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<document id="7830770EDC89F11FDC66B66661DC3E5A" ID-DOI="10.11646/zootaxa.5301.4.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="8036117" ID-ZooBank="01F6333F-807B-448A-80C8-EC1F8CE030EA" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1686723262927" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2023" docId="002087EAFFA9947664A8FE8FFCEAFD61" docLanguage="en" docName="zootaxa.5301.4.6.pdf" docOrigin="Zootaxa 5301 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5301.4.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paraphloeostiba tonkinensis" docType="treatment" docVersion="2" lastPageNumber="499" masterDocId="FC19FF92FFA4947B643FFFA9FFE2FFA3" masterDocTitle="New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines" masterLastPageNumber="500" masterPageNumber="486" pageNumber="499" updateTime="1686723785076" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="45A2A7EC6A1239D93A57B9D226147F30">New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines</mods:title>
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<treatment id="002087EAFFA9947664A8FE8FFCEAFD61" LSID="urn:lsid:plazi:treatment:002087EAFFA9947664A8FE8FFCEAFD61" httpUri="http://treatment.plazi.org/id/002087EAFFA9947664A8FE8FFCEAFD61" lastPageNumber="499" pageId="13" pageNumber="499">
<subSubSection id="C0936577FFA9947664A8FE8FFD25FEE3" box="[151,711,294,321]" pageId="13" pageNumber="499" type="nomenclature">
<paragraph id="883636FCFFA9947664A8FE8FFD25FEE3" blockId="13.[151,711,294,357]" box="[151,711,294,321]" pageId="13" pageNumber="499">
<heading id="D37E8190FFA9947664A8FE8FFD25FEE3" bold="true" box="[151,711,294,321]" fontSize="11" level="1" pageId="13" pageNumber="499" reason="1">
<taxonomicName id="4F894D7FFFA9947664A8FE8FFD25FEE3" ID-CoL="4D8B7" authority="(Cameron, 1940)" baseAuthorityName="Cameron" baseAuthorityYear="1940" box="[151,711,294,321]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="499" phylum="Arthropoda" rank="species" species="tonkinensis">
<emphasis id="BAFDEAEEFFA9947664A8FE8FFD25FEE3" bold="true" box="[151,711,294,321]" pageId="13" pageNumber="499">
<emphasis id="BAFDEAEEFFA9947664A8FE8FFE0DFEE2" bold="true" box="[151,495,294,321]" italics="true" pageId="13" pageNumber="499">Paraphloeostiba tonkinensis</emphasis>
(
<bibRefCitation id="EC184B0DFFA9947665C1FE8FFD5DFEE3" author="Cameron, M." box="[510,703,294,320]" pageId="13" pageNumber="499" pagination="181 - 184" refId="ref8312" refString="Cameron, M. (1940) New species of Oriental Staphylinidae (Col.). The Entomologist's Monthly Magazine, 76, 181 - 184." type="journal article" year="1940">Cameron, 1940</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C0936577FFA9947664A8FEE2FEBAFEC6" box="[151,344,331,357]" pageId="13" pageNumber="499" type="description">
<paragraph id="883636FCFFA9947664A8FEE2FEBAFEC6" blockId="13.[151,711,294,357]" box="[151,344,331,357]" pageId="13" pageNumber="499">
(
<figureCitation id="10B22A79FFA9947664A0FEE2FF1FFEC6" box="[159,253,331,357]" captionStart="FIGURES 3336" captionStartId="10.[151,264,843,867]" captionTargetBox="[159,1429,187,811]" captionTargetId="figure-15@10.[151,1436,181,819]" captionTargetPageId="10" captionText="FIGURES 3336. Habitus of Paraphloeostiba: 33—P. apicalis (Mindanao), 34—P. obesa (Borneo), 35—P. subopaca (Tamil Nadu), 36—P. tonkinensis (Leyte). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8036127" httpUri="https://zenodo.org/record/8036127/files/figure.png" pageId="13" pageNumber="499">Figs. 36</figureCitation>
,
<figureCitation id="10B22A79FFA994766536FEE2FEB2FEC6" box="[265,336,331,357]" captionStart="FIGURES 3742" captionStartId="12.[151,264,1810,1834]" captionTargetBox="[166,1417,195,1777]" captionTargetId="figure-15@12.[151,1436,181,1786]" captionTargetPageId="12" captionText="FIGURES 3742. Aedeagus of Paraphloeostiba (Figs. 3738: P. apicalis (Mindanao), Figs. 3940: P. obesa (Sabah), Figs. 4142: P. subopaca (Tamil Nadu), Figs. 4344: P. tonkinensis (Leyte)): 37, 39, 41, 43—parameral view, 38, 40, 42, 44—lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8036129" httpUri="https://zenodo.org/record/8036129/files/figure.png" pageId="13" pageNumber="499">4142</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C0936577FFA9947664A8FE26FDBAFE65" pageId="13" pageNumber="499" type="reference_group">
<paragraph id="883636FCFFA9947664A8FE26FD8DFE04" blockId="13.[151,623,399,454]" box="[151,623,399,423]" pageId="13" pageNumber="499">
<treatmentCitationGroup id="A89911D2FFA9947664A8FE26FD8DFE04" box="[151,623,399,423]" pageId="13" pageNumber="499">
<treatmentCitation id="092810EDFFA9947664A8FE26FD8DFE04" author="Cameron, M." box="[151,623,399,423]" page="181" pageId="13" pageNumber="499" year="1940">
<taxonomicName id="4F894D7FFFA9947664A8FE26FD8DFE04" ID-CoL="4GGGT" authority="Cameron, 1940: 181" authorityName="Cameron" authorityPageNumber="181" authorityYear="1940" box="[151,623,399,423]" class="Insecta" family="Staphylinidae" genus="Phloeonomus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="499" phylum="Arthropoda" rank="species" species="tonkinensis">
<emphasis id="BAFDEAEEFFA9947664A8FE26FE7BFE04" box="[151,409,399,423]" italics="true" pageId="13" pageNumber="499">Phloeonomus tonkinensis</emphasis>
<bibRefCitation id="EC184B0DFFA99476659FFE26FD8DFE04" author="Cameron, M." box="[416,623,399,423]" pageId="13" pageNumber="499" pagination="181 - 184" refId="ref8312" refString="Cameron, M. (1940) New species of Oriental Staphylinidae (Col.). The Entomologist's Monthly Magazine, 76, 181 - 184." type="journal article" year="1940">Cameron, 1940: 181</bibRefCitation>
</taxonomicName>
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</paragraph>
<paragraph id="883636FCFFA9947664A8FE07FDBAFE65" blockId="13.[151,623,399,454]" box="[151,600,430,454]" pageId="13" pageNumber="499">
<treatmentCitationGroup id="A89911D2FFA9947664A8FE07FDBAFE65" box="[151,600,430,454]" pageId="13" pageNumber="499">
<treatmentCitation id="092810EDFFA9947664A8FE07FDBAFE65" author="Steel, W. O." box="[151,600,430,454]" page="167" pageId="13" pageNumber="499" year="1960">
<taxonomicName id="4F894D7FFFA9947664A8FE07FDBAFE65" authority=": Steel 1960: 167" authorityName="Steel" authorityPageNumber="167" authorityYear="1960" box="[151,600,430,454]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="499" phylum="Arthropoda" rank="species" species="tonkinesis">
<emphasis id="BAFDEAEEFFA9947664A8FE07FE49FE65" box="[151,427,430,454]" italics="true" pageId="13" pageNumber="499">Paraphloeostiba tonkinesis</emphasis>
:
<bibRefCitation id="EC184B0DFFA994766586FE07FDBAFE65" author="Steel, W. O." box="[441,600,430,454]" pageId="13" pageNumber="499" pagination="141 - 172" refId="ref9011" refString="Steel, W. O. (1960) Three new omaliine genera from Asia and Australasia previously confused with Phloeonomus Thomson (Coleoptera: Staphylinidae). The Transactions of the Royal Entomological Society of London, 112, 141 - 172. https: // doi. org / 10.1111 / j. 1365 - 2311.1960. tb 00497. x" type="journal article" year="1960">Steel 1960: 167</bibRefCitation>
</taxonomicName>
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</paragraph>
</subSubSection>
<subSubSection id="C0936577FFA9947664A8FE5DFDEAFD91" pageId="13" pageNumber="499" type="materials_examined">
<paragraph id="883636FCFFA9947664A8FE5DFDEAFD91" blockId="13.[151,1437,500,707]" pageId="13" pageNumber="499">
<emphasis id="BAFDEAEEFFA9947664A8FE5DFE62FDAD" bold="true" box="[151,384,500,526]" pageId="13" pageNumber="499">Material examined.</emphasis>
<materialsCitation id="38E13CA1FFA9947665B8FE5DFDE6FD91" collectingDate="1991-02-22" collectionCode="SMNS" collectorName="Visca N Baybay. &amp; W. Shawaller" country="Philippines" elevation="350" location="Leyte" pageId="13" pageNumber="499" specimenCount="1" stateProvince="Southern Leyte">
<emphasis id="BAFDEAEEFFA9947665B8FE5DFDA1FDAD" bold="true" box="[391,579,500,526]" pageId="13" pageNumber="499">
<collectingCountry id="F09E766CFFA9947665B8FE5DFDDCFDAD" box="[391,574,500,526]" name="Philippines" pageId="13" pageNumber="499">PHILIPPINES</collectingCountry>
:
</emphasis>
2 ♁♁:
<collectingRegion id="4A4DF81EFFA9947666A9FE5CFD3BFDAC" box="[662,729,501,527]" country="Philippines" name="Southern Leyte" pageId="13" pageNumber="499">Leyte</collectingRegion>
,
<collectorName id="257C532AFFA9947666DCFE5DFC7CFDAD" box="[739,926,500,527]" pageId="13" pageNumber="499">Visca N Baybay.</collectorName>
<date id="FC37103CFFA99476679BFE5DFBC7FDAD" box="[932,1061,500,526]" pageId="13" pageNumber="499" value="1991-02-22">
<collectingDate id="EC73E9D4FFA99476679BFE5DFBC7FDAD" box="[932,1061,500,526]" pageId="13" pageNumber="499" value="1991-02-22">22.02.1991</collectingDate>
</date>
.
<elevation id="03A4D1CFFFA99476600FFE5DFB08FDAD" box="[1072,1258,500,527]" metricMagnitude="2" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="13" pageNumber="499" unit="m" value="350.0" valueMax="500.0" valueMin="200.0">
<quantity id="4F719B19FFA99476600FFE5DFB4CFDAC" box="[1072,1198,500,527]" metricMagnitude="2" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="13" pageNumber="499" unit="m" value="350.0" valueMax="500.0" valueMin="200.0">200500 m</quantity>
a.s.l.
</elevation>
, primary forest.
<collectorName id="257C532AFFA9947664A8FDB0FECFFD91" box="[151,301,536,563]" pageId="13" pageNumber="499">W. Shawaller</collectorName>
et al. leg. (
<collectionCode id="EE98AE39FFA994766595FDB1FE1EFD91" box="[426,508,536,562]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="13" pageNumber="499">SMNS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C0936577FFA9947664F8FD95FCEAFD61" pageId="13" pageNumber="499" type="discussion">
<paragraph id="883636FCFFA9947664F8FD95FCEAFD61" blockId="13.[151,1437,500,707]" pageId="13" pageNumber="499">
<emphasis id="BAFDEAEEFFA9947664F8FD95FED9FDF5" bold="true" box="[199,315,572,598]" pageId="13" pageNumber="499">Remarks.</emphasis>
<taxonomicName id="4F894D7FFFA994766500FD95FDB8FDF5" authorityName="Cameron" authorityYear="1940" box="[319,602,572,598]" class="Insecta" family="Staphylinidae" genus="Phloeonomus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="499" phylum="Arthropoda" rank="species" species="tonkinensis">
<emphasis id="BAFDEAEEFFA994766500FD95FDB8FDF5" box="[319,602,572,598]" italics="true" pageId="13" pageNumber="499">Phloeonomus tonkinensis</emphasis>
</taxonomicName>
was origianlly described from “
<collectingCountry id="F09E766CFFA99476678DFD95FBEAFDF5" box="[946,1032,572,598]" name="Vietnam" pageId="13" pageNumber="499">Tonkin</collectingCountry>
:
<collectingRegion id="4A4DF81EFFA99476602FFD94FB9CFDF5" box="[1040,1150,572,599]" country="Vietnam" name="Hoa Binh" pageId="13" pageNumber="499">Hoa Binh</collectingRegion>
”.
<bibRefCitation id="EC184B0DFFA9947660AFFD95FAF5FDF5" author="Steel, W. O." box="[1168,1303,572,598]" pageId="13" pageNumber="499" pagination="141 - 172" refId="ref9011" refString="Steel, W. O. (1960) Three new omaliine genera from Asia and Australasia previously confused with Phloeonomus Thomson (Coleoptera: Staphylinidae). The Transactions of the Royal Entomological Society of London, 112, 141 - 172. https: // doi. org / 10.1111 / j. 1365 - 2311.1960. tb 00497. x" type="journal article" year="1960">Steel (1960)</bibRefCitation>
redescribed it as
<taxonomicName id="4F894D7FFFA9947664F2FDC9FE6AFDD9" authorityName="Steel" authorityYear="1960" box="[205,392,608,634]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="499" phylum="Arthropoda" rank="genus">
<emphasis id="BAFDEAEEFFA9947664F2FDC9FE6AFDD9" box="[205,392,608,634]" italics="true" pageId="13" pageNumber="499">Paraphloeostiba</emphasis>
</taxonomicName>
, and provided figures of the aedeagus, the female genital segment and the spermatheca. The species was known from
<collectingCountry id="F09E766CFFA994766589FD2DFDF7FD3D" box="[438,533,644,670]" name="Vietnam" pageId="13" pageNumber="499">Vietnam</collectingCountry>
and
<collectingCountry id="F09E766CFFA994766672FD2DFD5DFD3D" box="[589,703,644,670]" name="Singapore" pageId="13" pageNumber="499">Singapore</collectingCountry>
(
<bibRefCitation id="EC184B0DFFA9947666F1FD2DFC8CFD3D" author="Herman, L. H." box="[718,878,644,670]" pageId="13" pageNumber="499" pagination="1 - 7" refId="ref8478" refString="Herman, L. H. (2001) Catalogue of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millenium. Bulletin of the American Museum of Natural History, 265 (1 - 7), i - vi + 1 - 4218. https: // doi. org / 10.1206 / 0003 - 0090.265.1.1" type="journal article" year="2001">Herman 2001</bibRefCitation>
). It is here recorded from
<collectingCountry id="F09E766CFFA9947660A9FD2DFAF6FD3D" box="[1174,1300,644,670]" name="Philippines" pageId="13" pageNumber="499">Philippines</collectingCountry>
for the first time. Habitus as in
<figureCitation id="10B22A79FFA99476654FFD01FE21FD61" box="[368,451,680,706]" captionStart="FIGURES 3336" captionStartId="10.[151,264,843,867]" captionTargetBox="[159,1429,187,811]" captionTargetId="figure-15@10.[151,1436,181,819]" captionTargetPageId="10" captionText="FIGURES 3336. Habitus of Paraphloeostiba: 33—P. apicalis (Mindanao), 34—P. obesa (Borneo), 35—P. subopaca (Tamil Nadu), 36—P. tonkinensis (Leyte). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8036127" httpUri="https://zenodo.org/record/8036127/files/figure.png" pageId="13" pageNumber="499">Fig. 36</figureCitation>
. Aedeagus as in
<figureCitation id="10B22A79FFA994766643FD01FCE6FD61" box="[636,772,680,706]" captionStart="FIGURES 3742" captionStartId="12.[151,264,1810,1834]" captionTargetBox="[166,1417,195,1777]" captionTargetId="figure-15@12.[151,1436,181,1786]" captionTargetPageId="12" captionText="FIGURES 3742. Aedeagus of Paraphloeostiba (Figs. 3738: P. apicalis (Mindanao), Figs. 3940: P. obesa (Sabah), Figs. 4142: P. subopaca (Tamil Nadu), Figs. 4344: P. tonkinensis (Leyte)): 37, 39, 41, 43—parameral view, 38, 40, 42, 44—lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8036129" httpUri="https://zenodo.org/record/8036129/files/figure.png" pageId="13" pageNumber="499">Figs. 4142</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
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