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<mods:title id="F303A9AC336EEEE74638B6968D2F1F5A">New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines</mods:title>
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<heading id="D37E8190FFAF947064A8FBABFD83FBBF" bold="true" box="[151,609,1026,1053]" fontSize="11" level="1" pageId="11" pageNumber="497" reason="1">
<taxonomicName id="4F894D7FFFAF947064A8FBABFD83FBBF" ID-CoL="75QXP" authority="(Fauvel, 1904)" baseAuthorityName="Fauvel" baseAuthorityYear="1904" box="[151,609,1026,1053]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="497" phylum="Arthropoda" rank="species" species="obesa">
<emphasis id="BAFDEAEEFFAF947064A8FBABFD83FBBF" bold="true" box="[151,609,1026,1053]" pageId="11" pageNumber="497">
<emphasis id="BAFDEAEEFFAF947064A8FBABFE4AFBBE" bold="true" box="[151,424,1026,1053]" italics="true" pageId="11" pageNumber="497">Paraphloeostiba obesa</emphasis>
(
<bibRefCitation id="EC184B0DFFAF94706587FBABFDBBFBBF" author="Fauvel, A." box="[440,601,1026,1052]" pageId="11" pageNumber="497" pagination="76 - 112" refId="ref8378" refString="Fauvel, A. (1904) Staphylinides exotiques nouveaux. 2 e Partie. Revue d'Entomologie, 23, 76 - 112." type="journal article" year="1904">Fauvel, 1904</bibRefCitation>
)
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<paragraph id="883636FCFFAF947064A8FB8FFF1BFBE2" blockId="11.[151,609,1026,1089]" box="[151,249,1062,1089]" pageId="11" pageNumber="497">
(
<figureCitation id="10B22A79FFAF947064A0FB8FFF13FBE2" box="[159,241,1062,1089]" captionStart="FIGURES 33–36" captionStartId="10.[151,264,843,867]" captionTargetBox="[159,1429,187,811]" captionTargetId="figure-15@10.[151,1436,181,819]" captionTargetPageId="10" captionText="FIGURES 33–36. Habitus of Paraphloeostiba: 33—P. apicalis (Mindanao), 34—P. obesa (Borneo), 35—P. subopaca (Tamil Nadu), 36—P. tonkinensis (Leyte). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8036127" httpUri="https://zenodo.org/record/8036127/files/figure.png" pageId="11" pageNumber="497">Fig. 34</figureCitation>
)
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<treatmentCitation id="092810EDFFAF947064A8FBC2FDE5FB20" author="Fauvel, A." box="[151,519,1131,1155]" page="89" pageId="11" pageNumber="497" year="1904">
<taxonomicName id="4F894D7FFFAF947064A8FBC2FDE5FB20" ID-CoL="3MCPM" authority="Fauvel, 1904: 89" authorityName="Fauvel" authorityPageNumber="89" authorityYear="1904" box="[151,519,1131,1155]" class="Insecta" family="Staphylinidae" genus="Homalium" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="497" phylum="Arthropoda" rank="species" species="obesum">
<emphasis id="BAFDEAEEFFAF947064A8FBC2FEB7FB20" box="[151,341,1131,1155]" italics="true" pageId="11" pageNumber="497">Homalium obesum</emphasis>
<bibRefCitation id="EC184B0DFFAF94706563FBC2FDE5FB20" author="Fauvel, A." box="[348,519,1131,1155]" pageId="11" pageNumber="497" pagination="76 - 112" refId="ref8378" refString="Fauvel, A. (1904) Staphylinides exotiques nouveaux. 2 e Partie. Revue d'Entomologie, 23, 76 - 112." type="journal article" year="1904">Fauvel, 1904: 89</bibRefCitation>
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<treatmentCitationGroup id="A89911D2FFAF947064A8FB23FBE2FB01" box="[151,1024,1162,1186]" pageId="11" pageNumber="497">
<treatmentCitation id="092810EDFFAF947064A8FB23FCD8FB01" author="Bernhauer, M. &amp; Schubert, K." box="[151,826,1162,1186]" page="49" pageId="11" pageNumber="497" year="1910">
<taxonomicName id="4F894D7FFFAF947064A8FB23FCD8FB01" authority=": Bernhauer &amp; Schubert 1910: 49" authorityName="Bernhauer &amp; Schubert" authorityPageNumber="49" authorityYear="1910" box="[151,826,1162,1186]" class="Insecta" family="Staphylinidae" genus="Phyllodrepa" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="497" phylum="Arthropoda" rank="species" species="obesa" subGenus="Hapalaraea">
<emphasis id="BAFDEAEEFFAF947064A8FB23FEF1FB01" box="[151,275,1162,1186]" italics="true" pageId="11" pageNumber="497">Phyllodrepa</emphasis>
(
<emphasis id="BAFDEAEEFFAF9470651DFB23FE7EFB01" box="[290,412,1162,1186]" italics="true" pageId="11" pageNumber="497">Hapalaraea</emphasis>
)
<emphasis id="BAFDEAEEFFAF94706594FB23FE04FB01" box="[427,486,1162,1186]" italics="true" pageId="11" pageNumber="497">obesa</emphasis>
:
<bibRefCitation id="EC184B0DFFAF947065CBFB23FCD8FB01" author="Bernhauer, M. &amp; Schubert, K." box="[500,826,1162,1186]" pageId="11" pageNumber="497" pagination="1 - 86" refId="ref8074" refString="Bernhauer, M. &amp; Schubert, K. (1910) Staphylinidae I. In: Schenkling, S. (Ed.), Coleopterorum Catalogus. 5 (19). Junk, Berlin, pp. 1 - 86. https: // doi. org / 10.1007 / 978 - 94 - 011 - 9697 - 0 _ 1" type="book chapter" year="1910">Bernhauer &amp; Schubert 1910: 49</bibRefCitation>
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,
<treatmentCitation id="092810EDFFAF9470677BFB23FBE2FB01" author="Cameron, M." box="[836,1024,1162,1186]" page="98" pageId="11" pageNumber="497" year="1928">
<bibRefCitation id="EC184B0DFFAF9470677BFB23FBE2FB01" author="Cameron, M." box="[836,1024,1162,1186]" pageId="11" pageNumber="497" pagination="90 - 110" refId="ref8211" refString="Cameron, M. (1928) Fauna sumatrensis. Staphylinidae (Col.). Entomologische Mitteilungen, 17 (2), 90 - 110." type="journal article" year="1928">Cameron 1928: 98</bibRefCitation>
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<treatmentCitationGroup id="A89911D2FFAF947064A8FB01FCF0FB63" box="[151,786,1192,1216]" pageId="11" pageNumber="497">
<treatmentCitation id="092810EDFFAF947064A8FB01FDD2FB63" author="Steel, W. O." box="[151,560,1192,1216]" page="152" pageId="11" pageNumber="497" year="1960">
<taxonomicName id="4F894D7FFFAF947064A8FB01FDD2FB63" ID-CoL="75QXP" authority=": Steel 1960: 152" authorityName=": Steel" authorityPageNumber="152" authorityYear="1960" box="[151,560,1192,1216]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="497" phylum="Arthropoda" rank="species" species="obesa">
<emphasis id="BAFDEAEEFFAF947064A8FB01FE63FB63" box="[151,385,1192,1216]" italics="true" pageId="11" pageNumber="497">Paraphloeostiba obesa</emphasis>
:
<bibRefCitation id="EC184B0DFFAF947065B0FB01FDD2FB63" author="Steel, W. O." box="[399,560,1192,1216]" pageId="11" pageNumber="497" pagination="141 - 172" refId="ref9011" refString="Steel, W. O. (1960) Three new omaliine genera from Asia and Australasia previously confused with Phloeonomus Thomson (Coleoptera: Staphylinidae). The Transactions of the Royal Entomological Society of London, 112, 141 - 172. https: // doi. org / 10.1111 / j. 1365 - 2311.1960. tb 00497. x" type="journal article" year="1960">Steel 1960: 152</bibRefCitation>
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,
<treatmentCitation id="092810EDFFAF94706604FB01FCF0FB63" author="Hammond, P. M." box="[571,786,1192,1216]" page="195" pageId="11" pageNumber="497" year="1984">
<bibRefCitation id="EC184B0DFFAF94706604FB01FCF0FB63" author="Hammond, P. M." box="[571,786,1192,1216]" pageId="11" pageNumber="497" pagination="187 - 218" refId="ref8542" refString="Hammond, P. M. (1984) An annotated check-list of Staphylinidae (Insecta: Coleoptera) recorded from Borneo. The Sarawak Museum Journal, 33 (54), 187 - 218." type="journal article" year="1984">Hammond 1984: 195</bibRefCitation>
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<paragraph id="883636FCFFAF947064A8FB6EFD47FB7C" blockId="11.[151,1024,1131,1278]" box="[151,677,1223,1247]" pageId="11" pageNumber="497">
<treatmentCitationGroup id="A89911D2FFAF947064A8FB6EFD47FB7C" box="[151,677,1223,1247]" pageId="11" pageNumber="497">
<treatmentCitation id="092810EDFFAF947064A8FB6EFD47FB7C" author="Bernhauer, M." box="[151,677,1223,1247]" page="135" pageId="11" pageNumber="497" year="1915">
<taxonomicName id="4F894D7FFFAF947064A8FB6EFD47FB7C" ID-CoL="4GGFZ" authority="Bernhauer, 1915: 135" authorityName="Bernhauer" authorityPageNumber="135" authorityYear="1915" box="[151,677,1223,1247]" class="Insecta" family="Staphylinidae" genus="Phloeonomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="497" phylum="Arthropoda" rank="species" species="densepunctatus">
<emphasis id="BAFDEAEEFFAF947064A8FB6EFE21FB7C" box="[151,451,1223,1247]" italics="true" pageId="11" pageNumber="497">Phloeonomus densepunctatus</emphasis>
<bibRefCitation id="EC184B0DFFAF947065F5FB6EFD47FB7C" author="Bernhauer, M." box="[458,677,1223,1247]" pageId="11" pageNumber="497" pagination="134 - 158" refId="ref8043" refString="Bernhauer, M. (1915) Neue Staphyliniden der indo-malaiischen Fauna, insbesondere der Sunda-Insel Borneo. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 65, 134 - 158" type="journal article" year="1915">Bernhauer, 1915: 135</bibRefCitation>
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<paragraph id="883636FCFFAF947064A8FB4FFC80FB5D" blockId="11.[151,1024,1131,1278]" box="[151,866,1254,1278]" pageId="11" pageNumber="497">
<treatmentCitationGroup id="A89911D2FFAF947064A8FB4FFC80FB5D" box="[151,866,1254,1278]" pageId="11" pageNumber="497">
<treatmentCitation id="092810EDFFAF947064A8FB4FFC80FB5D" author="Scheerpeltz, O. von" box="[151,866,1254,1278]" page="1049" pageId="11" pageNumber="497" year="1933">
<taxonomicName id="4F894D7FFFAF947064A8FB4FFC80FB5D" ID-CoL="4GGFZ" authority=": Scheerpeltz 1933: 1049" authorityName="Scheerpeltz" authorityPageNumber="1049" authorityYear="1933" box="[151,866,1254,1278]" class="Insecta" family="Staphylinidae" genus="Phloeonomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="497" phylum="Arthropoda" rank="species" species="densepunctatus" subGenus="Phloeonomus">
<emphasis id="BAFDEAEEFFAF947064A8FB4FFEC3FB5D" box="[151,289,1254,1278]" italics="true" pageId="11" pageNumber="497">Phloeonomus</emphasis>
(
<emphasis id="BAFDEAEEFFAF9470650FFB4FFE58FB5D" box="[304,442,1254,1278]" italics="true" pageId="11" pageNumber="497">Phloeonomus</emphasis>
)
<emphasis id="BAFDEAEEFFAF947065F6FB4FFD87FB5D" box="[457,613,1254,1278]" italics="true" pageId="11" pageNumber="497">densepunctatus</emphasis>
:
<bibRefCitation id="EC184B0DFFAF9470664DFB4FFC80FB5D" author="Scheerpeltz, O. von" box="[626,866,1254,1278]" pageId="11" pageNumber="497" pagination="989 - 1500" refId="ref8685" refString="Scheerpeltz, O. von (1933) Staphylinidae VII (pars 129). In: Schenkling S. (Ed.), Coleopterorum Catalogus. Vol. VI. Staphylinidae II. W. Junk, Berlin, pp. 989 - 1500." type="book chapter" year="1933">Scheerpeltz 1933: 1049</bibRefCitation>
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<paragraph id="883636FCFFAF947064A8FA85FD81FAC9" blockId="11.[151,1437,1324,1602]" pageId="11" pageNumber="497">
<emphasis id="BAFDEAEEFFAF947064A8FA85FE67FAE5" bold="true" box="[151,389,1324,1350]" pageId="11" pageNumber="497">Material examined.</emphasis>
<materialsCitation id="38E13CA1FFAF947065AEFA85FDBCFAC9" collectingDate="1996-02-12" collectionCode="SMNS" collectorName="W. Schawaller" country="Malaysia" elevation="500" location="Kinabalu National Park" municipality="Borneo" pageId="11" pageNumber="497" specimenCount="1" stateProvince="Sabah">
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<collectingCountry id="F09E766CFFAF947065AEFA85FDC9FAE5" box="[401,555,1324,1350]" name="Malaysia" pageId="11" pageNumber="497">MALAYSIA</collectingCountry>
:
</emphasis>
1 ♁:
<collectingMunicipality id="6852AC86FFAF94706641FA85FD31FAE5" box="[638,723,1324,1350]" pageId="11" pageNumber="497">Borneo</collectingMunicipality>
,
<collectingRegion id="4A4DF81EFFAF947066DBFA85FCC9FAE5" box="[740,811,1324,1350]" country="Malaysia" name="Sabah" pageId="11" pageNumber="497">Sabah</collectingRegion>
,
<location id="8D566027FFAF94706704FA85FBB3FAE5" LSID="urn:lsid:plazi:treatment:002087EAFFAF947064A8FBABFF58F9E1:8D566027FFAF94706704FA85FBB3FAE5" box="[827,1105,1324,1350]" country="Malaysia" municipality="Borneo" name="Kinabalu National Park" pageId="11" pageNumber="497" stateProvince="Sabah">Kinabalu National Park</location>
: Poring.
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a.s.l.
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29.11-
<date id="FC37103CFFAF947064A8FAF9FEFBFAC9" box="[151,281,1360,1386]" pageId="11" pageNumber="497" value="1996-02-12">
<collectingDate id="EC73E9D4FFAF947064A8FAF9FEFBFAC9" box="[151,281,1360,1386]" pageId="11" pageNumber="497" value="1996-02-12">02.12.1996</collectingDate>
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.
<collectorName id="257C532AFFAF9470651CFAF9FE27FAC9" box="[291,453,1360,1386]" pageId="11" pageNumber="497">W. Schawaller</collectorName>
leg. (
<collectionCode id="EE98AE39FFAF9470663BFAF9FDB4FAC9" box="[516,598,1360,1386]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="11" pageNumber="497">SMNS</collectionCode>
)
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.
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<subSubSection id="C0936577FFAF947064F8FADDFF58F9E1" pageId="11" pageNumber="497" type="discussion">
<paragraph id="883636FCFFAF947064F8FADDFF58F9E1" blockId="11.[151,1437,1324,1602]" pageId="11" pageNumber="497">
<emphasis id="BAFDEAEEFFAF947064F8FADDFED9FA2D" bold="true" box="[199,315,1396,1422]" pageId="11" pageNumber="497">Remarks.</emphasis>
<taxonomicName id="4F894D7FFFAF94706577FADDFDC3FA2D" authorityName="Fauvel" authorityYear="1904" box="[328,545,1396,1422]" class="Insecta" family="Staphylinidae" genus="Homalium" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="497" phylum="Arthropoda" rank="species" species="obesum">
<emphasis id="BAFDEAEEFFAF94706577FADDFDC3FA2D" box="[328,545,1396,1422]" italics="true" pageId="11" pageNumber="497">Homalium obesum</emphasis>
</taxonomicName>
was originallly described from “Sumatra: région de Deli, dans des tabacs importés”.
<bibRefCitation id="EC184B0DFFAF9470652AFA31FE43FA11" author="Steel, W. O." box="[277,417,1432,1458]" pageId="11" pageNumber="497" pagination="141 - 172" refId="ref9011" refString="Steel, W. O. (1960) Three new omaliine genera from Asia and Australasia previously confused with Phloeonomus Thomson (Coleoptera: Staphylinidae). The Transactions of the Royal Entomological Society of London, 112, 141 - 172. https: // doi. org / 10.1111 / j. 1365 - 2311.1960. tb 00497. x" type="journal article" year="1960">Steel (1960)</bibRefCitation>
redescribed it, designated the
<typeStatus id="5732885EFFAF947066C0FA31FC8BFA11" box="[767,873,1432,1458]" pageId="11" pageNumber="497">lectotype</typeStatus>
, provided figures of the aedeagus and the female accessory sclerite, and recorded it from Java,
<collectingCountry id="F09E766CFFAF947066A2FA15FD1FFA75" box="[669,765,1468,1494]" name="Malaysia" pageId="11" pageNumber="497">Sarawak</collectingCountry>
and Luzon.
<taxonomicName id="4F894D7FFFAF947067B2FA15FA7EFA74" authority="Bernhauer, 1915" authorityName="Bernhauer" authorityYear="1915" box="[909,1436,1468,1495]" class="Insecta" family="Staphylinidae" genus="Phloeonomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="497" phylum="Arthropoda" rank="species" species="densepunctatus">
<emphasis id="BAFDEAEEFFAF947067B2FA15FB38FA75" box="[909,1242,1468,1494]" italics="true" pageId="11" pageNumber="497">Phloeonomus densepunctatus</emphasis>
<bibRefCitation id="EC184B0DFFAF947060DDFA15FA7EFA74" author="Bernhauer, M." box="[1250,1436,1468,1495]" pageId="11" pageNumber="497" pagination="134 - 158" refId="ref8043" refString="Bernhauer, M. (1915) Neue Staphyliniden der indo-malaiischen Fauna, insbesondere der Sunda-Insel Borneo. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 65, 134 - 158" type="journal article" year="1915">Bernhauer, 1915</bibRefCitation>
</taxonomicName>
was originally described from “
<collectingCountry id="F09E766CFFAF947065CCFA49FDB8FA59" box="[499,602,1504,1530]" name="Malaysia" pageId="11" pageNumber="497">Sarawak</collectingCountry>
: Mt. Matang”.
<bibRefCitation id="EC184B0DFFAF947066C0FA49FC6AFA59" author="Steel, W. O." box="[767,904,1504,1530]" pageId="11" pageNumber="497" pagination="141 - 172" refId="ref9011" refString="Steel, W. O. (1960) Three new omaliine genera from Asia and Australasia previously confused with Phloeonomus Thomson (Coleoptera: Staphylinidae). The Transactions of the Royal Entomological Society of London, 112, 141 - 172. https: // doi. org / 10.1111 / j. 1365 - 2311.1960. tb 00497. x" type="journal article" year="1960">Steel (1960)</bibRefCitation>
synonymized it with
<taxonomicName id="4F894D7FFFAF94706049FA48FB36FA59" authorityName=": Steel" authorityYear="1960" box="[1142,1236,1504,1530]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="497" phylum="Arthropoda" rank="species" species="obesa">
<emphasis id="BAFDEAEEFFAF94706049FA48FB65FA59" box="[1142,1159,1505,1530]" italics="true" pageId="11" pageNumber="497">P</emphasis>
.
<emphasis id="BAFDEAEEFFAF947060ACFA49FB36FA59" box="[1171,1236,1504,1530]" italics="true" pageId="11" pageNumber="497">obesa</emphasis>
</taxonomicName>
.
<taxonomicName id="4F894D7FFFAF947060DDFA49FF3AF9BD" authorityName=": Steel" authorityYear="1960" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="497" phylum="Arthropoda" rank="species" species="obesa">
<emphasis id="BAFDEAEEFFAF947060DDFA49FF3AF9BD" italics="true" pageId="11" pageNumber="497">Paraphloeostiba obesa</emphasis>
</taxonomicName>
is known from
<collectingCountry id="F09E766CFFAF947065ABF9ADFDE6F9BD" box="[404,516,1540,1566]" name="Indonesia" pageId="11" pageNumber="497">Indonesia</collectingCountry>
,
<collectingCountry id="F09E766CFFAF9470662DF9ADFD9BF9BD" box="[530,633,1540,1566]" name="Malaysia" pageId="11" pageNumber="497">Malaysia</collectingCountry>
(including
<collectingRegion id="4A4DF81EFFAF947066C0F9ADFCA7F9BD" box="[767,837,1540,1566]" country="Malaysia" name="Sabah" pageId="11" pageNumber="497">Sabah</collectingRegion>
) and
<collectingCountry id="F09E766CFFAF947067B7F9ADFBE4F9BD" box="[904,1030,1540,1566]" name="Philippines" pageId="11" pageNumber="497">Philippines</collectingCountry>
(
<bibRefCitation id="EC184B0DFFAF94706028F9ADFB5BF9BD" author="Herman, L. H." box="[1047,1209,1540,1566]" pageId="11" pageNumber="497" pagination="1 - 7" refId="ref8478" refString="Herman, L. H. (2001) Catalogue of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millenium. Bulletin of the American Museum of Natural History, 265 (1 - 7), i - vi + 1 - 4218. https: // doi. org / 10.1206 / 0003 - 0090.265.1.1" type="journal article" year="2001">Herman 2001</bibRefCitation>
). Habitus as in
<figureCitation id="10B22A79FFAF9470614FF9ADFF54F9E1" captionStart="FIGURES 33–36" captionStartId="10.[151,264,843,867]" captionTargetBox="[159,1429,187,811]" captionTargetId="figure-15@10.[151,1436,181,819]" captionTargetPageId="10" captionText="FIGURES 33–36. Habitus of Paraphloeostiba: 33—P. apicalis (Mindanao), 34—P. obesa (Borneo), 35—P. subopaca (Tamil Nadu), 36—P. tonkinensis (Leyte). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8036127" httpUri="https://zenodo.org/record/8036127/files/figure.png" pageId="11" pageNumber="497">Fig. 34</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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