263 lines
35 KiB
XML
263 lines
35 KiB
XML
<document id="9C6603E67026425D740CEF95D9B3F461" ID-CLB-Dataset="26730" ID-DOI="10.11646/zootaxa.4609.1.13" ID-GBIF-Dataset="cf66c1d6-d78a-4ee9-a888-c7be00c85421" ID-ISSN="1175-5326" ID-Zenodo-Dep="3150403" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1558533835904" checkinUser="plazi" docAuthor="Mora-Aguilar, Eder F. & Delgado, Leonardo" docDate="2019" docId="1E0A87E8357FFFD144ADFAFCFF3BF9A3" docLanguage="en" docName="zootaxa.4609.1.13.pdf" docOrigin="Zootaxa 4609 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhyparus chimalapensis Mora-Aguilar & Delgado 2019, new species" docType="treatment" docVersion="8" lastPageNumber="197" masterDocId="E233FF90357FFFD0443AFFBFFF87FFAA" masterDocTitle="A new Mexican species of Rhyparus Westwood (Coleoptera: Scarabaeidae: Aphodiinae), with new records and a key to the Mexican and Guatemalan species" masterLastPageNumber="200" masterPageNumber="196" pageNumber="196" updateTime="1698731538934" updateUser="ExternalLinkService">
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<mods:mods id="8B16EE5EF67FE85C09698594279C005B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="632A049D0C884EA732F15075E18A1338">
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<mods:title id="84C7B267CA171BA40D5B223E7C85501D">A new Mexican species of Rhyparus Westwood (Coleoptera: Scarabaeidae: Aphodiinae), with new records and a key to the Mexican and Guatemalan species</mods:title>
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<mods:namePart id="3EE55D7CA0994044CC91078F4694C876">Mora-Aguilar, Eder F.</mods:namePart>
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<mods:name id="7EB3C91B5FD65852480DD6194D5CB510" type="personal">
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<mods:roleTerm id="5EDB8959A6861A26D796E6586F3C0973">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="464CEAABF3672FCC5C8AE276CB1F6D34">Delgado, Leonardo</mods:namePart>
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<mods:typeOfResource id="C38B023BFC6BD2F282B3D8D219FC07FB">text</mods:typeOfResource>
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<mods:title id="24A19952F7553771175E4985275E8790">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="1B982DC86343329D93BEB8455E87AA55">
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<mods:date id="4B8D9A0D6614016ED40B4D2B1F19302F">2019</mods:date>
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<mods:number id="5C67D87E12C69A43B49ABAE4D49EC292">2019-05-22</mods:number>
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<mods:number id="EFB2B97ABA3F6FD8A1185E0A91E28C03">4609</mods:number>
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<mods:identifier id="4E172B060325991AA3EA7499FEDF8BFB" type="DOI">10.11646/zootaxa.4609.1.13</mods:identifier>
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<treatment id="1E0A87E8357FFFD144ADFAFCFF3BF9A3" ID-DOI="http://doi.org/10.5281/zenodo.5941235" ID-GBIF-Taxon="157239505" ID-Zenodo-Dep="5941235" LSID="urn:lsid:plazi:treatment:1E0A87E8357FFFD144ADFAFCFF3BF9A3" httpUri="http://treatment.plazi.org/id/1E0A87E8357FFFD144ADFAFCFF3BF9A3" lastPageId="1" lastPageNumber="197" pageId="0" pageNumber="196">
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<subSubSection id="DEB96575357FFFD044ADFAFCFBA6FAF6" box="[151,1057,1347,1374]" pageId="0" pageNumber="196" type="nomenclature">
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<paragraph id="961C36FE357FFFD044ADFAFCFBA6FAF6" blockId="0.[151,1437,1347,2021]" box="[151,1057,1347,1374]" pageId="0" pageNumber="196">
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<emphasis id="A4D7EAEC357FFFD044ADFAFCFC23FAF7" bold="true" box="[151,932,1347,1374]" pageId="0" pageNumber="196">
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<taxonomicName id="51A34D7D357FFFD044ADFAFCFC80FAF7" authority="Mora-Aguilar & Delgado" authorityName="Mora-Aguilar & Delgado" authorityYear="2019" box="[151,775,1347,1374]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="196" phylum="Arthropoda" rank="species" species="chimalapensis" status="sp. nov.">
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<emphasis id="A4D7EAEC357FFFD044ADFAFCFE46FAF4" bold="true" box="[151,449,1347,1374]" italics="true" pageId="0" pageNumber="196">Rhyparus chimalapensis</emphasis>
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Mora-Aguilar & Delgado
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</taxonomicName>
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,
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<taxonomicNameLabel id="BFE45797357FFFD0472EFAFCFC23FAF7" box="[788,932,1347,1373]" pageId="0" pageNumber="196" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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(
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<figureCitation id="0E982A7B357FFFD04789FAFAFB9FFAF6" box="[947,1048,1348,1373]" captionStart="FIGURES 1–9" captionStartId="2.[151,270,1941,1965]" captionTargetBox="[151,1399,151,1912]" captionTargetId="figure@2.[151,1410,151,1921]" captionTargetPageId="2" captionText="FIGURES 1–9. Morphological characters of Rhyparus chimalapensis and R. opacus. Rhyparus chimalapensis holotype female: 1, dorsal habitus; 2, lateral habitus; 3, head; 4, pronotum; 5, elytral apex; 6, ventral habitus. Rhyparus opacus male: 7, dorsal habitus; 8, head; 9, elytral apex." httpUri="https://zenodo.org/record/3150407/files/figure.png" pageId="0" pageNumber="196">Figs. 1–6</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="DEB96575357FFFD044ADFAD6FE6EFA63" pageId="0" pageNumber="196" type="materials_examined">
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<paragraph id="961C36FE357FFFD044ADFAD6FE6EFA63" blockId="0.[151,1437,1347,2021]" pageId="0" pageNumber="196">
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<emphasis id="A4D7EAEC357FFFD044ADFAD6FEBBFA2A" bold="true" box="[151,316,1384,1409]" pageId="0" pageNumber="196">Type material.</emphasis>
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<materialsCitation id="26CB3CA3357FFFD04578FAD6FA1EFA0F" ID-GBIF-Occurrence="2243136214" collectorName="L. Delgado." country="Mexico" county="San Miguel Chimalapa" elevation="1600" latitude="16.6836" location="El Reten" longLatPrecision="7" longitude="-94.2623" municipality="San Antonio" pageId="0" pageNumber="196" specimenCount="1" stateProvince="Oaxaca" typeStatus="holotype">
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<typeStatus id="4918885C357FFFD04578FAD6FE22FA2B" box="[322,421,1385,1409]" pageId="0" pageNumber="196" type="holotype">Holotype</typeStatus>
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female labeled: “
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<collectingCountry id="EEB4766E357FFFD04667FAD6FD4BFA2B" box="[605,716,1385,1409]" name="Mexico" pageId="0" pageNumber="196">MEXICO</collectingCountry>
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:
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<collectingRegion id="5467F81C357FFFD046ECFAD6FCAEFA2B" box="[726,809,1385,1409]" country="Mexico" name="Oaxaca" pageId="0" pageNumber="196">Oaxaca</collectingRegion>
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,
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<collectingCounty id="7F7D4E72357FFFD04709FAD6FBAEFA2B" box="[819,1065,1385,1409]" pageId="0" pageNumber="196">San Miguel Chimalapa</collectingCounty>
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,
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<collectingMunicipality id="7678AC84357FFFD0400EFAD6FB3DFA2B" box="[1076,1210,1385,1409]" pageId="0" pageNumber="196">San Antonio</collectingMunicipality>
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,
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<location id="937C6025357FFFD040FEFAD6FAA4FA2B" LSID="urn:lsid:plazi:treatment:1E0A87E8357FFFD144ADFAFCFF3BF9A3:937C6025357FFFD040FEFAD6FAA4FA2B" box="[1220,1315,1385,1409]" country="Mexico" county="San Miguel Chimalapa" latitude="16.6836" longLatPrecision="7" longitude="-94.2623" municipality="San Antonio" name="El Reten" pageId="0" pageNumber="196" stateProvince="Oaxaca">El Retén</location>
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,
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<location id="937C6025357FFFD04116FAD6FA1FFA2B" LSID="urn:lsid:plazi:treatment:1E0A87E8357FFFD144ADFAFCFF3BF9A3:937C6025357FFFD04116FAD6FA1FFA2B" box="[1324,1432,1385,1409]" country="Mexico" county="San Miguel Chimalapa" latitude="16.6836" longLatPrecision="7" longitude="-94.2623" municipality="San Antonio" name="El Gringo" pageId="0" pageNumber="196" stateProvince="Oaxaca">El Gringo</location>
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, VI–VII-2013,
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<geoCoordinate id="F3975039357FFFD04508FA33FE29FA0E" box="[306,430,1420,1444]" direction="north" orientation="latitude" pageId="0" pageNumber="196" precision="5" value="16.6836">16.6836º N</geoCoordinate>
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, -
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<geoCoordinate id="F3975039357FFFD045FAFA33FDC6FA0E" box="[448,577,1420,1444]" direction="west" orientation="longitude" pageId="0" pageNumber="196" precision="5" value="-94.2623">94.2623º W</geoCoordinate>
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, Alt.
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<elevation id="1D8ED1CD357FFFD04640FA33FD53FA0E" box="[634,724,1420,1444]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="0" pageNumber="196" unit="m" value="1600.0">
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<quantity id="515B9B1B357FFFD04640FA33FD53FA0E" box="[634,724,1420,1444]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="0" pageNumber="196" unit="m" value="1600.0">1,600 m</quantity>
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</elevation>
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, T. intercepción, bosque mesófilo, E. Mora-A y
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<collectorName id="3B565328357FFFD040E2FA32FADFFA0F" box="[1240,1368,1421,1445]" pageId="0" pageNumber="196">L. Delgado.</collectorName>
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cols”
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</materialsCitation>
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.
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<materialsCitation id="26CB3CA3357FFFD044ADFA0EFE62FA63" ID-GBIF-Occurrence="2243136215" box="[151,485,1457,1481]" collectionCode="IEXA" country="Mexico" pageId="0" pageNumber="196" specimenCount="1" stateProvince="Oaxaca" typeStatus="holotype">
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<typeStatus id="4918885C357FFFD044ADFA0EFF7DFA63" box="[151,250,1457,1481]" pageId="0" pageNumber="196" type="holotype">Holotype</typeStatus>
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is deposited in
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<collectionCode id="F0B2AE3B357FFFD04598FA0EFE62FA63" box="[418,485,1457,1481]" pageId="0" pageNumber="196">IEXA</collectionCode>
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</materialsCitation>
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.
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="DEB96575357FFFD144FDFA6BFCAFFCBC" lastPageId="1" lastPageNumber="197" pageId="0" pageNumber="196" type="description">
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<paragraph id="961C36FE357FFFD144FDFA6BFCAFFCBC" blockId="0.[151,1437,1347,2021]" lastBlockId="1.[151,1437,154,1545]" lastPageId="1" lastPageNumber="197" pageId="0" pageNumber="196">
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<emphasis id="A4D7EAEC357FFFD044FDFA6BFED5FA46" bold="true" box="[199,338,1492,1516]" pageId="0" pageNumber="196">Description.</emphasis>
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<typeStatus id="4918885C357FFFD04560FA6AFE3AFA47" box="[346,445,1493,1517]" pageId="0" pageNumber="196" type="holotype">Holotype</typeStatus>
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female. Length
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<quantity id="515B9B1B357FFFD0464AFA6BFD42FA46" box="[624,709,1492,1516]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="0" pageNumber="196" unit="mm" value="3.6">3.6 mm</quantity>
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, width 1.0 mm. Narrow, elongate, sides parallel; color dark brown, almost black, alutaceous (
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<figureCitation id="0E982A7B357FFFD04590FA46FDADF9BA" box="[426,554,1528,1553]" captionStart="FIGURES 1–9" captionStartId="2.[151,270,1941,1965]" captionTargetBox="[151,1399,151,1912]" captionTargetId="figure@2.[151,1410,151,1921]" captionTargetPageId="2" captionText="FIGURES 1–9. Morphological characters of Rhyparus chimalapensis and R. opacus. Rhyparus chimalapensis holotype female: 1, dorsal habitus; 2, lateral habitus; 3, head; 4, pronotum; 5, elytral apex; 6, ventral habitus. Rhyparus opacus male: 7, dorsal habitus; 8, head; 9, elytral apex." httpUri="https://zenodo.org/record/3150407/files/figure.png" pageId="0" pageNumber="196">Figs. 1–2, 6</figureCitation>
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).
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<emphasis id="A4D7EAEC357FFFD04607FA46FDF0F9BB" box="[573,631,1529,1553]" italics="true" pageId="0" pageNumber="196">Head</emphasis>
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: clypeus with double anterior edge, upper edge with four teeth more-or-less triangular and equally spaced; clypeal margin arcuately emarginate between the teeth; surface of clypeus with scattered, fine punctures separated by one puncture diameter. Disc of frons convex, limited with a distinct narrow furrow from the surrounding surface, with four low obtuse frontal ridges, the middle two separated by about one-third or less the distance to the lateral ridge; surface with shallow, scattered, some setigerous punctures between ridges (
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<figureCitation id="0E982A7B357FFFD040EEF936FA91F90B" box="[1236,1302,1673,1697]" captionStart="FIGURES 1–9" captionStartId="2.[151,270,1941,1965]" captionTargetBox="[151,1399,151,1912]" captionTargetId="figure@2.[151,1410,151,1921]" captionTargetPageId="2" captionText="FIGURES 1–9. Morphological characters of Rhyparus chimalapensis and R. opacus. Rhyparus chimalapensis holotype female: 1, dorsal habitus; 2, lateral habitus; 3, head; 4, pronotum; 5, elytral apex; 6, ventral habitus. Rhyparus opacus male: 7, dorsal habitus; 8, head; 9, elytral apex." httpUri="https://zenodo.org/record/3150407/files/figure.png" pageId="0" pageNumber="196">Fig. 3</figureCitation>
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).
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<emphasis id="A4D7EAEC357FFFD04111F936FA12F90A" box="[1323,1429,1673,1696]" italics="true" pageId="0" pageNumber="196">Pronotum</emphasis>
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: subquadrate; with eight low, evenly spaced, interrupted, and higher posteriorly longitudinal ridges. Apical ridges shorter than flattened bridge; middle pair of ridges obsolete over flattened bridge, uninterrupted to basal margin, almost parallel; each basal ridge with a longitudinal furrow in the inner side; the next pair of ridges completely severed by large and deep transverse deep pits, then continuing to basal margin paralleling the first pair; deep transverse pits narrowly constricted by an oblique projection between second and third ridges. In dorsal view third pair of ridges sinuate with depressions at anterior and basal fourths, basal fourth sinuate, narrow and seemingly forming the posterior pronotal angle opposite elytral humerus; in dorsal view fourth pair of ridges visible as two lateral rounded projections, from the side appearing as a sinuate ridge; basal margin finely carinate with an adjacent line of punctures (
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<figureCitation id="0E982A7B357FFFD047D8F816FBA3F86B" box="[994,1060,1961,1985]" captionStart="FIGURES 1–9" captionStartId="2.[151,270,1941,1965]" captionTargetBox="[151,1399,151,1912]" captionTargetId="figure@2.[151,1410,151,1921]" captionTargetPageId="2" captionText="FIGURES 1–9. Morphological characters of Rhyparus chimalapensis and R. opacus. Rhyparus chimalapensis holotype female: 1, dorsal habitus; 2, lateral habitus; 3, head; 4, pronotum; 5, elytral apex; 6, ventral habitus. Rhyparus opacus male: 7, dorsal habitus; 8, head; 9, elytral apex." httpUri="https://zenodo.org/record/3150407/files/figure.png" pageId="0" pageNumber="196">Fig. 4</figureCitation>
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).
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<emphasis id="A4D7EAEC357FFFD0400DF816FBFDF86B" box="[1079,1146,1961,1985]" italics="true" pageId="0" pageNumber="196">Elytra</emphasis>
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: subparallel, slightly wider at basal third, longer than wide (1.00: 0.41); costae smooth and shiny, sutural costae slightly elevated, punctate; intervals flat, dull. Base of elytra finely carinate. Costae straight and parallel with a row of separated, very fine, appressed setae along each side; first pair slightly divergent posteriorly; fourth costa curving outward apically and swelling into a large, semitriangular, bulbous, inner apex of bulbous narrowly separated from first costa, and continuing downward in straight angle to join with sutural costa. Apices of three enclosed costae and opposite surface of fourth costae covered with a small light colored mass of glandular material. Intervals with a row of moderately coarse, deep punctures along each side, rows separated by about two puncture diameters, striae slightly crenulate, contiguous to costae, punctures separated in each row by one puncture diameter; apex of first interval flattened (
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<figureCitation id="0E982A7B357EFFD14705FECDFC45FE23" box="[831,962,369,394]" captionStart="FIGURES 1–9" captionStartId="2.[151,270,1941,1965]" captionTargetBox="[151,1399,151,1912]" captionTargetId="figure@2.[151,1410,151,1921]" captionTargetPageId="2" captionText="FIGURES 1–9. Morphological characters of Rhyparus chimalapensis and R. opacus. Rhyparus chimalapensis holotype female: 1, dorsal habitus; 2, lateral habitus; 3, head; 4, pronotum; 5, elytral apex; 6, ventral habitus. Rhyparus opacus male: 7, dorsal habitus; 8, head; 9, elytral apex." httpUri="https://zenodo.org/record/3150407/files/figure.png" pageId="1" pageNumber="197">Figs. 1–2, 5</figureCitation>
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).
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<emphasis id="A4D7EAEC357EFFD147E2FECDFB9BFE23" box="[984,1052,370,393]" italics="true" pageId="1" pageNumber="197">Venter</emphasis>
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: entire ventral surface feebly shiny but minutely alutaceous. Prosternal processes elevated, anterior half longitudinally carinate, with a shallow depression on each side. Mesostenum microreticulate with minute, close, appressed setae. Metasternum with a single row of six coarse punctures more-or-less parallel to midline furrow. Abdominal segments 2–5 slightly longer at sides, segments 3–5 with a marginal large puncture; median surface of segments with scattered fine punctures bearing very fine, moderately long setae. Pygidium with strong Y-shaped carina separating three depressed areas, the two lateral areas about five times as large as the median basal area, upper ends of carina with their apex swollen and enlarged; apical margin of pygidium roughly punctate (
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<figureCitation id="0E982A7B357EFFD14562FDD1FE1EFD2C" box="[344,409,622,646]" captionStart="FIGURES 1–9" captionStartId="2.[151,270,1941,1965]" captionTargetBox="[151,1399,151,1912]" captionTargetId="figure@2.[151,1410,151,1921]" captionTargetPageId="2" captionText="FIGURES 1–9. Morphological characters of Rhyparus chimalapensis and R. opacus. Rhyparus chimalapensis holotype female: 1, dorsal habitus; 2, lateral habitus; 3, head; 4, pronotum; 5, elytral apex; 6, ventral habitus. Rhyparus opacus male: 7, dorsal habitus; 8, head; 9, elytral apex." httpUri="https://zenodo.org/record/3150407/files/figure.png" pageId="1" pageNumber="197">Fig. 6</figureCitation>
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).
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<emphasis id="A4D7EAEC357EFFD14591FDD1FE5AFD2F" box="[427,477,622,645]" italics="true" pageId="1" pageNumber="197">Legs</emphasis>
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: profemur with anterior marginal groove, posterior margin slightly arcuate inward, surface covered with moderately coarse punctures. Mesofemur moderately constricted at apical fourth, posterior margin bisinuate, and surface alutaceous with scattered fine punctures. Metafemur flattened, long and slender; surface alutaceous with scattered, fine punctures. Protibia tridentate, two apical teeth rounded. Mesotibiae and metatibiae lacking triangular tooth on apical inner margin, with a small spur directed posteriorly.
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</paragraph>
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</subSubSection>
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<subSubSection id="DEB96575357EFFD144FDFC9DFCE6FBD4" pageId="1" pageNumber="197" type="discussion">
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<paragraph id="961C36FE357EFFD144FDFC9DFCE6FBD4" blockId="1.[151,1437,154,1545]" pageId="1" pageNumber="197">
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<emphasis id="A4D7EAEC357EFFD144FDFC9DFEB1FC90" bold="true" box="[199,310,802,826]" pageId="1" pageNumber="197">Remarks.</emphasis>
|
||
<taxonomicName id="51A34D7D357EFFD14506FC9DFDC6FC90" authorityName="Mora-Aguilar & Delgado" authorityYear="2019" box="[316,577,802,826]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="chimalapensis">
|
||
<emphasis id="A4D7EAEC357EFFD14506FC9DFDC6FC90" box="[316,577,802,826]" italics="true" pageId="1" pageNumber="197">Rhyparus chimalapensis</emphasis>
|
||
</taxonomicName>
|
||
belongs to the species group with all elytral intervals with two rows of punctures or less, along with
|
||
<taxonomicName id="51A34D7D357EFFD14567FCF9FD41FCF4" authority="Cartwright & Woodruff" authorityName="Cartwright & Woodruff" authorityYear="1969" box="[349,710,838,862]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="opacus">
|
||
<emphasis id="A4D7EAEC357EFFD14567FCF9FE41FCF7" box="[349,454,838,861]" italics="true" pageId="1" pageNumber="197">R. opacus</emphasis>
|
||
Cartwright & Woodruff
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="51A34D7D357EFFD146CDFCF9FB1DFCF7" authority="Cartwright & Woodruff. In" authorityName="Cartwright & Woodruff. In" authorityYear="1969" box="[759,1178,838,862]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="spangleri">
|
||
<emphasis id="A4D7EAEC357EFFD146CDFCF9FCFEFCF4" box="[759,889,838,862]" italics="true" pageId="1" pageNumber="197">R. spangleri</emphasis>
|
||
Cartwright & Woodruff. In
|
||
</taxonomicName>
|
||
the key of
|
||
<bibRefCitation id="F2324B0F357EFFD14134FCF9FECAFC28" author="Cartwright, O. L. & Woodruff, R. E." pageId="1" pageNumber="197" pagination="1 - 20" refId="ref2584" refString="Cartwright, O. L. & Woodruff, R. E. (1969) Ten Rhyparus from the Western Hemisphere (Coleoptera: Scarabaeidae: Aphidiinae). Smithsonian Contributions to Zoology, 21, 1 - 20. https: // doi. org / 10.5479 / si. 00810282.21" type="journal article" year="1969">Cartwright & Woodruff (1969)</bibRefCitation>
|
||
,
|
||
<taxonomicName id="51A34D7D357EFFD1456DFCD5FD8AFC28" authorityName="Mora-Aguilar & Delgado" authorityYear="2019" box="[343,525,874,898]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="chimalapensis">
|
||
<emphasis id="A4D7EAEC357EFFD1456DFCD5FEE0FC2B" box="[343,359,874,897]" italics="true" pageId="1" pageNumber="197">R</emphasis>
|
||
.
|
||
<emphasis id="A4D7EAEC357EFFD1454EFCD5FD8AFC28" box="[372,525,874,898]" italics="true" pageId="1" pageNumber="197">chimalapensis</emphasis>
|
||
</taxonomicName>
|
||
will key to
|
||
<taxonomicName id="51A34D7D357EFFD146B3FCD5FD75FC2B" authorityName="Cartwright & Woodruff" authorityYear="1969" box="[649,754,874,897]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="opacus">
|
||
<emphasis id="A4D7EAEC357EFFD146B3FCD5FD1EFC2B" box="[649,665,874,897]" italics="true" pageId="1" pageNumber="197">R</emphasis>
|
||
.
|
||
<emphasis id="A4D7EAEC357EFFD1469CFCD5FD75FC2B" box="[678,754,874,897]" italics="true" pageId="1" pageNumber="197">opacus</emphasis>
|
||
</taxonomicName>
|
||
. The new species is distinguished from
|
||
<taxonomicName id="51A34D7D357EFFD140A2FCD5FA86FC2B" authorityName="Cartwright & Woodruff" authorityYear="1969" box="[1176,1281,874,897]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="opacus">
|
||
<emphasis id="A4D7EAEC357EFFD140A2FCD5FB2FFC2B" box="[1176,1192,874,897]" italics="true" pageId="1" pageNumber="197">R</emphasis>
|
||
.
|
||
<emphasis id="A4D7EAEC357EFFD1408FFCD5FA86FC2B" box="[1205,1281,874,897]" italics="true" pageId="1" pageNumber="197">opacus</emphasis>
|
||
</taxonomicName>
|
||
by the following characters: frons with middle ridges separated by one-third of the distance to the lateral ridge (separated by one-half of the distance to the lateral ridge in
|
||
<taxonomicName id="51A34D7D357EFFD14610FC0DFD12FC63" authorityName="Cartwright & Woodruff" authorityYear="1969" box="[554,661,946,969]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="opacus">
|
||
<emphasis id="A4D7EAEC357EFFD14610FC0DFD12FC63" box="[554,661,946,969]" italics="true" pageId="1" pageNumber="197">R. opacus</emphasis>
|
||
</taxonomicName>
|
||
), pronotum with the two middle basal ridges almost parallel (divergent posteriorly in
|
||
<taxonomicName id="51A34D7D357EFFD1450AFC69FE1CFC47" authorityName="Cartwright & Woodruff" authorityYear="1969" box="[304,411,982,1005]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="opacus">
|
||
<emphasis id="A4D7EAEC357EFFD1450AFC69FE1CFC47" box="[304,411,982,1005]" italics="true" pageId="1" pageNumber="197">R. opacus</emphasis>
|
||
</taxonomicName>
|
||
), second elytral interval with most punctures separated by one puncture diameter on each row (separated by more than one puncture diameter in
|
||
<taxonomicName id="51A34D7D357EFFD14684FC45FCADFBBB" authorityName="Cartwright & Woodruff" authorityYear="1969" box="[702,810,1018,1041]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="opacus">
|
||
<emphasis id="A4D7EAEC357EFFD14684FC45FCADFBBB" box="[702,810,1018,1041]" italics="true" pageId="1" pageNumber="197">R. opacus</emphasis>
|
||
</taxonomicName>
|
||
), and apex of first elytral interval flattened (convex in
|
||
<taxonomicName id="51A34D7D357EFFD141BFFC45FF64FB9F" authorityName="Cartwright & Woodruff" authorityYear="1969" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="opacus">
|
||
<emphasis id="A4D7EAEC357EFFD141BFFC45FF64FB9F" italics="true" pageId="1" pageNumber="197">R. opacus</emphasis>
|
||
</taxonomicName>
|
||
) (
|
||
<figureCitation id="0E982A7B357EFFD144C0FBA1FEE7FB9C" box="[250,352,1054,1078]" captionStart="FIGURES 1–9" captionStartId="2.[151,270,1941,1965]" captionTargetBox="[151,1399,151,1912]" captionTargetId="figure@2.[151,1410,151,1921]" captionTargetPageId="2" captionText="FIGURES 1–9. Morphological characters of Rhyparus chimalapensis and R. opacus. Rhyparus chimalapensis holotype female: 1, dorsal habitus; 2, lateral habitus; 3, head; 4, pronotum; 5, elytral apex; 6, ventral habitus. Rhyparus opacus male: 7, dorsal habitus; 8, head; 9, elytral apex." httpUri="https://zenodo.org/record/3150407/files/figure.png" pageId="1" pageNumber="197">Figs. 1–9</figureCitation>
|
||
). The new species is also distinguished from
|
||
<taxonomicName id="51A34D7D357EFFD14705FBA1FC46FB9C" authorityName="Cartwright & Woodruff. In" authorityYear="1969" box="[831,961,1054,1078]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="spangleri">
|
||
<emphasis id="A4D7EAEC357EFFD14705FBA1FC46FB9C" box="[831,961,1054,1078]" italics="true" pageId="1" pageNumber="197">R. spangleri</emphasis>
|
||
</taxonomicName>
|
||
by the following characters: elytral intervals dull (shiny in
|
||
<taxonomicName id="51A34D7D357EFFD14510FBFDFE2BFBF0" authorityName="Cartwright & Woodruff. In" authorityYear="1969" box="[298,428,1090,1114]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="spangleri">
|
||
<emphasis id="A4D7EAEC357EFFD14510FBFDFE2BFBF0" box="[298,428,1090,1114]" italics="true" pageId="1" pageNumber="197">R. spangleri</emphasis>
|
||
</taxonomicName>
|
||
), punctures small and shallow (large and deep in
|
||
<taxonomicName id="51A34D7D357EFFD14782FBFDFBBCFBF0" authorityName="Cartwright & Woodruff. In" authorityYear="1969" box="[952,1083,1090,1114]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="spangleri">
|
||
<emphasis id="A4D7EAEC357EFFD14782FBFDFBBCFBF0" box="[952,1083,1090,1114]" italics="true" pageId="1" pageNumber="197">R. spangleri</emphasis>
|
||
</taxonomicName>
|
||
), and punctures separated by one puncture diameter (less than one puncture diameter in
|
||
<taxonomicName id="51A34D7D357EFFD146E6FBD9FCD7FBD4" box="[732,848,1126,1150]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="spagleri">
|
||
<emphasis id="A4D7EAEC357EFFD146E6FBD9FCD7FBD4" box="[732,848,1126,1150]" italics="true" pageId="1" pageNumber="197">R. spagleri</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DEB96575357EFFD144FDFB35FC12FB6F" pageId="1" pageNumber="197" type="etymology">
|
||
<paragraph id="961C36FE357EFFD144FDFB35FC12FB6F" blockId="1.[151,1437,154,1545]" pageId="1" pageNumber="197">
|
||
<emphasis id="A4D7EAEC357EFFD144FDFB35FECFFB08" bold="true" box="[199,328,1162,1186]" pageId="1" pageNumber="197">Etymology.</emphasis>
|
||
The specific epithet is derived from Chimalapas, the region of the state of
|
||
<collectingRegion id="5467F81C357EFFD14062FB35FB2FFB08" box="[1112,1192,1162,1186]" country="Mexico" name="Oaxaca" pageId="1" pageNumber="197">Oaxaca</collectingRegion>
|
||
where this species was collected, combined with the Latin suffix –ensis, meaning belonging to.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DEB96575357EFFD144FDFB6EFF3BF9A3" pageId="1" pageNumber="197" type="distribution">
|
||
<paragraph id="961C36FE357EFFD144FDFB6EFCF9FA9B" blockId="1.[151,1437,154,1545]" pageId="1" pageNumber="197">
|
||
<emphasis id="A4D7EAEC357EFFD144FDFB6EFEDDFB43" bold="true" box="[199,346,1233,1257]" pageId="1" pageNumber="197">Distribution.</emphasis>
|
||
<taxonomicName id="51A34D7D357EFFD14558FB6DFDEDFB40" authorityName="Mora-Aguilar & Delgado" authorityYear="2019" box="[354,618,1234,1258]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="chimalapensis">
|
||
<emphasis id="A4D7EAEC357EFFD14558FB6DFDEDFB40" box="[354,618,1234,1258]" italics="true" pageId="1" pageNumber="197">Rhyparus chimalapensis</emphasis>
|
||
</taxonomicName>
|
||
is only known to occur in the cloud forest of El Retén, in the Chimalapas region, located at the southeastern,
|
||
<collectingRegion id="5467F81C357EFFD14637FB49FDE7FAA4" box="[525,608,1270,1294]" country="Mexico" name="Oaxaca" pageId="1" pageNumber="197">Oaxaca</collectingRegion>
|
||
,
|
||
<collectingCountry id="EEB4766E357EFFD14650FB49FD3CFAA4" box="[618,699,1270,1294]" name="Mexico" pageId="1" pageNumber="197">Mexico</collectingCountry>
|
||
(
|
||
<figureCitation id="0E982A7B357EFFD146F3FB49FC9FFAA4" box="[713,792,1270,1294]" captionStart="FIGURE 10" captionStartId="3.[151,255,1214,1238]" captionTargetBox="[157,1426,157,1194]" captionTargetId="figure@3.[151,1435,151,1194]" captionTargetPageId="3" captionText="FIGURE 10. Geographical distribution of Mexican and Guatemalan species of Rhyparus. Rhyparus chimalapensis (black circle), R. costaricensis (black triangle), R. mexicanus (open triangle), R. opacus (open circle), and R. suspiciosus (open square). Elevational limits each 600 m; darker and punctate lines indicate higher elevation." httpUri="https://zenodo.org/record/3150411/files/figure.png" pageId="1" pageNumber="197">Fig. 10</figureCitation>
|
||
). The cloud forests at the
|
||
<typeStatus id="4918885C357EFFD14012FB49FBD1FAA4" box="[1064,1110,1270,1294]" pageId="1" pageNumber="197">type</typeStatus>
|
||
locality and neighboring areas (
|
||
<quantity id="515B9B1B357EFFD144A4FAA6FEC5FA98" box="[158,322,1305,1330]" metricMagnitude="3" metricUnit="m" metricValue="1.625" metricValueMax="1.8" metricValueMin="1.45" pageId="1" pageNumber="197" unit="m" value="1625.0" valueMax="1800.0" valueMin="1450.0">1,450–1,800 m</quantity>
|
||
) are highly conserved but not legally protected areas.
|
||
</paragraph>
|
||
<paragraph id="961C36FE357EFFD144ADFA81FD7CFA68" blockId="1.[151,1437,154,1545]" pageId="1" pageNumber="197">
|
||
The distribution of
|
||
<taxonomicName id="51A34D7D357EFFD14559FA81FDEFFAFC" authorityName="Mora-Aguilar & Delgado" authorityYear="2019" box="[355,616,1342,1366]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="chimalapensis">
|
||
<emphasis id="A4D7EAEC357EFFD14559FA81FDEFFAFC" box="[355,616,1342,1366]" italics="true" pageId="1" pageNumber="197">Rhyparus chimalapensis</emphasis>
|
||
</taxonomicName>
|
||
is isolated from the remaining Mexican species of this genus, especially from the more similar species
|
||
<taxonomicName id="51A34D7D357EFFD14599FADDFD8AFAD3" authorityName="Cartwright & Woodruff" authorityYear="1969" box="[419,525,1378,1401]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="opacus">
|
||
<emphasis id="A4D7EAEC357EFFD14599FADDFD8AFAD3" box="[419,525,1378,1401]" italics="true" pageId="1" pageNumber="197">R. opacus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="51A34D7D357EFFD1467BFADDFD42FAD0" authorityName="Cartwright & Woodruff. In" authorityYear="1969" box="[577,709,1378,1402]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="spangleri">
|
||
<emphasis id="A4D7EAEC357EFFD1467BFADDFD42FAD0" box="[577,709,1378,1402]" italics="true" pageId="1" pageNumber="197">R. spangleri</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="51A34D7D357EFFD146E8FADDFC0BFAD3" authorityName="Cartwright & Woodruff" authorityYear="1969" box="[722,908,1378,1402]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="opacus">
|
||
<emphasis id="A4D7EAEC357EFFD146E8FADDFC0BFAD3" box="[722,908,1378,1402]" italics="true" pageId="1" pageNumber="197">Rhyparus opacus</emphasis>
|
||
</taxonomicName>
|
||
is found northern of the Tehuantepec Isthmus, in the Sierra Madre Oriental of northwestern region of Oaxaca and central Veracruz states (
|
||
<figureCitation id="0E982A7B357EFFD14074FA39FB19FA34" box="[1102,1182,1414,1438]" captionStart="FIGURE 10" captionStartId="3.[151,255,1214,1238]" captionTargetBox="[157,1426,157,1194]" captionTargetId="figure@3.[151,1435,151,1194]" captionTargetPageId="3" captionText="FIGURE 10. Geographical distribution of Mexican and Guatemalan species of Rhyparus. Rhyparus chimalapensis (black circle), R. costaricensis (black triangle), R. mexicanus (open triangle), R. opacus (open circle), and R. suspiciosus (open square). Elevational limits each 600 m; darker and punctate lines indicate higher elevation." httpUri="https://zenodo.org/record/3150411/files/figure.png" pageId="1" pageNumber="197">Fig. 10</figureCitation>
|
||
).
|
||
<taxonomicName id="51A34D7D357EFFD1408BFA39FA03FA34" authorityName="Cartwright & Woodruff. In" authorityYear="1969" box="[1201,1412,1414,1438]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="spangleri">
|
||
<emphasis id="A4D7EAEC357EFFD1408BFA39FA03FA34" box="[1201,1412,1414,1438]" italics="true" pageId="1" pageNumber="197">Rhyparus spangleri</emphasis>
|
||
</taxonomicName>
|
||
is only known for a single male from Turrialba,
|
||
<collectingCountry id="EEB4766E357EFFD14645FA15FD70FA68" box="[639,759,1450,1474]" name="Costa Rica" pageId="1" pageNumber="197">Costa Rica</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="961C36FE357EFFD144FDFA71FF3BF9A3" blockId="1.[151,1437,154,1545]" pageId="1" pageNumber="197">
|
||
<emphasis id="A4D7EAEC357EFFD144FDFA71FEFBFA4F" bold="true" box="[199,380,1485,1510]" pageId="1" pageNumber="197">Natural history.</emphasis>
|
||
The only known specimen collected was caught in a flight intercept trap, located inside a cloud forest.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |