268 lines
35 KiB
XML
268 lines
35 KiB
XML
<document id="F1ACBC3B5CECB21E6CD3E26EEF40516C" ID-DOI="10.11646/zootaxa.5301.4.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="8036117" ID-ZooBank="01F6333F-807B-448A-80C8-EC1F8CE030EA" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1686723262927" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2023" docId="002087EAFFAC947164A8FBCBFE76F990" docLanguage="en" docName="zootaxa.5301.4.6.pdf" docOrigin="Zootaxa 5301 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5301.4.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paraphloeostiba ullrichi Shavrin 2023, sp.n." docType="treatment" docVersion="2" lastPageNumber="496" masterDocId="FC19FF92FFA4947B643FFFA9FFE2FFA3" masterDocTitle="New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines" masterLastPageNumber="500" masterPageNumber="486" pageNumber="494" updateTime="1686723785076" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
|
||
<mods:mods id="BEAF8EC1ACCAE7347BAB93F269CD2447" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="C5ED74CD24A9B10EBB56EC9E388EBBBA">
|
||
<mods:title id="156CA10E2A7CA38107477A91BCDA88BC">New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="424D3DB9096EDDB8216E03ED02DBA9F6" type="personal">
|
||
<mods:role id="2C0818216870971811BFF417C6DE348E">
|
||
<mods:roleTerm id="97E6C71119151AE49B80273963FDC27A">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="B8E59EBEC84E58818FDA6B91EDD391EF">Shavrin, Alexey V.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="30BA7887ED0133AA8154B75C6D539AD0">text</mods:typeOfResource>
|
||
<mods:relatedItem id="56AB2FEA2A882D016035615F76EB002D" type="host">
|
||
<mods:titleInfo id="9961B94D3F82D0B5D3FAA154168E2D3B">
|
||
<mods:title id="A0FAE29F839A9648BA4D3CD85C4EBE48">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="6EF920DBE922819710398C8DAA9E0F9F">
|
||
<mods:date id="19D948F371A5E2B1AE87F48D4573ED5F">2023</mods:date>
|
||
<mods:detail id="EA5CB5B606D40732AE75826A87C168B3" type="pubDate">
|
||
<mods:number id="90EC27B2B36E8D140040A24F932547F1">2023-06-13</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="E1949F244A064C3BB926C40243B3E08E" type="volume">
|
||
<mods:number id="05728B71A1C6792FE2E357E0B0296AF1">5301</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="58A0EED2B398B8C522CBC30A85100987" type="issue">
|
||
<mods:number id="3C02800C0C4F59A262FFC95E1788CDB2">4</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="FA6D67773AB8CDEAC46572FB22C5FE66" unit="page">
|
||
<mods:start id="810DEFA21AAFCC7D8462A25EC17BC59E">486</mods:start>
|
||
<mods:end id="131B655F20A236882D7467665AB832EB">500</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="51256DFD131CFB665E389ABCE7FF32CF">
|
||
<mods:url id="E48C4DF7F240BA07AE76747E3088B3AA">http://dx.doi.org/10.11646/zootaxa.5301.4.6</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="E006F5732753CC52BE32A01F540CB26A">journal article</mods:classification>
|
||
<mods:identifier id="6E292B4078B4BE4546F3C6D945775238" type="DOI">10.11646/zootaxa.5301.4.6</mods:identifier>
|
||
<mods:identifier id="B767DA39A8F0EFCB1BABA932A46C73E0" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="00F7584019C245806000A75FBA757AB0" type="Zenodo-Dep">8036117</mods:identifier>
|
||
<mods:identifier id="E68E792C462E59D7E3D307923F3D0DF8" type="ZooBank">01F6333F-807B-448A-80C8-EC1F8CE030EA</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="002087EAFFAC947164A8FBCBFE76F990" ID-DOI="http://doi.org/10.5281/zenodo.8036111" ID-Zenodo-Dep="8036111" LSID="urn:lsid:plazi:treatment:002087EAFFAC947164A8FBCBFE76F990" httpUri="http://treatment.plazi.org/id/002087EAFFAC947164A8FBCBFE76F990" lastPageId="10" lastPageNumber="496" pageId="8" pageNumber="494">
|
||
<subSubSection id="C0936577FFAC947364A8FBCBFDE2FBDE" box="[151,512,1122,1149]" pageId="8" pageNumber="494" type="nomenclature">
|
||
<paragraph id="883636FCFFAC947364A8FBCBFDE2FBDE" blockId="8.[151,512,1122,1185]" box="[151,512,1122,1149]" pageId="8" pageNumber="494">
|
||
<heading id="D37E8190FFAC947364A8FBCBFDE2FBDE" bold="true" box="[151,512,1122,1149]" fontSize="11" level="1" pageId="8" pageNumber="494" reason="1">
|
||
<emphasis id="BAFDEAEEFFAC947364A8FBCBFDE2FBDE" bold="true" box="[151,512,1122,1149]" pageId="8" pageNumber="494">
|
||
<taxonomicName id="4F894D7FFFAC947364A8FBCBFE5CFBDE" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[151,446,1122,1149]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="ullrichi" status="sp.n.">
|
||
<emphasis id="BAFDEAEEFFAC947364A8FBCBFE5CFBDE" bold="true" box="[151,446,1122,1149]" italics="true" pageId="8" pageNumber="494">Paraphloeostiba ullrichi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A1CE5795FFAC947365FAFBCAFDE2FBDE" box="[453,512,1123,1149]" pageId="8" pageNumber="494" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C0936577FFAC947364A8FB2EFEA8FB02" box="[151,330,1159,1185]" pageId="8" pageNumber="494" type="description">
|
||
<paragraph id="883636FCFFAC947364A8FB2EFEA8FB02" blockId="8.[151,512,1122,1185]" box="[151,330,1159,1185]" pageId="8" pageNumber="494">
|
||
(
|
||
<figureCitation id="10B22A79FFAC947364A0FB2EFF0DFB02" box="[159,239,1159,1185]" captionStart="FIGURES 1–6" captionStartId="2.[151,275,1541,1567]" captionTargetBox="[197,1385,250,1508]" captionTargetId="figure-45@2.[187,1400,237,1520]" captionTargetPageId="2" captionText="FIGURES 1–6. Habitus of Paraphloeostiba: 1—P. conjuncta (holotype), 2—P. coriacea, 3—P. flaveola (holotype), 4—P. gayndahensis (Papua New Guinea), 5—P. papuana (holotype), 6—P. ullrichi (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8036119" httpUri="https://zenodo.org/record/8036119/files/figure.png" pageId="8" pageNumber="494">Figs. 6</figureCitation>
|
||
,
|
||
<figureCitation id="10B22A79FFAC947364C4FB2EFEA0FB02" box="[251,322,1159,1185]" captionStart="FIGURES 26–32" captionStartId="9.[151,264,1422,1446]" captionTargetBox="[162,1423,195,1387]" captionTargetId="figure-24@9.[151,1436,181,1398]" captionTargetPageId="9" captionText="FIGURES 26–32. Aedeagus and abdominal segments of Paraphloeostiba ullrichi (paratypes): 26—aedeagus (parameral view), 27—aedeagus (lateral view), 28—male tergite VIII (dorsal view), 29—male sternite VIII (ventral view), 30—female tergite VIII (dorsal view), 31—female sternite VIII (ventral view), 32—female genital segment. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8036125" httpUri="https://zenodo.org/record/8036125/files/figure.png" pageId="8" pageNumber="494">26–32</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C0936577FFAC947364A8FB66FE48FA46" pageId="8" pageNumber="494" type="materials_examined">
|
||
<paragraph id="883636FCFFAC947364A8FB66FD86FA92" blockId="8.[151,1437,1230,2013]" pageId="8" pageNumber="494">
|
||
<emphasis id="BAFDEAEEFFAC947364A8FB66FEAAFB4B" bold="true" box="[151,328,1230,1257]" pageId="8" pageNumber="494">Type material.</emphasis>
|
||
<typeStatus id="5732885EFFAC9473656CFB66FE59FB4A" box="[339,443,1231,1257]" pageId="8" pageNumber="494">Holotype</typeStatus>
|
||
♁: ‘12 VIII 79 [handwritten in black] |
|
||
<collectingCountry id="F09E766CFFAC947367A9FB66FC2CFB4A" box="[918,974,1231,1257]" name="Papua New Guinea" pageId="8" pageNumber="494">PNG</collectingCountry>
|
||
/EHProv. | Umg. Kainantu | Onerunka` <printed>, ‘Papua | Nlle Guinée | W. G. Ullrich <printed>, ‘
|
||
<typeStatus id="5732885EFFAC94736715FB5AFC21FAAE" box="[810,963,1267,1293]" pageId="8" pageNumber="494">HOLOTYPE</typeStatus>
|
||
|
|
||
<taxonomicName id="4F894D7FFFAC947367EAFB5AFB72FAAE" authorityName="Steel" authorityYear="1960" box="[981,1168,1267,1293]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="494" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BAFDEAEEFFAC947367EAFB5AFB72FAAE" box="[981,1168,1267,1293]" italics="true" pageId="8" pageNumber="494">Paraphloeostiba</emphasis>
|
||
</taxonomicName>
|
||
|
|
||
<taxonomicName id="4F894D7FFFAC9473609DFB5AFB16FAAE" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[1186,1268,1267,1293]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="ullrichi" status="sp.n.">
|
||
<emphasis id="BAFDEAEEFFAC9473609DFB5AFB16FAAE" box="[1186,1268,1267,1293]" italics="true" pageId="8" pageNumber="494">ullrichi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="BAFDEAEEFFAC947360C4FB5AFAD0FAAE" bold="true" box="[1275,1330,1267,1293]" pageId="8" pageNumber="494">
|
||
<taxonomicNameLabel id="A1CE5795FFAC947360C4FB5AFAD0FAAE" box="[1275,1330,1267,1293]" pageId="8" pageNumber="494" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
| Shavrin A.V. des. 2023`<red, printed> (
|
||
<collectionCode id="EE98AE39FFAC947365C3FABEFDBAFA92" box="[508,600,1303,1329]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="8" pageNumber="494" type="Museum">MHNG</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="883636FCFFAC947364F8FA92FE48FA46" blockId="8.[151,1437,1230,2013]" pageId="8" pageNumber="494">
|
||
<typeStatus id="5732885EFFAC947364F8FA92FEDAFAF6" box="[199,312,1339,1365]" pageId="8" pageNumber="494">Paratypes</typeStatus>
|
||
: 3 ♁♁ (
|
||
<specimenCount id="9E8FFD75FFAC947365AFFA92FDD5FAF6" box="[400,567,1339,1365]" pageId="8" pageNumber="494" type="generic">two specimens</specimenCount>
|
||
dissected),
|
||
<specimenCount id="9E8FFD75FFAC94736684FA92FD1BFAF6" box="[699,761,1339,1365]" pageId="8" pageNumber="494" type="female">4 ♀♀</specimenCount>
|
||
(
|
||
<specimenCount id="9E8FFD75FFAC94736737FA92FC40FAF6" box="[776,930,1339,1365]" pageId="8" pageNumber="494" type="generic">one specimen</specimenCount>
|
||
dissected): same data as the holotype (2 ♁♁: cSh, 1 ♁,
|
||
<specimenCount id="9E8FFD75FFAC94736533FAF6FEACFADA" box="[268,334,1375,1401]" pageId="8" pageNumber="494" type="female">4 ♀♀</specimenCount>
|
||
:
|
||
<collectionCode id="EE98AE39FFAC94736563FAF6FE55FADA" box="[348,439,1375,1401]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="8" pageNumber="494" type="Museum">MHNG</collectionCode>
|
||
); 1 ♁ (head damaged, aedeagus missing),
|
||
<specimenCount id="9E8FFD75FFAC947367A6FAF6FC38FADA" box="[921,986,1375,1401]" pageId="8" pageNumber="494" type="female">7 ♀♀</specimenCount>
|
||
(
|
||
<specimenCount id="9E8FFD75FFAC947367D5FAF6FB65FADA" box="[1002,1159,1375,1401]" pageId="8" pageNumber="494" type="generic">one specimen</specimenCount>
|
||
without head): ‘PAPUA N.
|
||
<collectingCountry id="F09E766CFFAC94736485FA2AFEC6FA3E" box="[186,292,1411,1437]" name="Guinea-Bissau" pageId="8" pageNumber="494">GUINEA</collectingCountry>
|
||
| Onerunka VIII. 79 [handwritten in black] | nr Kainantu | W. G. Ullrich` <printed> (1 ♁,
|
||
<specimenCount id="9E8FFD75FFAC94736124FA2AFABFFA3E" box="[1307,1373,1411,1437]" pageId="8" pageNumber="494" type="female">2 ♀♀</specimenCount>
|
||
: cSh;
|
||
<specimenCount id="9E8FFD75FFAC947364A8FA0FFF38FA62" box="[151,218,1446,1473]" pageId="8" pageNumber="494" type="female">5 ♀♀</specimenCount>
|
||
:
|
||
<collectionCode id="EE98AE39FFAC947364D5FA0EFEA7FA62" box="[234,325,1447,1473]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="8" pageNumber="494" type="Museum">MHNG</collectionCode>
|
||
). All
|
||
<typeStatus id="5732885EFFAC947365BBFA0EFE0DFA62" box="[388,495,1447,1473]" pageId="8" pageNumber="494">paratypes</typeStatus>
|
||
with additional red printed label: ‘
|
||
<typeStatus id="5732885EFFAC94736741FA0EFBECFA62" box="[894,1038,1447,1473]" pageId="8" pageNumber="494">PARATYPE</typeStatus>
|
||
|
|
||
<taxonomicName id="4F894D7FFFAC94736018FA0EFB00FA62" authorityName="Steel" authorityYear="1960" box="[1063,1250,1447,1473]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="494" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BAFDEAEEFFAC94736018FA0EFB00FA62" box="[1063,1250,1447,1473]" italics="true" pageId="8" pageNumber="494">Paraphloeostiba</emphasis>
|
||
</taxonomicName>
|
||
|
|
||
<taxonomicName id="4F894D7FFFAC947360C5FA0EFAAEFA62" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[1274,1356,1447,1473]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="ullrichi" status="sp.n.">
|
||
<emphasis id="BAFDEAEEFFAC947360C5FA0EFAAEFA62" box="[1274,1356,1447,1473]" italics="true" pageId="8" pageNumber="494">ullrichi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="BAFDEAEEFFAC9473616AFA0EFA6FFA62" bold="true" box="[1365,1421,1447,1473]" pageId="8" pageNumber="494">
|
||
<taxonomicNameLabel id="A1CE5795FFAC9473616AFA0EFA6FFA62" box="[1365,1421,1447,1473]" pageId="8" pageNumber="494" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
| Shavrin A.V. des. 2023`.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C0936577FFAC947164F8FA47FE85FA07" lastPageId="10" lastPageNumber="496" pageId="8" pageNumber="494" type="description">
|
||
<paragraph id="883636FCFFAC947364F8FA47FE28F9D6" blockId="8.[151,1437,1230,2013]" pageId="8" pageNumber="494">
|
||
<emphasis id="BAFDEAEEFFAC947364F8FA47FEBBF9AB" bold="true" box="[199,345,1518,1544]" pageId="8" pageNumber="494">Description.</emphasis>
|
||
Measurements (n=16): HW: 0.34–0.39; HL: 0.18–0.24; OL: 0.12–0.13; TL: 0.02–0.00; AL (
|
||
<typeStatus id="5732885EFFAC947364A0F9BAFEEAF98E" box="[159,264,1555,1581]" pageId="8" pageNumber="494">holotype</typeStatus>
|
||
): 0.55; PL: 0.28–0.34; PWmax: 0.52–0.57; PWmin: 0.41–0.46; ESL: 0.49–0.54; EW: 0.62–0.66; MTbL (
|
||
<typeStatus id="5732885EFFAC947364A0F99EFEEAF9F2" box="[159,264,1591,1617]" pageId="8" pageNumber="494">holotype</typeStatus>
|
||
): 0.30; MTrL (
|
||
<typeStatus id="5732885EFFAC9473658FF99EFDFBF9F2" box="[432,537,1591,1617]" pageId="8" pageNumber="494">holotype</typeStatus>
|
||
): 0.13 (MTrL 1–4: 0.05; MTrL 5: 0.08); AW: 0.59–0.64; AedL: 0.30–0.32; BL: 1.40–2.34 (
|
||
<typeStatus id="5732885EFFAC94736529F9F2FE9CF9D6" box="[278,382,1627,1653]" pageId="8" pageNumber="494">holotype</typeStatus>
|
||
: 1.75).
|
||
</paragraph>
|
||
<paragraph id="883636FCFFAC947364F8F9D6FDA1F87E" blockId="8.[151,1437,1230,2013]" pageId="8" pageNumber="494">
|
||
Habitus as in
|
||
<figureCitation id="10B22A79FFAC94736552F9D6FE57F93A" box="[365,437,1663,1689]" captionStart="FIGURES 1–6" captionStartId="2.[151,275,1541,1567]" captionTargetBox="[197,1385,250,1508]" captionTargetId="figure-45@2.[187,1400,237,1520]" captionTargetPageId="2" captionText="FIGURES 1–6. Habitus of Paraphloeostiba: 1—P. conjuncta (holotype), 2—P. coriacea, 3—P. flaveola (holotype), 4—P. gayndahensis (Papua New Guinea), 5—P. papuana (holotype), 6—P. ullrichi (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8036119" httpUri="https://zenodo.org/record/8036119/files/figure.png" pageId="8" pageNumber="494">Fig. 6</figureCitation>
|
||
. Body yellow-brown (
|
||
<specimenCount id="9E8FFD75FFAC94736683F9D6FCB1F93A" box="[700,851,1663,1689]" pageId="8" pageNumber="494" type="generic" typeStatus="paratype">one paratype</specimenCount>
|
||
) to reddish-brown; mouthparts, antennae and legs yellow (antennomeres 7–11 sometimes slightly darker). Head with very sparse and fine punctation, middle portion of head usually without punctures; neck and scutellum without punctures; pronotum with sparse, moderately large and deep pucntation, sparser in middle portion; punctation of elytra sparse, distinctly larger and deeper than that on pronotum, sparser and finer in middle portions of each elytron; abdomen with fine and sparse punctation, sparser and finer in middle portion of each tergites. Head with very fine microsculpture, somewhat isodiametric in middle and sometimes indistinct in basal portions; neck with very fine transverse microreticulation; pronotum with fine transverse sculpture, indistinct in mediobasal portion in some
|
||
<typeStatus id="5732885EFFAC947367AFF8D2FC1DF836" box="[912,1023,1915,1941]" pageId="8" pageNumber="494">paratypes</typeStatus>
|
||
; scutellum with very fine transverse meshes; elytra without microsculpture except basal portion with fine transverse microreticulation in some
|
||
<typeStatus id="5732885EFFAC94736116F836FA7AF81A" box="[1321,1432,1951,1977]" pageId="8" pageNumber="494">paratypes</typeStatus>
|
||
; abdomen with fine transverse meshes.
|
||
</paragraph>
|
||
<caption id="DCF66674FFAD947264A8FA27FB52FA4E" ID-DOI="http://doi.org/10.5281/zenodo.8036125" ID-Zenodo-Dep="8036125" httpUri="https://zenodo.org/record/8036125/files/figure.png" pageId="9" pageNumber="495" startId="9.[151,264,1422,1446]" targetBox="[162,1423,195,1387]" targetPageId="9" targetType="figure">
|
||
<paragraph id="883636FCFFAD947264A8FA27FB52FA4E" blockId="9.[151,1437,1422,1518]" pageId="9" pageNumber="495">
|
||
<emphasis id="BAFDEAEEFFAD947264A8FA27FEB1FA05" bold="true" box="[151,339,1422,1446]" pageId="9" pageNumber="495">FIGURES 26–32.</emphasis>
|
||
Aedeagus and abdominal segments of
|
||
<taxonomicName id="4F894D7FFFAD947266E6FA27FC30FA05" authorityName="Shavrin" authorityYear="2023" box="[729,978,1422,1446]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="495" phylum="Arthropoda" rank="species" species="ullrichi">
|
||
<emphasis id="BAFDEAEEFFAD947266E6FA27FC30FA05" box="[729,978,1422,1446]" italics="true" pageId="9" pageNumber="495">Paraphloeostiba ullrichi</emphasis>
|
||
</taxonomicName>
|
||
(paratypes): 26—aedeagus (parameral view), 27—aedeagus (lateral view), 28—male tergite VIII (dorsal view), 29—male sternite VIII (ventral view), 30—female tergite VIII (dorsal view), 31—female sternite VIII (ventral view), 32—female genital segment. Scale bar: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="883636FCFFAD947264F8F9E9FAE6F8AC" blockId="9.[151,1437,1600,1986]" pageId="9" pageNumber="495">Head slightly elevated in middle portion, 1.6–1.8 times as broad as long, with short and deep anteocellar foveae, slightly longer than diameter of ocellus, reaching basal third of level of eyes. Apical maxillary palpomere narrow and short, about 1.3 times as long as preapical segment. Antennomeres 6–10 strongly transverse; antennomere 2 about as broad as basal antennomere, 3 distinctly narrower and shorter than 2, 4 shorter than 3, 5 slightly broader than 4, 6 about twice broader than 5, 7 distinctly broader and longer than 6, 8–10 slightly shorter than 7, apical antennomere about three times as long as 10, from middle gradually narrowed toward subacute apex.</paragraph>
|
||
<paragraph id="883636FCFFAD947264F8F8B0FA80F8D9" blockId="9.[151,1437,1600,1986]" pageId="9" pageNumber="495">Pronotum 1.6–1.8 times as broad as long, 1.4–1.5 times as broad as head, widest in middle, more narrowed posteriad than anteriad; posterior angles obtuse; anterior margin distinctly concave in middle; middle portion without paramedian impressions; laterobasal portions without or with with shallow and moderately wide impressions.</paragraph>
|
||
<paragraph id="883636FCFFAD947264F8F82DFEB4F861" blockId="9.[151,1437,1600,1986]" pageId="9" pageNumber="495">Elytra 1.2 times as broad as long, 1.5–1.7 as long as pronotum, distinctly broadened apicad; posterior margin slightly rounded.</paragraph>
|
||
<caption id="DCF66674FFAE947164A8FCE2FD29FC25" ID-DOI="http://doi.org/10.5281/zenodo.8036127" ID-Zenodo-Dep="8036127" httpUri="https://zenodo.org/record/8036127/files/figure.png" pageId="10" pageNumber="496" startId="10.[151,264,843,867]" targetBox="[159,1429,187,811]" targetPageId="10" targetType="figure">
|
||
<paragraph id="883636FCFFAE947164A8FCE2FD29FC25" blockId="10.[151,1437,843,903]" pageId="10" pageNumber="496">
|
||
<emphasis id="BAFDEAEEFFAE947164A8FCE2FEB7FCC0" bold="true" box="[151,341,843,867]" pageId="10" pageNumber="496">FIGURES 33–36.</emphasis>
|
||
Habitus of
|
||
<taxonomicName id="4F894D7FFFAE947165F2FCE2FD94FCC0" authorityName="Steel" authorityYear="1960" box="[461,630,843,867]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="496" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BAFDEAEEFFAE947165F2FCE2FD94FCC0" box="[461,630,843,867]" italics="true" pageId="10" pageNumber="496">Paraphloeostiba</emphasis>
|
||
</taxonomicName>
|
||
: 33—
|
||
<taxonomicName id="4F894D7FFFAE94716689FCE5FC43FCC0" authority="(Mindanao)" baseAuthorityName="Mindanao" box="[694,929,843,867]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="496" phylum="Arthropoda" rank="species" species="apicalis">
|
||
<emphasis id="BAFDEAEEFFAE94716689FCE5FCFDFCC0" box="[694,799,843,867]" italics="true" pageId="10" pageNumber="496">P. apicalis</emphasis>
|
||
(Mindanao)
|
||
</taxonomicName>
|
||
, 34—
|
||
<taxonomicName id="4F894D7FFFAE947167E1FCE5FB75FCC0" authority="(Borneo)" baseAuthorityName="Borneo" box="[990,1175,843,867]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="496" phylum="Arthropoda" rank="species" species="obesa">
|
||
<emphasis id="BAFDEAEEFFAE947167E1FCE5FBD0FCC0" box="[990,1074,843,867]" italics="true" pageId="10" pageNumber="496">P. obesa</emphasis>
|
||
(Borneo)
|
||
</taxonomicName>
|
||
, 35—
|
||
<taxonomicName id="4F894D7FFFAE947160EBFCE5FAB0FCC0" baseAuthorityName="Cameron" baseAuthorityYear="1924" box="[1236,1362,843,867]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="496" phylum="Arthropoda" rank="species" species="subopaca">
|
||
<emphasis id="BAFDEAEEFFAE947160EBFCE5FB01FCC0" box="[1236,1251,844,867]" italics="true" pageId="10" pageNumber="496">P</emphasis>
|
||
.
|
||
<emphasis id="BAFDEAEEFFAE947160CEFCE2FAB0FCC0" box="[1265,1362,843,867]" italics="true" pageId="10" pageNumber="496">subopaca</emphasis>
|
||
</taxonomicName>
|
||
(Tamil Nadu), 36—
|
||
<taxonomicName id="4F894D7FFFAE94716529FCD9FE46FC24" baseAuthorityName="Cameron" baseAuthorityYear="1940" box="[278,420,879,903]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="496" phylum="Arthropoda" rank="species" species="tonkinensis">
|
||
<emphasis id="BAFDEAEEFFAE94716529FCD9FEC7FC24" box="[278,293,880,903]" italics="true" pageId="10" pageNumber="496">P</emphasis>
|
||
.
|
||
<emphasis id="BAFDEAEEFFAE9471650DFCC6FE46FC24" box="[306,420,879,903]" italics="true" pageId="10" pageNumber="496">tonkinensis</emphasis>
|
||
</taxonomicName>
|
||
(Leyte). Scale bars: 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="883636FCFFAE947164F8FC70FD60FC50" blockId="10.[151,1437,985,1588]" box="[199,642,985,1012]" pageId="10" pageNumber="496">Abdomen slightly narrower than elytra.</paragraph>
|
||
<paragraph id="883636FCFFAE947164F8FC54FBFFFB04" blockId="10.[151,1437,985,1588]" pageId="10" pageNumber="496">
|
||
Male. Posterior margin of abdominal tergite VIII slightly concave (
|
||
<figureCitation id="10B22A79FFAE947167EFFC54FBC5FBB4" box="[976,1063,1021,1047]" captionStart="FIGURES 26–32" captionStartId="9.[151,264,1422,1446]" captionTargetBox="[162,1423,195,1387]" captionTargetId="figure-24@9.[151,1436,181,1398]" captionTargetPageId="9" captionText="FIGURES 26–32. Aedeagus and abdominal segments of Paraphloeostiba ullrichi (paratypes): 26—aedeagus (parameral view), 27—aedeagus (lateral view), 28—male tergite VIII (dorsal view), 29—male sternite VIII (ventral view), 30—female tergite VIII (dorsal view), 31—female sternite VIII (ventral view), 32—female genital segment. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8036125" httpUri="https://zenodo.org/record/8036125/files/figure.png" pageId="10" pageNumber="496">Fig. 28</figureCitation>
|
||
). Posterior margin of abdominal sternite VIII widely and deeply concave (
|
||
<figureCitation id="10B22A79FFAE94716646FB88FD32FB98" box="[633,720,1057,1083]" captionStart="FIGURES 26–32" captionStartId="9.[151,264,1422,1446]" captionTargetBox="[162,1423,195,1387]" captionTargetId="figure-24@9.[151,1436,181,1398]" captionTargetPageId="9" captionText="FIGURES 26–32. Aedeagus and abdominal segments of Paraphloeostiba ullrichi (paratypes): 26—aedeagus (parameral view), 27—aedeagus (lateral view), 28—male tergite VIII (dorsal view), 29—male sternite VIII (ventral view), 30—female tergite VIII (dorsal view), 31—female sternite VIII (ventral view), 32—female genital segment. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8036125" httpUri="https://zenodo.org/record/8036125/files/figure.png" pageId="10" pageNumber="496">Fig. 29</figureCitation>
|
||
). Aedeagus with small basal bulb, gradually narrowed toward narrow and long median lobe with rounded apex; parameres short, not reaching apex of median lobe, with two moderately long apical and three preapical setae; internal sac moderately narrow and short, with sclerotized narrow struncture in basal portion (
|
||
<figureCitation id="10B22A79FFAE947165F3FB24FDC2FB04" box="[460,544,1165,1191]" captionStart="FIGURES 26–32" captionStartId="9.[151,264,1422,1446]" captionTargetBox="[162,1423,195,1387]" captionTargetId="figure-24@9.[151,1436,181,1398]" captionTargetPageId="9" captionText="FIGURES 26–32. Aedeagus and abdominal segments of Paraphloeostiba ullrichi (paratypes): 26—aedeagus (parameral view), 27—aedeagus (lateral view), 28—male tergite VIII (dorsal view), 29—male sternite VIII (ventral view), 30—female tergite VIII (dorsal view), 31—female sternite VIII (ventral view), 32—female genital segment. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8036125" httpUri="https://zenodo.org/record/8036125/files/figure.png" pageId="10" pageNumber="496">Fig. 26</figureCitation>
|
||
). Lateral aspect of the aedeagus as in
|
||
<figureCitation id="10B22A79FFAE947167F8FB24FBFBFB04" box="[967,1049,1165,1191]" captionStart="FIGURES 26–32" captionStartId="9.[151,264,1422,1446]" captionTargetBox="[162,1423,195,1387]" captionTargetId="figure-24@9.[151,1436,181,1398]" captionTargetPageId="9" captionText="FIGURES 26–32. Aedeagus and abdominal segments of Paraphloeostiba ullrichi (paratypes): 26—aedeagus (parameral view), 27—aedeagus (lateral view), 28—male tergite VIII (dorsal view), 29—male sternite VIII (ventral view), 30—female tergite VIII (dorsal view), 31—female sternite VIII (ventral view), 32—female genital segment. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8036125" httpUri="https://zenodo.org/record/8036125/files/figure.png" pageId="10" pageNumber="496">Fig. 27</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="883636FCFFAE947164F8FB18FEF8FAB0" blockId="10.[151,1437,985,1588]" pageId="10" pageNumber="496">
|
||
Female. Posterior margin of abdominal tergite VIII truncate (
|
||
<figureCitation id="10B22A79FFAE94716742FB18FC31FB68" box="[893,979,1201,1227]" captionStart="FIGURES 26–32" captionStartId="9.[151,264,1422,1446]" captionTargetBox="[162,1423,195,1387]" captionTargetId="figure-24@9.[151,1436,181,1398]" captionTargetPageId="9" captionText="FIGURES 26–32. Aedeagus and abdominal segments of Paraphloeostiba ullrichi (paratypes): 26—aedeagus (parameral view), 27—aedeagus (lateral view), 28—male tergite VIII (dorsal view), 29—male sternite VIII (ventral view), 30—female tergite VIII (dorsal view), 31—female sternite VIII (ventral view), 32—female genital segment. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8036125" httpUri="https://zenodo.org/record/8036125/files/figure.png" pageId="10" pageNumber="496">Fig. 30</figureCitation>
|
||
). Posterior margin of abdominal sternite VIII widely rounded (
|
||
<figureCitation id="10B22A79FFAE947165B2FB7CFE02FB4C" box="[397,480,1237,1263]" captionStart="FIGURES 26–32" captionStartId="9.[151,264,1422,1446]" captionTargetBox="[162,1423,195,1387]" captionTargetId="figure-24@9.[151,1436,181,1398]" captionTargetPageId="9" captionText="FIGURES 26–32. Aedeagus and abdominal segments of Paraphloeostiba ullrichi (paratypes): 26—aedeagus (parameral view), 27—aedeagus (lateral view), 28—male tergite VIII (dorsal view), 29—male sternite VIII (ventral view), 30—female tergite VIII (dorsal view), 31—female sternite VIII (ventral view), 32—female genital segment. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8036125" httpUri="https://zenodo.org/record/8036125/files/figure.png" pageId="10" pageNumber="496">Fig. 31</figureCitation>
|
||
). Female genital segment as in
|
||
<figureCitation id="10B22A79FFAE9471677FFB7CFC70FB4C" box="[832,914,1237,1263]" captionStart="FIGURES 26–32" captionStartId="9.[151,264,1422,1446]" captionTargetBox="[162,1423,195,1387]" captionTargetId="figure-24@9.[151,1436,181,1398]" captionTargetPageId="9" captionText="FIGURES 26–32. Aedeagus and abdominal segments of Paraphloeostiba ullrichi (paratypes): 26—aedeagus (parameral view), 27—aedeagus (lateral view), 28—male tergite VIII (dorsal view), 29—male sternite VIII (ventral view), 30—female tergite VIII (dorsal view), 31—female sternite VIII (ventral view), 32—female genital segment. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8036125" httpUri="https://zenodo.org/record/8036125/files/figure.png" pageId="10" pageNumber="496">Fig. 32</figureCitation>
|
||
. Female acessory sclerite and spermatheca not recognized.
|
||
</paragraph>
|
||
<paragraph id="883636FCFFAE947164F8FAB4FE85FA07" blockId="10.[151,1437,985,1588]" pageId="10" pageNumber="496">
|
||
<emphasis id="BAFDEAEEFFAE947164F8FAB4FE52FA94" bold="true" box="[199,432,1309,1335]" pageId="10" pageNumber="496">Comparative notes.</emphasis>
|
||
Based on the glossy body with fine sculpture of the forebody and the general shape of the aedeagus,
|
||
<taxonomicName id="4F894D7FFFAE94716532FAEBFE9EFAF8" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[269,380,1345,1371]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="496" phylum="Arthropoda" rank="species" species="ullrichi" status="sp.n.">
|
||
<emphasis id="BAFDEAEEFFAE94716532FAEBFE9EFAF8" box="[269,380,1345,1371]" italics="true" pageId="10" pageNumber="496">P. ullrichi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="BAFDEAEEFFAE947165BBFAE8FE5EFAF8" bold="true" box="[388,444,1345,1371]" pageId="10" pageNumber="496">
|
||
<taxonomicNameLabel id="A1CE5795FFAE947165BBFAE8FE5EFAF8" box="[388,444,1345,1371]" pageId="10" pageNumber="496" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar to
|
||
<taxonomicName id="4F894D7FFFAE9471666CFAEBFC73FAF8" authority="Steel, 1960" authorityName="Steel" authorityYear="1960" box="[595,913,1345,1371]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="496" phylum="Arthropoda" rank="species" species="solomonensis">
|
||
<emphasis id="BAFDEAEEFFAE9471666CFAEBFCE5FAF8" box="[595,775,1345,1371]" italics="true" pageId="10" pageNumber="496">P. solomonensis</emphasis>
|
||
<bibRefCitation id="EC184B0DFFAE94716730FAE8FC73FAF8" author="Steel, W. O." box="[783,913,1345,1371]" pageId="10" pageNumber="496" pagination="141 - 172" refId="ref9011" refString="Steel, W. O. (1960) Three new omaliine genera from Asia and Australasia previously confused with Phloeonomus Thomson (Coleoptera: Staphylinidae). The Transactions of the Royal Entomological Society of London, 112, 141 - 172. https: // doi. org / 10.1111 / j. 1365 - 2311.1960. tb 00497. x" type="journal article" year="1960">Steel, 1960</bibRefCitation>
|
||
</taxonomicName>
|
||
, known from “
|
||
<collectingCountry id="F09E766CFFAE94716005FAE8FB1CFAF8" box="[1082,1278,1345,1371]" name="Solomon Islands" pageId="10" pageNumber="496">Solomon Islands</collectingCountry>
|
||
:
|
||
<collectingRegion id="4A4DF81EFFAE94716136FAE8FA7AFAF8" box="[1289,1432,1345,1371]" country="Solomon Islands" name="Guadalcanal" pageId="10" pageNumber="496">Guadalcanal</collectingRegion>
|
||
, Kukum...” (
|
||
<bibRefCitation id="EC184B0DFFAE94716524FACCFE75FADC" author="Steel, W. O." box="[283,407,1381,1407]" pageId="10" pageNumber="496" pagination="141 - 172" refId="ref9011" refString="Steel, W. O. (1960) Three new omaliine genera from Asia and Australasia previously confused with Phloeonomus Thomson (Coleoptera: Staphylinidae). The Transactions of the Royal Entomological Society of London, 112, 141 - 172. https: // doi. org / 10.1111 / j. 1365 - 2311.1960. tb 00497. x" type="journal article" year="1960">Steel 1960</bibRefCitation>
|
||
). It can be distinguished from it by the slightly broader apical portion of the median lobe and shorter parameres.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C0936577FFAE947164F8FA04FF12FA48" pageId="10" pageNumber="496" type="distribution">
|
||
<paragraph id="883636FCFFAE947164F8FA04FF12FA48" blockId="10.[151,1437,985,1588]" pageId="10" pageNumber="496">
|
||
<emphasis id="BAFDEAEEFFAE947164F8FA04FE83FA64" bold="true" box="[199,353,1453,1479]" pageId="10" pageNumber="496">Distribution.</emphasis>
|
||
<taxonomicName id="4F894D7FFFAE94716552FA04FD64FA64" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[365,646,1453,1479]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="496" phylum="Arthropoda" rank="species" species="ullrichi" status="sp.n.">
|
||
<emphasis id="BAFDEAEEFFAE94716552FA04FD64FA64" box="[365,646,1453,1479]" italics="true" pageId="10" pageNumber="496">Paraphloeostiba ullrichi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="BAFDEAEEFFAE947166ACFA04FD29FA64" bold="true" box="[659,715,1453,1479]" pageId="10" pageNumber="496">
|
||
<taxonomicNameLabel id="A1CE5795FFAE947166ACFA04FD29FA64" box="[659,715,1453,1479]" pageId="10" pageNumber="496" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is known from two localities in eastern part of
|
||
<collectingCountry id="F09E766CFFAE94716129FA07FF0EFA48" name="Papua New Guinea" pageId="10" pageNumber="496">Papua New Guinea</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C0936577FFAE947164F8FA5CFE76F990" pageId="10" pageNumber="496" type="etymology">
|
||
<paragraph id="883636FCFFAE947164F8FA5CFE76F990" blockId="10.[151,1437,985,1588]" pageId="10" pageNumber="496">
|
||
<emphasis id="BAFDEAEEFFAE947164F8FA5CFEACF9AC" bold="true" box="[199,334,1525,1551]" pageId="10" pageNumber="496">Etymology.</emphasis>
|
||
Patronymic, the species is named to honour W.G. Ullrich (Lübeck), collector of the
|
||
<typeStatus id="5732885EFFAE947160CEFA5FFAC3F9B3" box="[1265,1313,1526,1552]" pageId="10" pageNumber="496">type</typeStatus>
|
||
specimens in
|
||
<collectingCountry id="F09E766CFFAE9471648BF9B3FE72F990" box="[180,400,1561,1588]" name="Papua New Guinea" pageId="10" pageNumber="496">Papua New Guinea</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |