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<document id="EBAEFA730FD4AE2FAE33DE41A8466024" ID-DOI="10.5252/g2015n2a1" ID-ISSN="1638-9395" ID-Zenodo-Dep="5210219" ID-ZooBank="urn:lsid:zoobank.org:pub:85E655C9-759B-4A94-8FFB-43AE11567C1A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629163125226" checkinUser="felipe" docAuthor="Arcerito, Facundo René Meroi, Waisfeld, Beatriz &amp; Balseiro, Diego" docDate="2015" docId="480E87D9FF91EC11CAE9FF630AA9DC9A" docLanguage="en" docName="Geodiversitas.37.2.131-150.pdf" docOrigin="Geodiversitas 37 (2)" docSource="http://dx.doi.org/10.5252/g2015n2a1" docStyle="DocumentStyle:4DA100580BA89F561A6D37CADC50A8FD.3:Geodiversitas.2015-2017.journal_article.type2" docStyleId="4DA100580BA89F561A6D37CADC50A8FD" docStyleName="Geodiversitas.2015-2017.journal_article.type2" docStyleVersion="3" docTitle="Asaphellus isabelae Arcerito &amp; Waisfeld &amp; Balseiro 2015, n. sp." docType="treatment" docVersion="4" lastPageNumber="145" masterDocId="B437FFA1FF9CEC1FC90EFFB40902DA17" masterDocTitle="Diversification of Asaphellus Callaway, 1877 (Asaphidae: Trilobita) during the Tremadocian in South West Gondwana (Cordillera Oriental, Argentina)" masterLastPageNumber="150" masterPageNumber="131" pageNumber="144" updateTime="1699063343485" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="E843F03CF3698576C1FB04F8DD708A3A">Diversification of Asaphellus Callaway, 1877 (Asaphidae: Trilobita) during the Tremadocian in South West Gondwana (Cordillera Oriental, Argentina)</mods:title>
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<mods:namePart id="C53034EBF907BACB7E1B9BBCE47E43D0">Arcerito, Facundo René Meroi</mods:namePart>
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<mods:namePart id="3CFE122B418DB1815E0F6E6F30FCE31F">Waisfeld, Beatriz</mods:namePart>
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<mods:namePart id="A0CD7E301761C1673600AF77F104877B">Balseiro, Diego</mods:namePart>
</mods:name>
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<mods:title id="F4A8BE03CA6468877D26FDF78A6CE207">Geodiversitas</mods:title>
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<mods:date id="B6C74A3C36AF04B8423413CFD72FE741">2015</mods:date>
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<mods:number id="8F8D85C4B8111A37A47E0835CA7CA3E4">2015-06-26</mods:number>
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<mods:number id="80D2895F1440E3E6A5448D2E3FABAEF4">37</mods:number>
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<mods:url id="F73BDD434A160A31AB83E065FEE146D8">http://dx.doi.org/10.5252/g2015n2a1</mods:url>
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<mods:classification id="1734AF0ED0DFC872CB55DC41743EE53C">journal article</mods:classification>
<mods:identifier id="57E30B16A76E0BE0FBE194E49B78AB40" type="DOI">10.5252/g2015n2a1</mods:identifier>
<mods:identifier id="D985399A49D2A7E2AF994D117D8F66CD" type="ISSN">1638-9395</mods:identifier>
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<treatment id="480E87D9FF91EC11CAE9FF630AA9DC9A" ID-DOI="http://doi.org/10.5281/zenodo.5210276" ID-Zenodo-Dep="5210276" LSID="urn:lsid:plazi:treatment:480E87D9FF91EC11CAE9FF630AA9DC9A" httpUri="http://treatment.plazi.org/id/480E87D9FF91EC11CAE9FF630AA9DC9A" lastPageId="14" lastPageNumber="145" pageId="13" pageNumber="144">
<subSubSection id="88BD6544FF91EC12CAE9FF630DF7DAE5" box="[999,1269,215,242]" pageId="13" pageNumber="144" type="nomenclature">
<paragraph id="C01836CFFF91EC12CAE9FF630DF7DAE5" blockId="13.[999,1269,215,273]" box="[999,1269,215,242]" pageId="13" pageNumber="144">
<heading id="9B5081A3FF91EC12CAE9FF630DF7DAE5" box="[999,1269,215,242]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="144" reason="2">
<taxonomicName id="07A74D4CFF91EC12CAE9FF630DB1DAE5" authority="Arcerito &amp; Waisfeld &amp; Balseiro, 2015" authorityName="Arcerito &amp; Waisfeld &amp; Balseiro" authorityYear="2015" box="[999,1203,215,242]" class="Trilobita" family="Asaphidae" genus="Asaphellus" kingdom="Animalia" order="Asaphida" pageId="13" pageNumber="144" phylum="Arthropoda" rank="species" species="isabelae" status="sp. nov.">
<emphasis id="F2D3EADDFF91EC12CAE9FF630DB1DAE5" bold="true" box="[999,1203,215,242]" italics="true" pageId="13" pageNumber="144">Asaphellus isabelae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E9E057A6FF91EC12CDB4FF6C0DF7DAE5" box="[1210,1269,216,242]" pageId="13" pageNumber="144" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="88BD6544FF91EC12CD01FF430DCFDB06" box="[1039,1229,247,273]" pageId="13" pageNumber="144" type="description">
<paragraph id="C01836CFFF91EC12CD01FF430DCFDB06" blockId="13.[999,1269,215,273]" box="[1039,1229,247,273]" pageId="13" pageNumber="144">
(
<figureCitation id="589C2A4AFF91EC12CD18FF430D72DB06" box="[1046,1136,247,273]" captionStart="FIG" captionStartId="10.[132,143,1194,1212]" captionTargetBox="[132,1455,215,1150]" captionTargetId="figure-393@10.[132,1455,215,1156]" captionTargetPageId="10" captionText="FIG. 7. — A, B, Asaphellus sp. 2, pygidia in dorsal view: A, CEGH-UNC 25763; B, CEGH-UNC 25764; C-F, Asaphellus sp. 1, CEGH-UNC 25765-25768; C, pygidium, latex cast, dorsal view, CEGH-UNC 25765; D, pygidium, dorsal view, CEGH-UNC 25766; E, pygidium, dorsal view, CEGH-UNC 25765; F, pygidium and thorax, dorsal view,CEGH-UNC 25767;G, A. isabelae n. sp., articulated juvenile, latex cast, dorsal view,CEGH-UNC 25762. Scale bars: A-F, 0.5 cm; G, 0.25 cm. All specimens come from the Quebrada del Arenal, Santa Rosita Formation, Santa Victoria Group: A, B, G, from DLA 28; C, E, from DLA 25; D, F, from DLA 20." figureDoi="http://doi.org/10.5281/zenodo.5210227" httpUri="https://zenodo.org/record/5210227/files/figure.png" pageId="13" pageNumber="144">Figs 7G</figureCitation>
;
<figureCitation id="589C2A4AFF91EC12CD75FF430D9EDB06" box="[1147,1180,247,273]" captionStart="FIG" captionStartId="14.[240,251,1008,1026]" captionTargetBox="[161,1426,215,969]" captionTargetId="figure-546@14.[791,1436,197,728]" captionTargetPageId="14" captionText="FIG. 9. — A, reconstruction of A. stenorhachis (Harrington, 1938); B, reconstruction of A. isabelae n. sp. Drawing by Santiago Druetta." figureDoi="http://doi.org/10.5281/zenodo.5210229" httpUri="https://zenodo.org/record/5210229/files/figure.png" pageId="13" pageNumber="144">9B</figureCitation>
;
<figureCitation id="589C2A4AFF91EC12CDA6FF430DC7DB06" box="[1192,1221,247,273]" captionStart="FIG" captionStartId="16.[132,143,1894,1912]" captionTargetBox="[249,1337,216,1803]" captionTargetId="figure-188@16.[249,1339,215,1803]" captionTargetPageId="16" captionText="FIG. 10. — Asaphellus isabelae n. sp.: A, hypostoma, latex cast, dorsal view, CEGH-UNC 25749; B, hypostoma, dorsal view, CEGH-UNC 25752; C, right free cheek, dorsal view, CEGH-UNC 25750; D, left free cheek, latex cast, dorsal view, CEGH-UNC 25756; E, holotype cranidium, dorsal view, CEGH-UNC 25751; F, cranidium, latex cast, dorsal view, CEGH-UNC 25753; G, left free check, dorsal view, CEGH-UNC 25754; H-J, L, pygidia, dorsal view; H, CEGH-UNC 25755; I, CEGH-UNC 25758; J, CEGH-UNC 25757; L, CEGH-UNC 25760; K, pygidium, latex cast, dorsal view, CEGH-UNC 25759; M, almost complete moult, dorsal view, CEGH-UNC 25761. All specimens come from the Quebrada del Arenal, Santa Rosita Formation, Santa Victoria Group; A, C-E, G-J, L, M, come from DLA 28; B, F, K, come from DLA 30. Scale bars: A, B, 0.25 cm; C-M, 0.5 cm." figureDoi="http://doi.org/10.5281/zenodo.5210259" httpUri="https://zenodo.org/record/5210259/files/figure.png" pageId="13" pageNumber="144">10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="88BD6544FF91EC12CA23FE840D00DB75" pageId="13" pageNumber="144" type="reference_group">
<paragraph id="C01836CFFF91EC12CA23FE840D00DB75" blockId="13.[813,1455,304,354]" pageId="13" pageNumber="144">
<taxonomicName id="07A74D4CFF91EC12CA23FE840D11DB5F" ID-CoL="H4CK" baseAuthorityName="Kobayashi" baseAuthorityYear="1935" box="[813,1043,304,328]" class="Trilobita" family="Asaphidae" genus="Asaphellus" kingdom="Animalia" order="Asaphida" pageId="13" pageNumber="144" phylum="Arthropoda" rank="species" species="catamarcensis">
<emphasis id="F2D3EADDFF91EC12CA23FE840D11DB5F" box="[813,1043,304,328]" italics="true" pageId="13" pageNumber="144">Asaphellus catamarcensis</emphasis>
</taxonomicName>
–
<bibRefCitation id="A4364B3EFF91EC12CD23FE840C24DB5F" author="WAISFELD B. G. &amp; VACCARI N. E." box="[1069,1318,304,328]" pageId="13" pageNumber="144" pagination="295 - 410" refId="ref17349" refString="WAISFELD B. G. &amp; VACCARI N. E. 2003. - Trilobites, in BENEDETTO J. L. (ed.), Ordovician Fossils of Argentina. Secretaria de Ciencia y Tecnologia Universidad Nacional de Cordoba, Cordoba: 295 - 410." type="book chapter" year="2003">Waisfeld &amp; Vaccari 2003</bibRefCitation>
: pl. 22, fig. 5;
<emphasis id="F2D3EADDFF91EC12CA23FEFF0A53DB75" box="[813,849,331,354]" italics="true" pageId="13" pageNumber="144">non</emphasis>
<bibRefCitation id="A4364B3EFF91EC12CA59FEFE0AFCDB75" author="KOBAYASHI T." box="[855,1022,330,354]" pageId="13" pageNumber="144" pagination="59 - 67" refId="ref15587" refString="KOBAYASHI T. 1935. - On the Kainella Fauna of the Basal Ordovician Age found in Argentina. Japanese Journal of Geology and Geography 12: 59 - 67." type="journal article" year="1935">Kobayashi, 1935</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="88BD6544FF91EC12CA23FECF0AECD8F7" pageId="13" pageNumber="144" type="materials_examined">
<paragraph id="C01836CFFF91EC12CA23FECF0DF5DBD0" blockId="13.[811,1455,378,455]" pageId="13" pageNumber="144">
<materialsCitation id="70CF3C92FF91EC12CA23FECF0DF5DBD0" collectionCode="DLA" county="Type" location="Quebrada del Arenal" municipality="lower Tr" pageId="13" pageNumber="144" specimenCode="CEGH-UNC 25751, DLA 28" specimenCount="1" stateProvince="Victoria" typeStatus="holotype">
<typeStatus id="1F1C886DFF91EC12CA23FECF0A97DB86" box="[813,917,379,402]" pageId="13" pageNumber="144" type="holotype">HOLOTYPE</typeStatus>
. —
<specimenCode id="90019EB4FF91EC12CAB0FECE0D88DB85" box="[958,1162,378,402]" collectionCode="CEGH-UNC" pageId="13" pageNumber="144">CEGH-UNC 25751</specimenCode>
(
<figureCitation id="589C2A4AFF91EC12CD9BFECE0DE9DB85" box="[1173,1259,378,402]" captionStart="FIG" captionStartId="16.[132,143,1894,1912]" captionTargetBox="[249,1337,216,1803]" captionTargetId="figure-188@16.[249,1339,215,1803]" captionTargetPageId="16" captionText="FIG. 10. — Asaphellus isabelae n. sp.: A, hypostoma, latex cast, dorsal view, CEGH-UNC 25749; B, hypostoma, dorsal view, CEGH-UNC 25752; C, right free cheek, dorsal view, CEGH-UNC 25750; D, left free cheek, latex cast, dorsal view, CEGH-UNC 25756; E, holotype cranidium, dorsal view, CEGH-UNC 25751; F, cranidium, latex cast, dorsal view, CEGH-UNC 25753; G, left free check, dorsal view, CEGH-UNC 25754; H-J, L, pygidia, dorsal view; H, CEGH-UNC 25755; I, CEGH-UNC 25758; J, CEGH-UNC 25757; L, CEGH-UNC 25760; K, pygidium, latex cast, dorsal view, CEGH-UNC 25759; M, almost complete moult, dorsal view, CEGH-UNC 25761. All specimens come from the Quebrada del Arenal, Santa Rosita Formation, Santa Victoria Group; A, C-E, G-J, L, M, come from DLA 28; B, F, K, come from DLA 30. Scale bars: A, B, 0.25 cm; C-M, 0.5 cm." figureDoi="http://doi.org/10.5281/zenodo.5210259" httpUri="https://zenodo.org/record/5210259/files/figure.png" pageId="13" pageNumber="144">Fig. 10E</figureCitation>
).
<collectingCounty id="29794E43FF91EC12CDF7FECF0C2BDB84" box="[1273,1321,379,403]" pageId="13" pageNumber="144">Type</collectingCounty>
horizon
<specimenCode id="90019EB4FF91EC12CC73FECE0A44DBBA" collectionCode="DLA" pageId="13" pageNumber="144">DLA 28</specimenCode>
,
<collectingMunicipality id="207CACB5FF91EC12CA5EFE210AA1DBBA" box="[848,931,405,429]" pageId="13" pageNumber="144">lower Tr</collectingMunicipality>
2,
<location id="C5786014FF91EC12CAB6FE210D83DBBA" LSID="urn:lsid:plazi:treatment:480E87D9FF91EC11CAE9FF630AA9DC9A:C5786014FF91EC12CAB6FE210D83DBBA" box="[952,1153,405,429]" county="Type" municipality="lower Tr" name="Quebrada del Arenal" pageId="13" pageNumber="144" stateProvince="Victoria">
Quebrada del
<location id="C5786014FF91EC12CD4FFE210D83DBBA" LSID="urn:lsid:plazi:treatment:480E87D9FF91EC11CAE9FF630AA9DC9A:C5786014FF91EC12CD4FFE210D83DBBA" box="[1089,1153,405,429]" county="Type" municipality="lower Tr" name="Arenal creek" pageId="13" pageNumber="144" stateProvince="Victoria">Arenal</location>
</location>
creek,
<location id="C5786014FF91EC12CDCAFE210CA8DBBA" LSID="urn:lsid:plazi:treatment:480E87D9FF91EC11CAE9FF630AA9DC9A:C5786014FF91EC12CDCAFE210CA8DBBA" box="[1220,1450,405,429]" county="Type" municipality="lower Tr" name="lower Rupasca Member" pageId="13" pageNumber="144" stateProvince="Victoria">lower Rupasca Member</location>
,
<location id="C5786014FF91EC12CA22FE1B0D17DBD0" LSID="urn:lsid:plazi:treatment:480E87D9FF91EC11CAE9FF630AA9DC9A:C5786014FF91EC12CA22FE1B0D17DBD0" box="[812,1045,431,455]" county="Type" municipality="lower Tr" name="Santa Rosita Formation" pageId="13" pageNumber="144" stateProvince="Victoria">Santa Rosita Formation</location>
,
<location id="C5786014FF91EC12CD11FE1B0D57DBD0" LSID="urn:lsid:plazi:treatment:480E87D9FF91EC11CAE9FF630AA9DC9A:C5786014FF91EC12CD11FE1B0D57DBD0" box="[1055,1109,431,455]" county="Type" municipality="lower Tr" name="Santa" pageId="13" pageNumber="144" stateProvince="Victoria">Santa</location>
<collectingRegion id="0263F82DFF91EC12CD55FE1B0DA8DBD0" box="[1115,1194,431,455]" country="Australia" name="Victoria" pageId="13" pageNumber="144">Victoria</collectingRegion>
Group.
</materialsCitation>
</paragraph>
<paragraph id="C01836CFFF91EC12CA23FE6B0D49D851" blockId="13.[811,1456,479,582]" pageId="13" pageNumber="144">
MATERIAL. — 140 fragmentary specimens, including 1 partly complete individual, 20 cranidia, 48 pygidia, 28 free cheeks,
<specimenCount id="D6A1FD46FF91EC12CC33FE4E0CB2D806" box="[1341,1456,505,529]" pageId="13" pageNumber="144" type="generic">7 specimens</specimenCount>
with thoracic segments, and 12 hypostomas. Illustrated material
<collectionCode id="A6B6AE0AFF91EC12CA20FD9A0AB4D851" box="[814,950,558,582]" pageId="13" pageNumber="144">CEGH-UNC</collectionCode>
25749-25761.
</paragraph>
<paragraph id="C01836CFFF91EC12CA23FDEB0AECD8F7" blockId="13.[812,1457,606,736]" pageId="13" pageNumber="144">
OCCURRENCE AND AGE. —
<collectionCode id="A6B6AE0AFF91EC12CD36FDEA0D68D861" box="[1080,1130,606,630]" pageId="13" pageNumber="144">DLA</collectionCode>
28,
<collectionCode id="A6B6AE0AFF91EC12CD9AFDEA0DC4D861" box="[1172,1222,606,630]" pageId="13" pageNumber="144">DLA</collectionCode>
29,
<collectionCode id="A6B6AE0AFF91EC12CDFFFDEA0C21D861" box="[1265,1315,606,630]" pageId="13" pageNumber="144">DLA</collectionCode>
30, lower part of the Rupasca Member, Quebrada del Arenal,
<taxonomicName id="07A74D4CFF91EC12CC00FDCD0CAFD887" authority="Zone" authorityName="Zone" box="[1294,1453,632,656]" class="Trilobita" family="Remopleurididae" genus="Kainella" kingdom="Animalia" order="Asaphida" pageId="13" pageNumber="144" phylum="Arthropoda" rank="species" species="teiichii">
<emphasis id="F2D3EADDFF91EC12CC00FDCD0C6CD887" box="[1294,1390,632,656]" italics="true" pageId="13" pageNumber="144">K. teiichii</emphasis>
Zone
</taxonomicName>
, middle Tremadocian (Tr2); reworked boulder of the Cardonal Formation, Pascha-Incamayo area,
<taxonomicName id="07A74D4CFF91EC12CD98FD1A0C6CD8D2" authority="Zone" authorityName="Zone" box="[1174,1390,685,709]" class="Trilobita" family="Remopleurididae" genus="Kainella" kingdom="Animalia" order="Asaphida" pageId="13" pageNumber="144" phylum="Arthropoda" rank="species" species="meridionalis">
<emphasis id="F2D3EADDFF91EC12CD98FD1A0C29D8D2" box="[1174,1323,685,709]" italics="true" pageId="13" pageNumber="144">K. meridionalis</emphasis>
Zone
</taxonomicName>
, lower Tremadocian (Tr1).
</paragraph>
</subSubSection>
<subSubSection id="88BD6544FF91EC12CA23FD4C0C73D9B8" pageId="13" pageNumber="144" type="etymology">
<paragraph id="C01836CFFF91EC12CA23FD4C0CADD918" blockId="13.[813,1455,759,783]" box="[813,1455,759,783]" pageId="13" pageNumber="144">
ETYMOLOGY.
<emphasis id="F2D3EADDFF91EC12CAA4FD430AC1D918" bold="true" box="[938,963,759,783]" pageId="13" pageNumber="144">—</emphasis>
After FR Meroi Acerito’s grandmother Isabel Olivares.
</paragraph>
<paragraph id="C01836CFFF91EC12CA24FC820AA0D959" blockId="13.[810,1457,822,943]" box="[810,930,822,848]" pageId="13" pageNumber="144">D IAGNOSIS</paragraph>
<paragraph id="C01836CFFF91EC12CA23FCE10C73D9B8" blockId="13.[810,1457,822,943]" pageId="13" pageNumber="144">
Flat and large species of
<taxonomicName id="07A74D4CFF91EC12CD35FCE10DA4D978" authorityName=", Robison &amp; Pantoja-Alor" authorityYear="1968" box="[1083,1190,853,879]" class="Trilobita" family="Asaphidae" genus="Asaphellus" kingdom="Animalia" order="Asaphida" pageId="13" pageNumber="144" phylum="Arthropoda" rank="genus">
<emphasis id="F2D3EADDFF91EC12CD35FCE10DA4D978" box="[1083,1190,853,879]" italics="true" pageId="13" pageNumber="144">Asaphellus</emphasis>
</taxonomicName>
. Cuticle very thin, high degree of smoothness and effacement, except for the lightly impressed preglabellar furrow, pygidial axis indistinct.
</paragraph>
</subSubSection>
<subSubSection id="88BD6544FF91EC11CA23FC610A08DF5A" lastPageId="14" lastPageNumber="145" pageId="13" pageNumber="144" type="description">
<paragraph id="C01836CFFF91EC12CA23FC610ABFD9FA" blockId="13.[811,1458,981,2028]" box="[813,957,981,1007]" pageId="13" pageNumber="144">DESCRIPTION</paragraph>
<paragraph id="C01836CFFF91EC12CA23FC400D85DFDA" blockId="13.[811,1458,981,2028]" pageId="13" pageNumber="144">
Exoskeleton large, subelliptical and flat. Isopygous with flat cephalic and pygidial borders. Cephalon semicircular, with wide border, diminishing in width towards the genal angle. Doublure subparallel to the margin. Cranidium depressed or flat; axial furrow indistinct. Glabella almost indistinguishable, faint traces of convex anterior preglabellar furrow in some specimens (
<figureCitation id="589C2A4AFF91EC12CABEFB000D37DED9" box="[944,1077,1204,1230]" captionStart="FIG" captionStartId="16.[132,143,1894,1912]" captionTargetBox="[249,1337,216,1803]" captionTargetId="figure-188@16.[249,1339,215,1803]" captionTargetPageId="16" captionText="FIG. 10. — Asaphellus isabelae n. sp.: A, hypostoma, latex cast, dorsal view, CEGH-UNC 25749; B, hypostoma, dorsal view, CEGH-UNC 25752; C, right free cheek, dorsal view, CEGH-UNC 25750; D, left free cheek, latex cast, dorsal view, CEGH-UNC 25756; E, holotype cranidium, dorsal view, CEGH-UNC 25751; F, cranidium, latex cast, dorsal view, CEGH-UNC 25753; G, left free check, dorsal view, CEGH-UNC 25754; H-J, L, pygidia, dorsal view; H, CEGH-UNC 25755; I, CEGH-UNC 25758; J, CEGH-UNC 25757; L, CEGH-UNC 25760; K, pygidium, latex cast, dorsal view, CEGH-UNC 25759; M, almost complete moult, dorsal view, CEGH-UNC 25761. All specimens come from the Quebrada del Arenal, Santa Rosita Formation, Santa Victoria Group; A, C-E, G-J, L, M, come from DLA 28; B, F, K, come from DLA 30. Scale bars: A, B, 0.25 cm; C-M, 0.5 cm." figureDoi="http://doi.org/10.5281/zenodo.5210259" httpUri="https://zenodo.org/record/5210259/files/figure.png" pageId="13" pageNumber="144">Fig. 10E, F</figureCitation>
). Wide and subtriangular (exsag.) posterolateral projection. Posterior border and occipital ring reduced, poorly defined, and very close to posterior margin. Palpebral lobes of medium size, located at cranidial midlength. Anterior section of facial suture diverges gently forwards, bends smoothly adaxially, and converges medially to form an obtuse point sagittally in an ogive. Posterior section of facial suture curves backwards, bends downwards and slightly inwards at the level of the posterior border.
</paragraph>
<paragraph id="C01836CFFF91EC12CA4DFA670ADDDC5B" blockId="13.[811,1458,981,2028]" pageId="13" pageNumber="144">Free cheek long and relatively narrow, subtriangular with rounded external margin. Broad-based and long genal spine (55% of the total librigena length), slightly divergent from external margin.</paragraph>
<paragraph id="C01836CFFF91EC12CA4DF9E70DC8DCFB" blockId="13.[811,1458,981,2028]" pageId="13" pageNumber="144">Hypostoma elongated, oval in outline (sag.), with rounded posterior margin. Wide posterolateral border, with fine terraced lines. Median body widest (tr.) at midlength, anterior lobe convex (sag., tr.), posterior lobe short (sag.), crescentic, maculae transverse, sharply impressed.</paragraph>
<paragraph id="C01836CFFF91EC11CA4DF9460B99DE79" blockId="13.[811,1458,981,2028]" lastBlockId="14.[130,778,1076,1358]" lastPageId="14" lastPageNumber="145" pageId="13" pageNumber="144">
Thorax incompletely known; almost indistinguishable from
<taxonomicName id="07A74D4CFF91EC12CA25F8A60ABDDD3B" box="[811,959,1810,1836]" class="Trilobita" family="Asaphidae" genus="Asaphellus" kingdom="Animalia" order="Asaphida" pageId="13" pageNumber="144" phylum="Arthropoda" rank="species" species="stenorhachis">
<emphasis id="F2D3EADDFF91EC12CA25F8A60ABDDD3B" box="[811,959,1810,1836]" italics="true" pageId="13" pageNumber="144">A. stenorhachis</emphasis>
</taxonomicName>
but less convex. Axis weakly convex, about 35% of the total width (tr.), defined by well impressed and slightly backward-tapering furrows; articulating furrow narrow and deep; articulating half-ring short (tr.). Pleural region moderately wide (about 65% of thoracic maximum width); pleural furrow deeper and wider medially, runs toward the anterior margin of pleura, shallower and narrower adaxially, lightly impressed abaxially. Pleural extremities obliquely truncated and backward-facing, faint marked pleural fulcra.
</paragraph>
<caption id="94D86647FF92EC11C9FEFC440C47DE15" ID-DOI="http://doi.org/10.5281/zenodo.5210229" ID-Zenodo-Dep="5210229" box="[240,1349,1008,1026]" httpUri="https://zenodo.org/record/5210229/files/figure.png" pageId="14" pageNumber="145" startId="14.[240,251,1008,1026]" targetBox="[161,1426,215,969]" targetPageId="14">
<paragraph id="C01836CFFF92EC11C9FEFC440C47DE15" blockId="14.[240,1349,1008,1026]" box="[240,1349,1008,1026]" pageId="14" pageNumber="145">
FIG. 9. —
<emphasis id="F2D3EADDFF92EC11C84EFC44084FDE15" bold="true" box="[320,333,1008,1026]" pageId="14" pageNumber="145">A</emphasis>
, reconstruction of
<taxonomicName id="07A74D4CFF92EC11C8E4FC440A0BDE15" authority="(Harrington, 1938)" baseAuthorityName="Harrington" baseAuthorityYear="1938" box="[490,777,1008,1026]" class="Trilobita" family="Asaphidae" genus="Asaphellus" kingdom="Animalia" order="Asaphida" pageId="14" pageNumber="145" phylum="Arthropoda" rank="species" species="stenorhachis">
<emphasis id="F2D3EADDFF92EC11C8E4FC440B69DE15" box="[490,619,1008,1026]" italics="true" pageId="14" pageNumber="145">A. stenorhachis</emphasis>
(
<bibRefCitation id="A4364B3EFF92EC11CB7AFC440A01DE15" author="HARRINGTON H. J." box="[628,771,1008,1026]" pageId="14" pageNumber="145" pagination="109 - 289" refId="ref15300" refString="HARRINGTON H. J. 1938. - Sobre las faunas del Ordoviciano inferior del norte Argentino. Revista del Museo de La Plata, Seccion Paleontologia 4: 109 - 289." type="journal article" year="1938">Harrington, 1938</bibRefCitation>
)
</taxonomicName>
;
<emphasis id="F2D3EADDFF92EC11CA1EFC440A1FDE15" bold="true" box="[784,797,1008,1026]" pageId="14" pageNumber="145">B</emphasis>
, reconstruction of
<taxonomicName id="07A74D4CFF92EC11CAB4FC440D17DE15" authority="Arcerito &amp; Waisfeld &amp; Balseiro, 2015" authorityName="Arcerito &amp; Waisfeld &amp; Balseiro" authorityYear="2015" box="[954,1045,1008,1026]" class="Trilobita" family="Asaphidae" genus="Asaphellus" kingdom="Animalia" order="Asaphida" pageId="14" pageNumber="145" phylum="Arthropoda" rank="species" species="isabelae" status="sp. nov.">
<emphasis id="F2D3EADDFF92EC11CAB4FC440D17DE15" box="[954,1045,1008,1026]" italics="true" pageId="14" pageNumber="145">A. isabelae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E9E057A6FF92EC11CD14FC440D48DE15" box="[1050,1098,1008,1026]" pageId="14" pageNumber="145" rank="species">n. sp.</taxonomicNameLabel>
Drawing by Santiago Druetta.
</paragraph>
</caption>
<paragraph id="C01836CFFF92EC11C995FBC00A08DF5A" blockId="14.[130,778,1076,1358]" pageId="14" pageNumber="145">Pygidium of very low convexity, 57% wider than long (n=14), border and border furrow indistinct. Doublure evenly wide, about 10% length of pygidium (n = 11), subparallel to the pygidial margin along its extension. Anterior half-ring narrow (sag.), slightly convex forward.Articulating facet subtriangular, long (tr.). Anterior half-rib defined by narrow and shallow pleural furrow, remaining pleural furrows and ribs not visible.</paragraph>
</subSubSection>
<subSubSection id="88BD6544FF92EC11C98CFAC00AA9DC9A" pageId="14" pageNumber="145" type="discussion">
<paragraph id="C01836CFFF92EC11C98CFAC009EADF9B" blockId="14.[130,777,1396,2027]" box="[130,232,1396,1422]" pageId="14" pageNumber="145">REMARKS</paragraph>
<paragraph id="C01836CFFF92EC11C98AFA270B4FDC7B" blockId="14.[130,777,1396,2027]" pageId="14" pageNumber="145">
<taxonomicName id="07A74D4CFF92EC11C98AFA270844DFBA" authority="Arcerito &amp; Waisfeld &amp; Balseiro, 2015" authorityName="Arcerito &amp; Waisfeld &amp; Balseiro" authorityYear="2015" box="[132,326,1427,1453]" class="Trilobita" family="Asaphidae" genus="Asaphellus" kingdom="Animalia" order="Asaphida" pageId="14" pageNumber="145" phylum="Arthropoda" rank="species" species="isabelae" status="sp. nov.">
<emphasis id="F2D3EADDFF92EC11C98AFA270844DFBA" box="[132,326,1427,1453]" italics="true" pageId="14" pageNumber="145">Asaphellus isabelae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E9E057A6FF92EC11C841FA200889DFB9" box="[335,395,1428,1454]" pageId="14" pageNumber="145" rank="species">n. sp.</taxonomicNameLabel>
appears to represent an extreme in effacement, flatness, and reduction of the cuticle thickness within the group. The latter character is noteworthy, as even the sediment roughness is reproduced on the surface of the exoskeleton. Despite effacement complicates taxonomy, the extreme smoothness and flatness might be considered as an autoapomorphic character for this species.
</paragraph>
<paragraph id="C01836CFFF92EC11C995F9C608B8DDFC" blockId="14.[130,777,1396,2027]" pageId="14" pageNumber="145">
Such a combination of characters has seldom been recorded in other
<taxonomicName id="07A74D4CFF92EC11C83BF92608FFDCBB" box="[309,509,1682,1708]" class="Trilobita" family="Asaphidae" genus="Asaphellus" kingdom="Animalia" order="Asaphida" pageId="14" pageNumber="145" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="F2D3EADDFF92EC11C83BF92608A1DCBB" box="[309,419,1682,1708]" italics="true" pageId="14" pageNumber="145">Asaphellus</emphasis>
species.
</taxonomicName>
For example,
<taxonomicName id="07A74D4CFF92EC11CB97F9260821DCDB" authority="Yin" authorityName="Yin" class="Trilobita" family="Asaphidae" genus="Asaphellus" kingdom="Animalia" order="Asaphida" pageId="14" pageNumber="145" phylum="Arthropoda" rank="species" species="yanheensis">
<emphasis id="F2D3EADDFF92EC11CB97F92609F0DCDB" italics="true" pageId="14" pageNumber="145">Asaphellus yanheensis</emphasis>
Yin
</taxonomicName>
<emphasis id="F2D3EADDFF92EC11C822F9070840DCDB" box="[300,322,1715,1740]" italics="true" pageId="14" pageNumber="145">in</emphasis>
<bibRefCitation id="A4364B3EFF92EC11C842F9060B2CDCDB" author="YIN G. &amp; SHANJI L." box="[332,558,1714,1740]" pageId="14" pageNumber="145" pagination="134 - 213" refId="ref17484" refString="YIN G. &amp; SHANJI L. 1978. - Class Trilobita, in Guizhou Stratigraphic and Paleontologic Team (ed.), Palaeontological Atlas of Southwest China, Guizhou Province. Beijing Geological Publishing House 134 - 213." type="book chapter" year="1978">Yin &amp; Shanji, 1978</bibRefCitation>
(
<bibRefCitation id="A4364B3EFF92EC11CB31F9060BBFDCDB" author="PENG S." box="[575,701,1714,1740]" pageId="14" pageNumber="145" pagination="1 - 53" refId="ref16134" refString="PENG S. 1990. - Tremadocian stratigraphy and trilobite fauna of northwestern Hunan. 1, Trilobites from the Nantsinkwan Formation of the Yangtze Platform. Beringeria 2: 1 - 53." type="journal article" year="1990">Peng 1990</bibRefCitation>
: pl. 7, figs 6-13) from the Nantsinkwan Formation (Tremadocian, NW
<collectingRegion id="0263F82DFF92EC11C9B2F946080FDD1B" box="[188,269,1778,1804]" country="China" name="Hunan" pageId="14" pageNumber="145">Hunan</collectingRegion>
,
<collectingCountry id="B8B0765FFF92EC11C817F946085DDD1A" box="[281,351,1778,1805]" name="China" pageId="14" pageNumber="145">China</collectingCountry>
) exhibits a thorough effacement, with blurred furrows either in the cranidia or pygidia. Though the pygidium of the Chinese species exhibits great resemblance to
<taxonomicName id="07A74D4CFF92EC11C9ADF8E60811DD7C" authority="Arcerito &amp; Waisfeld &amp; Balseiro, 2015" authorityName="Arcerito &amp; Waisfeld &amp; Balseiro" authorityYear="2015" box="[163,275,1873,1900]" class="Trilobita" family="Asaphidae" genus="Asaphellus" kingdom="Animalia" order="Asaphida" pageId="14" pageNumber="145" phylum="Arthropoda" rank="species" species="isabelae" status="sp. nov.">
<emphasis id="F2D3EADDFF92EC11C9ADF8E60811DD7C" box="[163,275,1873,1900]" italics="true" pageId="14" pageNumber="145">A. isabelae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E9E057A6FF92EC11C812F8E60858DD7B" box="[284,346,1874,1900]" pageId="14" pageNumber="145" rank="species">n. sp.</taxonomicNameLabel>
, the cranidium of the former differs in several significant respects, such as a better defined anterior ogive, a median node that is placed more forwardly, a subparallel preocular facial sutures, and postocular sutures running straight backwards.
</paragraph>
<paragraph id="C01836CFFF92EC11CA4AFB800AA9DC9A" blockId="14.[811,1457,1076,1677]" pageId="14" pageNumber="145">
Other species also show effacement either on the cranidium or the pygidium.
<taxonomicName id="07A74D4CFF92EC11CD57FBE00A72DE98" authority="(Sheng 1958)" baseAuthorityName="Sheng" baseAuthorityYear="1958" class="Trilobita" family="Asaphidae" genus="Asaphellus" kingdom="Animalia" order="Asaphida" pageId="14" pageNumber="145" phylum="Arthropoda" rank="species" species="tungtzuensis">
<emphasis id="F2D3EADDFF92EC11CD57FBE00C5BDE79" box="[1113,1369,1108,1134]" italics="true" pageId="14" pageNumber="145">Asaphellus tungtzuensis</emphasis>
(
<bibRefCitation id="A4364B3EFF92EC11CC65FBE00A6ADE98" author="SHENG S." pageId="14" pageNumber="145" pagination="183 - 204" refId="ref16480" refString="SHENG S. 1958. - Ordovician Trilobites from southwest China. Acta Paleontologica Sinica 2: 183 - 204." type="journal article" year="1958">Sheng 1958</bibRefCitation>
)
</taxonomicName>
(
<bibRefCitation id="A4364B3EFF92EC11CA71FBC00AFFDE99" author="PENG S." box="[895,1021,1140,1166]" pageId="14" pageNumber="145" pagination="1 - 53" refId="ref16134" refString="PENG S. 1990. - Tremadocian stratigraphy and trilobite fauna of northwestern Hunan. 1, Trilobites from the Nantsinkwan Formation of the Yangtze Platform. Beringeria 2: 1 - 53." type="journal article" year="1990">Peng 1990</bibRefCitation>
: pl. 8, figs 1, 2) from the Tremadocian of north west
<collectingRegion id="0263F82DFF92EC11CADCFB200D20DEB9" box="[978,1058,1172,1198]" country="China" name="Hunan" pageId="14" pageNumber="145">Hunan</collectingRegion>
resembles
<taxonomicName id="07A74D4CFF92EC11CDAAFB200C18DEB9" authority="Arcerito &amp; Waisfeld &amp; Balseiro, 2015" authorityName="Arcerito &amp; Waisfeld &amp; Balseiro" authorityYear="2015" box="[1188,1306,1172,1198]" class="Trilobita" family="Asaphidae" genus="Asaphellus" kingdom="Animalia" order="Asaphida" pageId="14" pageNumber="145" phylum="Arthropoda" rank="species" species="isabelae" status="sp. nov.">
<emphasis id="F2D3EADDFF92EC11CDAAFB200C18DEB9" box="[1188,1306,1172,1198]" italics="true" pageId="14" pageNumber="145">A. isabelae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E9E057A6FF92EC11CC2AFB200C66DEB9" box="[1316,1380,1172,1198]" pageId="14" pageNumber="145" rank="species">n. sp.</taxonomicNameLabel>
in the general effacement of the cranidium, however, the pygidium shows traces of axial furrows delimiting a weak axis along its length. The cranidium of
<taxonomicName id="07A74D4CFF92EC11CD8EFB470DF0DF1A" authorityName=", Robison &amp; Pantoja-Alor" authorityYear="1968" box="[1152,1266,1267,1293]" class="Trilobita" family="Asaphidae" genus="Asaphellus" kingdom="Animalia" order="Asaphida" pageId="14" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="F2D3EADDFF92EC11CD8EFB470DF0DF1A" box="[1152,1266,1267,1293]" italics="true" pageId="14" pageNumber="145">Asaphellus</emphasis>
</taxonomicName>
cf.
<taxonomicName id="07A74D4CFF92EC11CC11FB400AC0DF39" authority="Chang, 1949" authorityName="Chang" authorityYear="1949" class="Trilobita" family="Asaphidae" genus="Asaphellus" kingdom="Animalia" order="Asaphida" pageId="14" pageNumber="145" phylum="Arthropoda" rank="species" species="trinodosus">
<emphasis id="F2D3EADDFF92EC11CC11FB400CADDF1A" box="[1311,1455,1267,1294]" italics="true" pageId="14" pageNumber="145">A. trinodosus</emphasis>
<bibRefCitation id="A4364B3EFF92EC11CA21FAA70AC0DF39" author="CHANG W. T." box="[815,962,1299,1326]" pageId="14" pageNumber="145" pagination="110 - 125" refId="ref14729" refString="CHANG W. T. 1949. - Ordovician trilobites from Kaiping, Hebei and the Qaidam Basin. Bulletin of the Geological Society of China. 29: 110 - 125." type="journal article" year="1949">Chang, 1949</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="A4364B3EFF92EC11CADFFAA70D93DF39" author="JELL P. A. &amp; STAIT B." box="[977,1169,1299,1326]" pageId="14" pageNumber="145" pagination="1 - 34" refId="ref15508" refString="JELL P. A. &amp; STAIT B. 1985. - Tremadoc trilobites from the Florentine Valley Formation, Tim Shea area, Tasmania. Memoirs of the Museum of Victoria 46: 1 - 34." type="journal article" year="1985">Jell &amp; Stait 1985</bibRefCitation>
) from the Florentine Valley Formation (Tremadocian,
<collectingRegion id="0263F82DFF92EC11CD77FA870DE4DF59" box="[1145,1254,1331,1358]" country="Australia" name="Tasmania" pageId="14" pageNumber="145">Tasmania</collectingRegion>
) is largely effaced; however, the pygidial axial furrows distinctly outline a convex axis. Among the Argentinean species of
<taxonomicName id="07A74D4CFF92EC11CC5BFAC70A48DFBA" authorityName=", Robison &amp; Pantoja-Alor" authorityYear="1968" class="Trilobita" family="Asaphidae" genus="Asaphellus" kingdom="Animalia" order="Asaphida" pageId="14" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="F2D3EADDFF92EC11CC5BFAC70A48DFBA" italics="true" pageId="14" pageNumber="145">Asaphellus</emphasis>
</taxonomicName>
, only
<taxonomicName id="07A74D4CFF92EC11CA9BFA270D3ADFBA" baseAuthorityName="Harrington" baseAuthorityYear="1938" box="[917,1080,1427,1453]" class="Trilobita" family="Asaphidae" genus="Asaphellus" kingdom="Animalia" order="Asaphida" pageId="14" pageNumber="145" phylum="Arthropoda" rank="species" species="stenorhachis">
<emphasis id="F2D3EADDFF92EC11CA9BFA270D3ADFBA" box="[917,1080,1427,1453]" italics="true" pageId="14" pageNumber="145">A. stenorhachis</emphasis>
</taxonomicName>
(see above) exhibits a general effacement though not to the degree of that of
<taxonomicName id="07A74D4CFF92EC11CC35FA070CB2DFDA" authority="Arcerito &amp; Waisfeld &amp; Balseiro, 2015" authorityName="Arcerito &amp; Waisfeld &amp; Balseiro" authorityYear="2015" box="[1339,1456,1459,1485]" class="Trilobita" family="Asaphidae" genus="Asaphellus" kingdom="Animalia" order="Asaphida" pageId="14" pageNumber="145" phylum="Arthropoda" rank="species" species="isabelae" status="sp. nov.">
<emphasis id="F2D3EADDFF92EC11CC35FA070CB2DFDA" box="[1339,1456,1459,1485]" italics="true" pageId="14" pageNumber="145">A. isabelae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E9E057A6FF92EC11CA20FA670A6EDFFA" box="[814,876,1491,1517]" pageId="14" pageNumber="145" rank="species">n. sp.</taxonomicNameLabel>
<taxonomicName id="07A74D4CFF92EC11CA7CFA670D6FDFFA" baseAuthorityName="Harrington" baseAuthorityYear="1938" box="[882,1133,1491,1517]" class="Trilobita" family="Asaphidae" genus="Asaphellus" kingdom="Animalia" order="Asaphida" pageId="14" pageNumber="145" phylum="Arthropoda" rank="species" species="stenorhachis">
<emphasis id="F2D3EADDFF92EC11CA7CFA670D6FDFFA" box="[882,1133,1491,1517]" italics="true" pageId="14" pageNumber="145">Asaphellus stenorhachis</emphasis>
</taxonomicName>
differs from the new species in the higher convexity mainly of the pygidium, a better definition of the pygidial axis in its anterior third, and the presence of some furrows, lightly impressed but discernible (for instance the border furrows in the cephalon and pygidium).
</paragraph>
</subSubSection>
</treatment>
</document>
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