treatments-xml/data/26/ED/70/26ED701B0AD45AD08CE6F9251E75751A.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1044.62593" ID-GBIF-Dataset="2b58bee7-f25a-42c9-8651-3060ada2112e" ID-PMC="PMC8222207" ID-Pensoft-Pub="1313-2970-1044-563" ID-Pensoft-UUID="8787843B77CB51DEB6FCF0B8F3A09AC3" ID-PubMed="34183886" ID-ZooBank="71DCAFC2B2BB475B91662BCDC83C7F07" ModsDocID="1313-2970-1044-563" checkinTime="1623875370007" checkinUser="pensoft" docAuthor="Neri, Paolo &amp; Toledano, Luca" docDate="2021" docId="26ED701B0AD45AD08CE6F9251E75751A" docLanguage="en" docName="ZooKeys 1044: 563-587" docOrigin="ZooKeys 1044" docPubDate="2021-06-16" docSource="http://dx.doi.org/10.3897/zookeys.1044.62593" docTitle="Bembidion (Peryphus) psuchrum Andrewes 1922, comb. nov." docType="treatment" docVersion="4" id="8787843B77CB51DEB6FCF0B8F3A09AC3" lastPageNumber="563" masterDocId="8787843B77CB51DEB6FCF0B8F3A09AC3" masterDocTitle="Geographic and taxonomic notes, addenda and corrigenda on the subtribe Bembidiina Stephens, 1827 of the 2017 ' Catalogue of Palaearctic Coleoptera' (Coleoptera, Carabidae, Bembidiina)" masterLastPageNumber="587" masterPageNumber="563" pageNumber="563" updateTime="1668150485866" updateUser="ExternalLinkService">
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<mods:title>Geographic and taxonomic notes, addenda and corrigenda on the subtribe Bembidiina Stephens, 1827 of the 2017 ' Catalogue of Palaearctic Coleoptera' (Coleoptera, Carabidae, Bembidiina)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Neri, Paolo</mods:namePart>
<mods:affiliation>Via Guido Rossa 21 &quot; San Lorenzo &quot;, 47121 Forli (FC), Italy</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Toledano, Luca</mods:namePart>
<mods:affiliation>Museo Civico di Storia Naturale di Verona, Lungadige Porta Vittoria 9, 37129 Verona, Italy</mods:affiliation>
<mods:nameIdentifier type="email">lucatole2@libero.it</mods:nameIdentifier>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2021</mods:date>
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<mods:number>2021-06-16</mods:number>
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<mods:detail type="volume">
<mods:number>1044</mods:number>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1044.62593</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1044.62593</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1044-563</mods:identifier>
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<treatment ID-GBIF-Taxon="182753952" LSID="urn:lsid:plazi:treatment:26ED701B0AD45AD08CE6F9251E75751A" httpUri="http://treatment.plazi.org/id/26ED701B0AD45AD08CE6F9251E75751A" lastPageNumber="563" pageId="0" pageNumber="563">
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<paragraph pageId="0" pageNumber="563">
<taxonomicName LSID="26ED701B-0AD4-5AD0-8CE6-F9251E75751A" authority="Andrewes, 1922" authorityName="Andrewes" authorityYear="1922" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bembidion (Peryphus) psuchrum" order="Coleoptera" pageId="0" pageNumber="563" phylum="Arthropoda" rank="species" species="psuchrum" status="comb. nov." subGenus="Peryphus">Bembidion (Peryphus) psuchrum Andrewes, 1922</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="563">comb. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Habitus of 1 Bembidion (Peryphanes) sanatum Bates, 1883, holotype (NHMUK), 5.60 mm 2 B. (Plataphus) pseudolucillum Netolitzky, 1938 from Kibune, Kyoto (NHMW), 4.20 mm 3 B. (Peryphus) psuchrum Andrewes, 1935 Inde, Himachal Pradesh, Mahari, 3500 m (CTVR), 3.50 mm 4 labels of the holotype of Bembidion sapporense Jedlicka (NMPC). Photographs: Luca Toledano." figureDoi="10.3897/zookeys.1044.62593.figures1-4" httpUri="https://binary.pensoft.net/fig/555046" pageId="0" pageNumber="563">Figures 3</figureCitation>
<figureCitation captionStart="Figures 5–9" captionStartId="F2" captionText="Figures 5 - 9. Median lobe of the aedeagi of 5 Bembidion (Peryphanes) sanatum Bates, 1883, holotype (NHMUK), 1.55 mm 6 B. (P.) dostali Kirschenhofer, 1984, paratype (NHMW), 1.53 mm 7 B. (P.) lulinense Habu, 1973 (drawing from Habu 1973), 1.41 mm 8 Bembidion (Peryphus) psuchrum Andrewes, 1935, holotype (NHMUK), 0.91 mm 9 Bembidion (Plataphus) pseudolucillum Netolitzky, paratype (NHMW), 0.80 mm." figureDoi="10.3897/zookeys.1044.62593.figures5-9" httpUri="https://binary.pensoft.net/fig/555047" pageId="0" pageNumber="563">, 8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="563" type="reference_group">
<paragraph pageId="0" pageNumber="563">
<taxonomicName authorityName="Neri &amp; Toledano" authorityYear="2021" baseAuthorityName="Neri &amp; Toledano" baseAuthorityYear="2021" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bembidion" order="Coleoptera" pageId="0" pageNumber="563" phylum="Arthropoda" rank="genus" status="comb. nov.">Bembidion</taxonomicName>
(
<emphasis italics="true" pageId="0" pageNumber="563">incertae sedis</emphasis>
) 
<taxonomicName authorityName="Neri &amp; Toledano" authorityYear="2021" baseAuthorityName="Neri &amp; Toledano" baseAuthorityYear="1922" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bembidion psuchrum" order="Coleoptera" pageId="0" pageNumber="563" phylum="Arthropoda" rank="species" species="psuchrum" status="comb. nov.">Bembidion psuchrum</taxonomicName>
Andrewes, 1922
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="563" type="materials_examined">
<paragraph pageId="0" pageNumber="563">Examined material.</paragraph>
<paragraph pageId="0" pageNumber="563">
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="W. Almora Divis." determinerName="H. E. Andrewes" elevation="3048" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<specimenCount type="male">1 ♂</specimenCount>
, 
<typeStatus>holotype</typeStatus>
of 
<taxonomicName class="Insecta" family="Carabidae" genus="Bembidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bembidium psuchrum" order="Coleoptera" pageId="0" pageNumber="563" phylum="Arthropoda" rank="species" species="psuchrum">
<emphasis italics="true" pageId="0" pageNumber="563">Bembidium psuchrum</emphasis>
</taxonomicName>
Andrewes, &quot; 
<specimenCount>♂</specimenCount>
1995 [no. dissection by 
<normalizedToken originalValue="Müller-Motzfeld">Mueller-Motzfeld</normalizedToken>
] // Holo / type [round, bordered in red] // Type [round, bordered in red] // 
<emphasis italics="true" pageId="0" pageNumber="563">Bembidium / psuchrum</emphasis>
Andr. / Type [handwritten] / 
<determinerName>H. E. Andrewes</determinerName>
det. [printed] // Brit. Mus. / 1923-24 // Bunderdhunga V. / 
<collectorName>W. Almora Divis.</collectorName>
/ 
<quantity metricMagnitude="3" metricUnit="m" metricValue="3.048" metricValueMax="3.6576" metricValueMin="2.4384" unit="ft" value="10000.0" valueMax="12000.0" valueMin="8000.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="3.048" metricValueMax="3.6576" metricValueMin="2.4384" unit="ft" value="10000.0" valueMax="12000.0" valueMin="8000.0">8000-12000 feet</elevation>
</quantity>
/ 
<normalizedToken originalValue="June’">June'</normalizedToken>
19 H.G.C. [printed] // 3120 [printed]&quot; (NHMUK)
</materialsCitation>
; 
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="G. Ledoux" country="Inde" elevation="3500" location="Mahari" specimenCount="1" specimenCount-male="1" stateProvince="Himachal Pradesh" typeStatus="holotype">
<specimenCount type="male">1 ♂</specimenCount>
, &quot; 
<collectingCountry name="India">Inde</collectingCountry>
/ 
<collectingRegion country="India" name="Himachal Pradesh">Himachal Pradesh</collectingRegion>
/ 
<location LSID="urn:lsid:plazi:treatment:26ED701B0AD45AD08CE6F9251E75751A:FB55D2E698C5490E307CF19F15A75DF6" country="Inde" name="Mahari" stateProvince="Himachal Pradesh">Mahari</location>
// 
<quantity metricMagnitude="3" metricUnit="m" metricValue="3.5" unit="m" value="3500.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="3.5" unit="m" value="3500.0">3500m</elevation>
</quantity>
/ 18.VIII.80 / 
<collectorName>G. Ledoux</collectorName>
&quot; (CTVR) (Fig. 
<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Habitus of 1 Bembidion (Peryphanes) sanatum Bates, 1883, holotype (NHMUK), 5.60 mm 2 B. (Plataphus) pseudolucillum Netolitzky, 1938 from Kibune, Kyoto (NHMW), 4.20 mm 3 B. (Peryphus) psuchrum Andrewes, 1935 Inde, Himachal Pradesh, Mahari, 3500 m (CTVR), 3.50 mm 4 labels of the holotype of Bembidion sapporense Jedlicka (NMPC). Photographs: Luca Toledano." figureDoi="10.3897/zookeys.1044.62593.figures1-4" httpUri="https://binary.pensoft.net/fig/555046" pageId="0" pageNumber="563">3</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="563" type="notes">
<paragraph pageId="0" pageNumber="563">Note.</paragraph>
<paragraph pageId="0" pageNumber="563">The holotype is in perfect condition and the slide with the aedeagus, on a label mounted on the same pin as the specimen, is clear and well preserved.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="563" type="type locality">
<paragraph pageId="0" pageNumber="563">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="563">
Kumaon, West Almora, Bunderdhunga, 8000-12000 ft. [India, Uttarakhand] (
<bibRefCitation author="Andrewes, HE" journalOrPublisher="Entomologist's Monthly Magazine" pageId="0" pageNumber="563" pagination="238 - 242" refId="B1" refString="Andrewes, HE, 1922. Notes on Oriental Carabidae. IV. Some new species of Bembidium from the Himalayas (cont.). Entomologist's Monthly Magazine 58: 238 - 242" title="Notes on Oriental Carabidae. IV. Some new species of Bembidium from the Himalayas (cont.)." volume="58" year="1922">Andrewes 1922</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="563" type="redescription">
<paragraph pageId="0" pageNumber="563">Redescription of the holotype.</paragraph>
<paragraph pageId="0" pageNumber="563">
Total body length 3.90 mm. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="563">Coloration</emphasis>
: head, pronotum and elytra black, metallic. Palps dark brown with last palpomere light. Legs with femora black metallic and apex slightly lighter, tibiae reddish and tarsi reddish with blackish metallic reflections. Antennae reddish black with metallic reflections. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="563">Head</emphasis>
: maximum width, including eyes, 0.82 mm; interocular distance 0.55 mm; frons and clypeus smooth with a few transverse lines, neck microsculptured at base; evident, smooth, deep frontal furrows, ending behind at the second supraorbital seta. Eyes moderately convex, temples slightly convex and oblique towards neck. Antennae long 2.05 mm. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="563">Pronotum</emphasis>
: length along mid line 0.84 mm; width of anterior margin 0.77 mm, maximum width 0.93 mm, width of base 0.61 mm; pronotal width/pronotal length ratio 1.11; very convex, barely transverse; sides entirely rebordered, narrowing with an evident sinuous curve towards the base with which they form a slightly acute corner; marginal gutter very narrow, of homogeneous width; whole surface glossy, laterobasal carina not very evident; median line evident, slightly widened at base; transverse anterior semilunar impression with a few punctures; basal transverse impression punctured between the subquadrate, not deep, basal foveae. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="563">Elytra</emphasis>
: length 2.37 mm, maximum overall elytral width, slightly beyond the middle, 1.59 mm; oval, moderately convex, evident and rounded shoulders; humeral margin reaching stria 5; totally glossy, faint trace of microsculpture only at the extreme apex. Striae with evident punctation, mainly on disc, barely visible at apex; the very faint punctation is visible also at apex. Evident apical stria.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="563" type="male genitalia">
<paragraph pageId="0" pageNumber="563">Male genitalia.</paragraph>
<paragraph pageId="0" pageNumber="563">
Medium sized (0.91 mm) aedeagus (Fig. 
<figureCitation captionStart="Figures 5–9" captionStartId="F2" captionText="Figures 5 - 9. Median lobe of the aedeagi of 5 Bembidion (Peryphanes) sanatum Bates, 1883, holotype (NHMUK), 1.55 mm 6 B. (P.) dostali Kirschenhofer, 1984, paratype (NHMW), 1.53 mm 7 B. (P.) lulinense Habu, 1973 (drawing from Habu 1973), 1.41 mm 8 Bembidion (Peryphus) psuchrum Andrewes, 1935, holotype (NHMUK), 0.91 mm 9 Bembidion (Plataphus) pseudolucillum Netolitzky, paratype (NHMW), 0.80 mm." figureDoi="10.3897/zookeys.1044.62593.figures5-9" httpUri="https://binary.pensoft.net/fig/555047" pageId="0" pageNumber="563">8</figureCitation>
), concave ventral margin with apex bent ventrally; central brush completely inside the median lobe, medium sized main sclerite, moderately wide and sinuate.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="563" type="conclusion">
<paragraph pageId="0" pageNumber="563">Conclusion.</paragraph>
<paragraph pageId="0" pageNumber="563">
The study of the holotype of the species, currently listed as 
<taxonomicName authorityName="Neri &amp; Toledano" authorityYear="2021" baseAuthorityName="Neri &amp; Toledano" baseAuthorityYear="2021" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bembidion" order="Coleoptera" pageId="0" pageNumber="563" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="563">Bembidion</emphasis>
</taxonomicName>
&quot;incertae sedis&quot; (
<bibRefCitation author="Marggi, W" journalOrPublisher="Brill, Leiden" pageId="0" pageNumber="563" refId="B29" refString="Marggi, W, Toledano, L, Neri, P, 2017. Carabidae: Bembidiini: Bembidiina: 294-342. In: Loebl I, Loebl D (Eds) Catalogue of Palaearctic Coleoptera (Vol. I). Archostemata - Myxophaga - Adephaga. Brill, Leiden" title="Carabidae: Bembidiini: Bembidiina: 294 - 342. In: Loebl I, Loebl D (Eds) Catalogue of Palaearctic Coleoptera (Vol. I). Archostemata - Myxophaga - Adephaga." year="2017">Marggi et al. 2017</bibRefCitation>
), suggests that the species could be assigned to the subgenus 
<taxonomicName class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bembidion (Peryphus)" order="Coleoptera" pageId="0" pageNumber="563" phylum="Arthropoda" rank="subGenus" subGenus="Peryphus">Bembidion Peryphus</taxonomicName>
Dejean, 1821.
</paragraph>
<caption doi="10.3897/zookeys.1044.62593.figures5-9" httpUri="https://binary.pensoft.net/fig/555047" pageId="0" pageNumber="563" start="Figures 5–9" startId="F2">
<paragraph pageId="0" pageNumber="563">
<emphasis bold="true" pageId="0" pageNumber="563">Figures 5-9.</emphasis>
Median lobe of the aedeagi of 
<emphasis bold="true" pageId="0" pageNumber="563">5</emphasis>
<taxonomicName authorityName="Bates" authorityYear="1883" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bembidion (Peryphanes) sanatum" order="Coleoptera" pageId="0" pageNumber="563" phylum="Arthropoda" rank="species" species="sanatum" subGenus="Peryphanes">Bembidion (Peryphanes) sanatum</taxonomicName>
Bates, 1883, holotype (NHMUK), 1.55 mm 
<emphasis bold="true" pageId="0" pageNumber="563">6</emphasis>
<taxonomicName lsidName="B. (P.) dostali" pageId="0" pageNumber="563" rank="species" species="dostali" subGenus="P.">B. (P.) dostali</taxonomicName>
Kirschenhofer, 1984, paratype (NHMW), 1.53 mm 
<emphasis bold="true" pageId="0" pageNumber="563">7</emphasis>
<taxonomicName lsidName="B. (P.) lulinense" pageId="0" pageNumber="563" rank="species" species="lulinense" subGenus="P.">B. (P.) lulinense</taxonomicName>
Habu, 1973 (drawing from Habu 1973), 1.41 mm 
<emphasis bold="true" pageId="0" pageNumber="563">8</emphasis>
<taxonomicName authorityName="Neri &amp; Toledano" authorityYear="2021" baseAuthorityName="Neri &amp; Toledano" baseAuthorityYear="1922" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bembidion (Peryphus) psuchrum" order="Coleoptera" pageId="0" pageNumber="563" phylum="Arthropoda" rank="species" species="psuchrum" subGenus="Peryphus">Bembidion (Peryphus) psuchrum</taxonomicName>
Andrewes, 1935, holotype (NHMUK), 0.91 mm 
<emphasis bold="true" pageId="0" pageNumber="563">9</emphasis>
<taxonomicName authorityName="Netolitzky" authorityYear="1938" baseAuthorityName="Neri &amp; Toledano" baseAuthorityYear="1938" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bembidion (Plataphus) pseudolucillum" order="Coleoptera" pageId="0" pageNumber="563" phylum="Arthropoda" rank="species" species="pseudolucillum" subGenus="Plataphus">Bembidion (Plataphus) pseudolucillum</taxonomicName>
Netolitzky, paratype (NHMW), 0.80 mm.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>