treatments-xml/data/43/73/87/437387DAFFE2FF8E224EFDA90DDFC4FA.xml
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<document ID-DOI="http://doi.org/10.5281/zenodo.6054091" ID-GBIF-Dataset="c1f56f9c-98ae-4dd9-9990-e87d4ce20c4a" ID-ISSN="2348-7372" ID-Zenodo-Dep="6054091" checkinTime="1480953219367" checkinUser="plazi" docAuthor="P. Girish Kumar, J. M. Carpenter &amp; P. M. Sureshan" docDate="2016" docId="437387DAFFE2FF8E224EFDA90DDFC4FA" docLanguage="en" docName="1. girish.pdf" docOrigin="Halteres 7 (1)" docStyle="DocumentStyle{}" docTitle="A taxonomic review of the genus Parancistrocerus Bequaert (Hymenoptera: Vespidae: Eumeninae) from the Indian subcontinent with the description of three new species" docType="key" docVersion="8" lastPageId="2" lastPageNumber="138" masterDocId="BF4AFFA2FFE3FF8C234BFFB30F49C361" masterDocTitle="A taxonomic review of the genus Parancistrocerus Bequaert (Hymenoptera: Vespidae: Eumeninae) from the Indian subcontinent with the description of three new species" masterLastPageNumber="156" masterPageNumber="36" pageId="1" pageNumber="137" updateTime="1644679175000" updateUser="ExternalLinkService">
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<mods:title>A taxonomic review of the genus Parancistrocerus Bequaert (Hymenoptera: Vespidae: Eumeninae) from the Indian subcontinent with the description of three new species</mods:title>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>P. Girish Kumar</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>J. M. Carpenter</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>P. M. Sureshan</mods:namePart>
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<mods:title>Halteres</mods:title>
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<mods:date>2016</mods:date>
<mods:detail type="volume">
<mods:number>7</mods:number>
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<mods:identifier type="DOI">10.5281/zenodo.192283</mods:identifier>
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<subSubSection box="[261,678,538,564]" pageId="1" pageNumber="137" type="nomenclature">
<paragraph blockId="1.[192,747,538,799]" box="[261,678,538,564]" pageId="1" pageNumber="137">
<emphasis bold="true" box="[261,678,538,564]" pageId="1" pageNumber="137">
Genus
<emphasis bold="true" box="[348,555,538,564]" italics="true" pageId="1" pageNumber="137">Parancistrocerus</emphasis>
Bequaert
</emphasis>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="137" type="reference_group">
<paragraph blockId="1.[192,747,538,799]" pageId="1" pageNumber="137">
<emphasis box="[192,396,571,596]" italics="true" pageId="1" pageNumber="137">Parancistrocerus</emphasis>
<bibRefCitation author="Bequaert, J." box="[441,662,570,597]" journalOrPublisher="Transactions of the American Entomological Society" pageId="1" pageNumber="137" pagination="57 - 117" part="51" refString="Bequaert, J. 1925. The genus Ancistrocerus (Hymenoptera, Vespidae) in North America, with a partial key to species. Transactions of the American Entomological Society 51: 57 - 117." title="The genus Ancistrocerus (Hymenoptera, Vespidae) in North America, with a partial key to species" type="journal article" year="1925">Bequaert, 1925</bibRefCitation>
: 64, subgenus of
<emphasis box="[444,607,605,631]" italics="true" pageId="1" pageNumber="137">Ancistrocerus</emphasis>
Wesmael. Type species:
<emphasis box="[466,695,638,664]" italics="true" pageId="1" pageNumber="137">Odynerus fulvipes</emphasis>
de Saussure, 1855 [=
<emphasis box="[573,594,673,699]" italics="true" pageId="1" pageNumber="137">O</emphasis>
.
<emphasis box="[639,746,671,698]" italics="true" pageId="1" pageNumber="137">“flavipes</emphasis>
Fabricius”
<emphasis box="[396,461,707,732]" italics="true" pageId="1" pageNumber="137">sensu</emphasis>
de
<bibRefCitation author="Saussure, H. F. de" box="[507,689,706,733]" journalOrPublisher="V. Masson &amp; Cherbuliez, Paris &amp; Geneva" pageId="1" pageNumber="137" pagination="1 - 3" refString="Saussure, H. F. de 1852 - 1858. Etudes sur la famille des vespides. Vols. 1 - 3. V. Masson &amp; Cherbuliez, Paris &amp; Geneva." title="1858. Etudes sur la famille des vespides. Vols" type="book chapter" year="1852">Saussure, 1852</bibRefCitation>
,
<emphasis italics="true" pageId="1" pageNumber="137">non Vespa flavipes</emphasis>
<bibRefCitation author="Fabricius, J. C. F." bookContentInfo="832 pp." box="[476,681,739,766]" journalOrPublisher="Kortii: Flensburgi et Lipsiae. xxviii" pageId="1" pageNumber="137" refString="Fabricius, J. C. F. 1775. Systema Entomolgiae, etc. Kortii: Flensburgi et Lipsiae. xxviii + 832 pp." title="Systema Entomolgiae, etc" type="book" year="1775">Fabricius, 1775</bibRefCitation>
], by original designation.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="137" type="diagnosis">
<paragraph blockId="1.[192,747,841,1238]" pageId="1" pageNumber="137">
<emphasis bold="true" box="[192,322,841,867]" pageId="1" pageNumber="137">Diagnosis:</emphasis>
T2 smooth basally, forming acarinarium beneath apex of T1 that is often full of mites (often concealed, tergum should be bent backwards to expose acarinarium); anterior face of pronotum with two close set, deeply impressed fovea, which may be approximated; propodeum with submarginal carina produced into pointed lamella apically and valvula enlarged and free posteriorly from submarginal carina; metanotum without tubercles; forewing with second submarginal cell not petiolate; metasoma sessile.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="137" type="distribution">
<paragraph blockId="1.[192,746,1279,1339]" pageId="1" pageNumber="137">
<emphasis bold="true" box="[192,355,1279,1305]" pageId="1" pageNumber="137">Distribution:</emphasis>
Nearctic, Neotropical, Oriental and Palaearctic regions.
</paragraph>
</subSubSection>
<subSubSection lastPageId="2" lastPageNumber="138" pageId="1" pageNumber="137" type="key">
<paragraph blockId="1.[192,747,1381,2047]" box="[317,718,1381,1407]" pageId="1" pageNumber="137">
<emphasis bold="true" box="[317,718,1381,1407]" pageId="1" pageNumber="137">Key to species and subspecies of</emphasis>
</paragraph>
<paragraph blockId="1.[192,747,1381,2047]" pageId="1" pageNumber="137">
<emphasis bold="true" box="[201,737,1415,1441]" pageId="1" pageNumber="137">
<taxonomicName authority="Bequaert" box="[201,531,1415,1441]" class="Insecta" family="Eumenidae" genus="Parancistrocerus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[201,408,1415,1441]" italics="true" pageId="1" pageNumber="137">Parancistrocerus</emphasis>
Bequaert
</taxonomicName>
from the Indian
</emphasis>
<emphasis bold="true" box="[389,550,1448,1474]" pageId="1" pageNumber="137">subcontinent</emphasis>
</paragraph>
<paragraph blockId="1.[192,747,1381,2047]" box="[243,695,1482,1508]" pageId="1" pageNumber="137">
(Modified from
<bibRefCitation author="Giordani Soika, A." box="[434,687,1482,1508]" journalOrPublisher="Annali del Museo civico di storia naturale &quot; G. Doria &quot; Vol. XC" pageId="1" pageNumber="137" pagination="1 - 348" refString="Giordani Soika, A. 1994. Ricerche sistematiche su alcuni generi di Eumenidi della Regione Orientale e della Papuasia. Annali del Museo civico di storia naturale &quot; G. Doria &quot; Vol. XC: 1 - 348." title="Ricerche sistematiche su alcuni generi di Eumenidi della Regione Orientale e della Papuasia" type="book chapter" year="1994">Giordani Soika, 1994</bibRefCitation>
)
</paragraph>
<paragraph blockId="1.[192,747,1381,2047]" pageId="1" pageNumber="137">
1. T2 with apical margin prolonged in middle (see Fig. 1 of
<bibRefCitation author="Giordani Soika, A." box="[421,687,1549,1575]" journalOrPublisher="Estratto dal Bollettino della Societa Entomologica Italiana" pageId="1" pageNumber="137" pagination="99 - 110" part="104" refString="Giordani Soika, A. 1972. Notulae Vespidologicae XXXII Nuovi Eumenidi Indomalesi. Estratto dal Bollettino della Societa Entomologica Italiana 104 (6 - 7): 99 - 110." title="Notulae Vespidologicae XXXII Nuovi Eumenidi Indomalesi" type="journal article" year="1972">Giordani Soika, 1972</bibRefCitation>
: 102 and Fig. 41 of
<bibRefCitation author="Giordani Soika, A." box="[462,743,1583,1609]" journalOrPublisher="Annali del Museo civico di storia naturale &quot; G. Doria &quot; Vol. XC" pageId="1" pageNumber="137" pagination="1 - 348" refString="Giordani Soika, A. 1994. Ricerche sistematiche su alcuni generi di Eumenidi della Regione Orientale e della Papuasia. Annali del Museo civico di storia naturale &quot; G. Doria &quot; Vol. XC: 1 - 348." title="Ricerche sistematiche su alcuni generi di Eumenidi della Regione Orientale e della Papuasia" type="book chapter" year="1994">Giordani Soika, 1994</bibRefCitation>
: 163)……………………………………...2
</paragraph>
<paragraph blockId="1.[192,747,1381,2047]" pageId="1" pageNumber="137">- T2 with apical margin normal, not prolonged in middle (Figs. 6, 12, 18, 27, 33 &amp; 39)…………………………………6</paragraph>
<paragraph blockId="1.[192,747,1381,2047]" pageId="1" pageNumber="137">
2. Superior carina of the propodeum well developed and distinct, especially at the top, where they clearly separate the horizontal dorsal face of the propodeum from the posterior face……
<taxonomicName authority="Giordani Soika" authorityName="Giordani Soika" class="Insecta" family="Eumenidae" genus="Parancistrocerus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="137" phylum="Arthropoda" rank="species" species="capocacciai">
<emphasis bold="true" box="[571,747,1886,1913]" italics="true" pageId="1" pageNumber="137">P. capocacciai</emphasis>
<emphasis bold="true" box="[240,432,1921,1947]" pageId="1" pageNumber="137">Giordani Soika</emphasis>
</taxonomicName>
</paragraph>
<paragraph blockId="1.[192,747,1381,2047]" lastBlockId="1.[841,1396,200,2048]" pageId="1" pageNumber="137">- Superior carina of propodeum not developed as above and irregular,no clear separation of dorsal and posterior surfaces of the propodeum………………………..3</paragraph>
<paragraph blockId="1.[841,1396,200,2048]" pageId="1" pageNumber="137">
3. Female unknown. Male: distance between clypeal teeth much shorter than basal width of clypeus, area between clypeal teeth more closely emarginate at apex; antennae with 11th article short, not longer than wide; apical antennal article very long, its apex reaches far beyond base of 11th article……………………..
<taxonomicName authority="Giordani Soika" authorityName="Giordani Soika" class="Insecta" family="Eumenidae" genus="Parancistrocerus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="137" phylum="Arthropoda" rank="species" species="irritatus">
<emphasis bold="true" pageId="1" pageNumber="137">
<emphasis bold="true" box="[1265,1396,470,496]" italics="true" pageId="1" pageNumber="137">P. irritatus</emphasis>
Giordani Soika
</emphasis>
</taxonomicName>
</paragraph>
<paragraph blockId="1.[841,1396,200,2048]" pageId="1" pageNumber="137">
- Male: distance between clypeal teeth slightly less than basal width of clypeus, area between clypeal teeth broadly emarginate at apex; antennae with 11th article long, about 1.50x as long as wide; apical antennal article short, its apex reaches to or slightly beyond base of 11th article……………………...
<taxonomicName authority="Giordani Soika" authorityName="Giordani Soika" class="Insecta" family="Eumenidae" genus="Parancistrocerus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="137" phylum="Arthropoda" rank="species" species="incorruptus">
<emphasis bold="true" pageId="1" pageNumber="137">
<emphasis bold="true" box="[1219,1396,773,800]" italics="true" pageId="1" pageNumber="137">P. incorruptus</emphasis>
Giordani Soika
</emphasis>
</taxonomicName>
………………………... 4
</paragraph>
<paragraph blockId="1.[841,1396,200,2048]" box="[841,1390,841,868]" pageId="1" pageNumber="137">4. T1 mostly red-ferruginous………………5</paragraph>
<paragraph blockId="1.[841,1396,200,2048]" pageId="1" pageNumber="137">
- T1 not red-ferruginous…………...ssp.
<emphasis bold="true" box="[889,1277,908,935]" pageId="1" pageNumber="137">
<emphasis bold="true" box="[889,1078,908,934]" italics="true" pageId="1" pageNumber="137">kalimpongensis</emphasis>
Giordani Soika
</emphasis>
</paragraph>
<paragraph blockId="1.[841,1396,200,2048]" pageId="1" pageNumber="137">
5. Propodeum mostly red-ferruginous; T2 without apical yellow band…………...ssp.
<taxonomicName authority="Giordani Soika" authorityName="Giordani Soika" box="[889,1230,1010,1036]" class="Insecta" family="Eumenidae" genus="Parancistrocerus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="137" phylum="Arthropoda" rank="species" species="incorruptus">
<emphasis bold="true" box="[889,1230,1010,1036]" pageId="1" pageNumber="137">
<emphasis bold="true" box="[889,1031,1010,1036]" italics="true" pageId="1" pageNumber="137">incorruptus</emphasis>
Giordani Soika
</emphasis>
</taxonomicName>
</paragraph>
<paragraph blockId="1.[841,1396,200,2048]" pageId="1" pageNumber="137">
- Propodeum black; T2 with apical yellow band………………………………….ssp.
<emphasis bold="true" box="[889,1180,1110,1137]" pageId="1" pageNumber="137">
<emphasis bold="true" box="[889,980,1110,1136]" italics="true" pageId="1" pageNumber="137">demens</emphasis>
Giordani Soika
</emphasis>
</paragraph>
<paragraph blockId="1.[841,1396,200,2048]" pageId="1" pageNumber="137">
6. T2 strongly reflexed at apex…………… ………......
<emphasis bold="true" box="[1022,1394,1178,1205]" pageId="1" pageNumber="137">
<emphasis bold="true" box="[1022,1189,1178,1205]" italics="true" pageId="1" pageNumber="137">P. assamensis</emphasis>
(Meade-Waldo)
</emphasis>
</paragraph>
<paragraph blockId="1.[841,1396,200,2048]" box="[841,1389,1212,1238]" pageId="1" pageNumber="137">- T2 not reflexed at apex………………….7</paragraph>
<paragraph blockId="1.[841,1396,200,2048]" pageId="1" pageNumber="137">7. S2 strongly and sharply lowered at base then widely and almost entirely depressed………………………………...8</paragraph>
<paragraph blockId="1.[841,1396,200,2048]" pageId="1" pageNumber="137">- S2 convex at base, very weakly depressed after……………………………………...9</paragraph>
<paragraph blockId="1.[841,1396,200,2048]" pageId="1" pageNumber="137">
8. Anterior face of T1 without median vertical carina on its upper half; clypeus of female more widely and deeply emarginate at apex…………...
<emphasis bold="true" pageId="1" pageNumber="137">
<emphasis bold="true" box="[1254,1396,1514,1542]" italics="true" pageId="1" pageNumber="137">P. rhipheus</emphasis>
(Cameron)
</emphasis>
</paragraph>
<paragraph blockId="1.[841,1396,200,2048]" pageId="1" pageNumber="137">
- Anterior face of T1 with median vertical carina on its upper half (Fig. 11); clypeus of female more narrowly and less deeply emarginate at apex (Fig. 8)......................... ……………..
<taxonomicName authority="Gusenleitner" authorityName="Gusenleitner" box="[1051,1384,1717,1744]" class="Insecta" family="Eumenidae" genus="Parancistrocerus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="137" phylum="Arthropoda" rank="species" species="holzschuhi">
<emphasis bold="true" box="[1051,1384,1717,1744]" pageId="1" pageNumber="137">
<emphasis bold="true" box="[1051,1215,1717,1744]" italics="true" pageId="1" pageNumber="137">P. holzschuhi</emphasis>
Gusenleitner
</emphasis>
</taxonomicName>
</paragraph>
<paragraph blockId="1.[841,1396,200,2048]" pageId="1" pageNumber="137">9. Vertical anterior face of T1 about as long as, or even longer than dorsal horizontal face……………………………………..10</paragraph>
<paragraph blockId="1.[841,1396,200,2048]" pageId="1" pageNumber="137">- Vertical anterior face of T1 distinctly shorter than dorsal horizontal face……..11</paragraph>
<paragraph blockId="1.[841,1396,200,2048]" lastBlockId="2.[192,747,200,1947]" lastPageId="2" lastPageNumber="138" pageId="1" pageNumber="137">
10. Width between apical teeth of clypeus equal to width between antennal sockets; upper carina of propodeum lamellate; punctures on T1 and T2 denser; markings on pronotum, mesopleuron, metanotum, T1, T2 and S2 ferruginous…………
<taxonomicName authority="Giordani Soika" authorityName="Giordani Soika" class="Insecta" family="Eumenidae" genus="Parancistrocerus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="138" phylum="Arthropoda" rank="species" species="feai">
<emphasis bold="true" pageId="2" pageNumber="138">
<emphasis bold="true" box="[669,746,233,260]" italics="true" pageId="2" pageNumber="138">P. feai</emphasis>
Giordani Soika
</emphasis>
</taxonomicName>
</paragraph>
<paragraph blockId="2.[192,747,200,1947]" pageId="2" pageNumber="138">
- Width between apical teeth of clypeus less than width between antennal sockets (Fig. 29); upper carina of propodeum not lamellate (Fig. 32); punctures on T1 and T2 fine, superficial; markings on pronotum, mesopleuron, metanotum, T1, T2 and S2 yellow……………...
<emphasis bold="true" pageId="2" pageNumber="138">
<taxonomicName authority="Girish Kumar &amp; Carpenter" authorityName="Girish Kumar &amp; Carpenter" class="Insecta" family="Eumenidae" genus="Parancistrocerus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="138" phylum="Arthropoda" rank="species" species="turensis" status="sp. n.">
<emphasis bold="true" box="[618,747,503,530]" italics="true" pageId="2" pageNumber="138">P. turensis</emphasis>
Girish Kumar &amp; Carpenter
</taxonomicName>
<taxonomicNameLabel box="[595,662,538,564]" pageId="2" pageNumber="138" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph blockId="2.[192,747,200,1947]" pageId="2" pageNumber="138">11. Dorsal face of propodeum absent mesally, not forming horizontal area behind midline of metanotum (Figs. 16 &amp; 38)………….12</paragraph>
<paragraph blockId="2.[192,747,200,1947]" pageId="2" pageNumber="138">- Dorsal face of propodeum forming horizontal area behind midline of metanotum (Figs. 5 &amp; 26)……………...13</paragraph>
<paragraph blockId="2.[192,747,200,1947]" pageId="2" pageNumber="138">
12. T2 with almost regular apical yellow band, about 2 irregular rows of punctures at median area of yellow band; punctures on T2 larger than that of alternative species; T1 with narrow apical yellow band, about 2-3 irregular rows of punctures at yellow band; clypeus of female black with broad yellow band at base and apex…………….. ……………….
<taxonomicName authority="Giordani Soika" authorityName="Giordani Soika" box="[424,741,1043,1070]" class="Insecta" family="Eumenidae" genus="Parancistrocerus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="138" phylum="Arthropoda" rank="species" species="vicinus">
<emphasis bold="true" box="[424,741,1043,1070]" pageId="2" pageNumber="138">
<emphasis bold="true" box="[424,543,1043,1070]" italics="true" pageId="2" pageNumber="138">P. vicinus</emphasis>
Giordani Soika
</emphasis>
</taxonomicName>
</paragraph>
<paragraph blockId="2.[192,747,200,1947]" pageId="2" pageNumber="138">
- T2 with wavy apical yellow band, about 4- 5 irregular rows of punctures at median area of yellow band; punctures on T2 smaller than that of alternative species; T1 with broad apical yellow band, about 4-6 irregular rows of punctures at yellow band; clypeus of female yellow except a brownish black spot at middle………….... ……………..
<emphasis bold="true" pageId="2" pageNumber="138">
<taxonomicName authority="Girish Kumar &amp; Carpenter" authorityName="Girish Kumar &amp; Carpenter" class="Insecta" family="Eumenidae" genus="Parancistrocerus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="138" phylum="Arthropoda" rank="species" species="jaferpaloti" status="sp. n.">
<emphasis bold="true" box="[402,560,1346,1373]" italics="true" pageId="2" pageNumber="138">P. jaferpaloti</emphasis>
Girish Kumar &amp; Carpenter
</taxonomicName>
<taxonomicNameLabel box="[410,476,1381,1407]" pageId="2" pageNumber="138" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph blockId="2.[192,747,200,1947]" pageId="2" pageNumber="138">
13. Occipital carina complete (Fig. 4); interspaces between punctures on mesopleuron and posterior portion of mesoscutum not carinate; body punctures less strong than that of alternative species; in female, cephalic fovea with a single deep pit…………..
<taxonomicName authority="Giordani Soika" authorityName="Giordani Soika" class="Insecta" family="Eumenidae" genus="Parancistrocerus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="138" phylum="Arthropoda" rank="species" species="androcles">
<emphasis bold="true" pageId="2" pageNumber="138">
<emphasis bold="true" box="[468,747,1616,1643]" italics="true" pageId="2" pageNumber="138">P. androcles androcles</emphasis>
(Meade-Waldo)
</emphasis>
</taxonomicName>
</paragraph>
<paragraph blockId="2.[192,747,200,1947]" pageId="2" pageNumber="138">
- Occipital carina not complete, evanescing at vertex (Fig. 25); interspaces between punctures on mesopleuron and posterior portion of mesoscutum carinate; body punctures stronger than that of alternative species; in female, cephalic fovea with two closely set pits………...
<emphasis bold="true" pageId="2" pageNumber="138">
<taxonomicName authority="Girish Kumar &amp; Carpenter" authorityName="Girish Kumar &amp; Carpenter" class="Insecta" family="Eumenidae" genus="Parancistrocerus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="138" phylum="Arthropoda" rank="species" species="loharbandensis" status="sp. n.">
<emphasis bold="true" box="[527,747,1885,1913]" italics="true" pageId="2" pageNumber="138">P. loharbandensis</emphasis>
Girish Kumar &amp; Carpenter
</taxonomicName>
<taxonomicNameLabel box="[595,662,1921,1947]" pageId="2" pageNumber="138" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>