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<document id="3DB4EAC021123ADE3260951315810284" ID-CLB-Dataset="286222" ID-DOI="10.1016/j.sajb.2020.08.029" ID-GBIF-Dataset="f1f8798f-0bdd-445d-af4d-c9c5ff3ba282" ID-ISSN="1727-9321" ID-Zenodo-Dep="10496806" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="jonas" IM.treatmentCitations_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1705063989156" checkinUser="felipe" docAuthor="Elliott, T. L. &amp; Muasya, A. M." docDate="2020" docId="351B0861B753FFF03041FEF678E6FA80" docLanguage="en" docName="SAfrJBot.135.296-316.pdf" docOrigin="South African Journal of Botany 135" docSource="http://dx.doi.org/10.1016/j.sajb.2020.08.029" docStyle="DocumentStyle:AD72C92759D651943977BAEA6E2218B4.1:SAfrJBot.2014-.journal_article" docStyleId="AD72C92759D651943977BAEA6E2218B4" docStyleName="SAfrJBot.2014-.journal_article" docStyleVersion="1" docTitle="Schoenus rigidus T. L. Elliott &amp; Muasya 2020, sp. nov." docType="treatment" docVersion="6" lastPageNumber="309" masterDocId="C9227019B75FFFFD3310FF8B7969FFE0" masterDocTitle="A taxonomic revision of the Epischoenus group of Schoenus (Cyperaceae, tribe Schoeneae)" masterLastPageNumber="316" masterPageNumber="296" pageNumber="308" updateTime="1706057461490" updateUser="ExternalLinkService">
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<mods:title id="439968ED60E48966EE5DA441107B847C">A taxonomic revision of the Epischoenus group of Schoenus (Cyperaceae, tribe Schoeneae)</mods:title>
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<mods:namePart id="F5F0CAC4A5591B9ECD1CBC3603743756">Elliott, T. L.</mods:namePart>
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<mods:namePart id="C1B2F6857D9DDC6DE11C02B6711CF820">Muasya, A. M.</mods:namePart>
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<mods:title id="F8793426753A965561FD6FEB28FDCFC3">South African Journal of Botany</mods:title>
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<mods:date id="E7A78AA181F6A2AC86B04EDB41C575A9">2020</mods:date>
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<mods:number id="61A8AF02C68CCC65D1BD9402A4C0CD88">2020-12-31</mods:number>
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<subSubSection id="F5A8EAFCB753FFF13041FEF67C25FE70" box="[849,1356,380,400]" pageId="12" pageNumber="308" type="nomenclature">
<paragraph id="BD0DB977B753FFF13041FEF67C25FE70" blockId="12.[818,1489,158,1990]" box="[849,1356,380,400]" pageId="12" pageNumber="308">
<emphasis id="8FC66565B753FFF13041FEF67A0AFE70" bold="true" box="[849,867,381,400]" pageId="12" pageNumber="308">5.</emphasis>
<taxonomicName id="7AB2C2F4B753FFF1307EFEF77D99FE70" ID-CoL="9V7FW" authority="T.L.Elliott &amp; Muasya" authorityName="T. L. Elliott &amp; Muasya" authorityYear="2020" box="[878,1264,380,400]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="12" pageNumber="308" phylum="Tracheophyta" rank="species" species="rigidus" status="sp. nov.">
<emphasis id="8FC66565B753FFF1307EFEF77D72FE70" bold="true" box="[878,1051,380,400]" italics="true" pageId="12" pageNumber="308">Schoenus rigidus</emphasis>
T.L.Elliott &amp; Muasya
</taxonomicName>
,
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<taxonomicNameLabel id="94F5D81EB753FFF137EFFEF77C25FE70" box="[1279,1356,380,400]" pageId="12" pageNumber="308" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
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<subSubSection id="F5A8EAFCB753FFF13645FEF67DBCFE1F" pageId="12" pageNumber="308" type="materials_examined">
<paragraph id="BD0DB977B753FFF13645FEF67DBCFE1F" blockId="12.[818,1489,158,1990]" pageId="12" pageNumber="308">
<materialsCitation id="0DDAB32AB753FFF13645FEF67D59FE04" collectingDate="1944-09-28" collectionCode="BOL" collectorName="Fourcade" country="South Africa" location="Montagu Pass" pageId="12" pageNumber="308" specimenCode="BOL_5450" specimenCount="1" stateProvince="Western Cape Province" typeStatus="holotype">
Type:
<collectingCountry id="C5A5F9E7B753FFF13687FEF67A05FE4C" name="South Africa" pageId="12" pageNumber="308">South Africa</collectingCountry>
,
<collectingRegion id="7F767795B753FFF13069FE127D0FFE4C" box="[889,1126,409,428]" country="South Africa" name="Western Cape" pageId="12" pageNumber="308">Western Cape Province</collectingRegion>
, 3322 (Oudtshoorn):
<location id="B86DEFACB753FFF13654FE127CA6FE4C" LSID="urn:lsid:plazi:treatment:351B0861B753FFF03041FEF678E6FA80:B86DEFACB753FFF13654FE127CA6FE4C" box="[1348,1487,409,428]" country="South Africa" name="Montagu Pass" pageId="12" pageNumber="308" stateProvince="Western Cape Province">Montagu Pass</location>
<locationDeviation id="ACBFA055B753FFF13022FE3E7D00FE28" box="[818,1129,437,456]" pageId="12" pageNumber="308">between Toll house and bridge</locationDeviation>
, (CD),
<date id="C90C9FB7B753FFF137A9FE3E7C5EFE28" box="[1209,1335,437,456]" pageId="12" pageNumber="308" value="1944-09-28">
<collectingDate id="D948665FB753FFF137A9FE3E7C5EFE28" box="[1209,1335,437,456]" pageId="12" pageNumber="308" value="1944-09-28">28 Sep 1944</collectingDate>
</date>
,
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<collectorName id="1047DCA1B753FFF13653FE3F7CF1FE28" box="[1347,1432,436,456]" pageId="12" pageNumber="308">Fourcade</collectorName>
6472
</emphasis>
(
<collectionCode id="DBA321B2B753FFF13028FE5A7A0EFE04" box="[824,871,465,484]" pageId="12" pageNumber="308">BOL</collectionCode>
! [
<specimenCode id="ED14110CB753FFF1306BFE5A7A8AFE04" box="[891,995,465,484]" pageId="12" pageNumber="308">BOL_5450</specimenCode>
],
<typeStatus id="620907D5B753FFF130E9FE5A7D40FE04" box="[1017,1065,465,484]" pageId="12" pageNumber="308">holo</typeStatus>
.;
</materialsCitation>
<materialsCitation id="0DDAB32AB753FFF1372EFE5A7AB8FDE0" collectingDate="1944-09-28" collectionCode="BOL, K, NBG" collectorName="Fourcade" country="South Africa" location="Montagu Pass" pageId="12" pageNumber="308" specimenCode="BOL_6107, NBG_0181112" specimenCount="1" stateProvince="Western Cape Province">
<collectionCode id="DBA321B2B753FFF1372EFE5A7D02FE04" box="[1086,1131,465,484]" pageId="12" pageNumber="308">BOL</collectionCode>
! [
<specimenCode id="ED14110CB753FFF13790FE5A7D81FE04" box="[1152,1256,465,484]" pageId="12" pageNumber="308">BOL_6107</specimenCode>
],
<collectionCode id="DBA321B2B753FFF137EEFE5A7C7BFE04" box="[1278,1298,465,484]" pageId="12" pageNumber="308">K</collectionCode>
! [2 sheets],
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! [
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],
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<materialsCitation id="0DDAB32AB753FFF130C5FE667DB9FE1F" box="[981,1232,492,512]" collectingDate="1944-09-28" collectionCode="PRE" collectorName="Fourcade" country="South Africa" location="Montagu Pass" pageId="12" pageNumber="308" specimenCode="PRE_0239816" specimenCount="1" stateProvince="Western Cape Province" typeStatus="isotype">
<collectionCode id="DBA321B2B753FFF130C5FE667D68FDE0" box="[981,1025,493,512]" pageId="12" pageNumber="308">PRE</collectionCode>
! [
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],
<typeStatus id="620907D5B753FFF137B7FE677DAFFE1F" box="[1191,1222,492,511]" pageId="12" pageNumber="308">iso</typeStatus>
.)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F5A8EAFCB753FFF13041FD837D69FA54" pageId="12" pageNumber="308" type="description">
<paragraph id="BD0DB977B753FFF13041FD837D69FA54" blockId="12.[818,1489,158,1990]" pageId="12" pageNumber="308">
Caespitose, perennial graminoid, aphyllopodic (
<figureCitation id="2589A5F2B753FFF13622FD837CACFDFC" box="[1330,1477,520,540]" captionStart="Fig" captionStartId="5.[100,125,892,906]" captionTargetBox="[314,1274,148,861]" captionTargetId="figure-207@5.[314,1274,148,861]" captionTargetPageId="5" captionText="Fig. 3. Examples of plants from five species of the Epischoenus group found in southern Africa, showing the corresponding inflorescences beneath each plant: A and F—S. dregeanus; B and G—S.complanatus; C and H—S. quadrangularis; D and I—S.selinae; and E and J—S.rigidus. The black scale bar adjacent to the inflorescence in L represents 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10496812" httpUri="https://zenodo.org/record/10496812/files/figure.png" pageId="12" pageNumber="308">Fig. 3E and 3J</figureCitation>
).
<emphasis id="8FC66565B753FFF13022FDA87A02FDD7" box="[818,875,547,567]" italics="true" pageId="12" pageNumber="308">Culms</emphasis>
(340
<emphasis id="8FC66565B753FFF1308EFDAF7AC7FDD7" box="[926,942,548,567]" italics="true" pageId="12" pageNumber="308"></emphasis>
)498
<emphasis id="8FC66565B753FFF130CAFDAF7A83FDD7" box="[986,1002,548,567]" italics="true" pageId="12" pageNumber="308"></emphasis>
1166(1370) £ (0.3
<emphasis id="8FC66565B753FFF137BAFDAF7DD3FDD7" box="[1194,1210,548,567]" italics="true" pageId="12" pageNumber="308"></emphasis>
)
<quantity id="7A4A1492B753FFF137D1FDAF7C54FDD8" box="[1217,1341,548,568]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="12" pageNumber="308" unit="mm" value="0.7" valueMax="0.8" valueMin="0.6">
0.6
<emphasis id="8FC66565B753FFF137CFFDAF7D86FDD7" box="[1247,1263,548,567]" italics="true" pageId="12" pageNumber="308"></emphasis>
0.8 mm
</quantity>
.
<emphasis id="8FC66565B753FFF13656FDA87CEAFDD7" box="[1350,1411,547,567]" italics="true" pageId="12" pageNumber="308">Leaves</emphasis>
usually absent, rudimentary if present, 27
<emphasis id="8FC66565B753FFF1376FFDCB7DE6FDB3" box="[1151,1167,576,595]" italics="true" pageId="12" pageNumber="308"></emphasis>
30 £
<quantity id="7A4A1492B753FFF137D5FDCB7C56FDB4" box="[1221,1343,576,596]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="12" pageNumber="308" unit="mm" value="0.65" valueMax="0.8" valueMin="0.5">
0.5
<emphasis id="8FC66565B753FFF137F3FDCB7D9AFDB3" box="[1251,1267,576,595]" italics="true" pageId="12" pageNumber="308"></emphasis>
0.8 mm
</quantity>
, straight, margins serrate.
<emphasis id="8FC66565B753FFF130A0FDD07A91FD8F" box="[944,1016,603,623]" italics="true" pageId="12" pageNumber="308">Sheaths</emphasis>
reddish-black, longitudinally striate, open, central vein terminating in a mucro up to several mm long (when rudimentary leaf not present), firm.
<emphasis id="8FC66565B753FFF13784FD187DA2FD47" box="[1172,1227,659,679]" italics="true" pageId="12" pageNumber="308">Ligule</emphasis>
usually absent, (0.6
<emphasis id="8FC66565B753FFF1369EFD1F7CF7FD47" box="[1422,1438,660,679]" italics="true" pageId="12" pageNumber="308"></emphasis>
)1.6 2.8(3.0) mm if present.
<emphasis id="8FC66565B753FFF13735FD247DF4FD23" box="[1061,1181,687,707]" italics="true" pageId="12" pageNumber="308">
In
<emphasis id="8FC66565B753FFF13727FD3B7D2AFD23" bold="true" box="[1079,1091,688,707]" italics="true" pageId="12" pageNumber="308">fl</emphasis>
orescence
</emphasis>
an erect pseudolateral panicle, (13
<emphasis id="8FC66565B753FFF13042FD477A0BFD3F" box="[850,866,716,735]" italics="true" pageId="12" pageNumber="308"></emphasis>
)1721(24) £ 3.05.0(8.0) mm, proximal rachis length (3.5
<emphasis id="8FC66565B753FFF136A7FD477CAEFD3F" box="[1463,1479,716,735]" italics="true" pageId="12" pageNumber="308"></emphasis>
) 4.87.0(9.0) mm, proximal primary inflorescence bract appressed to central axis but subsequent bracts not (
<figureCitation id="2589A5F2B753FFF137A4FC887D9EFCF7" box="[1204,1271,771,791]" captionStart="Fig" captionStartId="5.[100,125,892,906]" captionTargetBox="[314,1274,148,861]" captionTargetId="figure-207@5.[314,1274,148,861]" captionTargetPageId="5" captionText="Fig. 3. Examples of plants from five species of the Epischoenus group found in southern Africa, showing the corresponding inflorescences beneath each plant: A and F—S. dregeanus; B and G—S.complanatus; C and H—S. quadrangularis; D and I—S.selinae; and E and J—S.rigidus. The black scale bar adjacent to the inflorescence in L represents 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10496812" httpUri="https://zenodo.org/record/10496812/files/figure.png" pageId="12" pageNumber="308">Figs. 3J</figureCitation>
and
<figureCitation id="2589A5F2B753FFF13638FC8F7C2AFCF7" box="[1320,1347,772,791]" captionStart="Fig" captionStartId="7.[100,125,1009,1023]" captionTargetId="figure-461@7.[218,1370,148,978]" captionTargetPageId="7" captionText="Fig. 7. Diagrammatic representations of the inflorescences of 11 species in the Epischoenus group, with spikelets illustrated in dark grey, primary inflorescence bracts in white and chartaceous (membranaceous) lateral extensions of the primary inflorescence bracts in light grey. Illustrations are not drawn to exact scale and stippling represents hairs. Inflorescences of the following species are shown: A—S. dregeanus; B—S. complanatus; C—S. quadrangularis; D—S. selinae; E—S. rigidus; F—S. crinitus; G—S. adnatus; H—S. gracillimus; I—S. schonlandii; J—S. lucidus; and K—S.neovillosus." figureDoi="http://doi.org/10.5281/zenodo.10558713" httpUri="https://zenodo.org/record/10558713/files/figure.png" pageId="12" pageNumber="308">7E</figureCitation>
).
<emphasis id="8FC66565B753FFF13643FC897D4AFCD2" italics="true" pageId="12" pageNumber="308">
Proximal primary in
<emphasis id="8FC66565B753FFF1306CFC947AE1FCD2" bold="true" box="[892,904,799,818]" italics="true" pageId="12" pageNumber="308">fl</emphasis>
orescence bracts
</emphasis>
(15
<emphasis id="8FC66565B753FFF1375AFC947D33FCD2" box="[1098,1114,799,818]" italics="true" pageId="12" pageNumber="308"></emphasis>
)2025(32) mm long, narrow, lateral chartaceous extensions at base, involute, apex acute, exceeding length of inflorescence slightly.
<emphasis id="8FC66565B753FFF13761FCDD7DC2FC8A" box="[1137,1195,854,874]" italics="true" pageId="12" pageNumber="308">Spikes</emphasis>
2, (8
<emphasis id="8FC66565B753FFF137F0FCDC7D99FC8A" box="[1248,1264,855,874]" italics="true" pageId="12" pageNumber="308"></emphasis>
)1013(18) mm long, aggregated into clusters along rachis, mostly overlapping.
<emphasis id="8FC66565B753FFF1366EFCF97CA6FC66" box="[1406,1487,882,902]" italics="true" pageId="12" pageNumber="308">Spikelets</emphasis>
linear-lanceolate to lanceolate, 1
<emphasis id="8FC66565B753FFF1377CFC047D15FC42" box="[1132,1148,911,930]" italics="true" pageId="12" pageNumber="308"></emphasis>
4 spikelets per spike, (5.7
<emphasis id="8FC66565B753FFF13661FC047CE8FC42" box="[1393,1409,911,930]" italics="true" pageId="12" pageNumber="308"></emphasis>
)6.67.4 (8.9) £ (1.1
<emphasis id="8FC66565B753FFF130BEFC207AD7FC5E" box="[942,958,939,958]" italics="true" pageId="12" pageNumber="308"></emphasis>
)1.41.5(2.3) mm, pedicellate, colour reddish-brown, texture firm and opaque, slightly concealed by lateral chartaceous extensions of primary inflorescence bracts, varying number of sterile glumes at spikelet base (
<figureCitation id="2589A5F2B753FFF1370EFC747D09FBF2" box="[1054,1120,1023,1042]" captionStart="Fig" captionStartId="7.[100,125,1948,1962]" captionTargetBox="[178,1409,1751,1915]" captionTargetId="figure-584@7.[175,1412,1748,1918]" captionTargetPageId="7" captionText="Fig. 8. Examples of spikelets from 11 species of the Epischoenus group found in southern Africa: A—S.dregeanus; B—S.complanatus; C—S. quadrangularis; D—S. selinae; E—S.rigidus; F—S.crinitus; G—S.adnatus; H—S.gracillimus; I—S.schonlandii; J—S.lucidus; and K—S.neovillosus. The black scale bar beside K represents 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10496820" httpUri="https://zenodo.org/record/10496820/files/figure.png" pageId="12" pageNumber="308">Fig. 8E</figureCitation>
).
<emphasis id="8FC66565B753FFF1377FFC767C09FBF1" box="[1135,1376,1021,1041]" italics="true" pageId="12" pageNumber="308">Proximal spikelet prophyll</emphasis>
sometimes present, (0.7
<emphasis id="8FC66565B753FFF130BDFB907AD4FBCE" box="[941,957,1051,1070]" italics="true" pageId="12" pageNumber="308"></emphasis>
)1.52.6(3.7) mm, mucros absent.
<emphasis id="8FC66565B753FFF1360CFB927C0FFBCD" box="[1308,1382,1049,1069]" italics="true" pageId="12" pageNumber="308">Rachilla</emphasis>
(0.6)1.6 2.9(7.1) mm long.
<emphasis id="8FC66565B753FFF130FEFBBE7D5AFBA9" box="[1006,1075,1077,1097]" italics="true" pageId="12" pageNumber="308">Glumes</emphasis>
59 per spikelet, texture opaque and firm, sometimes with narrow hyaline margins, lower glumes varying in length, proximal glume (0.7)2.02.9(3.9) mm long, subproximal glume (1.1)1.72.9(5.3) mm long, upper glumes longer than basal ones, apex acuminate to obtuse.
<emphasis id="8FC66565B753FFF137AAFB2E7C2CFB59" box="[1210,1349,1189,1209]" italics="true" pageId="12" pageNumber="308">Glume mucros</emphasis>
often absent, proximal mucro 00.2(1.1) mm long, subproximal mucro
<quantity id="7A4A1492B753FFF136A7FB497AEDFB11" metricMagnitude="-4" metricUnit="m" metricValue="3.0" metricValueMax="6.0" metricValueMin="0.0" pageId="12" pageNumber="308" unit="mm" value="0.3" valueMax="0.6" valueMin="0.0">0 0.6 mm</quantity>
long.
<emphasis id="8FC66565B753FFF130C1FB567D48FB11" box="[977,1057,1245,1265]" italics="true" pageId="12" pageNumber="308">Stamens</emphasis>
3 per floret, anthers 2.83.3(3.7) mm long.
<emphasis id="8FC66565B753FFF1307CFB727ADFFAED" box="[876,950,1273,1293]" italics="true" pageId="12" pageNumber="308">Stigmas</emphasis>
3-branched, vestigial stigmas absent (i.e. yet to be observed).
<emphasis id="8FC66565B753FFF130B1FA9F7D57FAC8" box="[929,1086,1300,1320]" italics="true" pageId="12" pageNumber="308">Perianth bristles</emphasis>
absent (i.e. yet to be observed). Suprafloral axis sometimes thickening and strengthening, eventually curving.
<emphasis id="8FC66565B753FFF1309EFAC77AA0FA80" box="[910,969,1356,1376]" italics="true" pageId="12" pageNumber="308">Nutlet</emphasis>
2.32.7 £
<quantity id="7A4A1492B753FFF13755FAC67DA8FA81" box="[1093,1217,1357,1377]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.4" metricValueMin="1.1" pageId="12" pageNumber="308" unit="mm" value="1.25" valueMax="1.4" valueMin="1.1">1.11.4 mm</quantity>
, broad elliptic, trigonous, stipitate, shiny, yellowish in colour when young and turning reddish-brown with maturity; nutlet beak
<quantity id="7A4A1492B753FFF137CDFA0E7C30FA78" box="[1245,1369,1413,1432]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="12" pageNumber="308" unit="mm" value="0.2" valueMax="0.3" valueMin="0.1">0.10.3 mm</quantity>
(
<figureCitation id="2589A5F2B753FFF1367EFA0E7CACFA78" box="[1390,1477,1413,1432]" captionStart="Fig" captionStartId="9.[100,125,1925,1939]" captionTargetBox="[177,1411,1721,1889]" captionTargetId="figure-722@9.[174,1414,1718,1894]" captionTargetPageId="9" captionText="Fig. 10. Examples of nutlets from 11 species of the Epischoenus group found in southern Africa: A—S. dregeanus; B—S.complanatus; C—S. quadrangularis; D—S. selinae; E—S.rigidus; F—S.crinitus; G—S.adnatus; H—S.gracillimus; I—S.schonlandii; J—S.lucidus; and K—S.neovillosus. The black scale bar beside K represents 1 mm. Fully formed, mature nutlets of S.crinitus and S. neovillosus have yet to be observed." figureDoi="http://doi.org/10.5281/zenodo.10496826" httpUri="https://zenodo.org/record/10496826/files/figure.png" pageId="12" pageNumber="308">Fig. 10E</figureCitation>
). (
<figureCitation id="2589A5F2B753FFF1302AFA2A7A10FA54" box="[826,889,1441,1460]" captionStart="Fig" captionStartId="5.[100,125,892,906]" captionTargetBox="[314,1274,148,861]" captionTargetId="figure-207@5.[314,1274,148,861]" captionTargetPageId="5" captionText="Fig. 3. Examples of plants from five species of the Epischoenus group found in southern Africa, showing the corresponding inflorescences beneath each plant: A and F—S. dregeanus; B and G—S.complanatus; C and H—S. quadrangularis; D and I—S.selinae; and E and J—S.rigidus. The black scale bar adjacent to the inflorescence in L represents 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10496812" httpUri="https://zenodo.org/record/10496812/files/figure.png" pageId="12" pageNumber="308">Figs. 3</figureCitation>
,
<figureCitation id="2589A5F2B753FFF13096FA2A7AFBFA54" box="[902,914,1441,1460]" captionStart="Fig" captionStartId="7.[100,125,1009,1023]" captionTargetId="figure-461@7.[218,1370,148,978]" captionTargetPageId="7" captionText="Fig. 7. Diagrammatic representations of the inflorescences of 11 species in the Epischoenus group, with spikelets illustrated in dark grey, primary inflorescence bracts in white and chartaceous (membranaceous) lateral extensions of the primary inflorescence bracts in light grey. Illustrations are not drawn to exact scale and stippling represents hairs. Inflorescences of the following species are shown: A—S. dregeanus; B—S. complanatus; C—S. quadrangularis; D—S. selinae; E—S. rigidus; F—S. crinitus; G—S. adnatus; H—S. gracillimus; I—S. schonlandii; J—S. lucidus; and K—S.neovillosus." figureDoi="http://doi.org/10.5281/zenodo.10558713" httpUri="https://zenodo.org/record/10558713/files/figure.png" pageId="12" pageNumber="308">7</figureCitation>
,
<figureCitation id="2589A5F2B753FFF1308FFA2A7AC2FA54" box="[927,939,1441,1460]" captionStart="Fig" captionStartId="7.[100,125,1948,1962]" captionTargetBox="[178,1409,1751,1915]" captionTargetId="figure-584@7.[175,1412,1748,1918]" captionTargetPageId="7" captionText="Fig. 8. Examples of spikelets from 11 species of the Epischoenus group found in southern Africa: A—S.dregeanus; B—S.complanatus; C—S. quadrangularis; D—S. selinae; E—S.rigidus; F—S.crinitus; G—S.adnatus; H—S.gracillimus; I—S.schonlandii; J—S.lucidus; and K—S.neovillosus. The black scale bar beside K represents 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10496820" httpUri="https://zenodo.org/record/10496820/files/figure.png" pageId="12" pageNumber="308">8</figureCitation>
and
<figureCitation id="2589A5F2B753FFF130CFFA2A7A91FA54" box="[991,1016,1441,1460]" captionStart="Fig" captionStartId="9.[100,125,1925,1939]" captionTargetBox="[177,1411,1721,1889]" captionTargetId="figure-722@9.[174,1414,1718,1894]" captionTargetPageId="9" captionText="Fig. 10. Examples of nutlets from 11 species of the Epischoenus group found in southern Africa: A—S. dregeanus; B—S.complanatus; C—S. quadrangularis; D—S. selinae; E—S.rigidus; F—S.crinitus; G—S.adnatus; H—S.gracillimus; I—S.schonlandii; J—S.lucidus; and K—S.neovillosus. The black scale bar beside K represents 1 mm. Fully formed, mature nutlets of S.crinitus and S. neovillosus have yet to be observed." figureDoi="http://doi.org/10.5281/zenodo.10496826" httpUri="https://zenodo.org/record/10496826/files/figure.png" pageId="12" pageNumber="308">10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F5A8EAFCB753FFF13041FA377D8CF94F" pageId="12" pageNumber="308" type="biology_ecology">
<paragraph id="BD0DB977B753FFF13041FA377D59FA0C" blockId="12.[818,1489,158,1990]" pageId="12" pageNumber="308">
<emphasis id="8FC66565B753FFF13041FA377ADAFA30" box="[849,947,1468,1488]" italics="true" pageId="12" pageNumber="308">Flowering:</emphasis>
May to October, but flowering specimens have also been collected in February
</paragraph>
<paragraph id="BD0DB977B753FFF13041FA7F7D8CF94F" blockId="12.[818,1489,158,1990]" pageId="12" pageNumber="308">
<emphasis id="8FC66565B753FFF13041FA7F7D8AF9E8" box="[849,1251,1524,1544]" italics="true" pageId="12" pageNumber="308">
Distribution and ecology:
<taxonomicName id="7AB2C2F4B753FFF13756FA7F7D8AF9E8" authorityName="T. L. Elliott &amp; Muasya" authorityYear="2020" box="[1094,1251,1524,1544]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="12" pageNumber="308" phylum="Tracheophyta" rank="species" species="rigidus">Schoenus rigidus</taxonomicName>
</emphasis>
is known to occur relatively close to the southern coast of
<collectingCountry id="C5A5F9E7B753FFF1379DF99A7C6CF9C4" box="[1165,1285,1553,1572]" name="South Africa" pageId="12" pageNumber="308">South Africa</collectingCountry>
, with a reported distribution ranging from the Grahamstown area in the east to near George in the west (
<figureCitation id="2589A5F2B753FFF130E5F9C37D5DF9BB" box="[1013,1076,1608,1627]" captionStart="Fig" captionStartId="13.[269,294,723,737]" captionTargetBox="[101,769,148,692]" captionTargetId="figure-1006@13.[161,543,165,501]" captionTargetPageId="13" captionText="Fig. 15. Documented locations of S.rigidus." figureDoi="http://doi.org/10.5281/zenodo.10496840" httpUri="https://zenodo.org/record/10496840/files/figure.png" pageId="12" pageNumber="308">Fig. 15</figureCitation>
). Based on herbarium label information,
<taxonomicName id="7AB2C2F4B753FFF136D0F9CC7A1BF997" authorityName="T. L. Elliott &amp; Muasya" authorityYear="2020" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="12" pageNumber="308" phylum="Tracheophyta" rank="species" species="rigidus">
<emphasis id="8FC66565B753FFF136D0F9CC7A1BF997" italics="true" pageId="12" pageNumber="308">S. rigidus</emphasis>
</taxonomicName>
has been collected from both wet and dry habitats on mountainous slopes, flat areas and in forests. This species has a broad elevational tolerance, ranging from
<quantity id="7A4A1492B753FFF1377DF9177D89F94F" box="[1133,1248,1692,1711]" metricMagnitude="2" metricUnit="m" metricValue="7.25" metricValueMax="13.75" metricValueMin="0.75" pageId="12" pageNumber="308" unit="m" value="725.0" valueMax="1375.0" valueMin="75.0">
75
<emphasis id="8FC66565B753FFF13795F9177DFCF94F" box="[1157,1173,1692,1711]" italics="true" pageId="12" pageNumber="308"></emphasis>
1375 m
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="F5A8EAFCB753FFF13041F93C7D23F826" pageId="12" pageNumber="308" type="diagnosis">
<paragraph id="BD0DB977B753FFF13041F93C7D23F826" blockId="12.[818,1489,158,1990]" pageId="12" pageNumber="308">
<emphasis id="8FC66565B753FFF13041F93C7AD8F92B" box="[849,945,1719,1739]" italics="true" pageId="12" pageNumber="308">Diagnosis:</emphasis>
Other southern African
<taxonomicName id="7AB2C2F4B753FFF1378CF93C7C54F92B" box="[1180,1341,1719,1739]" pageId="12" pageNumber="308">
<emphasis id="8FC66565B753FFF1378CF93C7D9BF92B" box="[1180,1266,1719,1739]" italics="true" pageId="12" pageNumber="308">Schoenus</emphasis>
species
</taxonomicName>
closely resembling
<taxonomicName id="7AB2C2F4B753FFF1307AF9587AA8F907" authorityName="T. L. Elliott &amp; Muasya" authorityYear="2020" box="[874,961,1747,1767]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="12" pageNumber="308" phylum="Tracheophyta" rank="species" species="rigidus">
<emphasis id="8FC66565B753FFF1307AF9587AA8F907" box="[874,961,1747,1767]" italics="true" pageId="12" pageNumber="308">S. rigidus</emphasis>
</taxonomicName>
include
<taxonomicName id="7AB2C2F4B753FFF13708F9587D06F907" authorityName="T. L. Elliott, Muthaphuli &amp; Muasya" authorityYear="2020" box="[1048,1135,1747,1767]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="12" pageNumber="308" phylum="Tracheophyta" rank="species" species="selinae">
<emphasis id="8FC66565B753FFF13708F9587D06F907" box="[1048,1135,1747,1767]" italics="true" pageId="12" pageNumber="308">S. selinae</emphasis>
</taxonomicName>
and
<emphasis id="8FC66565B753FFF137B3F9587CA6F907" box="[1187,1487,1747,1767]" italics="true" pageId="12" pageNumber="308">
<taxonomicName id="7AB2C2F4B753FFF137B3F9587C4FF907" box="[1187,1318,1747,1767]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="12" pageNumber="308" phylum="Tracheophyta" rank="species" species="schonlandii">S. schonlandii</taxonomicName>
.
<taxonomicName id="7AB2C2F4B753FFF13621F9587CA6F907" authorityName="T. L. Elliott, Muthaphuli &amp; Muasya" authorityYear="2020" box="[1329,1487,1747,1767]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="12" pageNumber="308" phylum="Tracheophyta" rank="species" species="selinae">Schoenus selinae</taxonomicName>
</emphasis>
also has lateral chartaceous extensions on the primary inflorescence bracts, but compared to
<taxonomicName id="7AB2C2F4B753FFF13732F8817D11F8FE" authorityName="T. L. Elliott, Muthaphuli &amp; Muasya" authorityYear="2020" box="[1058,1144,1802,1822]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="12" pageNumber="308" phylum="Tracheophyta" rank="species" species="selinae">
<emphasis id="8FC66565B753FFF13732F8817D11F8FE" box="[1058,1144,1802,1822]" italics="true" pageId="12" pageNumber="308">S. selinae</emphasis>
</taxonomicName>
, the inflorescences of
<taxonomicName id="7AB2C2F4B753FFF13644F8817CC3F8FE" authorityName="T. L. Elliott &amp; Muasya" authorityYear="2020" box="[1364,1450,1802,1822]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="12" pageNumber="308" phylum="Tracheophyta" rank="species" species="rigidus">
<emphasis id="8FC66565B753FFF13644F8817CC3F8FE" box="[1364,1450,1802,1822]" italics="true" pageId="12" pageNumber="308">S. rigidus</emphasis>
</taxonomicName>
are more laterally branched. In addition, the chartaceous extensions of the primary inflorescence bracts tend to cover only the bases of the spikes in
<taxonomicName id="7AB2C2F4B753FFF13086F8D57A99F892" authorityName="T. L. Elliott &amp; Muasya" authorityYear="2020" box="[918,1008,1886,1906]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="12" pageNumber="308" phylum="Tracheophyta" rank="species" species="rigidus">
<emphasis id="8FC66565B753FFF13086F8D57A99F892" box="[918,1008,1886,1906]" italics="true" pageId="12" pageNumber="308">S. rigidus</emphasis>
</taxonomicName>
, whereas they cover more of the entire spike length in
<emphasis id="8FC66565B753FFF1309EF8F17DDBF86E" box="[910,1202,1914,1934]" italics="true" pageId="12" pageNumber="308">
<taxonomicName id="7AB2C2F4B753FFF1309EF8F17A8DF86E" authorityName="T. L. Elliott, Muthaphuli &amp; Muasya" authorityYear="2020" box="[910,996,1914,1934]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="12" pageNumber="308" phylum="Tracheophyta" rank="species" species="selinae">S. selinae</taxonomicName>
.
<taxonomicName id="7AB2C2F4B753FFF130FCF8F17DDBF86E" box="[1004,1202,1914,1934]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="12" pageNumber="308" phylum="Tracheophyta" rank="species" species="schonlandii">Schoenus schonlandii</taxonomicName>
</emphasis>
is a species partially overlapping in distribution; however, it lacks chartaceous extensions on the primary inflorescence bracts.
</paragraph>
</subSubSection>
<caption id="E9CDE9FFB752FFF0321DFD587B32FD01" ID-DOI="http://doi.org/10.5281/zenodo.10496840" ID-Zenodo-Dep="10496840" box="[269,603,721,737]" httpUri="https://zenodo.org/record/10496840/files/figure.png" pageId="13" pageNumber="309" startId="13.[269,294,723,737]" targetBox="[101,769,148,692]" targetPageId="13" targetType="figure">
<paragraph id="BD0DB977B752FFF0321DFD587B32FD01" blockId="13.[269,603,721,737]" box="[269,603,721,737]" pageId="13" pageNumber="309">
<emphasis id="8FC66565B752FFF0321DFD58782FFD01" bold="true" box="[269,326,723,737]" pageId="13" pageNumber="309">Fig. 15.</emphasis>
Documented locations of
<taxonomicName id="7AB2C2F4B752FFF03102FD5A7B3FFD01" authorityName="T. L. Elliott &amp; Muasya" authorityYear="2020" box="[530,598,721,737]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="13" pageNumber="309" phylum="Tracheophyta" rank="species" species="rigidus">S. rigidus</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="F5A8EAFCB752FFF03394FC9478E6FA80" pageId="13" pageNumber="309" type="materials_examined">
<paragraph id="BD0DB977B752FFF03394FC9478D2FCD2" blockId="13.[100,771,799,1990]" box="[132,443,799,818]" pageId="13" pageNumber="309">Additional collections examined</paragraph>
<paragraph id="BD0DB977B752FFF03394FCB078E6FA80" blockId="13.[100,771,799,1990]" pageId="13" pageNumber="309">
<materialsCitation id="0DDAB32AB752FFF03394FCB07BA1FC8A" collectingDate="1944-11-12" collectionCode="BOL" collectorName="Esterhuysen" country="South Africa" county="Willowmore" location="Smutsberg" municipality="Kouga Mts." pageId="13" pageNumber="309" specimenCount="1" stateProvince="Eastern Cape">
<collectingCountry id="C5A5F9E7B752FFF03394FCB07868FCAE" box="[132,257,827,846]" name="South Africa" pageId="13" pageNumber="309">South Africa</collectingCountry>
.
<collectingRegion id="7F767795B752FFF0321DFCB078CEFCAE" box="[269,423,827,846]" country="South Africa" name="Eastern Cape" pageId="13" pageNumber="309">EASTERN CAPE</collectingRegion>
:
<emphasis id="8FC66565B752FFF032A3FCB07BEAFCAE" bold="true" box="[435,643,827,846]" pageId="13" pageNumber="309">
3323 (
<collectingCounty id="546CC1FBB752FFF032E4FCB07B11FCAE" box="[500,632,827,846]" pageId="13" pageNumber="309">Willowmore</collectingCounty>
):
</emphasis>
<collectingMunicipality id="5D69230DB752FFF0319DFCB77B96FCAF" box="[653,767,828,847]" pageId="13" pageNumber="309">Kouga Mts.</collectingMunicipality>
, near
<location id="B86DEFACB752FFF03386FCDC7996FC8A" LSID="urn:lsid:plazi:treatment:351B0861B753FFF03041FEF678E6FA80:B86DEFACB752FFF03386FCDC7996FC8A" box="[150,255,855,874]" country="South Africa" county="Willowmore" municipality="Kouga Mts." name="Smutsberg" pageId="13" pageNumber="309" stateProvince="Eastern Cape">Smutsberg</location>
, (CA),
<date id="C90C9FB7B752FFF0325AFCDC78A0FC8A" box="[330,457,855,875]" pageId="13" pageNumber="309" value="1944-11-12">
<collectingDate id="D948665FB752FFF0325AFCDC78A0FC8A" box="[330,457,855,875]" pageId="13" pageNumber="309" value="1944-11-12">12 Nov 1944</collectingDate>
</date>
,
<emphasis id="8FC66565B752FFF032C3FCDD7BE0FC8A" box="[467,649,854,874]" italics="true" pageId="13" pageNumber="309">
<collectorName id="1047DCA1B752FFF032C3FCDD7B2AFC8A" box="[467,579,854,874]" pageId="13" pageNumber="309">Esterhuysen</collectorName>
10,781
</emphasis>
(
<collectionCode id="DBA321B2B752FFF03186FCDC7BABFC8A" box="[662,706,855,874]" country="South Africa" lsid="urn:lsid:biocol.org:col:15865" name="University of Cape Town" pageId="13" pageNumber="309" type="Herbarium">BOL</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="0DDAB32AB752FFF031C1FCDC7832FC42" collectingDate="1949-05-31" collectionCode="BOL" collectorName="Rhodes University Collection" country="South Africa" location="Paradise Kloof" municipality="Grahamstown" pageId="13" pageNumber="309" specimenCount="1" stateProvince="EASTERN CAPE">
<emphasis id="8FC66565B752FFF031C1FCDC7860FC66" bold="true" pageId="13" pageNumber="309">
3326 (
<collectingMunicipality id="5D69230DB752FFF0337BFCF87996FC66" box="[107,255,883,902]" pageId="13" pageNumber="309">Grahamstown</collectingMunicipality>
):
</emphasis>
<location id="B86DEFACB752FFF03201FCF878F4FC66" LSID="urn:lsid:plazi:treatment:351B0861B753FFF03041FEF678E6FA80:B86DEFACB752FFF03201FCF878F4FC66" box="[273,413,883,902]" country="South Africa" municipality="Grahamstown" name="Paradise Kloof" pageId="13" pageNumber="309" stateProvince="EASTERN CAPE">Paradise Kloof</location>
, (BC),
<date id="C90C9FB7B752FFF032F7FCF87B01FC66" box="[487,616,883,902]" pageId="13" pageNumber="309" value="1949-05-31">
<collectingDate id="D948665FB752FFF032F7FCF87B01FC66" box="[487,616,883,902]" pageId="13" pageNumber="309" value="1949-05-31">31 May 1949</collectingDate>
</date>
,
<emphasis id="8FC66565B752FFF03162FCF97872FC42" italics="true" pageId="13" pageNumber="309">
<collectorName id="1047DCA1B752FFF03162FCF9798FFC42" pageId="13" pageNumber="309">Rhodes University Collection</collectorName>
2910
</emphasis>
(
<collectionCode id="DBA321B2B752FFF03237FC04783CFC42" box="[295,341,911,930]" country="South Africa" lsid="urn:lsid:biocol.org:col:15865" name="University of Cape Town" pageId="13" pageNumber="309" type="Herbarium">BOL</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="0DDAB32AB752FFF03274FC047B63FC5E" collectingDate="1949-09-25" collectionCode="BOL" collectorName="Rhodes University" country="South Africa" location="Howieson's Poort" pageId="13" pageNumber="309" specimenCount="1" stateProvince="EASTERN CAPE">
kloof leading into
<location id="B86DEFACB752FFF03104FC047BA9FC42" LSID="urn:lsid:plazi:treatment:351B0861B753FFF03041FEF678E6FA80:B86DEFACB752FFF03104FC047BA9FC42" box="[532,704,911,930]" country="South Africa" name="Howieson's Poort" pageId="13" pageNumber="309" stateProvince="EASTERN CAPE">Howieson's Poort</location>
, (BC),
<date id="C90C9FB7B752FFF03374FC207989FC5E" box="[100,224,939,958]" pageId="13" pageNumber="309" value="1949-09-25">
<collectingDate id="D948665FB752FFF03374FC207989FC5E" box="[100,224,939,958]" pageId="13" pageNumber="309" value="1949-09-25">25 Sep 1949</collectingDate>
</date>
,
<emphasis id="8FC66565B752FFF033FAFC2178A0FC5E" box="[234,457,938,958]" italics="true" pageId="13" pageNumber="309">
<collectorName id="1047DCA1B752FFF033FAFC2178FAFC5E" box="[234,403,938,958]" pageId="13" pageNumber="309">Rhodes University</collectorName>
3102
</emphasis>
(
<collectionCode id="DBA321B2B752FFF032C6FC207B6DFC5E" box="[470,516,939,958]" country="South Africa" lsid="urn:lsid:biocol.org:col:15865" name="University of Cape Town" pageId="13" pageNumber="309" type="Herbarium">BOL</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="0DDAB32AB752FFF03104FC207B97FC3A" collectingDate="1949-09-25" collectionCode="GRA" collectorName="Martin" country="South Africa" location="Osmunda Kloof" pageId="13" pageNumber="309" specimenCount="1" stateProvince="EASTERN CAPE">
<location id="B86DEFACB752FFF03104FC207BC6FC5E" LSID="urn:lsid:plazi:treatment:351B0861B753FFF03041FEF678E6FA80:B86DEFACB752FFF03104FC207BC6FC5E" box="[532,687,939,958]" country="South Africa" name="Osmunda Kloof" pageId="13" pageNumber="309" stateProvince="EASTERN CAPE">Osmunda Kloof</location>
; marshy area near head of kloof, (BC),
<date id="C90C9FB7B752FFF032BBFC4C7B47FC3A" box="[427,558,967,986]" pageId="13" pageNumber="309" value="1949-09-25">
<collectingDate id="D948665FB752FFF032BBFC4C7B47FC3A" box="[427,558,967,986]" pageId="13" pageNumber="309" value="1949-09-25">25 Sep 1949</collectingDate>
</date>
,
<emphasis id="8FC66565B752FFF0312DFC4D7BDFFC3A" box="[573,694,966,986]" italics="true" pageId="13" pageNumber="309">
<collectorName id="1047DCA1B752FFF0312DFC4D7B14FC3A" box="[573,637,966,986]" pageId="13" pageNumber="309">Martin</collectorName>
3102
</emphasis>
(
<collectionCode id="DBA321B2B752FFF031D7FC4C7B91FC3A" box="[711,760,967,986]" country="South Africa" lsid="urn:lsid:biocol.org:col:14985" name="Albany Museum" pageId="13" pageNumber="309" type="Herbarium">GRA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="0DDAB32AB752FFF03374FC687873FBF2" collectingDate="2017-11-08" collectionCode="BOL" collectorName="Elliott &amp; Betz TE" country="South Africa" location="Osmunda Kloof" pageId="13" pageNumber="309" specimenCount="1" stateProvince="EASTERN CAPE">
<location id="B86DEFACB752FFF03374FC687868FC16" LSID="urn:lsid:plazi:treatment:351B0861B753FFF03041FEF678E6FA80:B86DEFACB752FFF03374FC687868FC16" box="[100,257,995,1014]" country="South Africa" name="Osmunda Kloof" pageId="13" pageNumber="309" stateProvince="EASTERN CAPE">Osmunda Kloof</location>
, S-facing slope, (BC),
<date id="C90C9FB7B752FFF032E1FC687B07FC16" box="[497,622,995,1014]" pageId="13" pageNumber="309" value="2017-11-08">
<collectingDate id="D948665FB752FFF032E1FC687B07FC16" box="[497,622,995,1014]" pageId="13" pageNumber="309" value="2017-11-08">8 Nov 2017</collectingDate>
</date>
,
<emphasis id="8FC66565B752FFF0316DFC6979BEFBF1" italics="true" pageId="13" pageNumber="309">
<collectorName id="1047DCA1B752FFF0316DFC697BDDFC16" box="[637,692,994,1014]" pageId="13" pageNumber="309">Elliott</collectorName>
&amp;
<collectorName id="1047DCA1B752FFF031CBFC697915FBF1" pageId="13" pageNumber="309">Betz TE</collectorName>
2016_257
</emphasis>
(
<collectionCode id="DBA321B2B752FFF033F8FC74787DFBF2" box="[232,276,1023,1042]" country="South Africa" lsid="urn:lsid:biocol.org:col:15865" name="University of Cape Town" pageId="13" pageNumber="309" type="Herbarium">BOL</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="0DDAB32AB752FFF03238FC767B06FBF2" box="[296,623,1021,1042]" collectionCode="BOL" collectorName="Elliott &amp; Betz TE" country="South Africa" pageId="13" pageNumber="309" specimenCount="1" stateProvince="EASTERN CAPE">
<emphasis id="8FC66565B752FFF03238FC767B45FBF1" box="[296,556,1021,1041]" italics="true" pageId="13" pageNumber="309">
<collectorName id="1047DCA1B752FFF03238FC767836FBF1" box="[296,351,1021,1041]" pageId="13" pageNumber="309">Elliott</collectorName>
&amp;
<collectorName id="1047DCA1B752FFF03296FC7678B8FBF1" box="[390,465,1021,1041]" pageId="13" pageNumber="309">Betz TE</collectorName>
2016_258
</emphasis>
(
<collectionCode id="DBA321B2B752FFF0312DFC747B00FBF2" box="[573,617,1023,1042]" country="South Africa" lsid="urn:lsid:biocol.org:col:15865" name="University of Cape Town" pageId="13" pageNumber="309" type="Herbarium">BOL</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="0DDAB32AB752FFF0316DFC76787FFBCE" collectionCode="BOL" collectorName="Elliott &amp; Betz TE" country="South Africa" pageId="13" pageNumber="309" specimenCount="1" stateProvince="EASTERN CAPE">
<emphasis id="8FC66565B752FFF0316DFC7679BEFBCD" italics="true" pageId="13" pageNumber="309">
<collectorName id="1047DCA1B752FFF0316DFC767BDDFBF1" box="[637,692,1021,1041]" pageId="13" pageNumber="309">Elliott</collectorName>
&amp;
<collectorName id="1047DCA1B752FFF031CBFC767915FBCD" pageId="13" pageNumber="309">Betz TE</collectorName>
2016_259
</emphasis>
(
<collectionCode id="DBA321B2B752FFF033F4FB907879FBCE" box="[228,272,1051,1070]" country="South Africa" lsid="urn:lsid:biocol.org:col:15865" name="University of Cape Town" pageId="13" pageNumber="309" type="Herbarium">BOL</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="0DDAB32AB752FFF0320FFB917B6CFB85" collectingDate="1921-06" collectingDateMax="1982-09-13" collectingDateMin="1921-06" collectionCode="NU" collectorName="Fourcade &amp; Getliffe" country="South Africa" county="Humansdorp" location="Storms River" municipality="Witte-Els Bosch" pageId="13" pageNumber="309" specimenCount="1" stateProvince="EASTERN CAPE">
<emphasis id="8FC66565B752FFF0320FFB91789BFBCD" bold="true" box="[287,498,1050,1069]" pageId="13" pageNumber="309">
3424 (
<collectingCounty id="546CC1FBB752FFF0324DFB91788EFBCD" box="[349,487,1050,1069]" pageId="13" pageNumber="309">Humansdorp</collectingCounty>
):
</emphasis>
<collectingMunicipality id="5D69230DB752FFF032E9FB917BFCFBCD" box="[505,661,1050,1069]" pageId="13" pageNumber="309">Witte-Els Bosch</collectingMunicipality>
, (AA),
<date id="C90C9FB7B752FFF031F2FB9079FFFBA9" pageId="13" pageNumber="309" value="1921-06">
<collectingDate id="D948665FB752FFF031F2FB9079FFFBA9" pageId="13" pageNumber="309" value="1921-06">Jun 1921</collectingDate>
</date>
,
<emphasis id="8FC66565B752FFF033B0FBBE7843FBA9" box="[160,298,1077,1097]" italics="true" pageId="13" pageNumber="309">
<collectorName id="1047DCA1B752FFF033B0FBBE799CFBA9" box="[160,245,1077,1097]" pageId="13" pageNumber="309">Fourcade</collectorName>
1374
</emphasis>
(BOL [2 sheets], K); P.E. to
<location id="B86DEFACB752FFF03123FBBD7BD8FBA9" LSID="urn:lsid:plazi:treatment:351B0861B753FFF03041FEF678E6FA80:B86DEFACB752FFF03123FBBD7BD8FBA9" box="[563,689,1078,1097]" country="South Africa" county="Humansdorp" municipality="Witte-Els Bosch" name="Storms River" pageId="13" pageNumber="309" stateProvince="EASTERN CAPE">Storms River</location>
,
<location id="B86DEFACB752FFF031AAFBBD79EEFB85" LSID="urn:lsid:plazi:treatment:351B0861B753FFF03041FEF678E6FA80:B86DEFACB752FFF031AAFBBD79EEFB85" country="South Africa" county="Humansdorp" municipality="Witte-Els Bosch" name="Witelsbos" pageId="13" pageNumber="309" stateProvince="EASTERN CAPE">Witelsbos</location>
, (AA),
<date id="C90C9FB7B752FFF033C3FBD97825FB85" box="[211,332,1106,1125]" pageId="13" pageNumber="309" value="1982-09-13">
<collectingDate id="D948665FB752FFF033C3FBD97825FB85" box="[211,332,1106,1125]" pageId="13" pageNumber="309" value="1982-09-13">13 Sep 1982</collectingDate>
</date>
,
<emphasis id="8FC66565B752FFF03246FBDA78A5FB85" box="[342,460,1105,1125]" italics="true" pageId="13" pageNumber="309">
<collectorName id="1047DCA1B752FFF03246FBDA78F1FB85" box="[342,408,1105,1125]" pageId="13" pageNumber="309">Getliffe</collectorName>
1155
</emphasis>
(
<collectionCode id="DBA321B2B752FFF032C9FBD97897FB85" box="[473,510,1106,1125]" country="Thailand" name="Department of Microbiology, Faculty of Science" pageId="13" pageNumber="309">NU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="0DDAB32AB752FFF0311EFBD878CAFB61" collectingDate="1955-07-03" collectionCode="BOL" collectorName="Levyns" country="South Africa" location="Storms River" pageId="13" pageNumber="309" specimenCount="1" stateProvince="EASTERN CAPE">
near
<location id="B86DEFACB752FFF03150FBD97BD7FB85" LSID="urn:lsid:plazi:treatment:351B0861B753FFF03041FEF678E6FA80:B86DEFACB752FFF03150FBD97BD7FB85" box="[576,702,1106,1125]" country="South Africa" name="Storms River" pageId="13" pageNumber="309" stateProvince="EASTERN CAPE">Storms River</location>
, (BA),
<date id="C90C9FB7B752FFF03374FBE579A4FB61" box="[100,205,1134,1153]" pageId="13" pageNumber="309" value="1955-07-03">
<collectingDate id="D948665FB752FFF03374FBE579A4FB61" box="[100,205,1134,1153]" pageId="13" pageNumber="309" value="1955-07-03">3 Jul 1955</collectingDate>
</date>
,
<emphasis id="8FC66565B752FFF033CAFBE67808FB61" box="[218,353,1133,1153]" italics="true" pageId="13" pageNumber="309">
<collectorName id="1047DCA1B752FFF033CAFBE67870FB61" box="[218,281,1133,1153]" pageId="13" pageNumber="309">Levyns</collectorName>
10,245
</emphasis>
(
<collectionCode id="DBA321B2B752FFF03260FBE578F7FB61" box="[368,414,1134,1153]" country="South Africa" lsid="urn:lsid:biocol.org:col:15865" name="University of Cape Town" pageId="13" pageNumber="309" type="Herbarium">BOL</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="0DDAB32AB752FFF032BFFBE5780DFB7D" collectingDate="1956-07-24" collectionCode="BOL" collectorName="Levyns" country="South Africa" location="Storms River" pageId="13" pageNumber="309" specimenCount="1" stateProvince="EASTERN CAPE">
<location id="B86DEFACB752FFF032BFFBE57B47FB61" LSID="urn:lsid:plazi:treatment:351B0861B753FFF03041FEF678E6FA80:B86DEFACB752FFF032BFFBE57B47FB61" box="[431,558,1134,1153]" country="South Africa" name="Storms River" pageId="13" pageNumber="309" stateProvince="EASTERN CAPE">Storms River</location>
Bridge, (BA),
<date id="C90C9FB7B752FFF031D7FBE579FFFB7D" pageId="13" pageNumber="309" value="1956-07-24">
<collectingDate id="D948665FB752FFF031D7FBE579FFFB7D" pageId="13" pageNumber="309" value="1956-07-24">24 Jul 1956</collectingDate>
</date>
,
<emphasis id="8FC66565B752FFF033B0FB02784DFB7D" box="[160,292,1161,1181]" italics="true" pageId="13" pageNumber="309">
<collectorName id="1047DCA1B752FFF033B0FB0279B6FB7D" box="[160,223,1161,1181]" pageId="13" pageNumber="309">Levyns</collectorName>
10,483
</emphasis>
(
<collectionCode id="DBA321B2B752FFF03220FB017837FB7D" box="[304,350,1162,1181]" country="South Africa" lsid="urn:lsid:biocol.org:col:15865" name="University of Cape Town" pageId="13" pageNumber="309" type="Herbarium">BOL</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="0DDAB32AB752FFF0327DFB0179D1FB59" collectingDate="1979-04-17" collectionCode="PRE" collectorName="de Kock" country="South Africa" location="Humansdorp" pageId="13" pageNumber="309" specimenCount="1" stateProvince="EASTERN CAPE">
<location id="B86DEFACB752FFF0327DFB017884FB7D" LSID="urn:lsid:plazi:treatment:351B0861B753FFF03041FEF678E6FA80:B86DEFACB752FFF0327DFB017884FB7D" box="[365,493,1162,1181]" country="South Africa" name="Humansdorp" pageId="13" pageNumber="309" stateProvince="EASTERN CAPE">Humansdorp</location>
, (BB),
<date id="C90C9FB7B752FFF03126FB017BC6FB7D" box="[566,687,1162,1181]" pageId="13" pageNumber="309" value="1979-04-17">
<collectingDate id="D948665FB752FFF03126FB017BC6FB7D" box="[566,687,1162,1181]" pageId="13" pageNumber="309" value="1979-04-17">17 Apr 1979</collectingDate>
</date>
,
<emphasis id="8FC66565B752FFF031A9FB027912FB59" italics="true" pageId="13" pageNumber="309">
<collectorName id="1047DCA1B752FFF031A9FB027A6BFB7D" box="[697,770,1161,1181]" pageId="13" pageNumber="309">de Kock</collectorName>
20
</emphasis>
(
<collectionCode id="DBA321B2B752FFF03398FB2D79DAFB59" box="[136,179,1190,1209]" country="South Africa" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="13" pageNumber="309" type="Herbarium">PRE</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="0DDAB32AB752FFF033D1FB2D79F7FB11" collectingDate="1944-11-21" collectingDateMax="1956-07-25" collectingDateMin="1944-11-21" collectionCode="BOL" collectorName="Esterhuysen &amp; Levyns" country="South Africa" location="Montagu Pass" municipality="Oudtshoorn" pageId="13" pageNumber="309" specimenCount="1" stateProvince="Western Cape">
<collectingRegion id="7F767795B752FFF033D1FB2D7836FB59" box="[193,351,1190,1209]" country="South Africa" name="Western Cape" pageId="13" pageNumber="309">WESTERN CAPE</collectingRegion>
:
<emphasis id="8FC66565B752FFF03278FB2D7B46FB59" bold="true" box="[360,559,1190,1209]" pageId="13" pageNumber="309">
3322 (
<collectingMunicipality id="5D69230DB752FFF032B5FB2D7B4DFB59" box="[421,548,1190,1209]" pageId="13" pageNumber="309">Oudtshoorn</collectingMunicipality>
):
</emphasis>
<location id="B86DEFACB752FFF03125FB2D7BD4FB59" LSID="urn:lsid:plazi:treatment:351B0861B753FFF03041FEF678E6FA80:B86DEFACB752FFF03125FB2D7BD4FB59" box="[565,701,1190,1209]" country="South Africa" municipality="Oudtshoorn" name="Montagu Pass" pageId="13" pageNumber="309" stateProvince="Western Cape">Montagu Pass</location>
, (CD),
<date id="C90C9FB7B752FFF03374FB49798BFB35" box="[100,226,1218,1237]" pageId="13" pageNumber="309" value="1944-11-21">
<collectingDate id="D948665FB752FFF03374FB49798BFB35" box="[100,226,1218,1237]" pageId="13" pageNumber="309" value="1944-11-21">21 Nov 1944</collectingDate>
</date>
,
<emphasis id="8FC66565B752FFF033FCFB4A78C8FB35" box="[236,417,1217,1237]" italics="true" pageId="13" pageNumber="309">
<collectorName id="1047DCA1B752FFF033FCFB4A7835FB35" box="[236,348,1217,1237]" pageId="13" pageNumber="309">Esterhuysen</collectorName>
10,864
</emphasis>
(BOL, K);
<date id="C90C9FB7B752FFF03114FB497B1DFB35" box="[516,628,1218,1237]" pageId="13" pageNumber="309" value="1956-07-25">
<collectingDate id="D948665FB752FFF03114FB497B1DFB35" box="[516,628,1218,1237]" pageId="13" pageNumber="309" value="1956-07-25">25 Jul 1956</collectingDate>
</date>
,
<emphasis id="8FC66565B752FFF0316DFB4A7A68FB35" box="[637,769,1217,1237]" italics="true" pageId="13" pageNumber="309">
<collectorName id="1047DCA1B752FFF0316DFB4A7BD5FB35" box="[637,700,1217,1237]" pageId="13" pageNumber="309">Levyns</collectorName>
10,502
</emphasis>
(
<collectionCode id="DBA321B2B752FFF0337AFB5579F1FB11" box="[106,152,1246,1265]" country="South Africa" lsid="urn:lsid:biocol.org:col:15865" name="University of Cape Town" pageId="13" pageNumber="309" type="Herbarium">BOL</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="0DDAB32AB752FFF033BDFB5579F5FAED" collectingDate="1956-08-03" collectionCode="BOL" collectorName="Levyns" country="South Africa" location="Forest Reserve" municipality="George" pageId="13" pageNumber="309" specimenCount="1" stateProvince="WESTERN CAPE">
<collectingMunicipality id="5D69230DB752FFF033BDFB55799AFB11" box="[173,243,1246,1265]" pageId="13" pageNumber="309">George</collectingMunicipality>
,
<location id="B86DEFACB752FFF03213FB5578FEFB11" LSID="urn:lsid:plazi:treatment:351B0861B753FFF03041FEF678E6FA80:B86DEFACB752FFF03213FB5578FEFB11" box="[259,407,1246,1265]" country="South Africa" municipality="George" name="Forest Reserve" pageId="13" pageNumber="309" stateProvince="WESTERN CAPE">Forest Reserve</location>
, (CD),
<date id="C90C9FB7B752FFF032FDFB557B00FB11" box="[493,617,1246,1265]" pageId="13" pageNumber="309" value="1956-08-03">
<collectingDate id="D948665FB752FFF032FDFB557B00FB11" box="[493,617,1246,1265]" pageId="13" pageNumber="309" value="1956-08-03">3 Aug 1956</collectingDate>
</date>
,
<emphasis id="8FC66565B752FFF03168FB567A68FB11" box="[632,769,1245,1265]" italics="true" pageId="13" pageNumber="309">
<collectorName id="1047DCA1B752FFF03168FB567BDEFB11" box="[632,695,1245,1265]" pageId="13" pageNumber="309">Levyns</collectorName>
10,575
</emphasis>
(
<collectionCode id="DBA321B2B752FFF0337AFB7179FFFAED" box="[106,150,1274,1293]" country="South Africa" lsid="urn:lsid:biocol.org:col:15865" name="University of Cape Town" pageId="13" pageNumber="309" type="Herbarium">BOL</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="0DDAB32AB752FFF033B5FB717B8CFAED" box="[165,741,1273,1293]" collectingDate="1925-04" collectionCode="GRA" collectorName="Phillips" country="South Africa" location="Noetzie" municipality="Knysna" pageId="13" pageNumber="309" specimenCount="1" stateProvince="WESTERN CAPE">
<emphasis id="8FC66565B752FFF033B5FB717855FAEC" bold="true" box="[165,316,1273,1293]" pageId="13" pageNumber="309">
3423 (
<collectingMunicipality id="5D69230DB752FFF033F2FB72785BFAEC" box="[226,306,1273,1292]" pageId="13" pageNumber="309">Knysna</collectingMunicipality>
):
</emphasis>
<location id="B86DEFACB752FFF03253FB7178E7FAED" LSID="urn:lsid:plazi:treatment:351B0861B753FFF03041FEF678E6FA80:B86DEFACB752FFF03253FB7178E7FAED" box="[323,398,1274,1293]" country="South Africa" municipality="Knysna" name="Noetzie" pageId="13" pageNumber="309" stateProvince="WESTERN CAPE">Noetzie</location>
, (AA),
<date id="C90C9FB7B752FFF032CAFB717B5FFAED" box="[474,566,1274,1293]" pageId="13" pageNumber="309" value="1925-04">
<collectingDate id="D948665FB752FFF032CAFB717B5FFAED" box="[474,566,1274,1293]" pageId="13" pageNumber="309" value="1925-04">Apr 1925</collectingDate>
</date>
,
<emphasis id="8FC66565B752FFF03150FB727BC8FAED" box="[576,673,1273,1293]" italics="true" pageId="13" pageNumber="309">
<collectorName id="1047DCA1B752FFF03150FB727BEAFAED" box="[576,643,1273,1293]" pageId="13" pageNumber="309">Phillips</collectorName>
8A
</emphasis>
(
<collectionCode id="DBA321B2B752FFF031BEFB717BB6FAED" box="[686,735,1274,1293]" country="South Africa" lsid="urn:lsid:biocol.org:col:14985" name="Albany Museum" pageId="13" pageNumber="309" type="Herbarium">GRA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="0DDAB32AB752FFF031FFFB717BACFAC9" collectingDate="1946-07" collectionCode="BOL" collectorName="Levyns" country="South Africa" location="Brackenhill Falls" pageId="13" pageNumber="309" specimenCount="1" stateProvince="WESTERN CAPE">
in forest near
<location id="B86DEFACB752FFF033C2FA9E7818FAC8" LSID="urn:lsid:plazi:treatment:351B0861B753FFF03041FEF678E6FA80:B86DEFACB752FFF033C2FA9E7818FAC8" box="[210,369,1301,1320]" country="South Africa" name="Brackenhill Falls" pageId="13" pageNumber="309" stateProvince="WESTERN CAPE">Brackenhill Falls</location>
, (AA),
<date id="C90C9FB7B752FFF032AAFA9E7B62FAC8" box="[442,523,1301,1320]" pageId="13" pageNumber="309" value="1946-07">
<collectingDate id="D948665FB752FFF032AAFA9E7B62FAC8" box="[442,523,1301,1320]" pageId="13" pageNumber="309" value="1946-07">Jul 1946</collectingDate>
</date>
,
<emphasis id="8FC66565B752FFF03104FA9F7BEEFAC8" box="[532,647,1300,1320]" italics="true" pageId="13" pageNumber="309">
<collectorName id="1047DCA1B752FFF03104FA9F7B3AFAC8" box="[532,595,1300,1320]" pageId="13" pageNumber="309">Levyns</collectorName>
7861
</emphasis>
(
<collectionCode id="DBA321B2B752FFF03182FA9D7BA9FAC9" box="[658,704,1302,1321]" country="South Africa" lsid="urn:lsid:biocol.org:col:15865" name="University of Cape Town" pageId="13" pageNumber="309" type="Herbarium">BOL</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="0DDAB32AB752FFF031DEFA9D7B5BFAA5" collectingDate="1946-07-13" collectionCode="BOL" collectorName="Levyns" country="South Africa" location="Sparrebosch" pageId="13" pageNumber="309" specimenCount="1" stateProvince="WESTERN CAPE">
<location id="B86DEFACB752FFF031DEFA9D79D8FAA4" LSID="urn:lsid:plazi:treatment:351B0861B753FFF03041FEF678E6FA80:B86DEFACB752FFF031DEFA9D79D8FAA4" country="South Africa" name="Sparrebosch" pageId="13" pageNumber="309" stateProvince="WESTERN CAPE">Sparrebosch</location>
, (AA),
<date id="C90C9FB7B752FFF033EFFAB9781BFAA4" box="[255,370,1329,1349]" pageId="13" pageNumber="309" value="1946-07-13">
<collectingDate id="D948665FB752FFF033EFFAB9781BFAA4" box="[255,370,1329,1349]" pageId="13" pageNumber="309" value="1946-07-13">13 Jul 1946</collectingDate>
</date>
,
<emphasis id="8FC66565B752FFF0326DFABB789BFAA4" box="[381,498,1328,1348]" italics="true" pageId="13" pageNumber="309">
<collectorName id="1047DCA1B752FFF0326DFABB78D5FAA4" box="[381,444,1328,1348]" pageId="13" pageNumber="309">Levyns</collectorName>
7868
</emphasis>
(
<collectionCode id="DBA321B2B752FFF032EFFAB97B44FAA5" box="[511,557,1330,1349]" country="South Africa" lsid="urn:lsid:biocol.org:col:15865" name="University of Cape Town" pageId="13" pageNumber="309" type="Herbarium">BOL</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="0DDAB32AB752FFF0312DFAB978E2FA80" collectingDate="1951-07-30" collectionCode="BOL" collectorName="Levyns" country="South Africa" location="Noetzie" pageId="13" pageNumber="309" specimenCount="1" stateProvince="WESTERN CAPE">
near
<location id="B86DEFACB752FFF03160FABA7BD2FAA4" LSID="urn:lsid:plazi:treatment:351B0861B753FFF03041FEF678E6FA80:B86DEFACB752FFF03160FABA7BD2FAA4" box="[624,699,1329,1348]" country="South Africa" name="Noetzie" pageId="13" pageNumber="309" stateProvince="WESTERN CAPE">Noetzie</location>
, (AA),
<date id="C90C9FB7B752FFF03374FAC679BBFA80" box="[100,210,1357,1376]" pageId="13" pageNumber="309" value="1951-07-30">
<collectingDate id="D948665FB752FFF03374FAC679BBFA80" box="[100,210,1357,1376]" pageId="13" pageNumber="309" value="1951-07-30">30 Jul 1951</collectingDate>
</date>
,
<emphasis id="8FC66565B752FFF033CBFAC77827FA80" box="[219,334,1356,1376]" italics="true" pageId="13" pageNumber="309">
<collectorName id="1047DCA1B752FFF033CBFAC77873FA80" box="[219,282,1356,1376]" pageId="13" pageNumber="309">Levyns</collectorName>
9642
</emphasis>
(
<collectionCode id="DBA321B2B752FFF03249FAC678ECFA80" box="[345,389,1357,1376]" country="South Africa" lsid="urn:lsid:biocol.org:col:15865" name="University of Cape Town" pageId="13" pageNumber="309" type="Herbarium">BOL</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>