461 lines
66 KiB
XML
461 lines
66 KiB
XML
<document id="F0BF16A49E8AD3E22CA810CB570E2AA1" ID-DOI="10.5281/zenodo.176880" ID-GBIF-Dataset="48069658-661a-4c81-bea3-59b725356587" ID-ISSN="1175-5326" ID-Zenodo-Dep="176880" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459842447241" checkinUser="plazi" docAuthor="Kodada, Ján, Jäch, Manfred A., Jr, Fedor Č Iampor & Ová, Zuzana Č Iamporová- Za Ť Ovi Č" docDate="2007" docId="4F6087CAFFF2FF8121B4D81C0A9F8241" docLanguage="en" docName="zt01481p068.pdf" docOrigin="Zootaxa 1481" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Geoparnus rhinoceros Kodada, Jäch, Jr & Ová, 2007, sp. nov." docType="treatment" docVersion="8" lastPageNumber="68" masterDocId="B359FFB2FFF3FF882123DD060932833B" masterDocTitle="Geoparnus rhinoceros sp. nov., a new edaphic dryopid with unusual sexual dimorphism (Coleoptera: Dryopidae)" masterLastPageNumber="68" masterPageNumber="59" pageNumber="60" updateTime="1698224410878" updateUser="plazi">
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<mods:mods id="18507EA792583917E918274F0944B96F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="34A6A12B7EA8EB56D2A094325BE3BD48">
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<mods:title id="FE42CA2C3FB1E9ECA13872A7718B7AD3">Geoparnus rhinoceros sp. nov., a new edaphic dryopid with unusual sexual dimorphism (Coleoptera: Dryopidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="FE06836CC7DE79E56DCBAD6B457A6DF2" type="personal">
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<mods:role id="C43E70FA36D35FC732792FB1908149EA">
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<mods:roleTerm id="4726EDE099AED80F5DD7CE41D5A87AA7">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7123381EAB41F0CBA8CF56FA6DCD4DBA">Kodada, Ján</mods:namePart>
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</mods:name>
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<mods:roleTerm id="A3F2AF86CE775E667C32283214A6435E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="48DB677A7A19E2D8C7CBAD565630C797">Jäch, Manfred A.</mods:namePart>
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</mods:name>
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<mods:name id="82899D794C6EB9EB9F4A71C056F5812B" type="personal">
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<mods:roleTerm id="269759EE7CBFA4C5E931A49248AECDE2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6ED3033E36A3B65A04D8B287CE4865C3">Jr, Fedor Č Iampor</mods:namePart>
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</mods:name>
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<mods:name id="2A64C7A0804886E00215F3994AF91D20" type="personal">
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<mods:role id="FB52D5FC187FD5D1825C7DD6F6FF4AE0">
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<mods:roleTerm id="84E5036FA88B465733340AE862DEF196">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7C84E8DDD24A1B570DBBBA89664D8B10">Ová, Zuzana Č Iamporová- Za Ť Ovi Č</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="DD4F3A7AA4A9F858AB11669F38A360EC">text</mods:typeOfResource>
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<mods:titleInfo id="320772F923D5CC65763784E7EFE41448">
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<mods:title id="27D5268E9A5DFF36CF4781F9B4A65B80">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="4FDE80DB32359300799BBBE06D58B676">
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<mods:date id="9CC61CB0D4F2D7D296658C95CEEA98C9">2007</mods:date>
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<mods:detail id="D6A39F28D53980B8FB88BCA4DF5080B8" type="volume">
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<mods:number id="E4728AA0E9FF895D925D208AFC72A112">1481</mods:number>
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<mods:end id="DDEC1E579B9918FE3E85FBC7B7D82616">68</mods:end>
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<mods:classification id="3A2443CE190E10D91DDF1C74E114397C">journal article</mods:classification>
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<mods:identifier id="24CC93F9C3B49DD3D364E83BC13A7035" type="DOI">10.5281/zenodo.176880</mods:identifier>
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<mods:identifier id="A43BE060A0CEE6C038F4D1A05545DBEB" type="GBIF-Dataset">48069658-661a-4c81-bea3-59b725356587</mods:identifier>
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<mods:identifier id="97C207AD4ECBBB28D654CBB8B8A74085" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="5032128C280F8AFA5DB21EE8A6BD8CF1" type="Zenodo-Dep">176880</mods:identifier>
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</mods:mods>
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<treatment id="4F6087CAFFF2FF8121B4D81C0A9F8241" ID-DOI="http://doi.org/10.5281/zenodo.6237917" ID-GBIF-Taxon="119360586" ID-Zenodo-Dep="6237917" LSID="urn:lsid:plazi:treatment:4F6087CAFFF2FF8121B4D81C0A9F8241" httpUri="http://treatment.plazi.org/id/4F6087CAFFF2FF8121B4D81C0A9F8241" lastPageId="9" lastPageNumber="68" pageId="1" pageNumber="60">
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<subSubSection id="8FD36557FFF2FF8921B4D81C0814866C" pageId="1" pageNumber="60" type="nomenclature">
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<paragraph id="C77636DCFFF2FF8921B4D81C0B39860F" blockId="1.[151,523,1306,1367]" box="[151,523,1306,1332]" pageId="1" pageNumber="60">
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<heading id="9C3E81B0FFF2FF8921B4D81C0B39860F" bold="true" box="[151,523,1306,1332]" fontSize="11" level="1" pageId="1" pageNumber="60" reason="1">
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<emphasis id="F5BDEACEFFF2FF8921B4D81C0B39860F" bold="true" box="[151,523,1306,1332]" pageId="1" pageNumber="60">
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||
<taxonomicName id="00C94D5FFFF2FF8921B4D81C0894860F" box="[151,422,1306,1332]" class="Insecta" family="Dryopidae" genus="Geoparnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="species" species="rhinoceros" status="sp. nov.">
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<emphasis id="F5BDEACEFFF2FF8921B4D81C0894860F" bold="true" box="[151,422,1306,1332]" italics="true" pageId="1" pageNumber="60">Geoparnus rhinoceros</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="EE8E57B5FFF2FF89208ED81C0B39860F" box="[429,523,1306,1332]" pageId="1" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="C77636DCFFF2FF8921B4D83B0814866C" blockId="1.[151,523,1306,1367]" box="[151,294,1341,1367]" pageId="1" pageNumber="60">
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(
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<figureCitation id="5FF22A59FFF2FF8921BCD83B09DE866C" box="[159,236,1341,1367]" captionStart="FIGURE 1" captionStartId="2.[151,255,2000,2024]" captionTargetBox="[215,1368,596,1969]" captionTargetId="figure@2.[196,1390,581,1998]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Geoparnus rhinoceros sp. nov., male." httpUri="https://zenodo.org/record/176881/files/figure.png" pageId="1" pageNumber="60">Figs. 1</figureCitation>
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–20)
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</paragraph>
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</subSubSection>
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||
<subSubSection id="8FD36557FFF2FF8921B4D88B0C178434" pageId="1" pageNumber="60" type="materials_examined">
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<paragraph id="C77636DCFFF2FF8921B4D88B0B0086F4" blockId="1.[151,1437,1421,2007]" pageId="1" pageNumber="60">
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<emphasis id="F5BDEACEFFF2FF8921B4D88B0873869C" bold="true" box="[151,321,1421,1447]" pageId="1" pageNumber="60">
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<typeStatus id="1872887EFFF2FF8921B4D88B09E7869C" box="[151,213,1421,1447]" pageId="1" pageNumber="60">Type</typeStatus>
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locality:
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</emphasis>
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forest floor debris in primary rain forest, close to a tributary of River
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<collectingCountry id="BFDE764CFFF2FF89255FD88B0D94869C" box="[1148,1190,1421,1447]" name="Spain" pageId="1" pageNumber="60">Sut</collectingCountry>
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, about
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<quantity id="00319B39FFF2FF8925DFD88B0C75869C" box="[1276,1351,1421,1447]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="60" unit="km" value="25.0">25 km</quantity>
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east of Kapit, Borneo,
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<collectingCountry id="BFDE764CFFF2FF89206FD8B3088686F4" box="[332,436,1461,1487]" name="Malaysia" pageId="1" pageNumber="60">Sarawak</collectingCountry>
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,
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<collectingCountry id="BFDE764CFFF2FF89209CD8B30B1C86F4" box="[447,558,1461,1487]" name="Malaysia" pageId="1" pageNumber="60">Malaysia</collectingCountry>
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.
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</paragraph>
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<paragraph id="C77636DCFFF2FF8921E5D8DB0C178434" blockId="1.[151,1437,1421,2007]" pageId="1" pageNumber="60">
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<emphasis id="F5BDEACEFFF2FF8921E5D8DB0B0E86CC" bold="true" box="[198,572,1501,1527]" pageId="1" pageNumber="60">
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Material examined:
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||
<typeStatus id="1872887EFFF2FF8920E9D8DB0B0E86CC" box="[458,572,1501,1527]" pageId="1" pageNumber="60" type="holotype">Holotype</typeStatus>
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||
</emphasis>
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||
ɗ (
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||
<collectionCode id="A1D8AE19FFF2FF89234FD8DB0B8C86CC" box="[620,702,1501,1527]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="1" pageNumber="60">NMW</collectionCode>
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||
): ”
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<collectingCountry id="BFDE764CFFF2FF8923FCD8DB0A7586CC" box="[735,839,1501,1527]" name="Malaysia" pageId="1" pageNumber="60">Sarawak</collectingCountry>
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(Borneo), ca
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<quantity id="00319B39FFF2FF8922D2D8DB0D0C86CC" box="[1009,1086,1501,1527]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="60" unit="km" value="25.0">25 km</quantity>
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E Kapit, 3. 1994, J. Kodada leg.”.
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<typeStatus id="1872887EFFF2FF8921FCDB03086E8524" box="[223,348,1541,1567]" pageId="1" pageNumber="60" type="paratype">
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||
<emphasis id="F5BDEACEFFF2FF8921FCDB03086E8524" bold="true" box="[223,348,1541,1567]" pageId="1" pageNumber="60">Paratypes</emphasis>
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||
</typeStatus>
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25 ɗɗ, 24 ΨΨ (
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<collectionCode id="A1D8AE19FFF2FF89233CDB030B5E8524" box="[543,620,1541,1567]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="1" pageNumber="60">NMW</collectionCode>
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,
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<collectionCode id="A1D8AE19FFF2FF89235ADB030B898524" box="[633,699,1541,1567]" pageId="1" pageNumber="60">CKB</collectionCode>
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||
): with the same locality data as
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||
<typeStatus id="1872887EFFF2FF89251BDB030D908524" box="[1080,1186,1541,1567]" pageId="1" pageNumber="60" type="holotype">holotype</typeStatus>
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||
; 4 ɗɗ, 5 ΨΨ (
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<collectionCode id="A1D8AE19FFF2FF89246EDB030CA28524" box="[1357,1424,1541,1567]" pageId="1" pageNumber="60">CKB</collectionCode>
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): "
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||
<collectingCountry id="BFDE764CFFF2FF8921BCDB2B0835857C" box="[159,263,1581,1607]" name="Malaysia" pageId="1" pageNumber="60">Sarawak</collectingCountry>
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||
(Borneo), Gunung Serapi, ca
|
||
<quantity id="00319B39FFF2FF89234CDB2B0B89857C" box="[623,699,1581,1607]" metricMagnitude="4" metricUnit="m" metricValue="1.9" pageId="1" pageNumber="60" unit="km" value="19.0">19 km</quantity>
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||
W Kuching, P. Bílek leg."; 2 ΨΨ (
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<collectionCode id="A1D8AE19FFF2FF89257ADB2B0DAE857C" box="[1113,1180,1581,1607]" pageId="1" pageNumber="60">CKB</collectionCode>
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||
): "
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||
<collectingCountry id="BFDE764CFFF2FF892599DB2B0C10857C" box="[1210,1314,1581,1607]" name="Malaysia" pageId="1" pageNumber="60">Sarawak</collectingCountry>
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||
(Borneo), Gunung Serapi, ca
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||
<quantity id="00319B39FFF2FF892055DB53088D8554" box="[374,447,1621,1647]" metricMagnitude="4" metricUnit="m" metricValue="1.9" pageId="1" pageNumber="60" unit="km" value="19.0">19 km</quantity>
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W Kuching, primary forest,
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<date id="B377101CFFF2FF892232DB530A488554" box="[785,890,1621,1647]" pageId="1" pageNumber="60" value="1994-03">III. 1994</date>
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, J. Kodada leg."; 106 ɗɗ, 97 ΨΨ (
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||
<collectionCode id="A1D8AE19FFF2FF892429DB530C598554" box="[1290,1387,1621,1647]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="1" pageNumber="60">MHNG</collectionCode>
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||
): "E
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||
<collectingCountry id="BFDE764CFFF2FF8921B4DB7B083585AC" box="[151,263,1661,1687]" name="Malaysia" pageId="1" pageNumber="60">Malaysia</collectingCountry>
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||
:
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<collectingCountry id="BFDE764CFFF2FF892032DB7B084B85AC" box="[273,377,1661,1687]" name="Malaysia" pageId="1" pageNumber="60">Sarawak</collectingCountry>
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, 1994 Santubong,
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<quantity id="00319B39FFF2FF892371DB7B0BAE85AC" box="[594,668,1661,1687]" metricMagnitude="4" metricUnit="m" metricValue="3.2" pageId="1" pageNumber="60" unit="km" value="32.0">32 km</quantity>
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N Kuching,
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<quantity id="00319B39FFF2FF892210DB7B0AAF85AC" box="[819,925,1661,1687]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="1" pageNumber="60" unit="m" value="50.0" valueMax="100.0" valueMin="0.0">0–100 m</quantity>
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||
,
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<date id="B377101CFFF2FF89228ADB7B0D2985AC" box="[937,1051,1661,1687]" pageId="1" pageNumber="60">11.–15.V.</date>
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, no 1a, Burckhardt & Löbl leg."; 46 ɗɗ, 51 ΨΨ (
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<collectionCode id="A1D8AE19FFF2FF89206CDBA308828584" box="[335,432,1701,1727]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="1" pageNumber="60">MHNG</collectionCode>
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||
): "E-
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||
<collectingCountry id="BFDE764CFFF2FF8920D7DBA30B568584" box="[500,612,1701,1727]" name="Malaysia" pageId="1" pageNumber="60">Malaysia</collectingCountry>
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||
:
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||
<collectingCountry id="BFDE764CFFF2FF89234CDBA30BE58584" box="[623,727,1701,1727]" name="Malaysia" pageId="1" pageNumber="60">Sarawak</collectingCountry>
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||
, 1994, Gn. Gading, E slope,
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<quantity id="00319B39FFF2FF89250EDBA30D5E8584" box="[1069,1132,1701,1727]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="60" unit="m" value="50.0">50 m</quantity>
|
||
,
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||
<quantity id="00319B39FFF2FF89255BDBA30D868584" box="[1144,1204,1701,1727]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="60" unit="km" value="9.0">9 km</quantity>
|
||
E Lundu,
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||
<date id="B377101CFFF2FF892411DBA30C408584" box="[1330,1394,1701,1727]" pageId="1" pageNumber="60">14.V.</date>
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||
, no 3a, Burckhardt & Löbl leg."; 8 ɗɗ, 6 ΨΨ (
|
||
<collectionCode id="A1D8AE19FFF2FF892399DBCB0A2385DC" LSID="urn:lsid:biocol.org:col:34840" box="[698,785,1741,1767]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="1" pageNumber="60">SMNS</collectionCode>
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||
): "Borneo:
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||
<collectingCountry id="BFDE764CFFF2FF892282DBCB0D3B85DC" box="[929,1033,1741,1767]" name="Malaysia" pageId="1" pageNumber="60">Sarawak</collectingCountry>
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||
, Belaga
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||
<date id="B377101CFFF2FF89255BDBCB0C0985DC" box="[1144,1339,1741,1767]" pageId="1" pageNumber="60" value="1990-03-14" valueMax="1990-03-16" valueMin="1990-03-14">14.–16. 3. 1990</date>
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||
leg. A. RIEDEL"; 5 ɗɗ, 2 ΨΨ (
|
||
<collectionCode id="A1D8AE19FFF2FF892090DBF30B3B8434" LSID="urn:lsid:biocol.org:col:34840" box="[435,521,1781,1807]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="1" pageNumber="60">SMNS</collectionCode>
|
||
): "Borneo:
|
||
<collectingCountry id="BFDE764CFFF2FF8923ACDBF30BC58434" box="[655,759,1781,1807]" name="Malaysia" pageId="1" pageNumber="60">Sarawak</collectingCountry>
|
||
, Kuching, Santubong
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<date id="B377101CFFF2FF8922D9DBF30D4C8434" box="[1018,1150,1781,1807]" pageId="1" pageNumber="60" value="1990-03-26">26. 3. 1990</date>
|
||
A. RIEDEL".
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8FD36557FFF2FF8A21E5DA1B08208269" lastPageId="2" lastPageNumber="61" pageId="1" pageNumber="60" type="diagnosis">
|
||
<paragraph id="C77636DCFFF2FF8A21E5DA1B0D0883E1" blockId="1.[151,1437,1421,2007]" lastBlockId="2.[151,1436,152,538]" lastPageId="2" lastPageNumber="61" pageId="1" pageNumber="60">
|
||
<emphasis id="F5BDEACEFFF2FF8921E5DA1B08CF840C" bold="true" box="[198,509,1821,1847]" pageId="1" pageNumber="60">Diagnosis (adult strage):</emphasis>
|
||
<taxonomicName id="00C94D5FFFF2FF892325DA1B0A3D840D" box="[518,783,1821,1846]" class="Insecta" family="Dryopidae" genus="Geoparnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="species" species="rhinoceros">
|
||
<emphasis id="F5BDEACEFFF2FF892325DA1B0A3D840D" box="[518,783,1821,1846]" italics="true" pageId="1" pageNumber="60">Geoparnus rhinoceros</emphasis>
|
||
</taxonomicName>
|
||
is a medium sized species (body length
|
||
<quantity id="00319B39FFF2FF8925DFDA1B0CBD840C" box="[1276,1423,1821,1847]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" metricValueMax="2.5" metricValueMin="1.7" pageId="1" pageNumber="60" unit="mm" value="2.1" valueMax="2.5" valueMin="1.7">1.7–2.5 mm</quantity>
|
||
), unique among all known
|
||
<taxonomicName id="00C94D5FFFF2FF8920E3DA430B0E8464" box="[448,572,1861,1887]" class="Insecta" family="Dryopidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="family">Dryopidae</taxonomicName>
|
||
by the horn-like process on the head of the male. From
|
||
<taxonomicName id="00C94D5FFFF2FF8925EADA430CAE8465" box="[1225,1436,1861,1886]" class="Insecta" family="Dryopidae" genus="Geoparnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="species" species="setifer">
|
||
<emphasis id="F5BDEACEFFF2FF8925EADA430CAE8465" box="[1225,1436,1861,1886]" italics="true" pageId="1" pageNumber="60">Geoparnus setifer</emphasis>
|
||
</taxonomicName>
|
||
it differs furthermore in the following most obvious characters: (a) body form ovoid, strongly convex, moderately constricted between pronotum and elytra (
|
||
<taxonomicName id="00C94D5FFFF2FF8923F4DA930A718495" box="[727,835,1941,1966]" class="Insecta" family="Dryopidae" genus="Geoparnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="species" species="setifer">
|
||
<emphasis id="F5BDEACEFFF2FF8923F4DA930A718495" box="[727,835,1941,1966]" italics="true" pageId="1" pageNumber="60">G. setifer</emphasis>
|
||
</taxonomicName>
|
||
is elongate-ovate, less convex); (b) flat-bottomed punctures on head and pronotum distinctly larger than a facet, shallowly impressed (in
|
||
<taxonomicName id="00C94D5FFFF2FF892581DABB0C3E84ED" box="[1186,1292,1981,2006]" class="Insecta" family="Dryopidae" genus="Geoparnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="species" species="setifer">
|
||
<emphasis id="F5BDEACEFFF2FF892581DABB0C3E84ED" box="[1186,1292,1981,2006]" italics="true" pageId="1" pageNumber="60">G. setifer</emphasis>
|
||
</taxonomicName>
|
||
these punctures are subequal to a diameter of a facet and deeply impressed); (c) scutellum concealed by elytra (in
|
||
<taxonomicName id="00C94D5FFFF1FF8A2469DD9E098A83E2" class="Insecta" family="Dryopidae" genus="Geoparnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="setifer">
|
||
<emphasis id="F5BDEACEFFF1FF8A2469DD9E098A83E2" italics="true" pageId="2" pageNumber="61">G. setifer</emphasis>
|
||
</taxonomicName>
|
||
scutellum small but well visible in dorsal view); (d) shape of male genitalia.
|
||
</paragraph>
|
||
<paragraph id="C77636DCFFF1FF8A21E5DDEE08208269" blockId="2.[151,1436,152,538]" pageId="2" pageNumber="61">
|
||
Members of
|
||
<taxonomicName id="00C94D5FFFF1FF8A2041DDEE08D7823A" box="[354,485,232,257]" class="Insecta" family="Dryopidae" genus="Geoparnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F5BDEACEFFF1FF8A2041DDEE08D7823A" box="[354,485,232,257]" italics="true" pageId="2" pageNumber="61">Geoparnus</emphasis>
|
||
</taxonomicName>
|
||
are characterized by combination of the following characters: (a) conspicuous stiff setae with ridged surface and multifurcate apex; (b) short and compact body form and the antennal configuration.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8FD36557FFF1FF8F21E5DC590BE58771" lastPageId="7" lastPageNumber="66" pageId="2" pageNumber="61" type="description">
|
||
<paragraph id="C77636DCFFF1FF8A21E5DC590B8B8121" blockId="2.[151,1436,152,538]" pageId="2" pageNumber="61">
|
||
<emphasis id="F5BDEACEFFF1FF8A21E5DC5908538242" bold="true" box="[198,353,351,377]" pageId="2" pageNumber="61">Description:</emphasis>
|
||
Habitus of male as in
|
||
<figureCitation id="5FF22A59FFF1FF8A2350DC660B898241" box="[627,699,352,378]" captionStart="FIGURE 1" captionStartId="2.[151,255,2000,2024]" captionTargetBox="[215,1368,596,1969]" captionTargetId="figure@2.[196,1390,581,1998]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Geoparnus rhinoceros sp. nov., male." httpUri="https://zenodo.org/record/176881/files/figure.png" pageId="2" pageNumber="61">Fig. 1</figureCitation>
|
||
. Body form ovoid, moderately constricted between pronotum and elytra, convex dorsally, very compact, strongly sclerotised; in males 1.54–1.92 (1.77±0.08) and in females 1.50–1.87 (1.72±0.08) times as long as wide (BL/EW); body length (BL): ɗɗ
|
||
<quantity id="00319B39FFF1FF8A2509DCB60DE682F1" box="[1066,1236,432,458]" metricMagnitude="-3" metricUnit="m" metricValue="2.065" metricValueMax="2.4" metricValueMin="1.73" pageId="2" pageNumber="61" unit="mm" value="2.065" valueMax="2.4" valueMin="1.73">1.73–2.40 mm</quantity>
|
||
(2.04±0.13), ΨΨ
|
||
<quantity id="00319B39FFF1FF8A21B4DCDE087082C9" box="[151,322,472,498]" metricMagnitude="-3" metricUnit="m" metricValue="2.13" metricValueMax="2.5" metricValueMin="1.76" pageId="2" pageNumber="61" unit="mm" value="2.13" valueMax="2.5" valueMin="1.76">1.76–2.50 mm</quantity>
|
||
(2.08±0.13). Colour black, clypeus, antennae, mouthparts and legs paler yellowish or reddishbrown. Immature specimens yellowish-brown.
|
||
</paragraph>
|
||
<caption id="93B66654FFF1FF8A21B4DAD60A1884D3" box="[151,810,2000,2024]" httpUri="https://zenodo.org/record/176881/files/figure.png" pageId="2" pageNumber="61" targetBox="[215,1368,596,1969]" targetPageId="2">
|
||
<paragraph id="C77636DCFFF1FF8A21B4DAD60A1884D3" blockId="2.[151,810,2000,2024]" box="[151,810,2000,2024]" pageId="2" pageNumber="61">
|
||
<emphasis id="F5BDEACEFFF1FF8A21B4DAD6082B84D3" bold="true" box="[151,281,2000,2024]" pageId="2" pageNumber="61">FIGURE 1.</emphasis>
|
||
Habitus of
|
||
<taxonomicName id="00C94D5FFFF1FF8A20B5DAD70BB784D3" box="[406,645,2001,2024]" class="Insecta" family="Dryopidae" genus="Geoparnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="rhinoceros" status="sp. nov.">
|
||
<emphasis id="F5BDEACEFFF1FF8A20B5DAD70BB784D3" box="[406,645,2001,2024]" italics="true" pageId="2" pageNumber="61">Geoparnus rhinoceros</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EE8E57B5FFF1FF8A23AFDAD60BD084D3" box="[652,738,2000,2024]" pageId="2" pageNumber="61" rank="species">
|
||
<emphasis id="F5BDEACEFFF1FF8A23AFDAD60BD084D3" bold="true" box="[652,738,2000,2024]" pageId="2" pageNumber="61">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, male.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="93B66654FFF0FF8B21B4DA340C1584B4" httpUri="https://zenodo.org/record/176882/files/figure.png" pageId="3" pageNumber="62" targetBox="[151,1436,194,1817]" targetPageId="3">
|
||
<paragraph id="C77636DCFFF0FF8B21B4DA340C1584B4" blockId="3.[151,1436,1842,1935]" pageId="3" pageNumber="62">
|
||
<emphasis id="F5BDEACEFFF0FF8B21B4DA3408708471" bold="true" box="[151,322,1842,1866]" pageId="3" pageNumber="62">FIGURES 2–9.</emphasis>
|
||
<taxonomicName id="00C94D5FFFF0FF8B206BDA350B058471" box="[328,567,1843,1866]" class="Insecta" family="Dryopidae" genus="Geoparnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="rhinoceros" status="sp. nov.">
|
||
<emphasis id="F5BDEACEFFF0FF8B206BDA350B058471" box="[328,567,1843,1866]" italics="true" pageId="3" pageNumber="62">Geoparnus rhinoceros</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EE8E57B5FFF0FF8B231EDA340BA18471" box="[573,659,1842,1866]" pageId="3" pageNumber="62" rank="species">
|
||
<emphasis id="F5BDEACEFFF0FF8B231EDA340BA18471" bold="true" box="[573,659,1842,1866]" pageId="3" pageNumber="62">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, 2) male head, dorsal view; 3) male head, lateral view; 4) pronotum, dorsal view; 5) hypomeron and part of prosternum, ventral view; 6) prothorax, ventral view; 7) meso- and metathorax, ventral view; 8) detail of flat-bottomed punctures with stiff setae, ventral view; 9) detail of stiff seta, lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="93B66654FFF7FF8C21B4DB8D0DEC85FF" httpUri="https://zenodo.org/record/176883/files/figure.png" pageId="4" pageNumber="63" targetBox="[151,1436,194,1650]" targetPageId="4">
|
||
<paragraph id="C77636DCFFF7FF8C21B4DB8D0DEC85FF" blockId="4.[151,1435,1675,1732]" pageId="4" pageNumber="63">
|
||
<emphasis id="F5BDEACEFFF7FF8C21B4DB8D086F8598" bold="true" box="[151,349,1675,1699]" pageId="4" pageNumber="63">FIGURES 10–14.</emphasis>
|
||
<taxonomicName id="00C94D5FFFF7FF8C2040DB8D0B638599" box="[355,593,1675,1698]" class="Insecta" family="Dryopidae" genus="Geoparnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="rhinoceros" status="sp. nov.">
|
||
<emphasis id="F5BDEACEFFF7FF8C2040DB8D0B638599" box="[355,593,1675,1698]" italics="true" pageId="4" pageNumber="63">Geoparnus rhinoceros</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EE8E57B5FFF7FF8C2374DB8D0B9E8598" box="[599,684,1675,1699]" pageId="4" pageNumber="63" rank="species">
|
||
<emphasis id="F5BDEACEFFF7FF8C2374DB8D0B9E8598" bold="true" box="[599,684,1675,1699]" pageId="4" pageNumber="63">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
; 10) elytron, lateral view; 11) male protibia, lateral view; 12) abdomen in male, ventral view; 13) fifth ventrite in male, ventral view; 14) fifth ventrite in female, ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="C77636DCFFF7FF8D21E4DBFE0A5983E1" blockId="4.[151,1437,1784,2010]" lastBlockId="5.[151,1437,152,498]" lastPageId="5" lastPageNumber="64" pageId="4" pageNumber="63">
|
||
Vestiture of body consists of several
|
||
<typeStatus id="1872887EFFF7FF8C235BDBFE0B848429" box="[632,694,1784,1810]" pageId="4" pageNumber="63">types</typeStatus>
|
||
of distinctly different setae: (a) short and longer hair-like setae (
|
||
<figureCitation id="5FF22A59FFF7FF8C21BCDA2608218401" box="[159,275,1824,1850]" captionStart="FIGURES 2 – 9" captionStartId="3.[151,269,1842,1866]" captionTargetBox="[151,1436,194,1817]" captionTargetId="figure@3.[151,1436,194,1818]" captionTargetPageId="3" captionText="FIGURES 2 – 9. Geoparnus rhinoceros sp. nov., 2) male head, dorsal view; 3) male head, lateral view; 4) pronotum, dorsal view; 5) hypomeron and part of prosternum, ventral view; 6) prothorax, ventral view; 7) meso- and metathorax, ventral view; 8) detail of flat-bottomed punctures with stiff setae, ventral view; 9) detail of stiff seta, lateral view." httpUri="https://zenodo.org/record/176882/files/figure.png" pageId="4" pageNumber="63">Figs. 2, 4</figureCitation>
|
||
,
|
||
<figureCitation id="5FF22A59FFF7FF8C2002DA2608738401" box="[289,321,1824,1850]" captionStart="FIGURES 10 – 14" captionStartId="4.[151,269,1675,1699]" captionTargetBox="[151,1436,194,1650]" captionTargetId="figure@4.[151,1436,194,1651]" captionTargetPageId="4" captionText="FIGURES 10 – 14. Geoparnus rhinoceros sp. nov.; 10) elytron, lateral view; 11) male protibia, lateral view; 12) abdomen in male, ventral view; 13) fifth ventrite in male, ventral view; 14) fifth ventrite in female, ventral view." httpUri="https://zenodo.org/record/176883/files/figure.png" pageId="4" pageNumber="63">12</figureCitation>
|
||
), and (b) conspicuous stiff shorter and longer setae with ridged surface and multifurcate apex (
|
||
<figureCitation id="5FF22A59FFF7FF8C21BCDA4E08268459" box="[159,276,1864,1890]" captionStart="FIGURES 2 – 9" captionStartId="3.[151,269,1842,1866]" captionTargetBox="[151,1436,194,1817]" captionTargetId="figure@3.[151,1436,194,1818]" captionTargetPageId="3" captionText="FIGURES 2 – 9. Geoparnus rhinoceros sp. nov., 2) male head, dorsal view; 3) male head, lateral view; 4) pronotum, dorsal view; 5) hypomeron and part of prosternum, ventral view; 6) prothorax, ventral view; 7) meso- and metathorax, ventral view; 8) detail of flat-bottomed punctures with stiff setae, ventral view; 9) detail of stiff seta, lateral view." httpUri="https://zenodo.org/record/176882/files/figure.png" pageId="4" pageNumber="63">Figs. 8, 9</figureCitation>
|
||
); all setae yellowish. Short hair-like setae adpressed, confined mainly to clypeus, pronotum, epipleura and ventral surface of thorax and abdomen, on elytra very scarce; longer hair-like setae concentrated mainly on anterolateral portion of clypeus; all hair-like setae arising from small sockets. Conspicuous stiff setae erect, arising either from large, flat-bottomed punctures (
|
||
<figureCitation id="5FF22A59FFF7FF8C22A8DAC60D7084E1" box="[907,1090,1984,2010]" captionStart="FIGURES 2 – 9" captionStartId="3.[151,269,1842,1866]" captionTargetBox="[151,1436,194,1817]" captionTargetId="figure@3.[151,1436,194,1818]" captionTargetPageId="3" captionText="FIGURES 2 – 9. Geoparnus rhinoceros sp. nov., 2) male head, dorsal view; 3) male head, lateral view; 4) pronotum, dorsal view; 5) hypomeron and part of prosternum, ventral view; 6) prothorax, ventral view; 7) meso- and metathorax, ventral view; 8) detail of flat-bottomed punctures with stiff setae, ventral view; 9) detail of stiff seta, lateral view." httpUri="https://zenodo.org/record/176882/files/figure.png" pageId="4" pageNumber="63">Figs. 2, 4, 7, 8</figureCitation>
|
||
,
|
||
<figureCitation id="5FF22A59FFF7FF8C2571DAC60DA584E1" box="[1106,1175,1984,2010]" captionStart="FIGURES 10 – 14" captionStartId="4.[151,269,1675,1699]" captionTargetBox="[151,1436,194,1650]" captionTargetId="figure@4.[151,1436,194,1651]" captionTargetPageId="4" captionText="FIGURES 10 – 14. Geoparnus rhinoceros sp. nov.; 10) elytron, lateral view; 11) male protibia, lateral view; 12) abdomen in male, ventral view; 13) fifth ventrite in male, ventral view; 14) fifth ventrite in female, ventral view." httpUri="https://zenodo.org/record/176883/files/figure.png" pageId="4" pageNumber="63">12-14</figureCitation>
|
||
) on dorsal portion of head, pronotum, meso- and metaventrite and partly on ventrites, or from small sockets on elytra, legs, pedicell, central portion of ventrites and lateral parts of pronotum.
|
||
</paragraph>
|
||
<paragraph id="C77636DCFFF6FF8D21E4DDEE0B8B82C9" blockId="5.[151,1437,152,498]" pageId="5" pageNumber="64">
|
||
Head: Clypeus with fine punctures and fine reticulation; anterior margin produced dorsad forming a hornlike process in male (
|
||
<figureCitation id="5FF22A59FFF6FF8D20ACDC1608CF8211" box="[399,509,272,298]" captionStart="FIGURES 2 – 9" captionStartId="3.[151,269,1842,1866]" captionTargetBox="[151,1436,194,1817]" captionTargetId="figure@3.[151,1436,194,1818]" captionTargetPageId="3" captionText="FIGURES 2 – 9. Geoparnus rhinoceros sp. nov., 2) male head, dorsal view; 3) male head, lateral view; 4) pronotum, dorsal view; 5) hypomeron and part of prosternum, ventral view; 6) prothorax, ventral view; 7) meso- and metathorax, ventral view; 8) detail of flat-bottomed punctures with stiff setae, ventral view; 9) detail of stiff seta, lateral view." httpUri="https://zenodo.org/record/176882/files/figure.png" pageId="5" pageNumber="64">Figs. 2, 3</figureCitation>
|
||
), arcuate and unproduced in female; surface around antennal insertion raised in male, nearly flat in female; frons and vertex with flat-bottomed large punctures, punctures ca. 3–4 times as wide as a diameter of a facet, punctures dense but distinctly separated from each other, interstices with fine punctures and short setae. Compound eyes moderately protruding from head outline, small, with few setae. Antennal insertion deep, scape short, pedicell enlarged, dorsally flat; antennal club moderately longer than scape and pedicell combined, nine-segmented.
|
||
</paragraph>
|
||
<paragraph id="C77636DCFFF6FF8D21B4DA7008E484E8" pageId="5" pageNumber="64">
|
||
<table id="B5C9C47CFFF6007721B4DA700CAE84E8" box="[151,1436,1910,2003]" gridcols="3" gridrows="3" pageId="5" pageNumber="64">
|
||
<tr id="79F9349EFFF6007721B4DA700CAE84B5" box="[151,1436,1910,1934]" gridrow="0" pageId="5" pageNumber="64">
|
||
<th id="3A285DE2FFF6007721B4DA700BF784B5" box="[151,709,1910,1934]" gridcol="0" gridrow="0" pageId="5" pageNumber="64">
|
||
<emphasis id="F5BDEACEFFF6FF8D21B4DA70085284B5" bold="true" box="[151,352,1910,1934]" pageId="5" pageNumber="64">FIGURES 15–20.</emphasis>
|
||
<taxonomicName id="00C94D5FFFF6FF8D204ADA710B6984B5" box="[361,603,1911,1934]" class="Insecta" family="Dryopidae" genus="Geoparnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="64" phylum="Arthropoda" rank="species" species="rhinoceros" status="sp. nov.">
|
||
<emphasis id="F5BDEACEFFF6FF8D204ADA710B6984B5" box="[361,603,1911,1934]" italics="true" pageId="5" pageNumber="64">Geoparnus rhinoceros</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EE8E57B5FFF6FF8D2347DA700B8F84B5" box="[612,701,1910,1934]" pageId="5" pageNumber="64" rank="species">
|
||
<emphasis id="F5BDEACEFFF6FF8D2347DA700B8F84B5" bold="true" box="[612,701,1910,1934]" pageId="5" pageNumber="64">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
;
|
||
</th>
|
||
<th id="3A285DE2FFF6007723EEDA700C1A84B5" box="[717,1320,1910,1934]" gridcol="1" gridrow="0" pageId="5" pageNumber="64">15 – 16) Aedeagus, ventral and lateral view; 17) ninth</th>
|
||
<th id="3A285DE2FFF60077240CDA700CAE84B5" box="[1327,1436,1910,1934]" gridcol="2" gridrow="0" pageId="5" pageNumber="64">sternite in</th>
|
||
</tr>
|
||
<tr id="79F9349EFFF6007721B4DA9F0CAE848B" box="[151,1436,1945,1968]" gridrow="1" pageId="5" pageNumber="64">
|
||
<th id="3A285DE2FFF6007721B4DA9F0BF7848B" box="[151,709,1945,1968]" gridcol="0" gridrow="1" pageId="5" pageNumber="64">male, ventral view; 18) ovipositor, lateral view; 19)</th>
|
||
<td id="3A285DE2FFF6007723EEDA9F0C1A848B" box="[717,1320,1945,1968]" gridcol="1" gridrow="1" pageId="5" pageNumber="64">eighth sternite in male, ventral view; 20) eighth sternite</td>
|
||
<td id="3A285DE2FFF60077240CDA9F0CAE848B" box="[1327,1436,1945,1968]" gridcol="2" gridrow="1" pageId="5" pageNumber="64">in female,</td>
|
||
</tr>
|
||
<tr id="79F9349EFFF6007721B4DABA0CAE84E8" box="[151,1436,1980,2003]" gridrow="2" pageId="5" pageNumber="64" rowspan-1="1" rowspan-2="1">
|
||
<th id="3A285DE2FFF6007721B4DABA0BF784E8" box="[151,709,1980,2003]" gridcol="0" gridrow="2" pageId="5" pageNumber="64">ventral view. Scale = 0.1 mm.</th>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="C77636DCFFF5FF8E21B4DFC30BA48057" blockId="6.[151,1437,709,1994]" pageId="6" pageNumber="65">
|
||
<emphasis id="F5BDEACEFFF5FF8E21B4DFC3086C81E6" bold="true" box="[151,350,709,733]" pageId="6" pageNumber="65">FIGURES 21–22.</emphasis>
|
||
21) Box & Whisker plots showing the variability of body length of males at all localities; statistically significant differences between two morphological forms were detected. Plots show medians (solid horizontal line), means (dash lines), 25th and 75th percentiles (boxes), 10th and 90th percentiles (whisker extending from boxes), and 5th and 95th percentiles (outliers); 22) Box & Whisker plots showing the variability of body length/ elytral width ratio of males; no distinct morphological forms were detected.
|
||
</paragraph>
|
||
<paragraph id="C77636DCFFF5FF8E21E4DE7E0D1485E1" blockId="6.[151,1437,709,1994]" pageId="6" pageNumber="65">
|
||
Thorax: Pronotal disc strongly convex; PL: ɗɗ
|
||
<quantity id="00319B39FFF5FF8E2260DE7E0ACA80A9" box="[835,1016,888,914]" metricMagnitude="-4" metricUnit="m" metricValue="6.95" metricValueMax="7.9" metricValueMin="6.0" pageId="6" pageNumber="65" unit="mm" value="0.695" valueMax="0.79" valueMin="0.6">0.60–0.79 mm</quantity>
|
||
(0.70±0.05), ΨΨ
|
||
<quantity id="00319B39FFF5FF8E25C4DE7E0CAE80A9" box="[1255,1436,888,914]" metricMagnitude="-4" metricUnit="m" metricValue="6.3" metricValueMax="7.6" metricValueMin="5.0" pageId="6" pageNumber="65" unit="mm" value="0.63" valueMax="0.76" valueMin="0.5">0.50–0.76 mm</quantity>
|
||
(0.68±0.05); PW: ɗɗ
|
||
<quantity id="00319B39FFF5FF8E20BADEA60B718081" box="[409,579,928,954]" metricMagnitude="-3" metricUnit="m" metricValue="1.185" metricValueMax="1.4" metricValueMin="0.97" pageId="6" pageNumber="65" unit="mm" value="1.185" valueMax="1.4" valueMin="0.97">0.97–1.40 mm</quantity>
|
||
(1.13±0.08), ΨΨ
|
||
<quantity id="00319B39FFF5FF8E2232DEA60A898081" box="[785,955,928,954]" metricMagnitude="-3" metricUnit="m" metricValue="1.125" metricValueMax="1.3" metricValueMin="0.95" pageId="6" pageNumber="65" unit="mm" value="1.125" valueMax="1.3" valueMin="0.95">0.95–1.30 mm</quantity>
|
||
(1.15±0.07); lateral sides arcuate, crenulate; anterior margin of pronotum straight; anterior corners deflected, short and acute, moderately protruding; posterior margin bisinuate; posterior corners short; punctation similar to that on vertex but less dense. Hypomeron widest posteriorly, with scattered larger punctures (
|
||
<figureCitation id="5FF22A59FFF5FF8E22A9D91E0AE08709" box="[906,978,1048,1074]" captionStart="FIGURES 2 – 9" captionStartId="3.[151,269,1842,1866]" captionTargetBox="[151,1436,194,1817]" captionTargetId="figure@3.[151,1436,194,1818]" captionTargetPageId="3" captionText="FIGURES 2 – 9. Geoparnus rhinoceros sp. nov., 2) male head, dorsal view; 3) male head, lateral view; 4) pronotum, dorsal view; 5) hypomeron and part of prosternum, ventral view; 6) prothorax, ventral view; 7) meso- and metathorax, ventral view; 8) detail of flat-bottomed punctures with stiff setae, ventral view; 9) detail of stiff seta, lateral view." httpUri="https://zenodo.org/record/176882/files/figure.png" pageId="6" pageNumber="65">Fig. 5</figureCitation>
|
||
); prosternum in front of coxae as long as prosternal process, strongly punctate (
|
||
<figureCitation id="5FF22A59FFF5FF8E2350D9460B888761" box="[627,698,1088,1114]" captionStart="FIGURES 2 – 9" captionStartId="3.[151,269,1842,1866]" captionTargetBox="[151,1436,194,1817]" captionTargetId="figure@3.[151,1436,194,1818]" captionTargetPageId="3" captionText="FIGURES 2 – 9. Geoparnus rhinoceros sp. nov., 2) male head, dorsal view; 3) male head, lateral view; 4) pronotum, dorsal view; 5) hypomeron and part of prosternum, ventral view; 6) prothorax, ventral view; 7) meso- and metathorax, ventral view; 8) detail of flat-bottomed punctures with stiff setae, ventral view; 9) detail of stiff seta, lateral view." httpUri="https://zenodo.org/record/176882/files/figure.png" pageId="6" pageNumber="65">Fig. 6</figureCitation>
|
||
); prosternal process longer than wide, lateral sides raised, apex subacute, surface densely punctate. Meso- and metaventrite very short (
|
||
<figureCitation id="5FF22A59FFF5FF8E22C6D96E0D1C87B9" box="[997,1070,1128,1154]" captionStart="FIGURES 2 – 9" captionStartId="3.[151,269,1842,1866]" captionTargetBox="[151,1436,194,1817]" captionTargetId="figure@3.[151,1436,194,1818]" captionTargetPageId="3" captionText="FIGURES 2 – 9. Geoparnus rhinoceros sp. nov., 2) male head, dorsal view; 3) male head, lateral view; 4) pronotum, dorsal view; 5) hypomeron and part of prosternum, ventral view; 6) prothorax, ventral view; 7) meso- and metathorax, ventral view; 8) detail of flat-bottomed punctures with stiff setae, ventral view; 9) detail of stiff seta, lateral view." httpUri="https://zenodo.org/record/176882/files/figure.png" pageId="6" pageNumber="65">Fig. 7</figureCitation>
|
||
), together shorter than prosternum, mesal portion deeply impressed, lateral portions with large punctures, interstices with fine reticulation. Elytra convex, without humeri; strongly declivitous laterally and posteriorly; lateral sides crenulate and visible in dorsal view to posterior margin of mesocoxa; apices moderately produced, acute; each elytron with nine rows of coarse, large, deeply impressed punctures, punctures about as coarse as those on pronotum but less sharply delimited and not flat-bottomed, often somewhat subquadrate; interstices on central portion usually less than half as wide as puncture diameter, laterally and posteriorly becoming narrower; intervals distinctly narrower than punctures in rows, feebly raised, with rows of stiff erect setae; anterior margin arcuate; scutellum concealed by elytra, inserted into cavity in sutural edge of each elytron. Epipleura anteriorly as wide as maximum width of profemur, narrowed posteriad, effaced near fifth ventrite, inflected at level of metacoxa, surface finely punctate. EL: ɗɗ
|
||
<quantity id="00319B39FFF5FF8E233FD8FE0BFA8529" box="[540,712,1528,1554]" metricMagnitude="-3" metricUnit="m" metricValue="1.36" metricValueMax="1.6" metricValueMin="1.12" pageId="6" pageNumber="65" unit="mm" value="1.36" valueMax="1.6" valueMin="1.12">1.12–1.60 mm</quantity>
|
||
(1.33±0.11), ΨΨ
|
||
<quantity id="00319B39FFF5FF8E22BFD8FE0D7A8529" box="[924,1096,1528,1554]" metricMagnitude="-3" metricUnit="m" metricValue="1.4249999999999998" metricValueMax="1.7" metricValueMin="1.15" pageId="6" pageNumber="65" unit="mm" value="1.4249999999999998" valueMax="1.7" valueMin="1.15">1.15–1.70 mm</quantity>
|
||
(1.39±0.11); EW: ɗɗ
|
||
<quantity id="00319B39FFF5FF8E2479D8FE08338501" metricMagnitude="-3" metricUnit="m" metricValue="1.125" metricValueMax="1.3" metricValueMin="0.95" pageId="6" pageNumber="65" unit="mm" value="1.125" valueMax="1.3" valueMin="0.95">0.95– 1.30 mm</quantity>
|
||
(1.16±0.09), ΨΨ
|
||
<quantity id="00319B39FFF5FF8E20F7DB260BB28501" box="[468,640,1568,1594]" metricMagnitude="-3" metricUnit="m" metricValue="1.225" metricValueMax="1.4" metricValueMin="1.05" pageId="6" pageNumber="65" unit="mm" value="1.225" valueMax="1.4" valueMin="1.05">1.05–1.40 mm</quantity>
|
||
(1.21±0.09).
|
||
<collectingCountry id="BFDE764CFFF5FF8E2200DB260A6C8501" box="[803,862,1568,1594]" name="India" pageId="6" pageNumber="65">Hind</collectingCountry>
|
||
wings fully absent. Foreleg longest, moderately shorter than body; middle and hind legs slightly shorter; protibia feebly longer than profemur, straight or slightly bent and moderately thickened in both sexes (
|
||
<figureCitation id="5FF22A59FFF5FF8E2214DB760AA385B1" box="[823,913,1648,1674]" captionStart="FIGURES 10 – 14" captionStartId="4.[151,269,1675,1699]" captionTargetBox="[151,1436,194,1650]" captionTargetId="figure@4.[151,1436,194,1651]" captionTargetPageId="4" captionText="FIGURES 10 – 14. Geoparnus rhinoceros sp. nov.; 10) elytron, lateral view; 11) male protibia, lateral view; 12) abdomen in male, ventral view; 13) fifth ventrite in male, ventral view; 14) fifth ventrite in female, ventral view." httpUri="https://zenodo.org/record/176883/files/figure.png" pageId="6" pageNumber="65">Fig. 11</figureCitation>
|
||
); meso- and metatibia similar in form but shorter, with a small ventrodistal tooth in male; combined length of protarsomeres and claws moderately exceeds half length of protibia; claws of foreleg in male wider than in female.
|
||
</paragraph>
|
||
<paragraph id="C77636DCFFF5FF8E21E4DBEE0A7384F1" blockId="6.[151,1437,709,1994]" pageId="6" pageNumber="65">
|
||
Abdominal intercoxal process (
|
||
<figureCitation id="5FF22A59FFF5FF8E2318DBEE0BA08439" box="[571,658,1768,1794]" captionStart="FIGURES 10 – 14" captionStartId="4.[151,269,1675,1699]" captionTargetBox="[151,1436,194,1650]" captionTargetId="figure@4.[151,1436,194,1651]" captionTargetPageId="4" captionText="FIGURES 10 – 14. Geoparnus rhinoceros sp. nov.; 10) elytron, lateral view; 11) male protibia, lateral view; 12) abdomen in male, ventral view; 13) fifth ventrite in male, ventral view; 14) fifth ventrite in female, ventral view." httpUri="https://zenodo.org/record/176883/files/figure.png" pageId="6" pageNumber="65">Fig. 12</figureCitation>
|
||
) moderately wider than prosternal process, apex subtruncate, lateral sides raised; first ventrite in middle slightly longer than second and third ventrite together; fourth ventrite about half as long as first one and nearly third as long as fifth ventrite; fifth ventrite more rounded and sides less declivous in male than in female (
|
||
<figureCitation id="5FF22A59FFF5FF8E2344DA660BCB8441" box="[615,761,1888,1914]" captionStart="FIGURES 10 – 14" captionStartId="4.[151,269,1675,1699]" captionTargetBox="[151,1436,194,1650]" captionTargetId="figure@4.[151,1436,194,1651]" captionTargetPageId="4" captionText="FIGURES 10 – 14. Geoparnus rhinoceros sp. nov.; 10) elytron, lateral view; 11) male protibia, lateral view; 12) abdomen in male, ventral view; 13) fifth ventrite in male, ventral view; 14) fifth ventrite in female, ventral view." httpUri="https://zenodo.org/record/176883/files/figure.png" pageId="6" pageNumber="65">Figs. 13, 14</figureCitation>
|
||
); surface of ventrites with large flat-bottomed punctures and micropunctures, setose. Eighth sternite with short median process in male (Fig. 19) and long in female (Fig. 20); ninth sternite narrow and long in male (Fig. 17).
|
||
</paragraph>
|
||
<caption id="93B66654FFF4FF8F21B4DF660AC3814C" box="[151,1009,608,632]" httpUri="https://zenodo.org/record/176884/files/figure.png" pageId="7" pageNumber="66" targetBox="[505,1084,190,586]" targetPageId="7">
|
||
<paragraph id="C77636DCFFF4FF8F21B4DF660AC3814C" blockId="7.[151,1009,608,632]" box="[151,1009,608,632]" pageId="7" pageNumber="66">
|
||
<emphasis id="F5BDEACEFFF4FF8F21B4DF6608148143" bold="true" box="[151,294,608,632]" pageId="7" pageNumber="66">FIGURE 23.</emphasis>
|
||
Known distribution of
|
||
<taxonomicName id="00C94D5FFFF4FF8F2302DF670A228143" box="[545,784,609,632]" class="Insecta" family="Dryopidae" genus="Geoparnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="rhinoceros" status="sp. nov.">
|
||
<emphasis id="F5BDEACEFFF4FF8F2302DF670A228143" box="[545,784,609,632]" italics="true" pageId="7" pageNumber="66">Geoparnus rhinoceros</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EE8E57B5FFF4FF8F2235DF660A5E8143" box="[790,876,608,632]" pageId="7" pageNumber="66" rank="species">
|
||
<emphasis id="F5BDEACEFFF4FF8F2235DF660A5E8143" bold="true" box="[790,876,608,632]" pageId="7" pageNumber="66">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
in Sarawak.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="C77636DCFFF4FF8F21E4DFA60C7580B9" blockId="7.[151,1437,672,1259]" pageId="7" pageNumber="66">Aedeagus (Figs. 15, 16): phallobasis robust, subcylindrical, moderately curved, 1.5–1.8 times as long as parameres, basally wider than apically (lateral view). Parameres moderately long, widest basally, gradually curved ventrad and gradually narrowed toward apices; apices subtruncate in lateral view. Penis distinctly shorter than parameres, widest basally, gradually narrowed apically; apex nearly acute (ventral view). Membranous ventral sac with fine longitudinal furrows; fibula absent. Ovipositor (Fig. 18) distinctly longer than abdomen; valvifer about 1.5–1.7 times as long as coxite; bursa copulatrix without sclerotised spinules.</paragraph>
|
||
<paragraph id="C77636DCFFF4FF8F21E4DE960A4D80E9" blockId="7.[151,1437,672,1259]" pageId="7" pageNumber="66">
|
||
<emphasis id="F5BDEACEFFF4FF8F21E4DE9608F68091" bold="true" box="[199,452,912,938]" pageId="7" pageNumber="66">Sexual dimorphism.</emphasis>
|
||
Males with horn-like process on clypeus and tibial teeth, on average shorter, narrower and less convex than females (see BL, EW in
|
||
<tableCitation id="8A4B0367FFF4FF8F23DEDEBE0A6380E9" box="[765,849,952,978]" captionStart="TABLE 1" captionStartId="7.[151,239,1312,1336]" captionText="TABLE 1. Variability of selected metric characters in males of Geoparnus rhinoceros with results of one way analysis of variance (BL—body length, EL—elytral length along suture, EW—maximum elytral width, PL—pronotal length along middle, PW—maximum pronotal width; first rows show ranges of measured values for specimens for different localities, in second rows mean and standard deviation for the respective measurements, n—number of measured specimens, F— Fisher`s value, H—the ANOVA on Ranks test statistics, P—probability)." pageId="7" pageNumber="66">Tabs. 1</tableCitation>
|
||
,
|
||
<tableCitation id="8A4B0367FFF4FF8F2243DEBE0A4380E9" box="[864,881,952,978]" captionStart="TABLE 2" captionStartId="8.[151,239,906,930]" captionText="TABLE 2. Variability of selected metric characters in females of Geoparnus rhinoceros with results of one way analysis of variance (BL – body length, EL – elytral length along suture, EW – maximum elytral width, PL – pronotal length along middle, PW – maximum pronotal width; first rows show ranges of measured values for specimens for different localities, in second rows mean and standard deviation for the respective measurements, n – number of measured specimens, F – Fisher`s value, H – the ANOVA on Ranks test statistics, P – probability)." pageId="7" pageNumber="66">2</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C77636DCFFF4FF8F21E4DEE60BE58771" blockId="7.[151,1437,672,1259]" pageId="7" pageNumber="66">
|
||
<emphasis id="F5BDEACEFFF4FF8F21E4DEE6081C80C1" bold="true" box="[199,302,992,1018]" pageId="7" pageNumber="66">Habitat.</emphasis>
|
||
Specimens were collected by sifting ground debris containing leaf litter, decaying twigs, remnants of epiphytes, bark and other plant material accumulated mainly around large, living or dead trees. Most specimens were found in primary lowland forest.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8FD36557FFF4FF8F21E4D95E0CAE8749" box="[199,1436,1112,1138]" pageId="7" pageNumber="66" type="distribution">
|
||
<paragraph id="C77636DCFFF4FF8F21E4D95E0CAE8749" blockId="7.[151,1437,672,1259]" box="[199,1436,1112,1138]" pageId="7" pageNumber="66">
|
||
<emphasis id="F5BDEACEFFF4FF8F21E4D95E08558749" bold="true" box="[199,359,1112,1138]" pageId="7" pageNumber="66">Distribution.</emphasis>
|
||
The species is known from several localities in central and southwestern
|
||
<collectingCountry id="BFDE764CFFF4FF8F25E1D95E0C158749" box="[1218,1319,1112,1138]" name="Malaysia" pageId="7" pageNumber="66">Sarawak</collectingCountry>
|
||
(
|
||
<figureCitation id="5FF22A59FFF4FF8F2414D95E0CBD8749" box="[1335,1423,1112,1138]" captionStart="FIGURE 23" captionStartId="7.[151,255,608,632]" captionTargetBox="[505,1084,190,586]" captionTargetId="figure@7.[505,1085,190,586]" captionTargetPageId="7" captionText="FIGURE 23. Known distribution of Geoparnus rhinoceros sp. nov. in Sarawak." httpUri="https://zenodo.org/record/176884/files/figure.png" pageId="7" pageNumber="66">Fig. 23</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8FD36557FFF4FF8121E4D9860A9F8241" lastPageId="9" lastPageNumber="68" pageId="7" pageNumber="66" type="etymology">
|
||
<paragraph id="C77636DCFFF4FF8F21E4D9860D6487A0" blockId="7.[151,1437,672,1259]" box="[199,1110,1152,1179]" pageId="7" pageNumber="66">
|
||
<emphasis id="F5BDEACEFFF4FF8F21E4D986086687A1" bold="true" box="[199,340,1152,1178]" pageId="7" pageNumber="66">Etymology.</emphasis>
|
||
Named in reference to the horn-like process on the male clypeus.
|
||
</paragraph>
|
||
<paragraph id="C77636DCFFF4FF8F21E4D9AE0C1C87D0" blockId="7.[151,1437,672,1259]" pageId="7" pageNumber="66">
|
||
<emphasis id="F5BDEACEFFF4FF8F21E4D9AE086187F9" bold="true" box="[199,339,1192,1218]" pageId="7" pageNumber="66">Variability.</emphasis>
|
||
Analysis of variance showed significant differences among populations within all measured characters (
|
||
<tableCitation id="8A4B0367FFF4FF8F203ED9D7084187D0" box="[285,371,1233,1259]" captionStart="TABLE 1" captionStartId="7.[151,239,1312,1336]" captionText="TABLE 1. Variability of selected metric characters in males of Geoparnus rhinoceros with results of one way analysis of variance (BL—body length, EL—elytral length along suture, EW—maximum elytral width, PL—pronotal length along middle, PW—maximum pronotal width; first rows show ranges of measured values for specimens for different localities, in second rows mean and standard deviation for the respective measurements, n—number of measured specimens, F— Fisher`s value, H—the ANOVA on Ranks test statistics, P—probability)." pageId="7" pageNumber="66">Tabs. 1</tableCitation>
|
||
,
|
||
<tableCitation id="8A4B0367FFF4FF8F20A2D9D708A087D0" box="[385,402,1233,1259]" captionStart="TABLE 2" captionStartId="8.[151,239,906,930]" captionText="TABLE 2. Variability of selected metric characters in females of Geoparnus rhinoceros with results of one way analysis of variance (BL – body length, EL – elytral length along suture, EW – maximum elytral width, PL – pronotal length along middle, PW – maximum pronotal width; first rows show ranges of measured values for specimens for different localities, in second rows mean and standard deviation for the respective measurements, n – number of measured specimens, F – Fisher`s value, H – the ANOVA on Ranks test statistics, P – probability)." pageId="7" pageNumber="66">2</tableCitation>
|
||
). The multiple comparison procedures revealed different results regarding sex.
|
||
</paragraph>
|
||
<caption id="93B66654FFF4FF8F21B4D8260A9386F9" pageId="7" pageNumber="66">
|
||
<paragraph id="C77636DCFFF4FF8F21B4D8260A9386F9" blockId="7.[151,1436,1312,1474]" pageId="7" pageNumber="66">
|
||
<emphasis id="F5BDEACEFFF4FF8F21B4D826083A8603" bold="true" box="[151,264,1312,1336]" pageId="7" pageNumber="66">TABLE 1.</emphasis>
|
||
Variability of selected metric characters in males of
|
||
<taxonomicName id="00C94D5FFFF4FF8F2214D8270D148603" box="[823,1062,1313,1336]" class="Insecta" family="Dryopidae" genus="Geoparnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="rhinoceros">
|
||
<emphasis id="F5BDEACEFFF4FF8F2214D8270D148603" box="[823,1062,1313,1336]" italics="true" pageId="7" pageNumber="66">Geoparnus rhinoceros</emphasis>
|
||
</taxonomicName>
|
||
with results of one way analysis of variance (BL—body length, EL—elytral length along suture, EW—maximum elytral width, PL—pronotal length along middle, PW—maximum pronotal width; first rows show ranges of measured values for specimens for different localities, in second rows mean and standard deviation for the respective measurements, n—number of measured specimens, F— Fisher`s value, H—the ANOVA on Ranks test statistics, P—probability).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="C77636DCFFF4FF8F21BFD8E50C7986C3" blockId="7.[156,1418,1507,2008]" box="[156,1355,1507,1528]" pageId="7" pageNumber="66">Locality BL [mm] EL [mm] EW [mm] BL/EW PL [mm] PW [mm] PL/PW</paragraph>
|
||
<paragraph id="C77636DCFFF4FF8F21BFDB0D0C56851B" blockId="7.[156,1418,1507,2008]" box="[156,1380,1547,1568]" pageId="7" pageNumber="66">Belaga 1.74–2.00 1.12–1.30 0.96–1.10 1.72–1.91 0.61–0.66 0.97–1.11 0.61–0.65</paragraph>
|
||
<paragraph id="C77636DCFFF4FF8F21BFDB350CB88573" blockId="7.[156,1418,1507,2008]" box="[156,1418,1587,1608]" pageId="7" pageNumber="66">n = 9 1.852 ± 0.104 1.187 ± 0.077 1.034 ± 0.044 1.791 ± 0.061 0.659 ± 0.036 1.049 ± 0.043 0.628 ± 0.013</paragraph>
|
||
<paragraph id="C77636DCFFF4FF8F21BFDB5D0C57854B" blockId="7.[156,1418,1507,2008]" box="[156,1381,1627,1648]" pageId="7" pageNumber="66">
|
||
<quantity id="00319B39FFF4FF8F21BFDB5D09EA854B" box="[156,216,1627,1648]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="66" unit="km" value="25.0">25 km</quantity>
|
||
E Kapit 1.76–2.07 1.12–1.30 1.00–1.14 1.73–1.90 0.64–0.76 0.97–1.12 0.62–0.72
|
||
</paragraph>
|
||
<paragraph id="C77636DCFFF4FF8F21BFDB850CB885A3" blockId="7.[156,1418,1507,2008]" box="[156,1418,1667,1688]" pageId="7" pageNumber="66">n = 20 1.926 ± 0.075 1.236 ± 0.052 1.075 ± 0.041 1.792 ± 0.053 0.690 ± 0.029 1.051 ± 0.038 0.657 ± 0.023</paragraph>
|
||
<paragraph id="C77636DCFFF4FF8F21BFDBAD0C5685FB" blockId="7.[156,1418,1507,2008]" box="[156,1380,1707,1728]" pageId="7" pageNumber="66">Gunung Gading 1.95–2.40 1.30–1.60 1.10–1.30 1.54–1.92 0.60–0.75 1.05–1.40 0.48–0.64</paragraph>
|
||
<paragraph id="C77636DCFFF4FF8F21BFDBD50CB885D3" blockId="7.[156,1418,1507,2008]" box="[156,1418,1747,1768]" pageId="7" pageNumber="66">n = 32 2.131 ± 0.105 1.445 ± 0.072 1.231 ± 0.061 1.733 ± 0.087 0.670 ± 0.045 1.180 ± 0.074 0.569 ± 0.041</paragraph>
|
||
<paragraph id="C77636DCFFF4FF8F21BFDBFD0C57842B" blockId="7.[156,1418,1507,2008]" box="[156,1381,1787,1808]" pageId="7" pageNumber="66">Santubong 1.89–2.27 1.19–1.48 1.06–1.29 1.56–1.88 0.66–0.79 1.06–1.25 0.58–0.69</paragraph>
|
||
<paragraph id="C77636DCFFF4FF8F21BFDA250CB88403" blockId="7.[156,1418,1507,2008]" box="[156,1418,1827,1848]" pageId="7" pageNumber="66">n = 31 2.072 ± 0.096 1.328 ± 0.069 1.168 ± 0.060 1.777 ± 0.087 0.744 ± 0.037 1.150 ± 0.054 0.648 ± 0.030</paragraph>
|
||
<paragraph id="C77636DCFFF4FF8F21BFDA4D0C56845B" blockId="7.[156,1418,1507,2008]" box="[156,1380,1867,1888]" pageId="7" pageNumber="66">Gunung Serapi 1.99–2.19 1.28–1.43 1.11–1.17 1.80–1.88 0.71–0.76 1.08–1.17 0.66–0.67</paragraph>
|
||
<paragraph id="C77636DCFFF4FF8F21BFDA750C57848B" blockId="7.[156,1418,1507,2008]" pageId="7" pageNumber="66">n = 3 2.092 ± 0.102 1.344 ± 0.078 1.136 ± 0.030 1.840 ± 0.041 0.748 ± 0.029 1.131 ± 0.049 0.662 ± 0.005 F = 24.04 H = 61.17 H = 59.20 H = 13.11 H = 45.13 H = 46.79 H = 58.58</paragraph>
|
||
<paragraph id="C77636DCFFF4FF802071DAC50BF98061" blockId="7.[156,1418,1507,2008]" lastBlockId="8.[151,1437,152,858]" lastPageId="8" lastPageNumber="67" pageId="7" pageNumber="66">
|
||
P <0.001 P <0.001 P <0.001 P = 0.011 P <0.001 P <0.001 P <0.001 In males pairwise multiple comparison tests of body length (BL), elytral length (EL) and maximum elytral width (EW) showed two morphological forms: (1) smaller and narrower males from central
|
||
<collectingCountry id="BFDE764CFFFBFF80241BDDC60CAF83E1" box="[1336,1437,192,218]" name="Malaysia" pageId="8" pageNumber="67">Sarawak</collectingCountry>
|
||
(Belaga, Kapit env.) and (2) larger and wider males from southwestern
|
||
<collectingCountry id="BFDE764CFFFBFF8022FDDDEE0D718239" box="[990,1091,232,258]" name="Malaysia" pageId="8" pageNumber="67">Sarawak</collectingCountry>
|
||
near Kuching (Gunung Gading, Santubong, Gunung Serapi). These groups are also evident from Box & Whisker plots of the data variability (BL—Fig. 21; plots of EL and EW not shown). Pairwise multiple analyses of maximum pronotal width (PW) and length (PL) suggest the same groups, with some exceptions: the conflicting comparisons always included males from Gunung Serapi, which is likely caused by insufficient data from this locality (n=3). Subsequent comparisons of the pooled data of both groups (using t-test) confirmed these two distinct morphological forms—significant differences were found within all tested metric characters (BL, EL, EW, PW with P<0.001, PL with P=0.008). Pairwise multiple analysis of BL/EW ratio showed no distinct groups (see also plot of the data variability Fig. 22). However the results of ANOVA (
|
||
<tableCitation id="8A4B0367FFFBFF8022E4DF2E0D228179" box="[967,1040,552,578]" captionStart="TABLE 1" captionStartId="7.[151,239,1312,1336]" captionText="TABLE 1. Variability of selected metric characters in males of Geoparnus rhinoceros with results of one way analysis of variance (BL—body length, EL—elytral length along suture, EW—maximum elytral width, PL—pronotal length along middle, PW—maximum pronotal width; first rows show ranges of measured values for specimens for different localities, in second rows mean and standard deviation for the respective measurements, n—number of measured specimens, F— Fisher`s value, H—the ANOVA on Ranks test statistics, P—probability)." pageId="8" pageNumber="67">Tab. 1</tableCitation>
|
||
) presented significant difference, this value of significance is the lowest within all characters tested (P=0.011). There was only one out of all ten pairwise comparisons of this character (population from Gunung Gading vs. Gunung Serapi) with significant difference. This suggests that ratio BL/EW vary rather slightly in males, and hence can be used as distinguishing character, but more data is needed to support this hypothesis. Concerning size of the clypeal horns, no disproportional development was observed as is known in many males of Scarabaeoidea (e.g. widely distributed palaearctic
|
||
<taxonomicName id="00C94D5FFFFBFF80203DDE1E0B468009" authority="Linnaeus" authorityName="Linnaeus" box="[286,628,792,818]" class="Insecta" family="Dynastidae" genus="Oryctes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="67" phylum="Arthropoda" rank="species" species="nasicornis">
|
||
<emphasis id="F5BDEACEFFFBFF80203DDE1E08CE800A" box="[286,508,792,817]" italics="true" pageId="8" pageNumber="67">Oryctes nasicornis</emphasis>
|
||
Linnaeus
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="00C94D5FFFFBFF8023A3DE1E0A3A8009" box="[640,776,792,818]" class="Insecta" family="Scarabaeidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="67" phylum="Arthropoda" rank="family">Dynastinae</taxonomicName>
|
||
), i.e. in the largest males the size of the horn is proportionally similar to the size in the smallest males.
|
||
</paragraph>
|
||
<caption id="93B66654FFFBFF8021B4DE8C0D228717" pageId="8" pageNumber="67">
|
||
<paragraph id="C77636DCFFFBFF8021B4DE8C0D228717" blockId="8.[151,1436,906,1068]" pageId="8" pageNumber="67">
|
||
<emphasis id="F5BDEACEFFFBFF8021B4DE8C083A8099" bold="true" box="[151,264,906,930]" pageId="8" pageNumber="67">TABLE 2.</emphasis>
|
||
Variability of selected metric characters in females of
|
||
<taxonomicName id="00C94D5FFFFBFF802272DE8C0D72809A" box="[849,1088,906,929]" class="Insecta" family="Dryopidae" genus="Geoparnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="67" phylum="Arthropoda" rank="species" species="rhinoceros">
|
||
<emphasis id="F5BDEACEFFFBFF802272DE8C0D72809A" box="[849,1088,906,929]" italics="true" pageId="8" pageNumber="67">Geoparnus rhinoceros</emphasis>
|
||
</taxonomicName>
|
||
with results of one way analysis of variance (BL – body length, EL – elytral length along suture, EW – maximum elytral width, PL – pronotal length along middle, PW – maximum pronotal width; first rows show ranges of measured values for specimens for different localities, in second rows mean and standard deviation for the respective measurements, n – number of measured specimens, F – Fisher`s value, H – the ANOVA on Ranks test statistics, P – probability).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="C77636DCFFFBFF8021BCD94B0C608759" blockId="8.[159,1425,1101,1602]" box="[159,1362,1101,1122]" pageId="8" pageNumber="67">Locality BL [mm] EL [mm] EW [mm] BL/EW PL [mm] PW [mm] PL/PW</paragraph>
|
||
<paragraph id="C77636DCFFFBFF8021BCD9730C5987B1" blockId="8.[159,1425,1101,1602]" box="[159,1387,1141,1162]" pageId="8" pageNumber="67">Belaga 1.91–2.04 1.25–1.33 1.09–1.18 1.68–1.85 0.66–0.71 0.98–1.15 0.62–0.70</paragraph>
|
||
<paragraph id="C77636DCFFFBFF8021BCD99B0CA38789" blockId="8.[159,1425,1101,1602]" box="[159,1425,1181,1202]" pageId="8" pageNumber="67">n = 6 1.973 ± 0.045 1.284 ± 0.026 1.119 ± 0.032 1.764 ± 0.052 0.689 ± 0.023 1.056 ± 0.054 0.654 ± 0.032</paragraph>
|
||
<paragraph id="C77636DCFFFBFF8021BCD9C30C5987E1" blockId="8.[159,1425,1101,1602]" box="[159,1387,1221,1242]" pageId="8" pageNumber="67">
|
||
<quantity id="00319B39FFFBFF8021BCD9C309EE87E1" box="[159,220,1221,1242]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="67" unit="km" value="25.0">25 km</quantity>
|
||
E Kapit 1.76–2.30 1.15–1.60 1.04–1.30 1.66–1.83 0.55–0.74 0.95–1.15 0.55–0.69
|
||
</paragraph>
|
||
<paragraph id="C77636DCFFFBFF8021BCD9EB0CB88639" blockId="8.[159,1425,1101,1602]" box="[159,1418,1261,1282]" pageId="8" pageNumber="67">n = 22 1.996 ± 0.125 1.343 ± 0.106 1.147 ± 0.065 1.740 ± 0.049 0.640 ± 0.054 1.058 ± 0.054 0.605 ±0.045</paragraph>
|
||
<paragraph id="C77636DCFFFBFF8021BCD8130C5E8611" blockId="8.[159,1425,1101,1602]" box="[159,1388,1301,1322]" pageId="8" pageNumber="67">Gunung Gading 1.95–2.50 1.35–1.70 1.20–1.40 1.50–1.79 0.50–0.75 1.15–1.30 0.44–0.61</paragraph>
|
||
<paragraph id="C77636DCFFFBFF8021BCD83B0CA28669" blockId="8.[159,1425,1101,1602]" box="[159,1424,1341,1362]" pageId="8" pageNumber="67">n = 30 2.167 ± 0.111 1.487 ± 0.079 1.308 ± 0.053 1.657 ± 0.079 0.666 ± 0.046 1.190 ± 0.038 0.560 ± 0.033</paragraph>
|
||
<paragraph id="C77636DCFFFBFF8021BCD8630C598641" blockId="8.[159,1425,1101,1602]" box="[159,1387,1381,1402]" pageId="8" pageNumber="67">Santubong 1.81–2.27 1.17–1.50 1.09–1.27 1.59–1.87 0.64–0.76 1.00–1.20 0.60–0.70</paragraph>
|
||
<paragraph id="C77636DCFFFBFF8021BCD88B0CA38699" blockId="8.[159,1425,1101,1602]" box="[159,1425,1421,1442]" pageId="8" pageNumber="67">n = 30 2.117 ± 0.109 1.386 ± 0.079 1.192 ± 0.047 1.776 ± 0.057 0.731 ± 0.034 1.107 ± 0.046 0.661 ± 0.021</paragraph>
|
||
<paragraph id="C77636DCFFFBFF8021BCD8B30C5986F1" blockId="8.[159,1425,1101,1602]" box="[159,1387,1461,1482]" pageId="8" pageNumber="67">Gunung Serapi 1.79–2.09 1.15–1.35 1.06–1.24 1.66–1.78 0.64–0.74 0.98–1.12 0.64–0.66</paragraph>
|
||
<paragraph id="C77636DCFFFBFF8021BCD8DB0C5E8521" blockId="8.[159,1425,1101,1602]" pageId="8" pageNumber="67">n = 6 1.943 ± 0.110 1.259 ± 0.075 1.141 ± 0.069 1.703 ± 0.040 0.685 ± 0.038 1.056 ± 0.056 0.648 ± 0.008 F = 12.12 F = 17.65 H = 61.65 F = 15.06 H = 41.05 H = 58.73 H = 24.88</paragraph>
|
||
<paragraph id="C77636DCFFFBFF802053DB2B0DEB84A7" blockId="8.[159,1425,1101,1602]" lastBlockId="8.[151,1436,1682,2028]" pageId="8" pageNumber="67">
|
||
P <0.001 P <0.001 P <0.001 P <0.001 P <0.001 P <0.001 P <0.001 Analyses of variance of female metric characters revealed different results. In spite of that ANOVA showed significant differences in all tested characters (
|
||
<tableCitation id="8A4B0367FFFBFF802234DBBC0A5185EF" box="[791,867,1722,1748]" captionStart="TABLE 2" captionStartId="8.[151,239,906,930]" captionText="TABLE 2. Variability of selected metric characters in females of Geoparnus rhinoceros with results of one way analysis of variance (BL – body length, EL – elytral length along suture, EW – maximum elytral width, PL – pronotal length along middle, PW – maximum pronotal width; first rows show ranges of measured values for specimens for different localities, in second rows mean and standard deviation for the respective measurements, n – number of measured specimens, F – Fisher`s value, H – the ANOVA on Ranks test statistics, P – probability)." pageId="8" pageNumber="67">Tab. 2</tableCitation>
|
||
), pairwise multiple comparisons and subsequent t-test did not propose the same groups (central
|
||
<collectingCountry id="BFDE764CFFFBFF80239FDBE40A1685C7" box="[700,804,1762,1788]" name="Malaysia" pageId="8" pageNumber="67">Sarawak</collectingCountry>
|
||
, soutwestern
|
||
<collectingCountry id="BFDE764CFFFBFF8022E0DBE40D1885C7" box="[963,1066,1762,1788]" name="Malaysia" pageId="8" pageNumber="67">Sarawak</collectingCountry>
|
||
) as analysis of male characters. These results suggest that females are more variable and the metric characters or proportions cannot be used to distinguish species. Higher intraspecific female variability was also supported by the pairwise test of the ratio BL/EW, where significant differences among populations were observed. Besides body proportions measurements of all tibiae were also analysed, however no well-defined groups were recognized.
|
||
</paragraph>
|
||
<paragraph id="C77636DCFFFBFF8121E5DAAC08158211" blockId="8.[151,1436,1682,2028]" lastBlockId="9.[151,1436,152,378]" lastPageId="9" lastPageNumber="68" pageId="8" pageNumber="67">
|
||
Regarding the results of the analysis of metric characters in males, it is possible that specimens from southwestern and central
|
||
<collectingCountry id="BFDE764CFFFBFF8020E3DAD40B1784D7" box="[448,549,2002,2028]" name="Malaysia" pageId="8" pageNumber="67">Sarawak</collectingCountry>
|
||
are discrete morphological forms, but concerning females these forms were not confirmed. It must also be noted, that all specimens in each of the populations were collected together from site of several square meters, and thus the differences could be due to idiosyncratic conditions of the sample rather than to a geographic cline (i.e. the particular conditions of the development of the specimens in the sample).
|
||
</paragraph>
|
||
<paragraph id="C77636DCFFFAFF8121E5DC3E0A9F8241" blockId="9.[151,1436,152,378]" pageId="9" pageNumber="68">Except variations in metric characters, in a few specimens elytral punctures are irregularly arranged when comparing left and right elytron and punctures are partly confluent.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |