treatments-xml/data/52/CA/A9/52CAA9DE4357E74F763604651AC8569B.xml
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.349.6164" ID-GBIF-Dataset="cd871e7e-7e75-4f2e-8c8b-7af9cb4803d4" ID-PMC="PMC3837407" ID-Pensoft-Pub="1313-2970-349-81" ID-PubMed="24294080" ID-ZBK="45895E0B79E74794A0155CD74FEDDF22" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-349-81" ModsDocOrigin="ZooKeys 349" ModsDocTitle="Definition and review of the lancangjiang species group of the termitophilous genus Orthogonius Macleay, 1825 (Coleoptera, Carabidae, Orthogoniini)" checkinTime="1451246687136" checkinUser="pensoft" docAuthor="Tian, Mingyi &amp; Deuve, Thierry" docDate="2013" docId="52CAA9DE4357E74F763604651AC8569B" docLanguage="en" docName="ZooKeys 349: 81-100" docOrigin="ZooKeys 349" docSource="http://dx.doi.org/10.3897/zookeys.349.6164" docTitle="Orthogonius euthyphallus Tian &amp; Deuve, sp. n." docType="treatment" docUuid="596B1C35-4776-4C08-8D97-CD945B431249" docUuidSource="ZooBank" docVersion="5" lastPageNumber="88" masterDocId="B203D41F1D41C83E8603FFE5D1499035" masterDocTitle="Definition and review of the lancangjiang species group of the termitophilous genus Orthogonius Macleay, 1825 (Coleoptera, Carabidae, Orthogoniini)" masterLastPageNumber="100" masterPageNumber="81" pageNumber="86" updateTime="1668156835216" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Definition and review of the lancangjiang species group of the termitophilous genus Orthogonius Macleay, 1825 (Coleoptera, Carabidae, Orthogoniini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tian, Mingyi</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Deuve, Thierry</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="volume">
<mods:number>349</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>81</mods:start>
<mods:end>100</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.349.6164</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.349.6164</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-349-81</mods:identifier>
<mods:identifier type="ZBK">45895E0B79E74794A0155CD74FEDDF22</mods:identifier>
<mods:identifier type="ZooBank">45895E0B79E74794A0155CD74FEDDF22</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152049511" LSID="urn:lsid:zoobank.org:act:596B1C35-4776-4C08-8D97-CD945B431249" httpUri="http://treatment.plazi.org/id/52CAA9DE4357E74F763604651AC8569B" lastPageId="7" lastPageNumber="88" pageId="5" pageNumber="86">
<subSubSection pageId="5" pageNumber="86" type="nomenclature">
<paragraph pageId="5" pageNumber="86">
<taxonomicName LSID="http://zoobank.org/596B1C35-4776-4C08-8D97-CD945B431249" authority="Tian &amp; Deuve" class="Insecta" family="Carabidae" genus="Orthogonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthogonius euthyphallus" order="Coleoptera" pageId="5" pageNumber="86" phylum="Arthropoda" rank="species" species="euthyphallus">Orthogonius euthyphallus Tian &amp; Deuve</taxonomicName>
<taxonomicNameLabel pageId="5" pageNumber="86">sp. n.</taxonomicNameLabel>
Figs 1, 4, 10-11, 14-15
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="86" type="diagnosis">
<paragraph pageId="5" pageNumber="86">Diagnosis.</paragraph>
<paragraph pageId="5" pageNumber="86">Medium to large sized, slender and elongate; head longer than wide; upper surface covered with dense punctures; antennae long; eyes moderate in size; labrum broad at front margin; pronotum quite narrow, but transverse in form; whole lateral expanded margins slightly reflexed; elytra not carinate on interval 7 in both sexes.</paragraph>
<paragraph pageId="5" pageNumber="86">Length: 14.0-16.0 mm; width: 5.5-6.5 mm. Habitus as in Fig. 4.</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="87" pageId="5" pageNumber="86" type="description">
<paragraph pageId="5" pageNumber="86">Description.</paragraph>
<paragraph pageId="5" pageNumber="86">Body elongate, strongly shiny.</paragraph>
<paragraph pageId="6" pageNumber="87">
<pageBreakToken pageId="6" pageNumber="87" start="start">Head</pageBreakToken>
and disc of pronotum black; elytra dark brown or brown; lateral expanded margins of pronotum, antennomere 1 and 3, mandibles, and tibiae dark brown; palps, legs excluding tibiae and antennae brown.
</paragraph>
<paragraph pageId="6" pageNumber="87">Macrosculpture: Surface of head, pronotum and elytra with dense punctures, head obscurely striate, pronotum and elytra without wrinkles.</paragraph>
<paragraph pageId="6" pageNumber="87">Microsculpture: Engraved meshes isodiametric on head and elytra, but moderate transverse on pronotum.</paragraph>
<paragraph pageId="6" pageNumber="87">Head longer than wide, HL/HW=1.15-1.18, eyes rather small but prominent, frons and vertex convex, frontal impressions large and deep; clypeus bisetose, basal portion unevenly convex; labrum broad at front margin, sexsetose; palps rather stout, subcylindrical, maxillary palpomere 3 as long as 4, palpomere 4 glabrous, palpomere 3 with two short setae at apex; labial palpomere 2 slightly longer than 3, bisetose on inner margin, with several additional setae at subapex and apex, palpomere 3 bearing a few tiny setae; ligula small, bisetose at apex; mentum without tooth, each of mentum and submentum bisetose, palpiger asetose. Antennae very long, extending to the middle of elytra; pubescent and slightly expanded from basal 1/3 of antennomere 4; antennomrere 3 almost as long as 4, and 1.32 times longer than 2; antennomere 1 unisetoae on subapex.</paragraph>
<paragraph pageId="6" pageNumber="87">Pronotum strongly transverse, PW/PL=1.55-1.63, disc moderately convex, fore and basal margins well beaded, sides evenly expanded, widest at about middle; lateral expanded margin well defined, flat and very smooth, slightly reflexed; both transversal impressions distinct, basal foveae small and deep; median line clear.</paragraph>
<paragraph pageId="6" pageNumber="87">Elytra elongate ovate, EL/EW=1.71-1.82; widest at a little behind middle, sides slightly expanded at middle, basal border complete, apex broadly sinuate, inner angle nearly rectangular and obtuse; striae very deep, intervals strongly convex, subequal in width in middle, interval 3 with basal and middle setiferous pores, the subapical one absent.</paragraph>
<paragraph pageId="6" pageNumber="87">Legs slender, fore tibia distinctly expanded at apex, apex obliquely truncate, outer margin slightly subserrate; middle and hind coxae smooth and glabrous; middle tibia strongly curve, slightly expanded at apex; hind tibia hardly expanded at apex, apical spurs long and sharp, hind tarsomere 3 much longer than 4, length ratio of tarsomeres 1-4 as 2.55, 1.79, 1.32 and 1.0; tarsomere 4 asymmetrically emarginated at apex, longer lobe as long as 1/3 of the whole joint; hind femur thin, slightly expanded medially, with two long setae posteriorly; tarsal claws strongly pectinate.</paragraph>
<paragraph pageId="6" pageNumber="87">Prosternal process well bordered at apex, abdominal ventrite VII of male distinctly and deeply emarginate at apical margin.</paragraph>
<paragraph pageId="6" pageNumber="87">Male genitalia (Figs 14-15): Slender and quite straight, slightly sinuate on ventral surface, apex blunt; the apical lamella as long as wide, broad at tip.</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="87" type="remarks">
<paragraph pageId="6" pageNumber="87">Remarks.</paragraph>
<paragraph pageId="6" pageNumber="87">
This new species differs from
<taxonomicName class="Insecta" family="Carabidae" genus="Orthogonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthogonius lancangjiang" order="Coleoptera" pageId="6" pageNumber="87" phylum="Arthropoda" rank="species" species="lancangjiang">Orthogonius lancangjiang</taxonomicName>
by the wider emargination on ventrite VII in males, which is gradually narrowed towards base (suddenly but somewhat obliquely narrowed towards base in
<taxonomicName class="Insecta" family="Carabidae" genus="Orthogonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthogonius lancangjiang" order="Coleoptera" pageId="6" pageNumber="87" phylum="Arthropoda" rank="species" species="lancangjiang">Orthogonius lancangjiang</taxonomicName>
), the median lobe of aedeagus is less straight, more sinuate ventrally, and apex thin and broad (thick and obtuse at apex in
<taxonomicName class="Insecta" family="Carabidae" genus="Orthogonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthogonius lancangjiang" order="Coleoptera" pageId="6" pageNumber="87" phylum="Arthropoda" rank="species" species="lancangjiang">Orthogonius lancangjiang</taxonomicName>
); and elytra with sparser punctures (densely punctate in
<taxonomicName class="Insecta" family="Carabidae" genus="Orthogonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthogonius lancangjiang" order="Coleoptera" pageId="6" pageNumber="87" phylum="Arthropoda" rank="species" species="lancangjiang">Orthogonius lancangjiang</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="88" type="material examined">
<paragraph pageId="7" pageNumber="88">
<pageBreakToken pageId="7" pageNumber="88" start="start">Material</pageBreakToken>
examined.
</paragraph>
<paragraph pageId="7" pageNumber="88">Holotype: male, &quot;S. Vietnam, 28-30.4.1994, 12 km N. Dalat, Lang Bian, Pacholatko &amp; Dembicky&quot;, &quot;Mus. Wien&quot;, in NHMV.</paragraph>
<paragraph pageId="7" pageNumber="88">Paratypes. 1 male, idem; 9 males and 3 females, &quot;S. Vietnam, 17-21.4.1995, 12 km N. Dalat, Lang Bian&quot;, &quot;12.03N108.27E, 1580-1750 m, Pacholatko &amp; Dembicky&quot;, and &quot;Mus. Wien&quot;, all in NHMV except 1 male and 1 female in SCAU.</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="88" type="etymology">
<paragraph pageId="7" pageNumber="88">Etymology.</paragraph>
<paragraph pageId="7" pageNumber="88">
The name of the new species is combined by the Greek prefix
<normalizedToken originalValue="“euthy-”">&quot;euthy-&quot;</normalizedToken>
, meaning straight, and word
<normalizedToken originalValue="“phallus”">&quot;phallus&quot;</normalizedToken>
, meaning penis, to refer to the straight median lobe of aedeagus.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="88" type="distribution">
<paragraph pageId="7" pageNumber="88">Distribution.</paragraph>
<paragraph pageId="7" pageNumber="88">Southern Vietnam (Fig. 1).</paragraph>
</subSubSection>
</treatment>
</document>