335 lines
34 KiB
XML
335 lines
34 KiB
XML
<document id="74AC99DD69688208F671FEE0F76DE03C" ID-DOI="10.11646/zootaxa.3941.3.7" ID-GBIF-Dataset="71fc4faa-0893-495f-8b83-dd18e0e0553e" ID-ISSN="1175-5326" ID-Zenodo-Dep="237243" ID-ZooBank="1F01CF04-B6BD-4A80-A779-F3A22BEA4564" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461026698890" checkinUser="plazi" docAuthor="Phan, Quoc Toan, Sasamoto, Akihiko & Hayashi, Fumio" docDate="2015" docId="2D280442E268FFEBFF436CF5FBF857FE" docLanguage="en" docName="zt03941p420.pdf" docOrigin="Zootaxa 3941 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Devadatta kompieri Phan, Sasamoto & Hayashi, 2015, sp. nov." docType="treatment" docUuid="B09F287C-621F-424F-8668-90B870E94CA0" docUuidSource="ZooBank" docVersion="8" lastPageNumber="418" masterDocId="D1117C3AE269FFEFFFD46C62FFC1560A" masterDocTitle="Description of two new species of the genus Devadatta from northern Vietnam and central Laos (Odonata: Devadattidae)" masterLastPageNumber="420" masterPageNumber="414" pageNumber="415" updateTime="1698572348918" updateUser="plazi">
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<mods:mods id="EBD3E03DA74BF99FF46C260E04AC045B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="E5FBA618CCDC79C2373CED9BB7C3F26C">
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<mods:title id="3BA8B878E33FE4F9BF665354B5826B3A">Description of two new species of the genus Devadatta from northern Vietnam and central Laos (Odonata: Devadattidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="0988BC5ABB19DB3CB1454ECBA85A2306" type="personal">
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<mods:roleTerm id="7EA80BA9727C8E511B2CDEC08D0063D0">Author</mods:roleTerm>
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<mods:namePart id="0B2C0F3EBCF2F3F01DDB7FEEF8532774">Phan, Quoc Toan</mods:namePart>
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<mods:roleTerm id="61167E16D3536FE2A7620BC5AAD536A0">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3D2996DBA103B63C7DDA7FB010C12395">Sasamoto, Akihiko</mods:namePart>
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</mods:name>
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<mods:name id="72BF940FBE74E6F04B96595FFD6C83D3" type="personal">
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<mods:role id="28EF6564817BBF96A1D155E01FB0821B">
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<mods:roleTerm id="CD45F2A9741B43BCE0D668B429C3B762">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C13B71C62A7304BCC282221B87029C47">Hayashi, Fumio</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="8B731D638BAD01FDD48B25AA5819EC27">text</mods:typeOfResource>
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<mods:title id="93ADE8145684C9271E9F3677B8BE2A89">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="F21EB662B51504584F06EBC43B18CC73">
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<mods:date id="C3DF18FC102BFA656C9E44E4A2FF75CC">2015</mods:date>
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<mods:detail id="8036596F879355F3EE15A6D5C23524E9" type="volume">
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<mods:number id="971BD0DA6F700B459EF0E95C75663201">3941</mods:number>
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</mods:detail>
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<mods:detail id="EE4A9FF4151B112ABC933FB1D1639CF0" type="issue">
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<mods:number id="82491C6EF2D463652CEFDF374524CBCA">3</mods:number>
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<mods:start id="8B22548FDC277F1BA1720B30CF94E04D">414</mods:start>
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<mods:classification id="D67C1E217E8B53EEC07DAF58D2B30018">journal article</mods:classification>
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<mods:identifier id="7193252C9F3EDD894AF34F910DF44CE9" type="DOI">10.11646/zootaxa.3941.3.7</mods:identifier>
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<treatment id="2D280442E268FFEBFF436CF5FBF857FE" ID-DOI="http://doi.org/10.5281/zenodo.6098415" ID-GBIF-Taxon="119606392" ID-Zenodo-Dep="6098415" LSID="urn:lsid:zoobank.org:act:B09F287C-621F-424F-8668-90B870E94CA0" httpUri="http://treatment.plazi.org/id/2D280442E268FFEBFF436CF5FBF857FE" lastPageId="4" lastPageNumber="418" pageId="1" pageNumber="415">
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<subSubSection id="ED9BE6DFE268FFEEFF436CF5FE5A56D9" pageId="1" pageNumber="415" type="nomenclature">
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<paragraph id="A53EB554E268FFEEFF436CF5FE2956B8" blockId="1.[151,488,151,211]" box="[151,488,151,178]" pageId="1" pageNumber="415">
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<heading id="FE760238E268FFEEFF436CF5FE2956B8" bold="true" box="[151,488,151,178]" fontSize="11" level="1" pageId="1" pageNumber="415" reason="1">
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<emphasis id="97F56946E268FFEEFF436CF5FE2956B8" bold="true" box="[151,488,151,178]" pageId="1" pageNumber="415">
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<taxonomicName id="6281CED7E268FFEEFF436CF5FE4556BB" box="[151,388,151,177]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="415" phylum="Arthropoda" rank="species" species="kompieri" status="sp. nov.">
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<emphasis id="97F56946E268FFEEFF436CF5FE4556BB" bold="true" box="[151,388,151,177]" italics="true" pageId="1" pageNumber="415">Devadatta kompieri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="8CC6D43DE268FFEEFE5F6CFAFE2956B8" box="[395,488,152,178]" pageId="1" pageNumber="415" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="A53EB554E268FFEEFF436CD8FE5A56D9" blockId="1.[151,488,151,211]" box="[151,411,186,211]" pageId="1" pageNumber="415">
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(
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<figureCitation id="3DBAA9D1E268FFEEFF4B6CD8FF2956D9" box="[159,232,186,211]" captionStart="FIGURE 1" captionStartId="1.[151,250,1171,1193]" captionTargetBox="[151,1436,426,1148]" captionTargetId="figure@1.[151,1436,407,1148]" captionTargetPageId="1" captionText="FIGURE 1. Devadatta spp. A, holotype of D. kompieri (male). B, holotype of D. yokoii (male). C, D. ducatrix of the specimen collected at Tu Le Community, Mu Cang Chai District, Yen Bai Province, northern Vietnam. The scale is marked in mm and the same for A and C." httpUri="https://zenodo.org/record/237244/files/figure.png" pageId="1" pageNumber="415">Figs. 1</figureCitation>
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A, 2A, 3A, 4A)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="ED9BE6DFE268FFEEFF436D60FD025769" pageId="1" pageNumber="415" type="materials_examined">
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<paragraph id="A53EB554E268FFEEFF436D60FD025769" blockId="1.[151,1436,258,355]" pageId="1" pageNumber="415">
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<emphasis id="97F56946E268FFEEFF436D60FEBE5711" bold="true" box="[151,383,258,283]" pageId="1" pageNumber="415">Material examined</emphasis>
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.
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||
<emphasis id="97F56946E268FFEEFE466D60FE3E5711" bold="true" box="[402,511,258,283]" pageId="1" pageNumber="415">
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<typeStatus id="7A3A0BF6E268FFEEFE466D60FE3E5711" box="[402,511,258,283]" pageId="1" pageNumber="415" type="holotype">Holotype</typeStatus>
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</emphasis>
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(
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<figureCitation id="3DBAA9D1E268FFEEFDC06D60FD985711" box="[532,601,258,283]" captionStart="FIGURE 1" captionStartId="1.[151,250,1171,1193]" captionTargetBox="[151,1436,426,1148]" captionTargetId="figure@1.[151,1436,407,1148]" captionTargetPageId="1" captionText="FIGURE 1. Devadatta spp. A, holotype of D. kompieri (male). B, holotype of D. yokoii (male). C, D. ducatrix of the specimen collected at Tu Le Community, Mu Cang Chai District, Yen Bai Province, northern Vietnam. The scale is marked in mm and the same for A and C." httpUri="https://zenodo.org/record/237244/files/figure.png" pageId="1" pageNumber="415">Fig. 1</figureCitation>
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A): male;
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<collectingCountry id="DD96F5C4E268FFEEFD0D6D60FCFA5711" box="[729,827,258,283]" name="Vietnam" pageId="1" pageNumber="415">Vietnam</collectingCountry>
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, Yen Bai Province, Mu Cang Chai District, Tu Le Community, “Nậm
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<collectingCountry id="DD96F5C4E268FFEEFEA76D44FE545735" box="[371,405,294,319]" name="Switzerland" pageId="1" pageNumber="415">Ch</collectingCountry>
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ậu” bridge;
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<date id="D13F9394E268FFEEFDCC6D45FD555735" box="[536,660,295,319]" pageId="1" pageNumber="415" value="2014-05-19">19-V-2014</date>
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; Quoc Toan Phan leg. (in
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<collectionCode id="C3902D91E268FFEEFC6C6D45FBCC5735" box="[952,1037,295,319]" pageId="1" pageNumber="415">TMUZ</collectionCode>
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).
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<emphasis id="97F56946E268FFEEFBCB6D44FB575735" bold="true" box="[1055,1174,294,319]" pageId="1" pageNumber="415">
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<typeStatus id="7A3A0BF6E268FFEEFBCB6D44FB575735" box="[1055,1174,294,319]" pageId="1" pageNumber="415" type="paratype">Paratypes</typeStatus>
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</emphasis>
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:
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<specimenCount id="B3877EDDE268FFEEFB726D45FB3E5735" box="[1190,1279,294,319]" pageId="1" pageNumber="415" type="male">4 males</specimenCount>
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, the same site and date as in
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<typeStatus id="7A3A0BF6E268FFEEFEE36D28FE585769" box="[311,409,330,355]" pageId="1" pageNumber="415" type="holotype">holotype</typeStatus>
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(
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<quantity id="627918B1E268FFEEFE7F6D29FE155769" box="[427,468,330,355]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="1" pageNumber="415" unit="in" value="1.0">1 in</quantity>
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<collectionCode id="C3902D91E268FFEEFE0F6D29FDEC5768" box="[475,557,331,354]" pageId="1" pageNumber="415">TMUZ</collectionCode>
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,
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<quantity id="627918B1E268FFEEFDEC6D29FDA55769" box="[568,612,330,355]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="1" pageNumber="415" unit="in" value="3.0">3 in</quantity>
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<collectionCode id="C3902D91E268FFEEFDBE6D29FD765769" box="[618,695,331,355]" pageId="1" pageNumber="415">PQTC</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<caption id="F1FEE5DCE268FFEEFF4368F1FEB452EC" httpUri="https://zenodo.org/record/237244/files/figure.png" pageId="1" pageNumber="415" targetBox="[151,1436,426,1148]" targetPageId="1">
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<paragraph id="A53EB554E268FFEEFF4368F1FEB452EC" blockId="1.[151,1436,1171,1254]" pageId="1" pageNumber="415">
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<emphasis id="97F56946E268FFEEFF4368F1FED352A2" bold="true" box="[151,274,1171,1193]" pageId="1" pageNumber="415">FIGURE 1.</emphasis>
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<taxonomicName id="6281CED7E268FFEEFECC68F1FE4352A3" box="[280,386,1171,1193]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="415" phylum="Arthropoda" rank="genus">
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<emphasis id="97F56946E268FFEEFECC68F1FE4352A3" box="[280,386,1171,1193]" italics="true" pageId="1" pageNumber="415">Devadatta</emphasis>
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</taxonomicName>
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spp.
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<emphasis id="97F56946E268FFEEFE6368F1FE0852A2" bold="true" box="[439,457,1171,1192]" pageId="1" pageNumber="415">A</emphasis>
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, holotype of
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||
<taxonomicName id="6281CED7E268FFEEFD8468F6FD0952A3" box="[592,712,1171,1193]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="415" phylum="Arthropoda" rank="species" species="kompieri">
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<emphasis id="97F56946E268FFEEFD8468F6FD0952A3" box="[592,712,1171,1193]" italics="true" pageId="1" pageNumber="415">D. kompieri</emphasis>
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</taxonomicName>
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||
(male).
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<emphasis id="97F56946E268FFEEFCC868F1FCEC52A2" bold="true" box="[796,813,1171,1192]" pageId="1" pageNumber="415">B</emphasis>
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||
, holotype of
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<taxonomicName id="6281CED7E268FFEEFC6068F6FBD152A3" box="[948,1040,1171,1193]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="415" phylum="Arthropoda" rank="species" species="yokoii">
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<emphasis id="97F56946E268FFEEFC6068F6FBD152A3" box="[948,1040,1171,1193]" italics="true" pageId="1" pageNumber="415">D. yokoii</emphasis>
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</taxonomicName>
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(male).
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<emphasis id="97F56946E268FFEEFBB068F1FBB752A3" bold="true" box="[1124,1142,1171,1193]" pageId="1" pageNumber="415">C</emphasis>
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,
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<taxonomicName id="6281CED7E268FFEEFB5768F6FB3752A3" box="[1155,1270,1171,1193]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="415" phylum="Arthropoda" rank="species" species="ducatrix">
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<emphasis id="97F56946E268FFEEFB5768F6FB3752A3" box="[1155,1270,1171,1193]" italics="true" pageId="1" pageNumber="415">D. ducatrix</emphasis>
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</taxonomicName>
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of the specimen collected at Tu Le Community, Mu Cang Chai District, Yen Bai Province, northern Vietnam. The scale is marked in mm and the same for A and C.
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</paragraph>
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</caption>
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||
<subSubSection id="ED9BE6DFE268FFEEFF13696FFD6B5364" pageId="1" pageNumber="415" type="etymology">
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<paragraph id="A53EB554E268FFEEFF13696FFD6B5364" blockId="1.[151,1437,1293,2002]" pageId="1" pageNumber="415">
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<emphasis id="97F56946E268FFEEFF13696FFE87532C" bold="true" box="[199,326,1293,1318]" pageId="1" pageNumber="415">Etymology</emphasis>
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. The new species is named after Mr. Tom Kompier, an enthusiastic entomologist, who has recommended several good study sites in northern
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<collectingCountry id="DD96F5C4E268FFEEFD0B6953FCFF5340" box="[735,830,1329,1354]" name="Vietnam" pageId="1" pageNumber="415">Vietnam</collectingCountry>
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to the first author and kindly supplied us with many interesting damselfly specimens from
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<collectingCountry id="DD96F5C4E268FFEEFD976937FD675364" box="[579,678,1365,1390]" name="Vietnam" pageId="1" pageNumber="415">Vietnam</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="ED9BE6DFE268FFEBFF13691BFBF857FE" lastPageId="4" lastPageNumber="418" pageId="1" pageNumber="415" type="description">
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<paragraph id="A53EB554E268FFEEFF13691BFF1A53D0" blockId="1.[151,1437,1293,2002]" pageId="1" pageNumber="415">
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||
<emphasis id="97F56946E268FFEEFF13691BFD9B5398" bold="true" box="[199,602,1401,1426]" pageId="1" pageNumber="415">
|
||
Description of the
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||
<typeStatus id="7A3A0BF6E268FFEEFE7A691BFDD45398" box="[430,533,1401,1426]" pageId="1" pageNumber="415" type="holotype">holotype</typeStatus>
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male
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||
</emphasis>
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. Head: Labrum, mandibles, and antennae black; postclypeus, frons and upper surface of head metallic dark blue; mouth parts bearing short brown setae; ocellus olive; occipital area glossy black.
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</paragraph>
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<paragraph id="A53EB554E268FFEEFF136987FE2C5060" blockId="1.[151,1437,1293,2002]" pageId="1" pageNumber="415">Thorax: Dorsal and lateral lobes of prothorax black, with pruinescence on part of basal and ventral areas. Synthorax generally black dorsally and laterally, lacking any pale maculation, with pale blue pruinescence over much of basal part of metepimeron and ventral surface. Legs brown with pruinescence on coxa; femur and tibia with strong but slender spines.</paragraph>
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<paragraph id="A53EB554E268FFEEFF136A17FEDF51DB" blockId="1.[151,1437,1293,2002]" pageId="1" pageNumber="415">
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Wings (
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||
<figureCitation id="3DBAA9D1E268FFEEFECB6A17FEA25084" box="[287,355,1653,1678]" captionStart="FIGURE 2" captionStartId="2.[151,250,1629,1651]" captionTargetBox="[222,1354,182,1597]" captionTargetId="figure@2.[211,1376,167,1608]" captionTargetPageId="2" captionText="FIGURE 2. Male fore- and hindwings of Devadatta kompieri. (A), D. yokoii (B), and D. ducatrix (C, same specimen shown in Fig. 1 C)." httpUri="https://zenodo.org/record/237245/files/figure.png" pageId="1" pageNumber="415">Fig. 2</figureCitation>
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||
A): Venation black, membrane hyaline with slight pale yellow tint in mature male. Fw is a little longer and narrower than Hw; both wings petiolated slightly proximal to anal crossing. Eleven antenodal veins in Fw and
|
||
<quantity id="627918B1E268FFEEFF276ADCFEED50DC" box="[243,300,1725,1750]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="1" pageNumber="415" unit="in" value="10.0">10 in</quantity>
|
||
Hw; subcostal veins proximal to subnodus with 4–5 crossveins in both wings; 38 postnodal veins in Fw and
|
||
<quantity id="627918B1E268FFEEFF216A80FEF150F0" box="[245,304,1761,1786]" metricMagnitude="-1" metricUnit="m" metricValue="7.62" pageId="1" pageNumber="415" unit="in" value="30.0">30 in</quantity>
|
||
Hw; arculus situated at 6th antenodal in Fw and 5th in Hw; cubital space in Fw with 3 crossveins before anal crossing, the same of Hw with 3 or 4 veins; quadrangle space with 3 or 4 crossveins in Fw, and 2 or 3 crossveins in Hw; first anal vein meeting wing margin before the level of nodus in Fw, almost equal with nodus in Hw; distal space between first anal vein and wing posterior margin rowed 2 or 3 cells in Fw and 1 or
|
||
<quantity id="627918B1E268FFEEFAEC6B2CFAA4516C" box="[1336,1381,1869,1894]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="1" pageNumber="415" unit="in" value="2.0">2 in</quantity>
|
||
Hw, respectively. Pterostigma dark brown, covering 5 to 6 underlying cells in Fw, and
|
||
<quantity id="627918B1E268FFEEFB856B13FBBE5180" box="[1105,1151,1905,1930]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="1" pageNumber="415" unit="in" value="5.0">5 in</quantity>
|
||
Hw. Wing tip with dark brown patch, covering a little more than half the area between distal end of pterostigma and apex in Fw, about onethird in Hw.
|
||
</paragraph>
|
||
<caption id="F1FEE5DCE26BFFEDFF436A3FFF335098" httpUri="https://zenodo.org/record/237245/files/figure.png" pageId="2" pageNumber="416" targetBox="[222,1354,182,1597]" targetPageId="2">
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<paragraph id="A53EB554E26BFFEDFF436A3FFF335098" blockId="2.[151,1436,1629,1682]" pageId="2" pageNumber="416">
|
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<emphasis id="97F56946E26BFFEDFF436A3FFED35079" bold="true" box="[151,274,1629,1651]" pageId="2" pageNumber="416">FIGURE 2.</emphasis>
|
||
Male fore- and hindwings of
|
||
<taxonomicName id="6281CED7E26BFFEDFDEB6A3FFCC85079" box="[575,777,1629,1651]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="416" phylum="Arthropoda" rank="species" species="kompieri">
|
||
<emphasis id="97F56946E26BFFEDFDEB6A3FFCC85079" box="[575,777,1629,1651]" italics="true" pageId="2" pageNumber="416">Devadatta kompieri</emphasis>
|
||
</taxonomicName>
|
||
. (
|
||
<emphasis id="97F56946E26BFFEDFCC96A3CFCEE5079" bold="true" box="[797,815,1630,1651]" pageId="2" pageNumber="416">A</emphasis>
|
||
),
|
||
<taxonomicName id="6281CED7E26BFFEDFC906A3DFC605079" box="[836,929,1629,1652]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="416" phylum="Arthropoda" rank="species" species="yokoii">
|
||
<emphasis id="97F56946E26BFFEDFC906A3DFC605079" box="[836,929,1629,1652]" italics="true" pageId="2" pageNumber="416">D. yokoii</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="97F56946E26BFFEDFC646A3CFC005079" bold="true" box="[944,961,1630,1651]" pageId="2" pageNumber="416">B</emphasis>
|
||
), and
|
||
<taxonomicName id="6281CED7E26BFFEDFBD46A3DFBB25079" box="[1024,1139,1629,1652]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="416" phylum="Arthropoda" rank="species" species="ducatrix">
|
||
<emphasis id="97F56946E26BFFEDFBD46A3DFBB25079" box="[1024,1139,1629,1652]" italics="true" pageId="2" pageNumber="416">D. ducatrix</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="97F56946E26BFFEDFB566A3FFB555079" bold="true" box="[1154,1172,1629,1651]" pageId="2" pageNumber="416">C</emphasis>
|
||
, same specimen shown in Fig. 1C).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A53EB554E26BFFEDFF136AD8FAF9518C" blockId="2.[151,1437,1722,2035]" pageId="2" pageNumber="416">
|
||
Abdomen: Blackish brown, with pruinescence laterally on S1 and S2 and ventral surface of all segments. Anal appendages (
|
||
<figureCitation id="3DBAA9D1E26BFFEDFEFC6ABFFEA850FC" box="[296,361,1757,1782]" captionStart="FIGURE 3" captionStartId="3.[151,250,1230,1252]" captionTargetBox="[164,1422,274,1197]" captionTargetId="figure@3.[151,1435,265,1209]" captionTargetPageId="3" captionText="FIGURE 3. Male caudal appendages of Devadatta kompieri (A), D. yokoii (B), and D. ducatrix (C, rearranged from Lieftinck 1969, p. 207, Fig. 2) in lateral, dorsal, and ventral views." httpUri="https://zenodo.org/record/237246/files/figure.png" pageId="2" pageNumber="416">Fig. 3</figureCitation>
|
||
A) blackish, with pruinescence; superior appendage a little more than twice as long as inferior in lateral view; superior appendage in dorsal view bent in the middle with distal portion turning obliquely inwards with inner margin expanded and tip obtusely swollen; apices of superior pair barely overlapping each other. Inferior appendage short and strongly tapering in lateral view; the tip sharply hooked inwards. Penile organ (
|
||
<figureCitation id="3DBAA9D1E26BFFEDFABB6B28FF64518C" captionStart="FIGURE 4" captionStartId="3.[151,250,1968,1990]" captionTargetBox="[160,1429,1340,1941]" captionTargetId="figure@3.[151,1436,1332,1946]" captionTargetPageId="3" captionText="FIGURE 4. Male penile organs of Devadatta kompieri (A), D. yokoii (B), and D. ducatrix (C, rearranged from Lieftinck 1969, p. 207, Fig. 2) in lateral and ventral views." httpUri="https://zenodo.org/record/237247/files/figure.png" pageId="2" pageNumber="416">Fig. 4</figureCitation>
|
||
A) is similar to that of
|
||
<taxonomicName id="6281CED7E26BFFEDFE706B0DFDE2518C" box="[420,547,1902,1926]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="416" phylum="Arthropoda" rank="species" species="ducatrix">
|
||
<emphasis id="97F56946E26BFFEDFE706B0DFDE2518C" box="[420,547,1902,1926]" italics="true" pageId="2" pageNumber="416">D. ducatrix</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="3DBAA9D1E26BFFEDFDE66B0FFDB7518C" box="[562,630,1901,1926]" captionStart="FIGURE 4" captionStartId="3.[151,250,1968,1990]" captionTargetBox="[160,1429,1340,1941]" captionTargetId="figure@3.[151,1436,1332,1946]" captionTargetPageId="3" captionText="FIGURE 4. Male penile organs of Devadatta kompieri (A), D. yokoii (B), and D. ducatrix (C, rearranged from Lieftinck 1969, p. 207, Fig. 2) in lateral and ventral views." httpUri="https://zenodo.org/record/237247/files/figure.png" pageId="2" pageNumber="416">Fig. 4</figureCitation>
|
||
C), lobes at apex of distal segment approximately semicircular.
|
||
</paragraph>
|
||
<paragraph id="A53EB554E26BFFEDFF136BF0FC6F51A0" blockId="2.[151,1437,1722,2035]" box="[199,942,1938,1963]" pageId="2" pageNumber="416">
|
||
<emphasis id="97F56946E26BFFEDFF136BF0FEB751A1" bold="true" box="[199,374,1938,1963]" pageId="2" pageNumber="416">Measurements</emphasis>
|
||
(mm). Hw 32, abdomen including appendages 37.
|
||
</paragraph>
|
||
<paragraph id="A53EB554E26BFFECFF136BD7FEC056DF" blockId="2.[151,1437,1722,2035]" lastBlockId="3.[151,1436,151,213]" lastPageId="3" lastPageNumber="417" pageId="2" pageNumber="416">
|
||
<emphasis id="97F56946E26BFFEDFF136BD7FDDF51C4" bold="true" box="[199,542,1973,1998]" pageId="2" pageNumber="416">
|
||
Variation in
|
||
<typeStatus id="7A3A0BF6E26BFFEDFEB06BD7FE0E51C4" box="[356,463,1973,1998]" pageId="2" pageNumber="416" type="paratype">paratype</typeStatus>
|
||
males
|
||
</emphasis>
|
||
. The anterior lobe of prothorax, synthorax, and anal appendages pruinose in mature male, but becoming matte black with increasing age. Details of wing venation as follows: in Fw, antenodal veins 10–13, postnodal 37–42, cubital space with 3–4; in Hw, antenodal 10–12, postnodal 29–39, cubital space with 4–6.
|
||
</paragraph>
|
||
<caption id="F1FEE5DCE26AFFECFF4368ACFD175308" httpUri="https://zenodo.org/record/237246/files/figure.png" pageId="3" pageNumber="417" targetBox="[164,1422,274,1197]" targetPageId="3">
|
||
<paragraph id="A53EB554E26AFFECFF4368ACFD175308" blockId="3.[151,1436,1230,1282]" pageId="3" pageNumber="417">
|
||
<emphasis id="97F56946E26AFFECFF4368ACFED252E9" bold="true" box="[151,275,1230,1252]" pageId="3" pageNumber="417">FIGURE 3.</emphasis>
|
||
Male caudal appendages of
|
||
<taxonomicName id="6281CED7E26AFFECFDE168ACFD3E52EE" box="[565,767,1230,1252]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="417" phylum="Arthropoda" rank="species" species="kompieri">
|
||
<emphasis id="97F56946E26AFFECFDE168ACFD3E52EE" box="[565,767,1230,1252]" italics="true" pageId="3" pageNumber="417">Devadatta kompieri</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="97F56946E26AFFECFCDA68ACFCE152E9" bold="true" box="[782,800,1230,1251]" pageId="3" pageNumber="417">A</emphasis>
|
||
),
|
||
<taxonomicName id="6281CED7E26AFFECFCE168ADFC5352EE" box="[821,914,1230,1252]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="417" phylum="Arthropoda" rank="species" species="yokoii">
|
||
<emphasis id="97F56946E26AFFECFCE168ADFC5352EE" box="[821,914,1230,1252]" italics="true" pageId="3" pageNumber="417">D. yokoii</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="97F56946E26AFFECFC7668ACFC7252E9" bold="true" box="[930,947,1230,1251]" pageId="3" pageNumber="417">B</emphasis>
|
||
), and
|
||
<taxonomicName id="6281CED7E26AFFECFC2768ADFBA752EE" box="[1011,1126,1230,1252]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="417" phylum="Arthropoda" rank="species" species="ducatrix">
|
||
<emphasis id="97F56946E26AFFECFC2768ADFBA752EE" box="[1011,1126,1230,1252]" italics="true" pageId="3" pageNumber="417">D. ducatrix</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="97F56946E26AFFECFBA168ACFB4652EE" bold="true" box="[1141,1159,1230,1252]" pageId="3" pageNumber="417">C</emphasis>
|
||
, rearranged from Lieftinck 1969, p. 207, Fig. 2) in lateral, dorsal, and ventral views.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="F1FEE5DCE26AFFECFF436BD2FD8751EE" httpUri="https://zenodo.org/record/237247/files/figure.png" pageId="3" pageNumber="417" targetBox="[160,1429,1340,1941]" targetPageId="3">
|
||
<paragraph id="A53EB554E26AFFECFF436BD2FD8751EE" blockId="3.[151,1436,1968,2020]" pageId="3" pageNumber="417">
|
||
<emphasis id="97F56946E26AFFECFF436BD2FED351CF" bold="true" box="[151,274,1968,1990]" pageId="3" pageNumber="417">FIGURE 4.</emphasis>
|
||
Male penile organs of
|
||
<taxonomicName id="6281CED7E26AFFECFE2F6BD2FD0451CC" box="[507,709,1968,1990]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="417" phylum="Arthropoda" rank="species" species="kompieri">
|
||
<emphasis id="97F56946E26AFFECFE2F6BD2FD0451CC" box="[507,709,1968,1990]" italics="true" pageId="3" pageNumber="417">Devadatta kompieri</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="97F56946E26AFFECFD076BD2FD2451CF" bold="true" box="[723,741,1968,1989]" pageId="3" pageNumber="417">A</emphasis>
|
||
),
|
||
<taxonomicName id="6281CED7E26AFFECFD2E6BD3FC9651CC" box="[762,855,1968,1990]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="417" phylum="Arthropoda" rank="species" species="yokoii">
|
||
<emphasis id="97F56946E26AFFECFD2E6BD3FC9651CC" box="[762,855,1968,1990]" italics="true" pageId="3" pageNumber="417">D. yokoii</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="97F56946E26AFFECFCB16BD2FCB751CF" bold="true" box="[869,886,1968,1989]" pageId="3" pageNumber="417">B</emphasis>
|
||
), and
|
||
<taxonomicName id="6281CED7E26AFFECFC626BD3FBE851CC" box="[950,1065,1968,1990]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="417" phylum="Arthropoda" rank="species" species="ducatrix">
|
||
<emphasis id="97F56946E26AFFECFC626BD3FBE851CC" box="[950,1065,1968,1990]" italics="true" pageId="3" pageNumber="417">D. ducatrix</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="97F56946E26AFFECFBEC6BD2FB8B51CC" bold="true" box="[1080,1098,1968,1990]" pageId="3" pageNumber="417">C</emphasis>
|
||
, rearranged from Lieftinck 1969, p. 207, Fig. 2) in lateral and ventral views.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A53EB554E26DFFEBFF136CF5FBC256BA" blockId="4.[151,1436,151,501]" box="[199,1027,151,176]" pageId="4" pageNumber="418">
|
||
<emphasis id="97F56946E26DFFEBFF136CF5FEB756BA" bold="true" box="[199,374,151,176]" pageId="4" pageNumber="418">Measurements</emphasis>
|
||
(mm). Hw 31–34; abdomen including appendages 32–37.
|
||
</paragraph>
|
||
<paragraph id="A53EB554E26DFFEBFF136CDEFE6356DF" blockId="4.[151,1436,151,501]" box="[199,418,188,213]" pageId="4" pageNumber="418">
|
||
<emphasis id="97F56946E26DFFEBFF136CDEFEDF56DF" bold="true" box="[199,286,188,213]" pageId="4" pageNumber="418">Female</emphasis>
|
||
. Unknown.
|
||
</paragraph>
|
||
<paragraph id="A53EB554E26DFFEBFF136CBDFE9457A6" blockId="4.[151,1436,151,501]" pageId="4" pageNumber="418">
|
||
<emphasis id="97F56946E26DFFEBFF136CBDFE5056F2" bold="true" box="[199,401,223,248]" pageId="4" pageNumber="418">Notes on biology</emphasis>
|
||
.
|
||
<taxonomicName id="6281CED7E26DFFEBFE776C82FD4756F2" box="[419,646,224,248]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="418" phylum="Arthropoda" rank="species" species="kompieri">
|
||
<emphasis id="97F56946E26DFFEBFE776C82FD4756F2" box="[419,646,224,248]" italics="true" pageId="4" pageNumber="418">Devadatta kompieri</emphasis>
|
||
</taxonomicName>
|
||
was found at a small spring-fed brooklet in pristine forest. Like its congeners,
|
||
<taxonomicName id="6281CED7E26DFFEBFEC26D67FE5D5716" box="[278,412,260,284]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="418" phylum="Arthropoda" rank="species" species="kompieri">
|
||
<emphasis id="97F56946E26DFFEBFEC26D67FE5D5716" box="[278,412,260,284]" italics="true" pageId="4" pageNumber="418">D. kompieri</emphasis>
|
||
</taxonomicName>
|
||
males occasionally showed up at sunny spots making short flights.
|
||
<taxonomicName id="6281CED7E26DFFEBFB456D66FAA85716" box="[1169,1385,260,284]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="418" phylum="Arthropoda" rank="species" species="ducatrix">
|
||
<emphasis id="97F56946E26DFFEBFB456D66FAA85716" box="[1169,1385,260,284]" italics="true" pageId="4" pageNumber="418">Devadatta ducatrix</emphasis>
|
||
</taxonomicName>
|
||
was also found at the same site. The first author noticed no differences in behaviour between
|
||
<taxonomicName id="6281CED7E26DFFEBFB6C6D4BFA83574A" box="[1208,1346,296,320]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="418" phylum="Arthropoda" rank="species" species="kompieri">
|
||
<emphasis id="97F56946E26DFFEBFB6C6D4BFA83574A" box="[1208,1346,296,320]" italics="true" pageId="4" pageNumber="418">D. kompieri</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="6281CED7E26DFFEBFA556D4BFF35576E" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="418" phylum="Arthropoda" rank="species" species="ducatrix">
|
||
<emphasis id="97F56946E26DFFEBFA556D4BFF35576E" italics="true" pageId="4" pageNumber="418">D. ducatrix</emphasis>
|
||
</taxonomicName>
|
||
during a short stay at the site. Other damselfly species observed at the same site were:
|
||
<taxonomicName id="6281CED7E26DFFEBFB096D2FFA5D576E" box="[1245,1436,332,356]" class="Insecta" family="Calopterygidae" genus="Mnais" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="418" phylum="Arthropoda" rank="species" species="andersoni">
|
||
<emphasis id="97F56946E26DFFEBFB096D2FFA5D576E" box="[1245,1436,332,356]" italics="true" pageId="4" pageNumber="418">Mnais andersoni</emphasis>
|
||
</taxonomicName>
|
||
McLachIan, 1873,
|
||
<taxonomicName id="6281CED7E26DFFEBFE546D12FD2C5782" authority="Ris, 1912" authorityName="Ris" authorityYear="1912" box="[384,749,367,392]" class="Insecta" family="Lestoideidae" genus="Philoganga" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="418" phylum="Arthropoda" rank="species" species="vetusta">
|
||
<emphasis id="97F56946E26DFFEBFE546D12FDA25782" box="[384,611,368,392]" italics="true" pageId="4" pageNumber="418">Philoganga vetusta</emphasis>
|
||
Ris, 1912
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6281CED7E26DFFEBFCD66D12FCB65782" box="[770,887,368,392]" class="Insecta" family="Calopterygidae" genus="Caliphaea" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="418" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="97F56946E26DFFEBFCD66D12FCB65782" box="[770,887,368,392]" italics="true" pageId="4" pageNumber="418">Caliphaea</emphasis>
|
||
</taxonomicName>
|
||
sp. allied to
|
||
<taxonomicName id="6281CED7E26DFFEBFBFB6D12FA9F5782" authority="Hagen, 1859" authorityName="Hagen" authorityYear="1859" box="[1071,1374,367,392]" class="Insecta" family="Calopterygidae" genus="Caliphaea" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="418" phylum="Arthropoda" rank="species" species="confusa">
|
||
<emphasis id="97F56946E26DFFEBFBFB6D12FB705782" box="[1071,1201,368,392]" italics="true" pageId="4" pageNumber="418">C. confusa</emphasis>
|
||
Hagen, 1859
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="6281CED7E26DFFEBFF436DF6FEEF57A6" box="[151,302,404,428]" class="Insecta" family="Megapodagrionidae" genus="Agriomorpha" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="418" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="97F56946E26DFFEBFF436DF6FEEF57A6" box="[151,302,404,428]" italics="true" pageId="4" pageNumber="418">Agriomorpha</emphasis>
|
||
</taxonomicName>
|
||
sp.
|
||
</paragraph>
|
||
<paragraph id="A53EB554E26DFFEBFF136DD5FBF857FE" blockId="4.[151,1436,151,501]" pageId="4" pageNumber="418">
|
||
<emphasis id="97F56946E26DFFEBFF136DD5FE0657DA" bold="true" box="[199,455,439,464]" pageId="4" pageNumber="418">Differential diagnosis</emphasis>
|
||
.
|
||
<taxonomicName id="6281CED7E26DFFEBFE026DDBFD9C57DA" box="[470,605,440,464]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="418" phylum="Arthropoda" rank="species" species="kompieri">
|
||
<emphasis id="97F56946E26DFFEBFE026DDBFD9C57DA" box="[470,605,440,464]" italics="true" pageId="4" pageNumber="418">D. kompieri</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="6281CED7E26DFFEBFD206DDBFCB557DA" box="[756,884,440,464]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="418" phylum="Arthropoda" rank="species" species="ducatrix">
|
||
<emphasis id="97F56946E26DFFEBFD206DDBFCB557DA" box="[756,884,440,464]" italics="true" pageId="4" pageNumber="418">D. ducatrix</emphasis>
|
||
</taxonomicName>
|
||
and also
|
||
<taxonomicName id="6281CED7E26DFFEBFC366DDBFB8857DA" box="[994,1097,440,464]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="418" phylum="Arthropoda" rank="species" species="yokoii">
|
||
<emphasis id="97F56946E26DFFEBFC366DDBFB8857DA" box="[994,1097,440,464]" italics="true" pageId="4" pageNumber="418">D. yokoii</emphasis>
|
||
</taxonomicName>
|
||
described in the next section, but the males can be distinguished from each other (see diagnosis under
|
||
<taxonomicName id="6281CED7E26DFFEBFC176DBFFBE857FE" box="[963,1065,476,500]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="418" phylum="Arthropoda" rank="species" species="yokoii">
|
||
<emphasis id="97F56946E26DFFEBFC176DBFFBE857FE" box="[963,1065,476,500]" italics="true" pageId="4" pageNumber="418">D. yokoii</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |