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<document id="DE608DB554CF93EE64B52A0CF7AAFD92" ID-DOI="10.11646/zootaxa.5301.4.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="8036117" ID-ZooBank="01F6333F-807B-448A-80C8-EC1F8CE030EA" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1686723262927" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2023" docId="002087EAFFAF947064A8FDBCFB3FFC12" docLanguage="en" docName="zootaxa.5301.4.6.pdf" docOrigin="Zootaxa 5301 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5301.4.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paraphloeostiba apicalis" docType="treatment" docVersion="2" lastPageNumber="497" masterDocId="FC19FF92FFA4947B643FFFA9FFE2FFA3" masterDocTitle="New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines" masterLastPageNumber="500" masterPageNumber="486" pageNumber="497" updateTime="1686723785076" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="1A94187B151F7F8E104A43C0FD756039">New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines</mods:title>
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<mods:namePart id="A22FB20B0976B2B69E30E9CD3CB9745D">Shavrin, Alexey V.</mods:namePart>
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<mods:title id="F48D437A9F29DBDF74EF480981C4E0FC">Zootaxa</mods:title>
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<mods:date id="C6C4299263998D39205BE6A1EFAE47DC">2023</mods:date>
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<mods:number id="6951E5DC0ED312AB4937815D73EE049A">2023-06-13</mods:number>
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<mods:classification id="AA7D83D6A87022439C34590B262F3D9E">journal article</mods:classification>
<mods:identifier id="AC67467776E0AC1C1929EF171F5ED9C0" type="DOI">10.11646/zootaxa.5301.4.6</mods:identifier>
<mods:identifier id="0ACEEB22081832784BF438B06ADDAA9B" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
<treatment id="002087EAFFAF947064A8FDBCFB3FFC12" LSID="urn:lsid:plazi:treatment:002087EAFFAF947064A8FDBCFB3FFC12" httpUri="http://treatment.plazi.org/id/002087EAFFAF947064A8FDBCFB3FFC12" lastPageNumber="497" pageId="11" pageNumber="497">
<subSubSection id="C0936577FFAF947064A8FDBCFD7BFD8C" box="[151,665,533,560]" pageId="11" pageNumber="497" type="nomenclature">
<paragraph id="883636FCFFAF947064A8FDBCFD7BFD8C" blockId="11.[151,665,533,595]" box="[151,665,533,560]" pageId="11" pageNumber="497">
<heading id="D37E8190FFAF947064A8FDBCFD7BFD8C" bold="true" box="[151,665,533,560]" fontSize="11" level="1" pageId="11" pageNumber="497" reason="1">
<taxonomicName id="4F894D7FFFAF947064A8FDBCFD7BFD8C" ID-CoL="6TQXL" authority="(Cameron, 1925)" baseAuthorityName="Cameron" baseAuthorityYear="1925" box="[151,665,533,560]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="497" phylum="Arthropoda" rank="species" species="apicalis">
<emphasis id="BAFDEAEEFFAF947064A8FDBCFD7BFD8C" bold="true" box="[151,665,533,560]" pageId="11" pageNumber="497">
<emphasis id="BAFDEAEEFFAF947064A8FDBCFE23FD93" bold="true" box="[151,449,533,560]" italics="true" pageId="11" pageNumber="497">Paraphloeostiba apicalis</emphasis>
(
<bibRefCitation id="EC184B0DFFAF947065EFFDBCFD73FD8C" author="Cameron, M." box="[464,657,533,559]" pageId="11" pageNumber="497" pagination="174 - 198" refId="ref8185" refString="Cameron, M. (1925) New Staphylinidae from the Dutch East Indies. Treubia, 6 (2), 174 - 198." type="journal article" year="1925">Cameron, 1925</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C0936577FFAF947064A8FD90FEBAFDF0" box="[151,344,569,595]" pageId="11" pageNumber="497" type="description">
<paragraph id="883636FCFFAF947064A8FD90FEBAFDF0" blockId="11.[151,665,533,595]" box="[151,344,569,595]" pageId="11" pageNumber="497">
(
<figureCitation id="10B22A79FFAF947064A0FD90FF1FFDF0" box="[159,253,569,595]" captionStart="FIGURES 33–36" captionStartId="10.[151,264,843,867]" captionTargetBox="[159,1429,187,811]" captionTargetId="figure-15@10.[151,1436,181,819]" captionTargetPageId="10" captionText="FIGURES 33–36. Habitus of Paraphloeostiba: 33—P. apicalis (Mindanao), 34—P. obesa (Borneo), 35—P. subopaca (Tamil Nadu), 36—P. tonkinensis (Leyte). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8036127" httpUri="https://zenodo.org/record/8036127/files/figure.png" pageId="11" pageNumber="497">Figs. 33</figureCitation>
,
<figureCitation id="10B22A79FFAF94706536FD90FEB2FDF0" box="[265,336,569,595]" captionStart="FIGURES 37–42" captionStartId="12.[151,264,1810,1834]" captionTargetBox="[166,1417,195,1777]" captionTargetId="figure-15@12.[151,1436,181,1786]" captionTargetPageId="12" captionText="FIGURES 37–42. Aedeagus of Paraphloeostiba (Figs. 37–38: P. apicalis (Mindanao), Figs. 39–40: P. obesa (Sabah), Figs. 41–42: P. subopaca (Tamil Nadu), Figs. 43–44: P. tonkinensis (Leyte)): 37, 39, 41, 43—parameral view, 38, 40, 42, 44—lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8036129" httpUri="https://zenodo.org/record/8036129/files/figure.png" pageId="11" pageNumber="497">37–38</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C0936577FFAF947064A8FDD7FDFDFD17" pageId="11" pageNumber="497" type="reference_group">
<paragraph id="883636FCFFAF947064A8FDD7FDACFD36" blockId="11.[151,590,637,692]" box="[151,590,637,662]" pageId="11" pageNumber="497">
<treatmentCitationGroup id="A89911D2FFAF947064A8FDD7FDACFD36" box="[151,590,637,662]" pageId="11" pageNumber="497">
<treatmentCitation id="092810EDFFAF947064A8FDD7FDACFD36" author="Cameron, M." box="[151,590,637,662]" page="175" pageId="11" pageNumber="497" year="1925">
<taxonomicName id="4F894D7FFFAF947064A8FDD7FDACFD36" ID-CoL="4GGFR" authority="Cameron, 1925: 175" authorityName="Cameron" authorityYear="1925" box="[151,590,637,662]" class="Insecta" family="Staphylinidae" genus="Phloeonomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="497" phylum="Arthropoda" rank="species" species="apicalis">
<emphasis id="BAFDEAEEFFAF947064A8FDD7FE95FD35" box="[151,375,638,662]" italics="true" pageId="11" pageNumber="497">Phloeonomus apicalis</emphasis>
<bibRefCitation id="EC184B0DFFAF94706541FDD7FDACFD36" author="Cameron, M." box="[382,590,637,662]" pageId="11" pageNumber="497" pagination="174 - 198" refId="ref8185" refString="Cameron, M. (1925) New Staphylinidae from the Dutch East Indies. Treubia, 6 (2), 174 - 198." type="journal article" year="1925">Cameron, 1925: 175</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="883636FCFFAF947064A8FD35FDFDFD17" blockId="11.[151,590,637,692]" box="[151,543,668,692]" pageId="11" pageNumber="497">
Other references see in
<bibRefCitation id="EC184B0DFFAF947065B9FD35FDFDFD17" author="Shavrin, A. V." box="[390,543,668,692]" pageId="11" pageNumber="497" pagination="301 - 329" refId="ref8834" refString="Shavrin, A. V. (2020) New species and records of Paraphloeostiba Steel, 1960 from China and Laos, and descriptions of four news species of related genera (Coleoptera: Staphylinidae: Omaliinae: Omaliini). Zootaxa, 4890 (3), 301 - 329. https: // doi. org / 10.11646 / zootaxa. 4890.3.1" type="journal article" year="2020">Shavrin (2020)</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="C0936577FFAF947064A8FD4BFE29FCCA" pageId="11" pageNumber="497" type="materials_examined">
<paragraph id="883636FCFFAF947064A8FD4BFE29FCCA" blockId="11.[151,1437,738,945]" pageId="11" pageNumber="497">
<emphasis id="BAFDEAEEFFAF947064A8FD4BFE63FD5F" bold="true" box="[151,385,738,764]" pageId="11" pageNumber="497">Material examined.</emphasis>
<materialsCitation id="38E13CA1FFAF947065B7FD4BFE33FC82" collectionCode="BMNH" collectorName="L. Dembicky" country="India" elevation="100" latitude="27.35" location="Dirang" longLatPrecision="1237" longitude="92.21667" pageId="11" pageNumber="497" specimenCount="1" specimenCount-female="1" stateProvince="Arunachal Pradesh">
<emphasis id="BAFDEAEEFFAF947065B7FD4BFD11FD5F" bold="true" box="[392,755,738,764]" pageId="11" pageNumber="497">
<collectingCountry id="F09E766CFFAF947065B7FD4BFE3DFD5F" box="[392,479,738,764]" name="India" pageId="11" pageNumber="497">INDIA</collectingCountry>
:
<collectingRegion id="4A4DF81EFFAF947065D6FD4BFD0DFD5F" box="[489,751,738,764]" country="India" name="Arunachal Pradesh" pageId="11" pageNumber="497">ARUNACHAL PRADESH</collectingRegion>
:
</emphasis>
<specimenCount id="9E8FFD75FFAF947066C5FD4AFCC4FD5E" box="[762,806,739,765]" pageId="11" pageNumber="497" type="female">1 ♀</specimenCount>
:
<location id="8D566027FFAF9470670CFD4BFC63FD5F" LSID="urn:lsid:plazi:treatment:002087EAFFAF947064A8FDBCFB3FFC12:8D566027FFAF9470670CFD4BFC63FD5F" box="[819,897,738,764]" country="India" latitude="27.35" longLatPrecision="1237" longitude="92.21667" name="Dirang" pageId="11" pageNumber="497" stateProvince="Arunachal Pradesh">Dirang</location>
vicinity,
<geoCoordinate id="EDBD503BFFAF947067D5FD4AFBA8FD5E" box="[1002,1098,739,765]" degrees="27" direction="north" minutes="21" orientation="latitude" pageId="11" pageNumber="497" precision="925" value="27.35">27°21’N</geoCoordinate>
<geoCoordinate id="EDBD503BFFAF9470606DFD4AFB50FD5E" box="[1106,1202,739,765]" degrees="92" direction="east" minutes="13" orientation="longitude" pageId="11" pageNumber="497" precision="925" value="92.21667">92°13’E</geoCoordinate>
. 1800 ±
<quantity id="4F719B19FFAF9470612CFD4AFAB8FD5E" box="[1299,1370,739,765]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="497" unit="m" value="100.0">
<elevation id="03A4D1CFFFAF9470612CFD4AFAB8FD5E" box="[1299,1370,739,765]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="497" unit="m" value="100.0">100 m</elevation>
</quantity>
a.s.l..
<collectorName id="257C532AFFAF947064A8FCAEFECFFC83" box="[151,301,774,801]" pageId="11" pageNumber="497">L. Dembický</collectorName>
leg. (
<collectionCode id="EE98AE39FFAF94706550FCAEFE2BFC82" box="[367,457,775,801]" pageId="11" pageNumber="497">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="38E13CA1FFAF947065E1FCAFFDFDFCE6" collectingDate="1996-09-05" collectionCode="SMNS" collectorName="Bolm" country="Philippines" elevation="1000" location="Busdi." municipality="Mindanao" pageId="11" pageNumber="497" specimenCount="1" specimenCount-female="1">
<emphasis id="BAFDEAEEFFAF947065E1FCAFFD79FC83" bold="true" box="[478,667,774,800]" pageId="11" pageNumber="497">
<collectingCountry id="F09E766CFFAF947065E1FCAFFD77FC83" box="[478,661,774,800]" name="Philippines" pageId="11" pageNumber="497">PHILIPPINES</collectingCountry>
:
</emphasis>
1 ♁,
<specimenCount id="9E8FFD75FFAF947066E1FCAEFCEEFC82" box="[734,780,775,801]" pageId="11" pageNumber="497" type="female">1 ♀</specimenCount>
:
<collectingMunicipality id="6852AC86FFAF94706725FCAFFC6CFC83" box="[794,910,774,800]" pageId="11" pageNumber="497">Mindanao</collectingMunicipality>
,
<locationDeviation id="99842FDEFFAF947067A4FCAEFB49FC83" box="[923,1195,774,801]" location="Malaybalay" pageId="11" pageNumber="497">
<quantity id="4F719B19FFAF947067A4FCAEFC06FC83" box="[923,996,774,801]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="497" unit="km" value="30.0">30 km</quantity>
E of Malaybalay
</locationDeviation>
,
<location id="8D566027FFAF94706087FCAFFAE3FC83" LSID="urn:lsid:plazi:treatment:002087EAFFAF947064A8FDBCFB3FFC12:8D566027FFAF94706087FCAFFAE3FC83" box="[1208,1281,774,800]" country="Philippines" municipality="Mindanao" name="Busdi." pageId="11" pageNumber="497">Busdi.</location>
<elevation id="03A4D1CFFFAF94706136FCAEFA7FFC83" box="[1289,1437,774,801]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="497" unit="m" value="1000.0">
<quantity id="4F719B19FFAF94706136FCAEFA82FC82" box="[1289,1376,775,801]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="497" unit="m" value="1000.0">1000 m</quantity>
a.s.l.
</elevation>
05-
<date id="FC37103CFFAF94706485FC83FEDCFCE6" box="[186,318,810,837]" pageId="11" pageNumber="497" value="1996-09-05">
<collectingDate id="EC73E9D4FFAF94706485FC83FEDCFCE6" box="[186,318,810,837]" pageId="11" pageNumber="497" value="1996-09-05">09.05.1996</collectingDate>
</date>
.
<collectorName id="257C532AFFAF94706577FC83FE64FCE7" box="[328,390,810,836]" pageId="11" pageNumber="497">Bolm</collectorName>
leg. (
<collectionCode id="EE98AE39FFAF947065FAFC82FDF5FCE6" box="[453,535,811,837]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="11" pageNumber="497">SMNS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="38E13CA1FFAF94706615FC82FE25FCCA" collectingDate="1996-05-17" collectionCode="SMNS" collectorName="Bolm" country="Philippines" elevation="1700" location="Bagongsilang." pageId="11" pageNumber="497" specimenCount="1" specimenCount-female="1">
<specimenCount id="9E8FFD75FFAF94706615FC82FDB4FCE6" box="[554,598,811,837]" pageId="11" pageNumber="497" type="female">1 ♀</specimenCount>
: same island,
<locationDeviation id="99842FDEFFAF947066CCFC82FBE3FCE6" box="[755,1025,810,837]" location="Maramag" pageId="11" pageNumber="497">
<quantity id="4F719B19FFAF947066CCFC82FCD8FCE7" box="[755,826,810,837]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="497" unit="km" value="30.0">30 km</quantity>
NW of Maramag
</locationDeviation>
,
<location id="8D566027FFAF94706033FC83FB4FFCE7" LSID="urn:lsid:plazi:treatment:002087EAFFAF947064A8FDBCFB3FFC12:8D566027FFAF94706033FC83FB4FFCE7" box="[1036,1197,810,836]" country="Philippines" name="Bagongsilang." pageId="11" pageNumber="497">Bagongsilang.</location>
13-
<date id="FC37103CFFAF947060EBFC83FAB8FCE6" box="[1236,1370,810,837]" pageId="11" pageNumber="497" value="1996-05-17">
<collectingDate id="EC73E9D4FFAF947060EBFC83FAB8FCE6" box="[1236,1370,810,837]" pageId="11" pageNumber="497" value="1996-05-17">17.05.1996</collectingDate>
</date>
.
<elevation id="03A4D1CFFFAF9470615BFC82FF0BFCCB" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="11" pageNumber="497" unit="m" value="1700.0">
<quantity id="4F719B19FFAF9470615BFC82FF4FFCCA" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="11" pageNumber="497" unit="m" value="1700.0">1700 m</quantity>
a.s.l.
</elevation>
<collectorName id="257C532AFFAF947064D0FCE7FECFFCCB" box="[239,301,846,872]" pageId="11" pageNumber="497">Bolm</collectorName>
leg. (
<collectionCode id="EE98AE39FFAF94706552FCE6FE5DFCCA" box="[365,447,847,873]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="11" pageNumber="497">SMNS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C0936577FFAF947064F8FCDAFB3FFC12" pageId="11" pageNumber="497" type="discussion">
<paragraph id="883636FCFFAF947064F8FCDAFB3FFC12" blockId="11.[151,1437,738,945]" pageId="11" pageNumber="497">
<emphasis id="BAFDEAEEFFAF947064F8FCDAFED9FC2E" bold="true" box="[199,315,883,909]" pageId="11" pageNumber="497">Remarks.</emphasis>
<taxonomicName id="4F894D7FFFAF9470657EFCDBFDB9FC2F" baseAuthorityName="Cameron" baseAuthorityYear="1925" box="[321,603,882,908]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="497" phylum="Arthropoda" rank="species" species="apicalis">
<emphasis id="BAFDEAEEFFAF9470657EFCDBFDB9FC2F" box="[321,603,882,908]" italics="true" pageId="11" pageNumber="497">Paraphloeostiba apicalis</emphasis>
</taxonomicName>
was known from
<collectingCountry id="F09E766CFFAF9470671DFCDBFC6DFC2F" box="[802,911,882,908]" name="Indonesia" pageId="11" pageNumber="497">Indonesia</collectingCountry>
and
<collectingCountry id="F09E766CFFAF947067FCFCDBFBA3FC2F" box="[963,1089,882,908]" name="Philippines" pageId="11" pageNumber="497">Philippines</collectingCountry>
(
<bibRefCitation id="EC184B0DFFAF94706070FCDAFB0EFC2E" author="Herman, L. H." box="[1103,1260,883,909]" pageId="11" pageNumber="497" pagination="1 - 7" refId="ref8478" refString="Herman, L. H. (2001) Catalogue of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millenium. Bulletin of the American Museum of Natural History, 265 (1 - 7), i - vi + 1 - 4218. https: // doi. org / 10.1206 / 0003 - 0090.265.1.1" type="journal article" year="2001">Herman 2001</bibRefCitation>
,
<bibRefCitation id="EC184B0DFFAF947060C8FCDBFA73FC2E" author="Shavrin, A. V." box="[1271,1425,882,909]" pageId="11" pageNumber="497" pagination="301 - 329" refId="ref8834" refString="Shavrin, A. V. (2020) New species and records of Paraphloeostiba Steel, 1960 from China and Laos, and descriptions of four news species of related genera (Coleoptera: Staphylinidae: Omaliinae: Omaliini). Zootaxa, 4890 (3), 301 - 329. https: // doi. org / 10.11646 / zootaxa. 4890.3.1" type="journal article" year="2020">Shavrin 2020</bibRefCitation>
). It is here recorded from
<collectingCountry id="F09E766CFFAF94706598FC3FFE03FC13" box="[423,481,918,944]" name="India" pageId="11" pageNumber="497">India</collectingCountry>
for the first time. Habitus as in
<figureCitation id="10B22A79FFAF9470677AFC3FFC75FC12" box="[837,919,918,945]" captionStart="FIGURES 33–36" captionStartId="10.[151,264,843,867]" captionTargetBox="[159,1429,187,811]" captionTargetId="figure-15@10.[151,1436,181,819]" captionTargetPageId="10" captionText="FIGURES 33–36. Habitus of Paraphloeostiba: 33—P. apicalis (Mindanao), 34—P. obesa (Borneo), 35—P. subopaca (Tamil Nadu), 36—P. tonkinensis (Leyte). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8036127" httpUri="https://zenodo.org/record/8036127/files/figure.png" pageId="11" pageNumber="497">Fig. 33</figureCitation>
. Aedeagus as in
<figureCitation id="10B22A79FFAF9470606EFC3FFB3BFC12" box="[1105,1241,918,945]" captionStart="FIGURES 37–42" captionStartId="12.[151,264,1810,1834]" captionTargetBox="[166,1417,195,1777]" captionTargetId="figure-15@12.[151,1436,181,1786]" captionTargetPageId="12" captionText="FIGURES 37–42. Aedeagus of Paraphloeostiba (Figs. 37–38: P. apicalis (Mindanao), Figs. 39–40: P. obesa (Sabah), Figs. 41–42: P. subopaca (Tamil Nadu), Figs. 43–44: P. tonkinensis (Leyte)): 37, 39, 41, 43—parameral view, 38, 40, 42, 44—lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8036129" httpUri="https://zenodo.org/record/8036129/files/figure.png" pageId="11" pageNumber="497">Figs. 37–38</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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