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<document ID-DOI="http://doi.org/10.5281/zenodo.6094633" ID-GBIF-Dataset="db290dd1-5814-46cd-bc8c-1f2e1a199756" ID-GBIF-Taxon="114926048" ID-ISSN="1175-5326" ID-Zenodo-Dep="6094633" ID-ZooBank="97FCBEF2-E95E-47A8-A959-408241A2574D" checkinTime="1447430318407" checkinUser="donat" docAuthor="S. O. Shattuck" docDate="2015" docId="104687953472B456FF17FF26FCE7FD2A" docLanguage="en" docName="29697.pdf" docOrigin="Zootaxa 3955 (2)" docTitle="Myrmecorhynchus emeryi Andre" docType="treatment" docVersion="5" lastPageId="6" lastPageNumber="289" masterDocId="EC7FFFED3476B450FF80FFB1FFBFF972" masterDocTitle="A review of the ant genus Myrmecorhynchus (Hymenoptera: Formicidae)" masterLastPageNumber="290" masterPageNumber="283" pageId="4" pageNumber="287" updateTime="1644953846168" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A review of the ant genus Myrmecorhynchus (Hymenoptera: Formicidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>S. O. Shattuck</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2015</mods:date>
<mods:detail type="volume">
<mods:number>3955</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>283</mods:start>
<mods:end>290</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.11646/zootaxa.3955.2.6</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="ZooBank">97FCBEF2-E95E-47A8-A959-408241A2574D</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6094633" ID-GBIF-Taxon="114926048" ID-Zenodo-Dep="6094633" LSID="urn:lsid:plazi:treatment:104687953472B456FF17FF26FCE7FD2A" httpUri="http://treatment.plazi.org/id/104687953472B456FF17FF26FCE7FD2A" lastPageId="6" lastPageNumber="289" pageId="4" pageNumber="287">
<subSubSection box="[151,554,151,177]" pageId="4" pageNumber="287" type="nomenclature">
<paragraph box="[151,554,151,177]" pageId="4" pageNumber="287">
<taxonomicName authority="Andre" authorityName="Andre" box="[151,554,151,177]" class="Insecta" family="Formicidae" genus="Myrmecorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="287" phylum="Arthropoda" rank="species" species="emeryi">
<heading bold="true" box="[151,554,151,177]" fontSize="11" level="1" pageId="4" pageNumber="287" reason="1">
<emphasis bold="true" box="[151,554,151,177]" pageId="4" pageNumber="287">
<emphasis bold="true" box="[151,467,151,177]" italics="true" pageId="4" pageNumber="287">Myrmecorhynchus emeryi</emphasis>
André
</emphasis>
</heading>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection box="[151,295,186,211]" pageId="4" pageNumber="287" type="description">
<paragraph box="[151,295,186,211]" pageId="4" pageNumber="287">
(
<figureCitation box="[159,231,186,211]" captionStart="FIGURE 2" captionStartId="2.[151,249,994,1016]" captionTargetBox="[211,1375,195,972]" captionTargetId="figure@2.[211,1376,195,972]" captionTargetPageId="2" captionText="FIGURE 2. Myrmecorhynchus emeryi worker, 9.5 km S Bungendore, New South Wales, S. O. Shattuck (ANIC 32 - 042179)." pageId="4" pageNumber="287">Figs 2</figureCitation>
,
<figureCitation box="[243,259,187,211]" captionStart="FIGURE 4" captionStartId="4.[151,249,1870,1892]" captionText="FIGURE 4. Myrmecorhynchus emeryi, major worker, Gerroa, New South Wales, B. B. Lowery (ANIC 32 - 010956)." pageId="4" pageNumber="287">4</figureCitation>
,
<figureCitation box="[271,285,186,211]" captionStart="FIGURE 5" captionStartId="5.[151,249,1411,1433]" captionTargetBox="[151,1436,401,1393]" captionTargetId="figure@5.[151,1436,401,1393]" captionTargetPageId="5" captionText="FIGURE 5. Myrmecorhynchus emeryi, minor worker, Mt. Remarkable, Flinders Ranges, South Australia, P. J. M. Greenslade (ANIC 32 - 010964)." pageId="4" pageNumber="287">
<number box="[271,285,186,211]" pageId="4" pageNumber="287" value="5.0">5</number>
</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="287" type="reference_group">
<paragraph box="[151,605,258,280]" pageId="4" pageNumber="287">
<treatmentCitation box="[151,605,258,280]" pageId="4" pageNumber="287">
<taxonomicName authority="Andre, 1896: 254" authorityName="Andre" authorityPageNumber="254" authorityYear="1896" box="[151,601,258,280]" class="Insecta" family="Formicidae" genus="Myrmecorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="287" phylum="Arthropoda" rank="species" species="emeryi">
<emphasis box="[151,414,258,280]" italics="true" pageId="4" pageNumber="287">Myrmecorhynchus emeryi</emphasis>
André,
<number box="[496,551,258,280]" pageId="4" pageNumber="287" value="1896.0">1896</number>
:
<number box="[560,601,258,280]" pageId="4" pageNumber="287" value="254.0">254</number>
</taxonomicName>
.
</treatmentCitation>
</paragraph>
<paragraph pageId="4" pageNumber="287">
<treatmentCitation box="[151,622,289,311]" pageId="4" pageNumber="287">
<taxonomicName authority="Clark, 1934: 43" authorityName="Clark" authorityPageNumber="43" authorityYear="1934" box="[151,618,289,311]" class="Insecta" family="Formicidae" genus="Myrmecorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="287" phylum="Arthropoda" rank="species" species="musgravei">
<emphasis box="[151,451,289,311]" italics="true" pageId="4" pageNumber="287">Myrmecorhynchus musgravei</emphasis>
Clark,
<number box="[526,580,289,311]" pageId="4" pageNumber="287" value="1934.0">1934</number>
:
<number box="[590,618,289,311]" pageId="4" pageNumber="287" value="43.0">43</number>
</taxonomicName>
.
</treatmentCitation>
<treatmentCitationLabel box="[627,819,289,311]" pageId="4" pageNumber="287">NEW SYNONYM</treatmentCitationLabel>
<treatmentCitation box="[151,619,320,342]" pageId="4" pageNumber="287">
<taxonomicName authority="Clark, 1934: 46" authorityName="Clark" authorityPageNumber="46" authorityYear="1934" box="[151,615,320,342]" class="Insecta" family="Formicidae" genus="Myrmecorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="287" phylum="Arthropoda" rank="species" species="rufithorax">
<emphasis box="[151,448,320,342]" italics="true" pageId="4" pageNumber="287">Myrmecorhynchus rufithorax</emphasis>
Clark,
<number box="[523,577,320,342]" pageId="4" pageNumber="287" value="1934.0">1934</number>
:
<number box="[587,615,320,342]" pageId="4" pageNumber="287" value="46.0">46</number>
</taxonomicName>
.
</treatmentCitation>
<treatmentCitationLabel box="[625,817,320,342]" pageId="4" pageNumber="287">NEW SYNONYM</treatmentCitationLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="287" type="nomenclature">
<paragraph pageId="4" pageNumber="287">
<emphasis bold="true" box="[151,227,387,411]" pageId="4" pageNumber="287">
<typeStatus box="[151,222,387,411]" pageId="4" pageNumber="287">Types</typeStatus>
.
</emphasis>
<materialsCitation ID-GBIF-Occurrence="1227780108" country="Australia" pageId="4" pageNumber="287" stateProvince="Victoria">
<taxonomicName box="[235,348,387,411]" class="Insecta" family="Formicidae" genus="Myrmecorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="287" phylum="Arthropoda" rank="species" species="emeryi">
<emphasis box="[235,348,387,411]" italics="true" pageId="4" pageNumber="287">M. emeryi</emphasis>
</taxonomicName>
:
<typeStatus box="[364,468,386,411]" pageId="4" pageNumber="287">Holotype</typeStatus>
worker from the Victorian Alps,
<collectingRegion box="[845,935,386,411]" pageId="4" pageNumber="287">Victoria</collectingRegion>
(Musee National d'Histoire Naturelle, Paris, examined)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="1227780104" country="Australia" pageId="4" pageNumber="287" stateProvince="Victoria">
<emphasis box="[292,454,423,447]" italics="true" pageId="4" pageNumber="287">
<taxonomicName box="[292,454,423,447]" class="Insecta" family="Formicidae" genus="Myrmecorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="287" phylum="Arthropoda" rank="species" species="musgravei">M. musgravei</taxonomicName>
</emphasis>
:
<typeStatus box="[478,582,422,447]" pageId="4" pageNumber="287">Holotype</typeStatus>
male from Lamington Nat. Park (as National Park), Queensland (A. Musgrave,
<date box="[279,498,458,483]" pageId="4" pageNumber="287" value="1926-12-19">
<number box="[279,307,459,483]" pageId="4" pageNumber="287" value="19.0">19</number>
December
<number box="[441,498,458,483]" pageId="4" pageNumber="287" value="1926.0">1926</number>
</date>
) (Australian Museum, Sydney)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="1227780107" country="Australia" pageId="4" pageNumber="287" stateProvince="Victoria">
<taxonomicName box="[875,1028,458,483]" class="Insecta" family="Formicidae" genus="Myrmecorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="287" phylum="Arthropoda" rank="species" species="rufithorax">
<emphasis box="[875,1028,458,483]" italics="true" pageId="4" pageNumber="287">M. rufithorax</emphasis>
</taxonomicName>
:
<typeStatus box="[1045,1137,459,483]" pageId="4" pageNumber="287">Syntype</typeStatus>
workers from Warburton,
<collectingRegion box="[151,241,494,519]" pageId="4" pageNumber="287">Victoria</collectingRegion>
(L. B. Thorn, in trees) (Museum
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="1227780092" box="[616,856,494,519]" country="Australia" pageId="4" pageNumber="287" stateProvince="Victoria">
<collectingRegion box="[616,708,494,519]" pageId="4" pageNumber="287">Victoria</collectingRegion>
, Melbourne)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="287" type="diagnosis">
<paragraph pageId="4" pageNumber="287">
<emphasis bold="true" box="[199,321,530,555]" pageId="4" pageNumber="287">Diagnosis.</emphasis>
Mesosomal dorsum and legs with abundant erect hairs. Body colour variable, ranging from reddish to reddish-brown.
</paragraph>
</subSubSection>
<caption box="[151,1321,1870,1892]" pageId="4" pageNumber="287">
<paragraph box="[151,1321,1870,1892]" pageId="4" pageNumber="287">
<emphasis bold="true" box="[151,275,1870,1892]" pageId="4" pageNumber="287">FIGURE 4.</emphasis>
<taxonomicName box="[281,544,1870,1892]" class="Insecta" family="Formicidae" genus="Myrmecorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="287" phylum="Arthropoda" rank="species" species="emeryi">
<emphasis box="[281,544,1870,1892]" italics="true" pageId="4" pageNumber="287">Myrmecorhynchus emeryi</emphasis>
</taxonomicName>
, major worker, Gerroa, New South Wales, B.B. Lowery (ANIC 32-010956).
</paragraph>
</caption>
<subSubSection lastPageId="5" lastPageNumber="288" pageId="4" pageNumber="287" type="discussion">
<paragraph lastPageId="5" lastPageNumber="288" pageId="4" pageNumber="287">
<emphasis bold="true" box="[199,335,1931,1956]" pageId="4" pageNumber="287">Comments.</emphasis>
<emphasis box="[346,641,1931,1956]" italics="true" pageId="4" pageNumber="287">
<taxonomicName box="[346,641,1931,1956]" class="Insecta" family="Formicidae" genus="Myrmecorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="287" phylum="Arthropoda" rank="species" species="emeryi">Myrmecorhynchus emeryi</taxonomicName>
</emphasis>
is the most widely distributed species in the genus, occurring from south-eastern Queensland (with a single record further north) south through New South Wales and Victoria to south-western Western Australia. It is found in a range of habitats including mallee, sandstone scrub and moist gully woodlands, dry to wet sclerophyll and rainforest. Nests are found in dead branches, twigs and vines, in soil most commonly at the base of tree but also under rocks or occasionally in moss. Nests have also been found in dead stumps (including a
<emphasis box="[383,539,223,248]" italics="true" pageId="5" pageNumber="288">
<taxonomicName box="[383,539,223,248]" class="Magnoliopsida" family="Xanthorrhoeaceae" genus="Xanthorrhoea" higherTaxonomySource="IPNI" kingdom="Pantae" order="Xanthorrhoeales" pageId="5" pageNumber="288" phylum="Magnoliophyta" rank="genus">Xanthorrhoea</taxonomicName>
</emphasis>
stump). Three females were found in a cavity in a small dead twig
<quantity box="[1313,1370,224,248]" metricMagnitude="0" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="288" unit="m" value="2.5">2.5m</quantity>
(
<quantity box="[1388,1428,223,248]" metricMagnitude="0" metricUnit="m" metricValue="2.4384" pageId="5" pageNumber="288" unit="ft" value="8.0">8 ft</quantity>
) above the ground. This would suggest that this species is pleometrotic, however additional observations are needed to confirm this. Workers are most commonly seen foraging on trees and low vegetation and have been found feeding at extra-floral nectaries of
<emphasis box="[539,740,332,356]" italics="true" pageId="5" pageNumber="288">
<taxonomicName box="[539,740,332,356]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" higherTaxonomySource="CoL" kingdom="Plantae" order="Fabales" pageId="5" pageNumber="288" phylum="Tracheophyta" rank="species" species="pycnantha">Acacia pycnantha</taxonomicName>
</emphasis>
.
</paragraph>
<caption pageId="5" pageNumber="288" targetBox="[151,1436,401,1393]" targetPageId="5">
<paragraph pageId="5" pageNumber="288">
<emphasis bold="true" box="[151,276,1411,1433]" pageId="5" pageNumber="288">FIGURE 5.</emphasis>
<emphasis box="[284,549,1411,1433]" italics="true" pageId="5" pageNumber="288">
<taxonomicName box="[284,549,1411,1433]" class="Insecta" family="Formicidae" genus="Myrmecorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="288" phylum="Arthropoda" rank="species" species="emeryi">Myrmecorhynchus emeryi</taxonomicName>
</emphasis>
, minor worker, Mt. Remarkable, Flinders Ranges, South Australia, P.J.M. Greenslade (ANIC32-010964).
</paragraph>
</caption>
<paragraph pageId="5" pageNumber="288">
When describing the taxon
<taxonomicName authority="Clark (1934)" authorityName="Clark" authorityYear="1934" box="[523,843,1508,1533]" class="Insecta" family="Formicidae" genus="Myrmecorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="288" phylum="Arthropoda" rank="species" species="rufithorax">
<emphasis box="[523,677,1508,1533]" italics="true" pageId="5" pageNumber="288">M. rufithorax</emphasis>
, Clark (
<number box="[777,835,1508,1533]" pageId="5" pageNumber="288" value="1934.0">1934</number>
)
</taxonomicName>
commented that this species was “near
<taxonomicName authority="Andre" authorityName="Andre" class="Insecta" family="Formicidae" genus="Myrmecorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="288" phylum="Arthropoda" rank="species" species="emeryi">
<emphasis box="[1318,1436,1509,1533]" italics="true" pageId="5" pageNumber="288">M. emeryi</emphasis>
Andre
</taxonomicName>
”, but gave no specific characters to separate these two taxa. With the presently available material, there seems little doubt that
<emphasis box="[430,588,1580,1605]" italics="true" pageId="5" pageNumber="288">
<taxonomicName box="[430,588,1580,1605]" class="Insecta" family="Formicidae" genus="Myrmecorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="288" phylum="Arthropoda" rank="species" species="rufithorax">M. rufithorax</taxonomicName>
</emphasis>
represents the pale form of a single variable species, with
<emphasis box="[1316,1436,1581,1605]" italics="true" pageId="5" pageNumber="288">
<taxonomicName box="[1316,1436,1581,1605]" class="Insecta" family="Formicidae" genus="Myrmecorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="288" phylum="Arthropoda" rank="species" species="emeryi">M. emeryi</taxonomicName>
</emphasis>
representing the darker form. Intermediately coloured individuals are abundant and no additional characters could be found to suggest that more than a single species is involved. Because of this these two taxa are here considered synonyms.
<emphasis box="[282,618,1688,1713]" italics="true" pageId="5" pageNumber="288">
<taxonomicName box="[282,618,1688,1713]" class="Insecta" family="Formicidae" genus="Myrmecorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="288" phylum="Arthropoda" rank="species" species="musgravei">Myrmecorhynchus musgravei</taxonomicName>
</emphasis>
, also described by Clark (
<number box="[931,990,1688,1713]" pageId="5" pageNumber="288" value="1934.0">1934</number>
), was based on an unassociated male. Clark mentions several characters which separate
<taxonomicName box="[724,880,1725,1749]" class="Insecta" family="Formicidae" genus="Myrmecorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="288" phylum="Arthropoda" rank="species" species="musgravei">
<emphasis box="[724,880,1725,1749]" italics="true" pageId="5" pageNumber="288">M. musgravei</emphasis>
</taxonomicName>
from
<taxonomicName box="[953,1069,1725,1749]" class="Insecta" family="Formicidae" genus="Myrmecorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="288" phylum="Arthropoda" rank="species" species="emeryi">
<emphasis box="[953,1069,1725,1749]" italics="true" pageId="5" pageNumber="288">M. emeryi</emphasis>
</taxonomicName>
, including differences in colour (testaceous in
<taxonomicName box="[322,481,1761,1785]" class="Insecta" family="Formicidae" genus="Myrmecorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="288" phylum="Arthropoda" rank="species" species="musgravei">
<emphasis box="[322,481,1761,1785]" italics="true" pageId="5" pageNumber="288">M. musgravei</emphasis>
</taxonomicName>
, reddish yellow in
<taxonomicName box="[717,836,1761,1785]" class="Insecta" family="Formicidae" genus="Myrmecorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="288" phylum="Arthropoda" rank="species" species="emeryi">
<emphasis box="[717,836,1761,1785]" italics="true" pageId="5" pageNumber="288">M. emeryi</emphasis>
</taxonomicName>
) and wing morphology (discoidal cell large in
<taxonomicName class="Insecta" family="Formicidae" genus="Myrmecorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="288" phylum="Arthropoda" rank="species" species="musgravei">
<emphasis italics="true" pageId="5" pageNumber="288">M. musgravei</emphasis>
</taxonomicName>
, small in
<taxonomicName box="[393,511,1797,1821]" class="Insecta" family="Formicidae" genus="Myrmecorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="288" phylum="Arthropoda" rank="species" species="emeryi">
<emphasis box="[393,511,1797,1821]" italics="true" pageId="5" pageNumber="288">M. emeryi</emphasis>
</taxonomicName>
). However the limited available male material makes it difficult to assess the taxonomic significance of these and other characters used by Clark (
<number box="[921,981,1832,1857]" pageId="5" pageNumber="288" value="1934.0">1934</number>
). Numerous worker collections from the region of the
<typeStatus box="[302,351,1869,1893]" pageId="5" pageNumber="288">type</typeStatus>
locality of
<emphasis box="[481,635,1869,1893]" italics="true" pageId="5" pageNumber="288">
<taxonomicName box="[481,635,1869,1893]" class="Insecta" family="Formicidae" genus="Myrmecorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="288" phylum="Arthropoda" rank="species" species="musgravei">M. musgravei</taxonomicName>
</emphasis>
in south-eastern Queensland suggest that a single species occurs there. There is no evidence for two distinct species in this region. Combined, the presently available information suggests that
<taxonomicName box="[200,354,1941,1965]" class="Insecta" family="Formicidae" genus="Myrmecorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="288" phylum="Arthropoda" rank="species" species="musgravei">
<emphasis box="[200,354,1941,1965]" italics="true" pageId="5" pageNumber="288">M. musgravei</emphasis>
</taxonomicName>
is the male of
<taxonomicName box="[519,632,1941,1965]" class="Insecta" family="Formicidae" genus="Myrmecorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="288" phylum="Arthropoda" rank="species" species="emeryi">
<emphasis box="[519,632,1941,1965]" italics="true" pageId="5" pageNumber="288">M. emeryi</emphasis>
</taxonomicName>
and not a separate species.
</paragraph>
<paragraph pageId="5" pageNumber="288">
The major worker of this species was described by Emery (
<number box="[864,924,1976,2001]" pageId="5" pageNumber="288" value="1898.0">1898</number>
), the queen and male by Clark (
<number box="[1281,1339,1976,2001]" pageId="5" pageNumber="288" value="1934.0">1934</number>
) and the larva by Wheeler (
<number box="[359,417,2012,2037]" pageId="5" pageNumber="288" value="1917.0">1917</number>
) and Wheeler &amp; Wheeler (
<number box="[721,781,2012,2037]" pageId="5" pageNumber="288" value="1968.0">1968</number>
).
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="289" type="materials_examined">
<paragraph pageId="6" pageNumber="289">
<materialsCitation ID-GBIF-Occurrence="1227780093" box="[199,1240,151,176]" country="Australia" pageId="6" pageNumber="289" stateProvince="Australian Capital Territory">
<emphasis bold="true" box="[199,554,151,176]" pageId="6" pageNumber="289">Material examined. Australia</emphasis>
:
<emphasis box="[570,890,151,176]" italics="true" pageId="6" pageNumber="289">
<collectingRegion box="[570,890,151,176]" pageId="6" pageNumber="289">Australian Capital Territory</collectingRegion>
</emphasis>
: Black Mountain (Greaves,T.)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780099" country="Australia" pageId="6" pageNumber="289" stateProvince="Australian Capital Territory">Black Mountain Reserve (Liepa,Z.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780098" box="[386,820,188,212]" country="Australia" pageId="6" pageNumber="289" stateProvince="Australian Capital Territory">Black Mt. (Greaves,T.; Lowery,B.B.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780090" box="[837,1252,188,212]" country="Australia" pageId="6" pageNumber="289" stateProvince="Australian Capital Territory">Black Mt., Canberra (Lowery,B.B.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780109" country="Australia" pageId="6" pageNumber="289" stateProvince="Australian Capital Territory">Mount Ainslie (Lowery,B.B.)</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="1227780112" box="[327,880,223,248]" country="Australia" pageId="6" pageNumber="289" stateProvince="Australian Capital Territory">
<emphasis box="[327,522,223,248]" italics="true" pageId="6" pageNumber="289">New South Wales</emphasis>
:
<quantity box="[537,598,223,248]" metricMagnitude="4" metricUnit="m" metricValue="1.609344" pageId="6" pageNumber="289" unit="mi" value="10.0">10mi</quantity>
. S Kiama (Lowery,B.B.)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780100" box="[892,1311,223,248]" country="Australia" pageId="6" pageNumber="289" stateProvince="Australian Capital Territory">
<quantity box="[892,955,223,248]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="289" unit="km" value="20.0">20km</quantity>
E Legume (Greenslade,P.J.M.)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780094" country="Australia" pageId="6" pageNumber="289" stateProvince="Australian Capital Territory">
<quantity box="[1323,1377,223,248]" metricMagnitude="3" metricUnit="m" metricValue="6.437376" pageId="6" pageNumber="289" unit="mi" value="4.0">4 mi</quantity>
. E of Berry (Lowery,B.B.)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780101" box="[411,852,260,284]" country="Australia" pageId="6" pageNumber="289" stateProvince="Australian Capital Territory">
<quantity box="[411,483,260,284]" metricMagnitude="3" metricUnit="m" metricValue="9.5" pageId="6" pageNumber="289" unit="km" value="9.5">9.5km</quantity>
S Bungendore (Shattuck,S.O.)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780096" box="[870,1319,260,284]" country="Australia" pageId="6" pageNumber="289" stateProvince="Australian Capital Territory">Berry Mountain, Berry (Lowery,B.B.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780091" country="Australia" pageId="6" pageNumber="289" stateProvince="Australian Capital Territory">Bilambil (Lowery,B.B.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780110" box="[332,893,295,320]" country="Australia" pageId="6" pageNumber="289" stateProvince="Australian Capital Territory">Blue Knob Mt., Nightcap Range (Lowery,B.B.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780097" box="[910,1237,295,320]" country="Australia" pageId="6" pageNumber="289" stateProvince="Australian Capital Territory">Bogan River (Armstrong,J.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780105" country="Australia" pageId="6" pageNumber="289" stateProvince="Australian Capital Territory">Bola Creek NP, Sydney (Lowery,B.B.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780103" box="[442,897,332,356]" country="Australia" pageId="6" pageNumber="289" stateProvince="Australian Capital Territory">Cambewarra Mountain (Lowery,B.B.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780102" country="Australia" pageId="6" pageNumber="289" stateProvince="Australian Capital Territory">Cool and Candle Crk., Kuring-gai Chase (Lowery,B.B.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780111" box="[333,693,367,392]" country="Australia" pageId="6" pageNumber="289" stateProvince="Australian Capital Territory">Cowan, Sydney (Lowery,B.B.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780095" box="[709,1296,367,392]" country="Australia" pageId="6" pageNumber="289" stateProvince="Australian Capital Territory">Dalrymple Forest, Pymble, Sydney (Lowery,B.B.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780114" country="Australia" pageId="6" pageNumber="289" stateProvince="Australian Capital Territory">Foxground (Lowery,B.B.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780129" box="[342,819,404,428]" country="Australia" pageId="6" pageNumber="289" stateProvince="Australian Capital Territory">Frenchs Forest, Sydney (Lowery,B.B.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780126" box="[844,1108,404,428]" country="Australia" pageId="6" pageNumber="289" stateProvince="Australian Capital Territory">Gerroa (Lowery,B.B.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780122" country="Australia" pageId="6" pageNumber="289" stateProvince="Australian Capital Territory">
Gerroa,
<quantity box="[1239,1300,404,428]" metricMagnitude="4" metricUnit="m" metricValue="1.609344" pageId="6" pageNumber="289" unit="mi" value="10.0">10mi</quantity>
. S Kiama (Lowery,B.B.)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780135" box="[335,960,439,464]" country="Australia" pageId="6" pageNumber="289" stateProvince="Australian Capital Territory">Gibraltar Ra. Nat. Pk. via Glen Innes (Monteith,G.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780127" box="[979,1431,439,464]" country="Australia" pageId="6" pageNumber="289" stateProvince="Australian Capital Territory">Hills to W of Mudgee (Lowery,B.B.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780119" box="[151,600,476,500]" country="Australia" pageId="6" pageNumber="289" stateProvince="Australian Capital Territory">Jerusalem Bay, Cowan (Lowery,B.B.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780118" box="[619,901,476,500]" country="Australia" pageId="6" pageNumber="289" stateProvince="Australian Capital Territory">Kyogle (Froggatt,W.W.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780123" box="[919,1378,476,500]" country="Australia" pageId="6" pageNumber="289" stateProvince="Australian Capital Territory">Mt. Saddleback, Kiama (Lowery,B.B.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780138" country="Australia" pageId="6" pageNumber="289" stateProvince="Australian Capital Territory">Mt. Warning (Lowery,B.B.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780117" box="[444,1095,511,536]" country="Australia" pageId="6" pageNumber="289" stateProvince="Australian Capital Territory">Riverview College, Lane Cove, Sydney (Lowery,B.B.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780128" country="Australia" pageId="6" pageNumber="289" stateProvince="Australian Capital Territory">Riverview College, Sydney (Lowery,B.B.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780133" box="[332,747,548,572]" country="Australia" pageId="6" pageNumber="289" stateProvince="Australian Capital Territory">Runnyford Ck., Nelligen (Liepa,Z.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780125" box="[764,1319,548,572]" country="Australia" pageId="6" pageNumber="289" stateProvince="Australian Capital Territory">St. Ives, Frenchs Forest, Sydney (Lowery,B.B.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780115" country="Australia" pageId="6" pageNumber="289" stateProvince="Australian Capital Territory">
Tooloom Plateau,
<quantity box="[252,316,583,608]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="289" unit="km" value="14.0">14km</quantity>
W Urbenville (Naumann,I.D.)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780124" box="[694,1091,583,608]" country="Australia" pageId="6" pageNumber="289" stateProvince="Australian Capital Territory">W of Mt. Warning (Lowery,B.B.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780116" box="[1107,1431,583,608]" country="Australia" pageId="6" pageNumber="289" stateProvince="Australian Capital Territory">Wallerawang (Lowery,B.B.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780121" box="[151,536,620,644]" country="Australia" pageId="6" pageNumber="289" stateProvince="Australian Capital Territory">Wentworth Falls (Wheeler,W.M.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780132" box="[551,868,620,644]" country="Australia" pageId="6" pageNumber="289" stateProvince="Australian Capital Territory">Wentworth Falls, Blue Mts</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="1227780139" box="[882,1048,620,644]" country="Australia" pageId="6" pageNumber="289" stateProvince="Australian Capital Territory">(Lowery,B.B.)</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="1227780136" country="Australia" pageId="6" pageNumber="289" stateProvince="Queensland">
<emphasis box="[1062,1196,620,644]" italics="true" pageId="6" pageNumber="289">
<collectingRegion box="[1062,1196,620,644]" pageId="6" pageNumber="289">Queensland</collectingRegion>
</emphasis>
:
<quantity box="[1214,1278,620,644]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="6" pageNumber="289" unit="km" value="13.0">13km</quantity>
NE Eungella (Ward,P.S.)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780134" box="[297,658,655,680]" country="Australia" pageId="6" pageNumber="289" stateProvince="Queensland">
<quantity box="[297,347,655,680]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="289" unit="km" value="1.0">1km</quantity>
SSE Eungella (Ward,P.S.)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780137" box="[674,1216,655,680]" country="Australia" pageId="6" pageNumber="289" stateProvince="Queensland">
<quantity box="[674,735,655,680]" metricMagnitude="4" metricUnit="m" metricValue="8.04672" pageId="6" pageNumber="289" unit="mi" value="50.0">50mi</quantity>
. W Mackay, Eungella NP (Lowery,B.B.)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780131" country="Australia" pageId="6" pageNumber="289" stateProvince="Queensland">
<quantity box="[1232,1282,655,680]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="289" unit="km" value="7.0">7km</quantity>
NNW North Tamborine (Ward,P.S.)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780120" box="[459,1210,692,716]" country="Australia" pageId="6" pageNumber="289" stateProvince="Queensland">Diggings Road, Eungella (Monteith,G.B. &amp; Thompson,G.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780130" country="Australia" pageId="6" pageNumber="289" stateProvince="Queensland">Lamington N.P. (Andersen,A.N.); Mt. Tamborine (Greaves,T.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780159" box="[702,1431,727,752]" country="Australia" pageId="6" pageNumber="289" stateProvince="Queensland">
Palm Grove N.P.,
<quantity box="[920,970,727,752]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="289" unit="km" value="8.0">8km</quantity>
ESE North Tamborine (Shattuck,S.O.)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780161" box="[151,432,764,788]" country="Australia" pageId="6" pageNumber="289" stateProvince="Queensland">Tomewin (Lowery,B.B.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780148" box="[447,915,764,788]" country="Australia" pageId="6" pageNumber="289" stateProvince="Queensland">Tomewin, Murwillumbah (Lowery,B.B.)</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="1227780167" country="Australia" pageId="6" pageNumber="289" stateProvince="Queensland">
<emphasis box="[929,1107,764,788]" italics="true" pageId="6" pageNumber="289">South Australia</emphasis>
: Belair Rec. Park, Adelaide (O'Dowd,D.J.)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780165" box="[332,822,799,824]" country="Australia" pageId="6" pageNumber="289" stateProvince="Queensland">Bridgewater (Greenslade,P.J.M. &amp; Hutson)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780144" box="[835,1383,799,824]" country="Australia" pageId="6" pageNumber="289" stateProvince="Queensland">Kangaroo Is., Breakneck R. (Greenslade,P.J.M.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780143" country="Australia" pageId="6" pageNumber="289" stateProvince="Queensland">Mt. Bold Reservoir, SE of Adelaide (Ward,P.S.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780147" box="[697,992,836,860]" country="Australia" pageId="6" pageNumber="289" stateProvince="Queensland">Mt. Lofty (Lowery,B.B.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780150" box="[1011,1377,836,860]" country="Australia" pageId="6" pageNumber="289" stateProvince="Queensland">Mt. Lofty summit (Gross,G.F.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780142" country="Australia" pageId="6" pageNumber="289" stateProvince="Queensland">Mt. Remarkable, Flinders Ranges (Greenslade,P.J.M.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780162" box="[727,1006,871,896]" country="Australia" pageId="6" pageNumber="289" stateProvince="Queensland">Sevenhill (Lowery,B.B.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780153" box="[1019,1338,871,896]" country="Australia" pageId="6" pageNumber="289" stateProvince="Queensland">Spring Gully (Lowery,B.B.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780152" country="Australia" pageId="6" pageNumber="289" stateProvince="Queensland">Streaky Bay (Lowery,B.B.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780140" box="[380,691,908,932]" country="Australia" pageId="6" pageNumber="289" stateProvince="Queensland">Upper Sturt (Herridge,J.A.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780155" box="[702,1178,908,932]" country="Australia" pageId="6" pageNumber="289" stateProvince="Queensland">
Worlds End,
<quantity box="[848,912,908,932]" metricMagnitude="4" metricUnit="m" metricValue="2.3" pageId="6" pageNumber="289" unit="km" value="23.0">23km</quantity>
S Burra (Lowery,B.B.)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="1227780141" country="Australia" pageId="6" pageNumber="289" stateProvince="Victoria">
<emphasis box="[1188,1276,909,932]" italics="true" pageId="6" pageNumber="289">
<collectingRegion box="[1188,1276,909,932]" pageId="6" pageNumber="289">Victoria</collectingRegion>
</emphasis>
:
<quantity box="[1292,1356,908,932]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="289" unit="km" value="15.0">15km</quantity>
WNW Yaapeet, Wyperfeld Natl. Pk (Andersen,A.N.)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780146" box="[723,1101,943,968]" country="Australia" pageId="6" pageNumber="289" stateProvince="Victoria">
<quantity box="[723,787,943,968]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="289" unit="km" value="20.0">20km</quantity>
S Ouyen (Shattuck,S.O.)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780151" box="[1121,1431,943,968]" country="Australia" pageId="6" pageNumber="289" stateProvince="Victoria">Harrietville (Lowery,B.B.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780168" box="[151,438,980,1004]" country="Australia" pageId="6" pageNumber="289" stateProvince="Victoria">Heathcote (Lowery,B.B.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780160" box="[450,757,980,1004]" country="Australia" pageId="6" pageNumber="289" stateProvince="Victoria">Hurstbridge (Lowery,B.B.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780154" box="[770,1061,980,1004]" country="Australia" pageId="6" pageNumber="289" stateProvince="Victoria">Leongatha (Lowery,B.B.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780164" box="[1074,1354,980,1004]" country="Australia" pageId="6" pageNumber="289" stateProvince="Victoria">Warrandyte (Greaves,T.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780157" country="Australia" pageId="6" pageNumber="289" stateProvince="Victoria">Woori Yallock (as Wooriyalock) (Thorn,L.B.)</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="1227780145" box="[608,1174,1015,1040]" country="Australia" pageId="6" pageNumber="289" stateProvince="Victoria">
<emphasis box="[608,811,1015,1040]" italics="true" pageId="6" pageNumber="289">Western Australia</emphasis>
:
<quantity box="[827,896,1015,1040]" metricMagnitude="4" metricUnit="m" metricValue="1.7702784" pageId="6" pageNumber="289" unit="mi" value="11.0">11 mi</quantity>
. E Pingrup (Greaves,T.)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780149" country="Australia" pageId="6" pageNumber="289" stateProvince="Victoria">
<quantity box="[1187,1251,1015,1040]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="289" unit="km" value="15.0">15km</quantity>
N of Norseman (Lowery,B.B.)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780194" box="[338,615,1052,1076]" country="Australia" pageId="6" pageNumber="289" stateProvince="Victoria">Corrigin (Lowery,B.B.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780178" box="[637,963,1052,1076]" country="Australia" pageId="6" pageNumber="289" stateProvince="Victoria">Grass Patch (Lowery,B.B.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780177" country="Australia" pageId="6" pageNumber="289" stateProvince="Victoria">
Junana Rock,
<quantity box="[1163,1212,1052,1076]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="289" unit="km" value="9.0">9km</quantity>
NW Mt. Ragged (Taylor,R.W.)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780195" box="[319,558,1087,1112]" country="Australia" pageId="6" pageNumber="289" stateProvince="Victoria">Pingrup (Greaves,T.)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1227780184" box="[569,852,1087,1112]" country="Australia" pageId="6" pageNumber="289" stateProvince="Victoria">Tingledale (Taylor,R.W.)</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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