDC2" bold="true" box="[240,348,674,698]" pageId="3" pageNumber="2">Thorax.</emphasis>
Pronotal basal margin absent in central region adjacent to scutellum. Pronotal surface with small punctures larger, more deeply impressed than those of head; punctures sparse, becoming denser laterally. Mesosternal process long, slightly dorsoventrally compressed, rounded at apex. Each elytron with striae nearly obsolete, only sutural and 3 discal striae at all visible, interstriae weakly punctate as pronotum; epipleural fold narrow, terminating just posterior to first abdominal sternite.
</paragraph>
<paragraph id="DC4136ADD41BFF9E1AC0FCF8FD9DDCC1" blockId="3.[192,1441,385,954]" pageId="3" pageNumber="2">
<emphasis id="EE8AEABFD41BFF9E1AC0FCF8FE9BDC21" bold="true" box="[240,378,833,857]" pageId="3" pageNumber="2">Abdomen.</emphasis>
Surface smooth with several short, fine hairs at apex.
<emphasis id="EE8AEABFD41BFF9E1E3CFCFBFB66DC22" box="[1036,1159,834,858]" italics="true" pageId="3" pageNumber="2">Genitalia:</emphasis>
Genital capsule length
<quantity id="1B069B48D41BFF9E1AF0FCD8FEFFDC01" box="[192,286,865,889]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="3" pageNumber="2" unit="mm" value="7.5">7.5 mm</quantity>
. Parameres (
<figureCitation id="44C52A28D41BFF9E1B8BFCDBFDFCDC01" box="[443,541,865,890]" captionStart="Figures 6–20" captionStartId="10.[192,288,1444,1466]" captionTargetBox="[274,1350,220,1414]" captionTargetId="figure-131@10.[256,1379,192,1440]" captionTargetPageId="10" captionText="Figures 6–20. Chrysina genitalia photographs (not to scale). 6–8) Chrysina falcifer Hawks, new species, male genital capsule. 6) Dorsal view. 7) Ventral view. 8) Apical view. 9–11) Chrysina galbina Hawks, new species, male genital capsule. 9) Dorsal view. 10) Ventral view. 11) Apical view. 12–13) Chrysina juxtaprasina Hawks, new species, male genital capsule. 12) Dorsal view. 13) Ventral view. 14–16) Chrysina paulseni Hawks, new species, male genital capsule. 14) Dorsal view. 15) Ventral view. 16) Apical view. 17, 18) Chrysina sagacita Hawks, new species, male genital capsule. 17) Dorsal view. 18) Ventral view. 19) Chrysina juxtaprasina Hawks, new species, female inferior plates. 20) Chrysina paulseni Hawks, new species, female inferior plates." figureDoi="http://doi.org/10.5281/zenodo.5172566" httpUri="https://zenodo.org/record/5172566/files/figure.png" pageId="3" pageNumber="2">Fig. 6–8</figureCitation>
) fused into dorsoventrally flattened, ventrally reflexed, narrow hook, apex weakly notched; each side near base with 1 short triangular spine; ventral keel absent. Ventral plates asymmetrical, narrow, apices acute.
</paragraph>
<paragraph id="DC4136ADD41BFF9E1AF0FC5BFE18DB42" blockId="3.[192,1441,993,1082]" pageId="3" pageNumber="2">
<emphasis id="EE8AEABFD41BFF9E1AF0FC5BFD91DC81" bold="true" box="[192,624,993,1018]" pageId="3" pageNumber="2">
Male
<typeStatus id="0345880FD41BFF9E1B3CFC5BFE68DC82" box="[268,393,994,1018]" pageId="3" pageNumber="2" type="paratype">paratype</typeStatus>
variation (n= 1).
</emphasis>
Length
<quantity id="1B069B48D41BFF9E18E9FC5BFCD0DC81" box="[729,817,994,1018]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="2" unit="mm" value="22.0">22 mm</quantity>
, width
<quantity id="1B069B48D41BFF9E19BDFC5BFC07DC81" box="[909,998,994,1018]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="2" unit="mm" value="11.0">11 mm</quantity>
. Male
<typeStatus id="0345880FD41BFF9E1E08FC5BFB47DC82" box="[1080,1190,994,1018]" pageId="3" pageNumber="2" type="paratype">paratype</typeStatus>
(
<figureCitation id="44C52A28D41BFF9E1E87FC5BFB1ADC82" box="[1207,1275,994,1018]" captionStart="Figures 1–5" captionStartId="9.[192,288,1527,1549]" captionTargetBox="[195,1435,197,1510]" captionTargetId="figure-53@9.[192,1440,192,1518]" captionTargetPageId="9" captionText="Figures 1–5. Chrysina adult dorsal habitus photographs (not to scale). 1) Chrysina falcifer Hawks, new species. 2) Chrysina galbina Hawks, new species. 3) Chrysina juxtaprasina Hawks, new species. 4) Chrysina paulseni Hawks, new species. 5) Chrysina sagacita Hawks, new species." figureDoi="http://doi.org/10.5281/zenodo.5172562" httpUri="https://zenodo.org/record/5172562/files/figure.png" pageId="3" pageNumber="2">Fig. 1</figureCitation>
) differs from the
<typeStatus id="0345880FD41BFF9E1ADEFBBBFEB6DB62" box="[238,343,1026,1050]" pageId="3" pageNumber="2" type="holotype">holotype</typeStatus>
in the following ways. Ratio of antennal club length to interocular distance = 1: 2.2. Tibiae and tarsi coppery orange.
</paragraph>
<paragraph id="DC4136ADD41BFF9E1AF0FBDBFE0ADA42" blockId="3.[192,1441,1121,1338]" pageId="3" pageNumber="2">
<emphasis id="EE8AEABFD41BFF9E1AF0FBDBFD71DB01" bold="true" box="[192,656,1121,1146]" pageId="3" pageNumber="2">
Female
<typeStatus id="0345880FD41BFF9E1B1DFBDBFE4BDB02" box="[301,426,1122,1146]" pageId="3" pageNumber="2" type="paratype">paratype</typeStatus>
variation (n= 1).
</emphasis>
Length
<quantity id="1B069B48D41BFF9E18C7FBDBFCAFDB01" box="[759,846,1122,1146]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="3" pageNumber="2" unit="mm" value="27.0">27 mm</quantity>
, width
<quantity id="1B069B48D41BFF9E1999FBDBFBE3DB01" box="[937,1026,1122,1146]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="2" unit="mm" value="14.0">14 mm</quantity>
. Female differs from the
<typeStatus id="0345880FD41BFF9E1F07FBDBFA41DB02" box="[1335,1440,1122,1146]" pageId="3" pageNumber="2" type="holotype">holotype</typeStatus>
in the following aspects. Female with clypeal margin, tibiae, tarsi, and elytral margin purplish red. Elytral margin thickened in basal 2/3. As is common in the genus, the female body is slightly more convex in lateral view, legs and tarsi are relatively slightly smaller, and the antennal club is shorter. Ratio of interocular distance to width of pronotum at base = 1: 2.1. Ratio of antennal club length to interocular distance = 1: 2.4. Inferior genital plates subcircular, strongly convex, irregularly emarginate along apical margin.
</paragraph>
</subSubSection>
<subSubSection id="94E46526D41BFF9E1AF0FAD8FE46DAA2" pageId="3" pageNumber="2" type="diagnosis">
<paragraph id="DC4136ADD41BFF9E1AF0FAD8FE46DAA2" blockId="3.[192,1440,1377,1498]" pageId="3" pageNumber="2">
<emphasis id="EE8AEABFD41BFF9E1AF0FAD8FEB1DA01" bold="true" box="[192,336,1377,1401]" pageId="3" pageNumber="2">Diagnosis.</emphasis>
<taxonomicName id="1BFE4D2ED41BFF9E1B69FADBFDDBDA02" authorityName="Hawks" authorityYear="2017" box="[345,570,1378,1402]" class="Insecta" family="Rutelidae" genus="Chrysina" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="falcifera">
<emphasis id="EE8AEABFD41BFF9E1B69FADBFDDBDA02" box="[345,570,1378,1402]" italics="true" pageId="3" pageNumber="2">Chrysina falcifera</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="1BFE4D2ED41BFF9E191BFADBFB63DA02" authority="(Waterhouse)" baseAuthorityName="Waterhouse" baseAuthorityYear="1871" box="[811,1154,1378,1402]" class="Insecta" family="Rutelidae" genus="Chrysina" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="marginata">
<emphasis id="EE8AEABFD41BFF9E191BFADBFC33DA01" box="[811,978,1378,1401]" italics="true" pageId="3" pageNumber="2">C. marginata</emphasis>
(Waterhouse)
</taxonomicName>
and other members of the marginata group (
<bibRefCitation id="B86F4B5CD41BFF9E1BE6FA3BFD94DAE2" author="Hawks, D. C." box="[470,629,1410,1434]" pageId="3" pageNumber="2" pagination="1 - 8" refId="ref5979" refString="Hawks, D. C. 2001. Taxonomic and nomenclatural changes in Chrysina and a synonymic checklist of species (Scarabaeidae: Rutelinae). Occasional Papers of the Consortium Coleopterorum 4: 1 - 8." type="journal article" year="2001">Hawks 2001</bibRefCitation>
). It differs most conspicuously in the form of the male genitalia. The parameres of all other species of the marginata group possess very long lateral processes (
<bibRefCitation id="B86F4B5CD41BFF9E1F7AFA1BFF1FDAA2" author="Hawks, D. C." pageId="3" pageNumber="2" pagination="21 - 29" refId="ref5937" refString="Hawks, D. C. 1999. A review of the Plusiotis marginata complex including the description of a new species (Coleoptera: Scarabaeidae: Rutelinae). Occasional Papers of the Consortium Coleopterorum 3: 21 - 29." type="journal article" year="1999">Hawks 1999</bibRefCitation>
;
<bibRefCitation id="B86F4B5CD41BFF9E1B3AFA7BFE7ADAA2" author="Curoe, D. J." box="[266,411,1474,1498]" pageId="3" pageNumber="2" pagination="2 - 4" refId="ref5904" refString="Curoe, D. J. 2011. A new species of Chrysina Kirby of the marginata complex (Coleoptera: Scarabaeidae: Rutelinae). Besoiro 20: 2 - 4." type="journal article" year="2011">Curoe 2011</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="94E46526D41BFF9E1AF0F9BBFA60D942" pageId="3" pageNumber="2" type="etymology">
<paragraph id="DC4136ADD41BFF9E1AF0F9BBFA60D942" blockId="3.[192,1440,1538,1594]" pageId="3" pageNumber="2">
<emphasis id="EE8AEABFD41BFF9E1AF0F9BBFEBDD962" bold="true" box="[192,348,1538,1562]" pageId="3" pageNumber="2">Etymology.</emphasis>
The specific epithet is derived from the Latin
<taxonomicName id="1BFE4D2ED41BFF9E19B3F9BBFC38D962" authorityName="Hawks" authorityYear="2017" box="[899,985,1538,1562]" class="Insecta" family="Rutelidae" genus="Chrysina" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="falcifer">
<emphasis id="EE8AEABFD41BFF9E19B3F9BBFC38D962" box="[899,985,1538,1562]" italics="true" pageId="3" pageNumber="2">falcifer</emphasis>
</taxonomicName>
, meaning sickle-bearing, in reference to the strongly falcate male genitalia. It is used as a feminine adjective in the nominative singular.
</paragraph>
</subSubSection>
<subSubSection id="94E46526D41BFF9E1AF0F9DBFD7ED9A2" pageId="3" pageNumber="2" type="discussion">
<paragraph id="DC4136ADD41BFF9E1AF0F9DBFD7ED9A2" blockId="3.[192,1440,1634,1754]" pageId="3" pageNumber="2">
<emphasis id="EE8AEABFD41BFF9E1AF0F9DBFEA5D902" bold="true" box="[192,324,1634,1658]" pageId="3" pageNumber="2">Remarks.</emphasis>
Savanilla de Pirrís is in
<collectingRegion id="1E3AF84FD41BFF9E1841F9DBFCB3D902" box="[625,850,1634,1658]" country="Costa Rica" name="San Jose" pageId="3" pageNumber="2">San José Province</collectingRegion>
,
<collectingCountry id="A4E9763DD41BFF9E196DF9DBFC03D902" box="[861,994,1634,1658]" name="Costa Rica" pageId="3" pageNumber="2">Costa Rica</collectingCountry>
, located in the foothills of the Pacific Coast Range near Rio Grande de Pirrís. To my knowledge, there are no recently collected specimens of this species, and it would be worthwhile to survey for it, and assess its habitat and potential conservation issues in this region of
<collectingCountry id="A4E9763DD41BFF9E1824F97BFD7AD9A2" box="[532,667,1730,1754]" name="Costa Rica" pageId="3" pageNumber="2">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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