283 lines
28 KiB
XML
283 lines
28 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.905.39592" ID-GBIF-Dataset="ee5a0817-7cfc-4c21-ba7d-3beabb06cccd" ID-PMC="PMC7004242" ID-Pensoft-Pub="1313-2970-905-1" ID-Pensoft-UUID="4C7E0CB428DF56BDB15B7AC8D6707961" ID-PubMed="32063726" ID-ZooBank="F4C766E4633A41039FFDE952718F41FB" ModsDocID="1313-2970-905-1" checkinTime="1579539902851" checkinUser="pensoft" docAuthor="Salata, Sebastian & Fisher, Brian L." docDate="2020" docId="3DAA2F51B5B205102782C5F093C90C0B" docLanguage="en" docName="ZooKeys 905: 1-235" docOrigin="ZooKeys 905" docSource="http://dx.doi.org/10.3897/zookeys.905.39592" docTitle="Pheidole fisaka Salata & Fisher 2020, sp. nov." docType="treatment" docUuid="2BA075C8-7EA3-444A-B35F-CE9BA0DC0798" docUuidSource="ZooBank" docVersion="5" id="4C7E0CB428DF56BDB15B7AC8D6707961" lastPageNumber="1" masterDocId="4C7E0CB428DF56BDB15B7AC8D6707961" masterDocTitle="Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups" masterLastPageNumber="235" masterPageNumber="1" pageNumber="1" updateTime="1668168337901" updateUser="ExternalLinkService">
|
|
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo>
|
|
<mods:title>Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Salata, Sebastian</mods:namePart>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Fisher, Brian L.</mods:namePart>
|
|
</mods:name>
|
|
<mods:typeOfResource>text</mods:typeOfResource>
|
|
<mods:relatedItem type="host">
|
|
<mods:titleInfo>
|
|
<mods:title>ZooKeys</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part>
|
|
<mods:date>2020</mods:date>
|
|
<mods:detail type="volume">
|
|
<mods:number>905</mods:number>
|
|
</mods:detail>
|
|
<mods:extent unit="page">
|
|
<mods:start>1</mods:start>
|
|
<mods:end>235</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:location>
|
|
<mods:url>http://dx.doi.org/10.3897/zookeys.905.39592</mods:url>
|
|
</mods:location>
|
|
<mods:classification>journal article</mods:classification>
|
|
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.905.39592</mods:identifier>
|
|
<mods:identifier type="Pensoft-Pub">1313-2970-905-1</mods:identifier>
|
|
<mods:identifier type="ZooBank">F4C766E4633A41039FFDE952718F41FB</mods:identifier>
|
|
<mods:identifier type="Pensoft-UUID">4C7E0CB428DF56BDB15B7AC8D6707961</mods:identifier>
|
|
</mods:mods>
|
|
<treatment ID-GBIF-Taxon="161777008" LSID="urn:lsid:zoobank.org:act:2BA075C8-7EA3-444A-B35F-CE9BA0DC0798" httpUri="http://treatment.plazi.org/id/3DAA2F51B5B205102782C5F093C90C0B" lastPageNumber="1" pageId="0" pageNumber="1">
|
|
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<taxonomicName LSID="http://zoobank.org/2BA075C8-7EA3-444A-B35F-CE9BA0DC0798" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole fisaka" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="fisaka" status="sp. nov.">Pheidole fisaka</taxonomicName>
|
|
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
|
|
<figureCitation captionStart="Figure 79" captionStartId="F79" captionText="Figure 79. Pheidole fisaka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923180) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0274930)." figureDoi="10.3897/zookeys.905.39592.figure79" httpUri="https://binary.pensoft.net/fig/373126" pageId="0" pageNumber="1">Figs 79A-F</figureCitation>
|
|
<figureCitation captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">, 84R</figureCitation>
|
|
<figureCitation captionStart="Figure 86" captionStartId="F86" captionText="Figure 86. Distribution. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U)." figureDoi="10.3897/zookeys.905.39592.figure86" httpUri="https://binary.pensoft.net/fig/373133" pageId="0" pageNumber="1">, 86R</figureCitation>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="type material">
|
|
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
|
|
.
|
|
</emphasis>
|
|
Madagascar. •1 major worker; Toamasina; Ankerana;
|
|
<geoCoordinate degrees="18.40062" direction="south" orientation="latitude" precision="1" value="-18.40062">-18.40062</geoCoordinate>
|
|
,
|
|
<geoCoordinate degrees="48.81311" direction="east" orientation="longitude" precision="1" value="48.81311">48.81311</geoCoordinate>
|
|
; alt. 865 m; 17 Jan 2012; B.L. Fisher et al. leg.; CASENT0274930 (CASC).
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Paratype</emphasis>
|
|
.
|
|
</emphasis>
|
|
Madagascar. •1w.; same data as for holotype; BLF27851, CASENT0923180 (CASC).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
|
|
<paragraph pageId="0" pageNumber="1">Other material.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
Madagascar. -
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Toamasina</emphasis>
|
|
: •1w., 1s.; Ankerana;
|
|
<geoCoordinate degrees="18.4061" direction="south" orientation="latitude" precision="5" value="-18.4061">-18.4061</geoCoordinate>
|
|
,
|
|
<geoCoordinate degrees="48.82029" direction="east" orientation="longitude" precision="1" value="48.82029">48.82029</geoCoordinate>
|
|
; alt. 725 m; 16 Jan 2012; B.L. Fisher et al. leg.; CASENT0273490, CASENT0273492 (CASC). •2w., 1m.; Ankerana;
|
|
<geoCoordinate degrees="18.40829" direction="south" orientation="latitude" precision="1" value="-18.40829">-18.40829</geoCoordinate>
|
|
,
|
|
<geoCoordinate degrees="48.82107" direction="east" orientation="longitude" precision="1" value="48.82107">48.82107</geoCoordinate>
|
|
; alt. 750 m; 21 Jan 2012; B.L. Fisher et al. leg.; CASENT0275257, CASENT0275258 (CASC). •3w., 1q.; Ankerana;
|
|
<geoCoordinate degrees="18.4104" direction="south" orientation="latitude" precision="5" value="-18.4104">-18.4104</geoCoordinate>
|
|
,
|
|
<geoCoordinate degrees="48.8189" direction="east" orientation="longitude" precision="5" value="48.8189">48.8189</geoCoordinate>
|
|
; alt. 855 m; 27 Jan 2012; B.L. Fisher et al. leg.; CASENT0274031, CASENT0274032, CASENT0274033 (CASC).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
|
|
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
|
|
.
|
|
</emphasis>
|
|
Small species: HL: 1.01-1.0; HW: 0.99-0.98, WL: 0.8-0.76; head in full-face view sub-rectangular, anterior of eyes straight, posterior of eyes convex; sides of the head with moderately dense, short, erect pilosity; antennal scrobes indistinct and not delimited by carinulae; scrobe surface shiny, foveolate with thick, longitudinal to irregular, and long rugae; inner hypostomal teeth distinct, moderately high, closely spaced, triangular, with rounded apex and wide base; outer hypostomal teeth thinner and narrower than inner hypostomal teeth, dentate; gaster finely shagreened; body yellowish brown.
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
|
|
.
|
|
</emphasis>
|
|
Head foveolate, genae smooth; promesonotum low, convex, short, with posterior declivity smoothly declining towards propodeum; propodeal spines minute; mesosoma foveolate, lateral sides of pronotum and propodeum with fading sculpture, katepisternum smooth.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="description">
|
|
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
|
|
Measurements (
|
|
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
|
|
= 2): HL: 1.01-1.0; HW: 0.99-0.98; SL: 0.44-0.45; EL: 0.13-0.13; WL: 0.8-0.76; PSL: 0.17-0.14; MTL: 0.440.44; PNW: 0.58-0.54; PTW: 0.16-0.14; PPW: 0.39-0.35; CI: 98.5-97.7; SI: 44.5-46.2; PSLI: 17.1-13.6; PPI: 40.6-40.5; PNI: 58.0-55.0; MTI: 44.6-45.3.
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
|
|
.
|
|
</emphasis>
|
|
In full-face view longer than wide, anterior of eyes straight, posterior of eyes convex; slightly widening posteriorly (Fig.
|
|
<figureCitation captionStart="Figure 79" captionStartId="F79" captionText="Figure 79. Pheidole fisaka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923180) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0274930)." figureDoi="10.3897/zookeys.905.39592.figure79" httpUri="https://binary.pensoft.net/fig/373126" pageId="0" pageNumber="1">79B</figureCitation>
|
|
). In lateral view sub-rectangular; ventral and dorsal faces relatively flat; dorsal face finely depressed posteriorly, forming shallow, transverse depression between frons and occipital lobes; inner hypostomal teeth visible. Sides of the head with moderately dense, short, erect pilosity; whole head with dense, short, suberect to erect pilosity. Antennal scrobes indistinct and not delimited by carinulae; scrobe surface shiny, foveolate with thick, longitudinal to irregular, and long rugae. Occipital lobes shiny, with thick, irregular rugae, interspaces with distinct, irregular rugulae; frons with moderately dense, thick and longitudinal rugae, interspaces smooth to rugo-foveolate in the posterior part; genae shiny, with dense and thin, irregular rugoreticulation; malar area with thin, longitudinal, dense rugoreticulation. Centre of clypeus shiny and smooth, lateral sides with longitudinal rugulae; median notch present, wide, and shallow; median longitudinal carina absent; lateral longitudinal carinae present. Scape, when laid back, reaching the midlength of head; pilosity suberect (Fig.
|
|
<figureCitation captionStart="Figure 79" captionStartId="F79" captionText="Figure 79. Pheidole fisaka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923180) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0274930)." figureDoi="10.3897/zookeys.905.39592.figure79" httpUri="https://binary.pensoft.net/fig/373126" pageId="0" pageNumber="1">79B, D</figureCitation>
|
|
). Inner hypostomal teeth distinct, moderately high, closely spaced, triangular, with rounded apex and wide base; outer hypostomal teeth thinner and narrower than inner hypostomal teeth, dentate (Fig.
|
|
<figureCitation captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">84R</figureCitation>
|
|
).
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
|
|
.
|
|
</emphasis>
|
|
In lateral view, promesonotum low and arched, posterior mesonotum steep, without tubercle-like projections; promesonotal groove absent; metanotal groove absent; propodeal spines short, triangular, with rounded apex and wide base; humeral area laterally well produced (Fig.
|
|
<figureCitation captionStart="Figure 79" captionStartId="F79" captionText="Figure 79. Pheidole fisaka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923180) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0274930)." figureDoi="10.3897/zookeys.905.39592.figure79" httpUri="https://binary.pensoft.net/fig/373126" pageId="0" pageNumber="1">79D</figureCitation>
|
|
). Surface shiny, with fine and moderately dense foveolae and additional rugoreticulation on the lateral sides of propodeum; dorsal surface of promesonotum with fading foveolae; katepisternum smooth. Pilosity moderately dense and long, erect (Fig.
|
|
<figureCitation captionStart="Figure 79" captionStartId="F79" captionText="Figure 79. Pheidole fisaka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923180) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0274930)." figureDoi="10.3897/zookeys.905.39592.figure79" httpUri="https://binary.pensoft.net/fig/373126" pageId="0" pageNumber="1">79D, F</figureCitation>
|
|
).
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
|
.
|
|
</emphasis>
|
|
Shiny and with fine foveolae; peduncle short, with indistinct horizontal lobes on its basal part; node relatively high, triangular, with rounded apex, in rear view node slightly dorsoventrally concave; pilosity moderately sparse and erect (Fig.
|
|
<figureCitation captionStart="Figure 79" captionStartId="F79" captionText="Figure 79. Pheidole fisaka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923180) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0274930)." figureDoi="10.3897/zookeys.905.39592.figure79" httpUri="https://binary.pensoft.net/fig/373126" pageId="0" pageNumber="1">79D, F</figureCitation>
|
|
).
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
|
|
.
|
|
</emphasis>
|
|
Shiny, finely shagreened; in dorsal view sides with moderately long, acute, and triangular projections; pilosity long, moderately long, and erect (Fig.
|
|
<figureCitation captionStart="Figure 79" captionStartId="F79" captionText="Figure 79. Pheidole fisaka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923180) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0274930)." figureDoi="10.3897/zookeys.905.39592.figure79" httpUri="https://binary.pensoft.net/fig/373126" pageId="0" pageNumber="1">79D, F</figureCitation>
|
|
).
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
|
.
|
|
</emphasis>
|
|
Shiny and finely shagreened; pilosity dense, moderately long, and erect (Fig.
|
|
<figureCitation captionStart="Figure 79" captionStartId="F79" captionText="Figure 79. Pheidole fisaka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923180) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0274930)." figureDoi="10.3897/zookeys.905.39592.figure79" httpUri="https://binary.pensoft.net/fig/373126" pageId="0" pageNumber="1">79D, F</figureCitation>
|
|
).
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
|
.
|
|
</emphasis>
|
|
Yellow; frons, occipital lobes; dorsal surface of mesosoma and gaster yellowish brown to brown (Fig.
|
|
<figureCitation captionStart="Figure 79" captionStartId="F79" captionText="Figure 79. Pheidole fisaka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923180) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0274930)." figureDoi="10.3897/zookeys.905.39592.figure79" httpUri="https://binary.pensoft.net/fig/373126" pageId="0" pageNumber="1">79D, F</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<caption doi="10.3897/zookeys.905.39592.figure79" httpUri="https://binary.pensoft.net/fig/373126" pageId="0" pageNumber="1" start="Figure 79" startId="F79">
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="1">Figure 79.</emphasis>
|
|
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole fisaka" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="fisaka">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole fisaka</emphasis>
|
|
</taxonomicName>
|
|
sp. nov., full-face view (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
|
), profile (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
|
), and dorsal view (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
|
|
) of paratype minor worker (CASENT0923180) and full-face view (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
|
), profile (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
|
|
), and dorsal view (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
|
|
) of holotype major worker (CASENT0274930).
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
|
|
Measurements (
|
|
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
|
|
= 2): HL: 0.43-0.43; HW: 0.4-0.39; SL: 0.38-0.37; EL: 0.09-0.08; WL: 0.48-0.48; PSL: 0.07-0.06; MTL: 0.28-0.29; PNW: 0.25-0.25; PTW: 0.06-0.06; PPW: 0.09-0.09; CI: 92.9-91.8; SI: 95.0-93.1; PSLI: 15.2-14.5; PPI: 68.5-63.4; PNI: 62.3-63.9; MTI: 70.5-73.0.
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
|
|
.
|
|
</emphasis>
|
|
Occipital margin straight or indistinctly concave; occipital carina absent (Fig.
|
|
<figureCitation captionStart="Figure 79" captionStartId="F79" captionText="Figure 79. Pheidole fisaka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923180) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0274930)." figureDoi="10.3897/zookeys.905.39592.figure79" httpUri="https://binary.pensoft.net/fig/373126" pageId="0" pageNumber="1">79A</figureCitation>
|
|
). Pilosity moderately dense, short, suberect to erect. Head foveolate; genae smooth. Clypeus foveolate; median longitudinal carina absent; absence of two lateral longitudinal carinae. Scape, when laid back, reaching the posterior head margin; pilosity erect (Fig.
|
|
<figureCitation captionStart="Figure 79" captionStartId="F79" captionText="Figure 79. Pheidole fisaka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923180) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0274930)." figureDoi="10.3897/zookeys.905.39592.figure79" httpUri="https://binary.pensoft.net/fig/373126" pageId="0" pageNumber="1">79A, C</figureCitation>
|
|
).
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
|
|
.
|
|
</emphasis>
|
|
In lateral view, promesonotum low, convex, short, with posterior declivity smoothly declining towards propodeum; promesonotal groove absent; metanotal groove indistinct; propodeal spines minute, triangular, with acute apex (Fig.
|
|
<figureCitation captionStart="Figure 79" captionStartId="F79" captionText="Figure 79. Pheidole fisaka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923180) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0274930)." figureDoi="10.3897/zookeys.905.39592.figure79" httpUri="https://binary.pensoft.net/fig/373126" pageId="0" pageNumber="1">79C</figureCitation>
|
|
). Sculpture foveolate; lateral sides of pronotum and propodeum with fading sculpture; katepisternum smooth. Pilosity moderately sparse, moderately long, and erect (Fig.
|
|
<figureCitation captionStart="Figure 79" captionStartId="F79" captionText="Figure 79. Pheidole fisaka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923180) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0274930)." figureDoi="10.3897/zookeys.905.39592.figure79" httpUri="https://binary.pensoft.net/fig/373126" pageId="0" pageNumber="1">79C, E</figureCitation>
|
|
).
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
|
.
|
|
</emphasis>
|
|
Peduncle very short and thin; with few short, erect setae (Fig.
|
|
<figureCitation captionStart="Figure 79" captionStartId="F79" captionText="Figure 79. Pheidole fisaka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923180) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0274930)." figureDoi="10.3897/zookeys.905.39592.figure79" httpUri="https://binary.pensoft.net/fig/373126" pageId="0" pageNumber="1">79C, E</figureCitation>
|
|
).
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
|
|
.
|
|
</emphasis>
|
|
Short, low and convex; with moderately short, erect setae (Fig.
|
|
<figureCitation captionStart="Figure 79" captionStartId="F79" captionText="Figure 79. Pheidole fisaka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923180) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0274930)." figureDoi="10.3897/zookeys.905.39592.figure79" httpUri="https://binary.pensoft.net/fig/373126" pageId="0" pageNumber="1">79C, E</figureCitation>
|
|
).
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
|
.
|
|
</emphasis>
|
|
With sparse, erect pilosity (Fig.
|
|
<figureCitation captionStart="Figure 79" captionStartId="F79" captionText="Figure 79. Pheidole fisaka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923180) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0274930)." figureDoi="10.3897/zookeys.905.39592.figure79" httpUri="https://binary.pensoft.net/fig/373126" pageId="0" pageNumber="1">79C, E</figureCitation>
|
|
).
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
|
.
|
|
</emphasis>
|
|
Yellow (Fig.
|
|
<figureCitation captionStart="Figure 79" captionStartId="F79" captionText="Figure 79. Pheidole fisaka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923180) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0274930)." figureDoi="10.3897/zookeys.905.39592.figure79" httpUri="https://binary.pensoft.net/fig/373126" pageId="0" pageNumber="1">79C, E</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="etymology">
|
|
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">Malagasy for flat, in reference for shape of the head in major workers.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
|
|
<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">The species was collected between 725-865 m in elevation, in rainforest. Nests were located in rotten logs.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="comments">
|
|
<paragraph pageId="0" pageNumber="1">Comments.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole fisaka" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="fisaka">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole fisaka</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. is most similar to
|
|
<taxonomicName lsidName="P. binara" pageId="0" pageNumber="1" rank="species" species="binara">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. binara</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
|
|
.
|
|
</emphasis>
|
|
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole fisaka" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="fisaka">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole fisaka</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. differs from
|
|
<taxonomicName lsidName="P. binara" pageId="0" pageNumber="1" rank="species" species="binara">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. binara</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. in smooth surface between rugae on frons and katepisternum and fading foveolae on propodeal dorsum.
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
|
|
.
|
|
</emphasis>
|
|
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole fisaka" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="fisaka">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole fisaka</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. differs from
|
|
<taxonomicName lsidName="P. binara" pageId="0" pageNumber="1" rank="species" species="binara">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. binara</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. in shorter and more convex promesonotum, and lateral sides of pronotum and propodeum with fading foveolae and smooth katepisternum.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |