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<document id="797F84F2574E0203B08C30D61707892F" ID-DOI="10.11646/zootaxa.5301.4.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="8036117" ID-ZooBank="01F6333F-807B-448A-80C8-EC1F8CE030EA" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1686723262927" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2023" docId="002087EAFFA3947364A8FB36FA85FBB2" docLanguage="en" docName="zootaxa.5301.4.6.pdf" docOrigin="Zootaxa 5301 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5301.4.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paraphloeostiba papuana" docType="treatment" docVersion="2" lastPageNumber="494" masterDocId="FC19FF92FFA4947B643FFFA9FFE2FFA3" masterDocTitle="New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines" masterLastPageNumber="500" masterPageNumber="486" pageNumber="493" updateTime="1686723785076" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3CE63805768C05C088E9E967F3B00FF6">New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines</mods:title>
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<treatment id="002087EAFFA3947364A8FB36FA85FBB2" LSID="urn:lsid:plazi:treatment:002087EAFFA3947364A8FB36FA85FBB2" httpUri="http://treatment.plazi.org/id/002087EAFFA3947364A8FB36FA85FBB2" lastPageId="8" lastPageNumber="494" pageId="7" pageNumber="493">
<subSubSection id="C0936577FFA3947C64A8FB36FD44FB19" box="[151,678,1183,1210]" pageId="7" pageNumber="493" type="nomenclature">
<paragraph id="883636FCFFA3947C64A8FB36FD44FB19" blockId="7.[151,678,1183,1247]" box="[151,678,1183,1210]" pageId="7" pageNumber="493">
<heading id="D37E8190FFA3947C64A8FB36FD44FB19" bold="true" box="[151,678,1183,1210]" fontSize="11" level="1" pageId="7" pageNumber="493" reason="1">
<taxonomicName id="4F894D7FFFA3947C64A8FB36FD44FB19" ID-CoL="762XP" authority="(Cameron, 1931)" baseAuthorityName="Cameron" baseAuthorityYear="1931" box="[151,678,1183,1210]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="493" phylum="Arthropoda" rank="species" species="papuana">
<emphasis id="BAFDEAEEFFA3947C64A8FB36FD44FB19" bold="true" box="[151,678,1183,1210]" pageId="7" pageNumber="493">
<emphasis id="BAFDEAEEFFA3947C64A8FB36FE2CFB19" bold="true" box="[151,462,1183,1210]" italics="true" pageId="7" pageNumber="493">Paraphloeostiba papuana</emphasis>
(
<bibRefCitation id="EC184B0DFFA3947C65E2FB09FD7CFB19" author="Cameron, M." box="[477,670,1184,1210]" pageId="7" pageNumber="493" pagination="355 - 364" refId="ref8275" refString="Cameron, M. (1931) Staphylinidae (Coleoptera) from New Guinea, in the South Australian Museum. Records of the South Australian Museum, 4 (3), 355 - 364." type="journal article" year="1931">Cameron, 1931</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C0936577FFA3947C64A8FB6DFEA8FB7C" box="[151,330,1220,1247]" pageId="7" pageNumber="493" type="description">
<paragraph id="883636FCFFA3947C64A8FB6DFEA8FB7C" blockId="7.[151,678,1183,1247]" box="[151,330,1220,1247]" pageId="7" pageNumber="493">
(
<figureCitation id="10B22A79FFA3947C64A0FB6DFF0DFB7C" box="[159,239,1220,1247]" captionStart="FIGURES 16" captionStartId="2.[151,275,1541,1567]" captionTargetBox="[197,1385,250,1508]" captionTargetId="figure-45@2.[187,1400,237,1520]" captionTargetPageId="2" captionText="FIGURES 16. Habitus of Paraphloeostiba: 1—P. conjuncta (holotype), 2—P. coriacea, 3—P. flaveola (holotype), 4—P. gayndahensis (Papua New Guinea), 5—P. papuana (holotype), 6—P. ullrichi (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8036119" httpUri="https://zenodo.org/record/8036119/files/figure.png" pageId="7" pageNumber="493">Figs. 5</figureCitation>
,
<figureCitation id="10B22A79FFA3947C64C4FB6DFEA1FB7C" box="[251,323,1220,1247]" captionStart="FIGURES 1525" captionStartId="6.[151,264,1579,1603]" captionTargetBox="[168,1422,200,1548]" captionTargetId="figure-15@6.[151,1436,181,1555]" captionTargetPageId="6" captionText="FIGURES 1525. Aedeagus and abdominal segments of Paraphloeostiba flaveola (Figs, 1521: holotype) and P. papuana (Figs. 2225): 15—aedeagus, parameral view, 16—aedeagus, (lateral view), 17—male tergite VIII, dorsal view, 18—male sternite VIII, ventral view, 19, 22—female tergite VIII, dorsal view, 20, 23—female sternite VIII, ventral view, 21, 24—female genital segment, ventral view, 25—spermatheca, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8036123" httpUri="https://zenodo.org/record/8036123/files/figure.png" pageId="7" pageNumber="493">2225</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C0936577FFA3947C64A8FAA1FDA9FAFD" pageId="7" pageNumber="493" type="reference_group">
<paragraph id="883636FCFFA3947C64A8FAA1FD82FA83" blockId="7.[151,808,1288,1374]" box="[151,608,1288,1313]" pageId="7" pageNumber="493">
<treatmentCitationGroup id="A89911D2FFA3947C64A8FAA1FD82FA83" box="[151,608,1288,1313]" pageId="7" pageNumber="493">
<treatmentCitation id="092810EDFFA3947C64A8FAA1FD82FA83" author="Cameron, M." box="[151,608,1288,1313]" page="356" pageId="7" pageNumber="493" year="1931">
<taxonomicName id="4F894D7FFFA3947C64A8FAA1FD82FA83" authority="Cameron, 1931: 356" authorityName="Cameron" authorityPageNumber="356" authorityYear="1931" box="[151,608,1288,1313]" class="Insecta" family="Staphylinidae" genus="Phloeonomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="493" phylum="Arthropoda" rank="species" species="papuanus">
<emphasis id="BAFDEAEEFFA3947C64A8FAA1FE68FA83" box="[151,394,1288,1312]" italics="true" pageId="7" pageNumber="493">Phloeonomus papuanus</emphasis>
<bibRefCitation id="EC184B0DFFA3947C65AFFAA0FD82FA83" author="Cameron, M." box="[400,608,1288,1313]" pageId="7" pageNumber="493" pagination="355 - 364" refId="ref8275" refString="Cameron, M. (1931) Staphylinidae (Coleoptera) from New Guinea, in the South Australian Museum. Records of the South Australian Museum, 4 (3), 355 - 364." type="journal article" year="1931">Cameron, 1931: 356</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="883636FCFFA3947C64A8FA8EFCCAFA9C" blockId="7.[151,808,1288,1374]" box="[151,808,1319,1343]" pageId="7" pageNumber="493">
<treatmentCitationGroup id="A89911D2FFA3947C64A8FA8EFCCAFA9C" box="[151,808,1319,1343]" pageId="7" pageNumber="493">
<treatmentCitation id="092810EDFFA3947C64A8FA8EFCCAFA9C" author="Scheerpeltz, O. von" box="[151,808,1319,1343]" page="1050" pageId="7" pageNumber="493" year="1933">
<taxonomicName id="4F894D7FFFA3947C64A8FA8EFCCAFA9C" authority=": Scheerpeltz 1933: 1050" authorityName="Scheerpeltz" authorityPageNumber="1050" authorityYear="1933" box="[151,808,1319,1343]" class="Insecta" family="Staphylinidae" genus="Phloeonomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="493" phylum="Arthropoda" rank="species" species="papuanus" subGenus="Phloeonomus">
<emphasis id="BAFDEAEEFFA3947C64A8FA8EFEC3FA9C" box="[151,289,1319,1343]" italics="true" pageId="7" pageNumber="493">Phloeonomus</emphasis>
(
<emphasis id="BAFDEAEEFFA3947C650FFA8EFE58FA9C" box="[304,442,1319,1343]" italics="true" pageId="7" pageNumber="493">Phloeonomus</emphasis>
)
<emphasis id="BAFDEAEEFFA3947C65F6FA81FDCEFA9C" box="[457,556,1320,1343]" italics="true" pageId="7" pageNumber="493">papuanus</emphasis>
:
<bibRefCitation id="EC184B0DFFA3947C6607FA8EFCCAFA9C" author="Scheerpeltz, O. von" box="[568,808,1319,1343]" pageId="7" pageNumber="493" pagination="989 - 1500" refId="ref8685" refString="Scheerpeltz, O. von (1933) Staphylinidae VII (pars 129). In: Schenkling S. (Ed.), Coleopterorum Catalogus. Vol. VI. Staphylinidae II. W. Junk, Berlin, pp. 989 - 1500." type="book chapter" year="1933">Scheerpeltz 1933: 1050</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="883636FCFFA3947C64A8FAEFFDA9FAFD" blockId="7.[151,808,1288,1374]" box="[151,587,1350,1374]" pageId="7" pageNumber="493">
<treatmentCitationGroup id="A89911D2FFA3947C64A8FAEFFDA9FAFD" box="[151,587,1350,1374]" pageId="7" pageNumber="493">
<treatmentCitation id="092810EDFFA3947C64A8FAEFFDA9FAFD" author="Steel, W. O." box="[151,587,1350,1374]" page="153" pageId="7" pageNumber="493" year="1960">
<taxonomicName id="4F894D7FFFA3947C64A8FAEFFDA9FAFD" ID-CoL="762XP" authority=": Steel 1960: 153" authorityName="Steel" authorityPageNumber="153" authorityYear="1960" box="[151,587,1350,1374]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="493" phylum="Arthropoda" rank="species" species="papuana">
<emphasis id="BAFDEAEEFFA3947C64A8FAEFFE7DFAFD" box="[151,415,1350,1374]" italics="true" pageId="7" pageNumber="493">Paraphloeostiba papuana</emphasis>
:
<bibRefCitation id="EC184B0DFFA3947C6593FAEFFDA9FAFD" author="Steel, W. O." box="[428,587,1350,1374]" pageId="7" pageNumber="493" pagination="141 - 172" refId="ref9011" refString="Steel, W. O. (1960) Three new omaliine genera from Asia and Australasia previously confused with Phloeonomus Thomson (Coleoptera: Staphylinidae). The Transactions of the Royal Entomological Society of London, 112, 141 - 172. https: // doi. org / 10.1111 / j. 1365 - 2311.1960. tb 00497. x" type="journal article" year="1960">Steel 1960: 153</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C0936577FFA3947C64A8FA25FEE7F9F9" pageId="7" pageNumber="493" type="materials_examined">
<paragraph id="883636FCFFA3947C64A8FA25FAC6F9B1" blockId="7.[151,1437,1420,2023]" pageId="7" pageNumber="493">
<emphasis id="BAFDEAEEFFA3947C64A8FA25FEA4FA05" bold="true" box="[151,326,1420,1446]" pageId="7" pageNumber="493">Type material.</emphasis>
<typeStatus id="5732885EFFA3947C6571FA25FE54FA05" box="[334,438,1420,1446]" pageId="7" pageNumber="493">Holotype</typeStatus>
of
<taxonomicName id="4F894D7FFFA3947C65E2FA25FC42FA05" authority="Cameron, 1931" authorityName="Cameron" authorityYear="1931" box="[477,928,1420,1446]" class="Insecta" family="Staphylinidae" genus="Phloeonomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="493" phylum="Arthropoda" rank="species" species="papuanus">
<emphasis id="BAFDEAEEFFA3947C65E2FA25FD09FA05" box="[477,747,1420,1446]" italics="true" pageId="7" pageNumber="493">Phloeonomus papuanus</emphasis>
<bibRefCitation id="EC184B0DFFA3947C66CDFA25FC42FA05" author="Cameron, M." box="[754,928,1420,1446]" pageId="7" pageNumber="493" pagination="355 - 364" refId="ref8275" refString="Cameron, M. (1931) Staphylinidae (Coleoptera) from New Guinea, in the South Australian Museum. Records of the South Australian Museum, 4 (3), 355 - 364." type="journal article" year="1931">Cameron, 1931</bibRefCitation>
</taxonomicName>
<specimenCount id="9E8FFD75FFA3947C6797FA24FC5FFA04" box="[936,957,1421,1447]" pageId="7" pageNumber="493" type="female"></specimenCount>
(
<figureCitation id="10B22A79FFA3947C67F2FA25FBF1FA04" box="[973,1043,1420,1447]" captionStart="FIGURES 16" captionStartId="2.[151,275,1541,1567]" captionTargetBox="[197,1385,250,1508]" captionTargetId="figure-45@2.[187,1400,237,1520]" captionTargetPageId="2" captionText="FIGURES 16. Habitus of Paraphloeostiba: 1—P. conjuncta (holotype), 2—P. coriacea, 3—P. flaveola (holotype), 4—P. gayndahensis (Papua New Guinea), 5—P. papuana (holotype), 6—P. ullrichi (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8036119" httpUri="https://zenodo.org/record/8036119/files/figure.png" pageId="7" pageNumber="493">Fig. 5</figureCitation>
): Type` &lt;round printed label with red margin&gt;, Mt. Lamington | N. E.
<collectingRegion id="4A4DF81EFFA3947C6672FA18FD73FA68" box="[589,657,1457,1483]" country="Indonesia" name="Papua Barat" pageId="7" pageNumber="493">Papua</collectingRegion>
|
<quantity id="4F719B19FFA3947C6693FA19FC99FA69" box="[684,891,1456,1483]" metricMagnitude="2" metricUnit="m" metricValue="4.2672" metricValueMax="4.572" metricValueMin="3.9624" pageId="7" pageNumber="493" unit="ft" value="1400.0" valueMax="1500.0" valueMin="1300.0">1300 to 1500 feet</quantity>
| C. T. McNamara` &lt;printed&gt;,
<taxonomicName id="4F894D7FFFA3947C60D0FA19FA6FFA69" authorityName="Heer" authorityYear="1839" box="[1263,1421,1456,1482]" class="Insecta" family="Staphylinidae" genus="Phloeonomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="493" phylum="Arthropoda" rank="genus">Phloeonomus</taxonomicName>
| papuanus | TYPE [red]
<collectingRegion id="4A4DF81EFFA3947C6596FA7DFE06FA4D" box="[425,484,1492,1518]" country="Myanmar" name="Chin" pageId="7" pageNumber="493">Cam</collectingRegion>
.`&lt;handwritten in black&gt;, NHMUK014663272 &lt;printed label with barcode on the right side&gt;,
<taxonomicName id="4F894D7FFFA3947C6518FA51FE00F9B1" authorityName="Steel" authorityYear="1960" box="[295,482,1528,1554]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="493" phylum="Arthropoda" rank="genus">
<emphasis id="BAFDEAEEFFA3947C6518FA51FE00F9B1" box="[295,482,1528,1554]" italics="true" pageId="7" pageNumber="493">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="4F894D7FFFA3947C65C9FA50FCFCF9B0" authority="(Cameron, 1952)" baseAuthorityName="Cameron" baseAuthorityYear="1952" box="[502,798,1528,1555]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="493" phylum="Arthropoda" rank="species" species="papuana">
<emphasis id="BAFDEAEEFFA3947C65C9FA50FDBAF9B1" box="[502,600,1529,1554]" italics="true" pageId="7" pageNumber="493">papuana</emphasis>
(
<bibRefCitation id="EC184B0DFFA3947C6659FA51FCF4F9B0" author="Cameron, M." box="[614,790,1528,1555]" pageId="7" pageNumber="493" pagination="241 - 256" refId="ref8340" refString="Cameron, M. (1952) Results of the Archibold Expeditions. New species of Staphylinidae (Col.) from New Guinea and Misool. Treubia, 21 (2), 241 - 256." type="journal article" year="1952">Cameron, 1952</bibRefCitation>
)
</taxonomicName>
| Shavrin A.V. det. 2023`&lt;printed&gt; (
<collectionCode id="EE98AE39FFA3947C6081FA51FAFAF9B1" box="[1214,1304,1528,1554]" pageId="7" pageNumber="493">BMNH</collectionCode>
).
</paragraph>
<paragraph id="883636FCFFA3947C64F8F9B5FEE7F9F9" blockId="7.[151,1437,1420,2023]" pageId="7" pageNumber="493">
<emphasis id="BAFDEAEEFFA3947C64F8F9B5FDD1F995" bold="true" box="[199,563,1564,1590]" pageId="7" pageNumber="493">Additional material examined.</emphasis>
<materialsCitation id="38E13CA1FFA3947C6605F9B5FEE3F9F9" collectionCode="BMNH" country="Papua New Guinea" location="PAPUA NEW GUINEA" pageId="7" pageNumber="493" specimenCount="1" specimenCount-female="1">
<emphasis id="BAFDEAEEFFA3947C6605F9B5FCBCF995" bold="true" box="[570,862,1564,1590]" pageId="7" pageNumber="493">
<collectingCountry id="F09E766CFFA3947C6605F9B5FCBBF995" box="[570,857,1564,1590]" name="Papua New Guinea" pageId="7" pageNumber="493">PAPUA NEW GUINEA</collectingCountry>
:
</emphasis>
<specimenCount id="9E8FFD75FFA3947C675AF9B5FC72F994" box="[869,912,1564,1591]" pageId="7" pageNumber="493" type="female">1 ♀</specimenCount>
: Finschhafen, New Guinea 1.4/[19]44 Helfer` (
<collectionCode id="EE98AE39FFA3947C64A0F9E9FF1BF9F9" box="[159,249,1600,1626]" pageId="7" pageNumber="493">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C0936577FFA3947364F8F9CDFBF9FC22" lastPageId="8" lastPageNumber="494" pageId="7" pageNumber="493" type="description">
<paragraph id="883636FCFFA3947C64F8F9CDFB16F965" blockId="7.[151,1437,1420,2023]" pageId="7" pageNumber="493">
<emphasis id="BAFDEAEEFFA3947C64F8F9CDFE97F9DD" bold="true" box="[199,373,1636,1662]" pageId="7" pageNumber="493">Redescription.</emphasis>
Measurements (n=2): HW: 0.380.42; HL: 0.200.22; OL: 0.15; TL: 0.02; AL (
<typeStatus id="5732885EFFA3947C60D6F9CDFAB0F9DD" box="[1257,1362,1636,1662]" pageId="7" pageNumber="493">holotype</typeStatus>
): 0.46; PL: 0.280.30; PWmax: 0.520.55; PWmin: 0.420.44; ESL: 0.470.52; EW: 0.600.62; MTbL (
<typeStatus id="5732885EFFA3947C60D8F921FAB2F901" box="[1255,1360,1672,1698]" pageId="7" pageNumber="493">holotype</typeStatus>
): 0.29; MTrL (
<typeStatus id="5732885EFFA3947C64D7F905FEB3F965" box="[232,337,1708,1734]" pageId="7" pageNumber="493">holotype</typeStatus>
): 0.15 (MTrL 14: 0.05; MTrL 5: 0.10); AW: 0.650.67; BL: 1.801.85 (
<typeStatus id="5732885EFFA3947C6040F905FB0AF965" box="[1151,1256,1708,1734]" pageId="7" pageNumber="493">holotype</typeStatus>
).
</paragraph>
<paragraph id="883636FCFFA3947364F8F979FEBEFF76" blockId="7.[151,1437,1420,2023]" lastBlockId="8.[151,1437,151,1041]" lastPageId="8" lastPageNumber="494" pageId="7" pageNumber="493">
Habitus as in
<figureCitation id="10B22A79FFA3947C6551F979FE54F948" box="[366,438,1744,1771]" captionStart="FIGURES 16" captionStartId="2.[151,275,1541,1567]" captionTargetBox="[197,1385,250,1508]" captionTargetId="figure-45@2.[187,1400,237,1520]" captionTargetPageId="2" captionText="FIGURES 16. Habitus of Paraphloeostiba: 1—P. conjuncta (holotype), 2—P. coriacea, 3—P. flaveola (holotype), 4—P. gayndahensis (Papua New Guinea), 5—P. papuana (holotype), 6—P. ullrichi (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8036119" httpUri="https://zenodo.org/record/8036119/files/figure.png" pageId="7" pageNumber="493">Fig. 5</figureCitation>
. Body yellow-brown, with darker lateral portions of head and abdomen in
<typeStatus id="5732885EFFA3947C610CF979FA7BF949" box="[1331,1433,1744,1770]" pageId="7" pageNumber="493">holotype</typeStatus>
; mouthparts, antennae and legs yellow (antenomeres 711 of specimen from Finschhafen slightly darker). Head with indistinct and very fine punctation, denser in middle; neck without punctures; pronotum with very dense and fine punctation, without distinct punctures in middle; scutellum without punctures; punctation of elytra dense, slightly larger and deeper than that on pronotum, finer along suture and in apical portions of each elytron; abdomen without visible punctures on abdominal tergites IVVI, with very indistinct, fine and sparse punctation on tergites VIIVIII. Head with coarse microscupture; neck with distinct transverse meshes; pronotum with very dense microreticulation, slightly coarser than that in middle portion of head; microsculpture of elytra dense, slightly more transverse than that on pronotum; abdomen with dense sculpture, slightly transverse on abdominal tergites IVVI and transverse on tergites VIIVIII.
</paragraph>
<paragraph id="883636FCFFAC947364F8FF76FE91FE72" blockId="8.[151,1437,151,1041]" pageId="8" pageNumber="494">Head with wide and moderately shallow postantennal depression, 1.9 times as broad as long, with very short (about as long as diameter of ocellus) and deep anteocellar foveae; postocular carina indistinct, acute, distance between posterior margin of eye and carina about as long as one nearest ommatidium. Apical maxillary palpomere sligthly more than twice as long as preapical segment. Antennomere 6 slightly and antennomeres 710 strongly transverse; antennomere 2 slightly narrower than basal antennomere, 3 slightly longer than 2, 4 slightly shorter than 4, 6 distinctly broader than 5, 79 significantly broader than 6, 10 slightly longer than 9, apical antennomere about twice as long as 10.</paragraph>
<paragraph id="883636FCFFAC947364F8FE72FC6EFD9E" blockId="8.[151,1437,151,1041]" pageId="8" pageNumber="494">Pronotum 1.8 times as broad as long, 1.3 times as broad as head, widest in middle, slightly more narrowed posteriad than anteriad; posterior angles obtuse; anterior margin slightly concave in middle; middle portion without paramedian impressions; laterobasal impressions wide and shallow.</paragraph>
<paragraph id="883636FCFFAC947364F8FDEEFE4EFD26" blockId="8.[151,1437,151,1041]" pageId="8" pageNumber="494">Elytra slightly convex, 1.11.2 times as broad as long, 1.61.7 times as long as pronotum slightly more narrowed posteriad than annteriad.</paragraph>
<paragraph id="883636FCFFAC947364F8FD26FD91FD0A" blockId="8.[151,1437,151,1041]" box="[199,627,655,681]" pageId="8" pageNumber="494">Abdomen slightly broader than elytra.</paragraph>
<paragraph id="883636FCFFAC947364F8FD1AFE9AFD6E" blockId="8.[151,1437,151,1041]" box="[199,376,691,717]" pageId="8" pageNumber="494">Male unknown.</paragraph>
<paragraph id="883636FCFFAC947364F8FD7EFB6AFCB6" blockId="8.[151,1437,151,1041]" pageId="8" pageNumber="494">
Female. Posterior margin of abdominal tergite VIII truncate (
<figureCitation id="10B22A79FFAC94736742FD7EFC31FD52" box="[893,979,727,753]" captionStart="FIGURES 1525" captionStartId="6.[151,264,1579,1603]" captionTargetBox="[168,1422,200,1548]" captionTargetId="figure-15@6.[151,1436,181,1555]" captionTargetPageId="6" captionText="FIGURES 1525. Aedeagus and abdominal segments of Paraphloeostiba flaveola (Figs, 1521: holotype) and P. papuana (Figs. 2225): 15—aedeagus, parameral view, 16—aedeagus, (lateral view), 17—male tergite VIII, dorsal view, 18—male sternite VIII, ventral view, 19, 22—female tergite VIII, dorsal view, 20, 23—female sternite VIII, ventral view, 21, 24—female genital segment, ventral view, 25—spermatheca, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8036123" httpUri="https://zenodo.org/record/8036123/files/figure.png" pageId="8" pageNumber="494">Fig. 22</figureCitation>
). Posterior margin of abdominal sternite VIII rounded (
<figureCitation id="10B22A79FFAC94736506FD52FE6CFCB6" box="[313,398,763,789]" captionStart="FIGURES 1525" captionStartId="6.[151,264,1579,1603]" captionTargetBox="[168,1422,200,1548]" captionTargetId="figure-15@6.[151,1436,181,1555]" captionTargetPageId="6" captionText="FIGURES 1525. Aedeagus and abdominal segments of Paraphloeostiba flaveola (Figs, 1521: holotype) and P. papuana (Figs. 2225): 15—aedeagus, parameral view, 16—aedeagus, (lateral view), 17—male tergite VIII, dorsal view, 18—male sternite VIII, ventral view, 19, 22—female tergite VIII, dorsal view, 20, 23—female sternite VIII, ventral view, 21, 24—female genital segment, ventral view, 25—spermatheca, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8036123" httpUri="https://zenodo.org/record/8036123/files/figure.png" pageId="8" pageNumber="494">Fig. 23</figureCitation>
). Female accessory sclerite as in
<figureCitation id="10B22A79FFAC9473673EFD52FCB6FCB6" box="[769,852,763,789]" captionStart="FIGURES 1525" captionStartId="6.[151,264,1579,1603]" captionTargetBox="[168,1422,200,1548]" captionTargetId="figure-15@6.[151,1436,181,1555]" captionTargetPageId="6" captionText="FIGURES 1525. Aedeagus and abdominal segments of Paraphloeostiba flaveola (Figs, 1521: holotype) and P. papuana (Figs. 2225): 15—aedeagus, parameral view, 16—aedeagus, (lateral view), 17—male tergite VIII, dorsal view, 18—male sternite VIII, ventral view, 19, 22—female tergite VIII, dorsal view, 20, 23—female sternite VIII, ventral view, 21, 24—female genital segment, ventral view, 25—spermatheca, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8036123" httpUri="https://zenodo.org/record/8036123/files/figure.png" pageId="8" pageNumber="494">Fig. 24</figureCitation>
. Spermatheca as in
<figureCitation id="10B22A79FFAC9473600EFD52FB66FCB6" box="[1073,1156,762,789]" captionStart="FIGURES 1525" captionStartId="6.[151,264,1579,1603]" captionTargetBox="[168,1422,200,1548]" captionTargetId="figure-15@6.[151,1436,181,1555]" captionTargetPageId="6" captionText="FIGURES 1525. Aedeagus and abdominal segments of Paraphloeostiba flaveola (Figs, 1521: holotype) and P. papuana (Figs. 2225): 15—aedeagus, parameral view, 16—aedeagus, (lateral view), 17—male tergite VIII, dorsal view, 18—male sternite VIII, ventral view, 19, 22—female tergite VIII, dorsal view, 20, 23—female sternite VIII, ventral view, 21, 24—female genital segment, ventral view, 25—spermatheca, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8036123" httpUri="https://zenodo.org/record/8036123/files/figure.png" pageId="8" pageNumber="494">Fig. 25</figureCitation>
.
</paragraph>
<paragraph id="883636FCFFAC947364F8FCB7FBF9FC22" blockId="8.[151,1437,151,1041]" pageId="8" pageNumber="494">
<emphasis id="BAFDEAEEFFAC947364F8FCB7FE53FC9A" bold="true" box="[199,433,798,825]" pageId="8" pageNumber="494">Comparative notes.</emphasis>
Based on the slightly convex elytra, the shape of the pronotum
<taxonomicName id="4F894D7FFFAC947360B9FCB6FAEAFC9B" baseAuthorityName="Cameron" baseAuthorityYear="1952" box="[1158,1288,799,825]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="papuana">
<emphasis id="BAFDEAEEFFAC947360B9FCB6FB75FC9B" box="[1158,1175,799,824]" italics="true" pageId="8" pageNumber="494">P</emphasis>
.
<emphasis id="BAFDEAEEFFAC94736099FCB6FAEAFC9B" box="[1190,1288,799,824]" italics="true" pageId="8" pageNumber="494">papuana</emphasis>
</taxonomicName>
is somewhat similar to
<taxonomicName id="4F894D7FFFAC94736532FCEAFE8DFCFE" box="[269,367,835,861]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="obesa">
<emphasis id="BAFDEAEEFFAC94736532FCEAFEFCFCFF" box="[269,286,835,860]" italics="true" pageId="8" pageNumber="494">P</emphasis>
.
<emphasis id="BAFDEAEEFFAC94736511FCEAFE8DFCFE" box="[302,367,835,861]" italics="true" pageId="8" pageNumber="494">obesa</emphasis>
</taxonomicName>
(see below), from which it can be distinguished by the smaller and slightly paler body, finer punctation of the pronotum, and different shape of the female accessory sclerite.
</paragraph>
</subSubSection>
<subSubSection id="C0936577FFAC947364F8FC23FA9FFC06" box="[199,1405,906,933]" pageId="8" pageNumber="494" type="distribution">
<paragraph id="883636FCFFAC947364F8FC23FA9FFC06" blockId="8.[151,1437,151,1041]" box="[199,1405,906,933]" pageId="8" pageNumber="494">
<emphasis id="BAFDEAEEFFAC947364F8FC23FE83FC07" bold="true" box="[199,353,906,932]" pageId="8" pageNumber="494">Distribution.</emphasis>
<taxonomicName id="4F894D7FFFAC94736558FC22FD69FC07" baseAuthorityName="Cameron" baseAuthorityYear="1952" box="[359,651,907,933]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="papuana">
<emphasis id="BAFDEAEEFFAC94736558FC22FD69FC07" box="[359,651,907,933]" italics="true" pageId="8" pageNumber="494">Paraphloeostiba papuana</emphasis>
</taxonomicName>
is known from two localities in eastern part of
<collectingCountry id="F09E766CFFAC947360A2FC22FA98FC06" box="[1181,1402,907,933]" name="Papua New Guinea" pageId="8" pageNumber="494">Papua New Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C0936577FFAC947364F8FC06FA85FBB2" pageId="8" pageNumber="494" type="discussion">
<paragraph id="883636FCFFAC947364F8FC06FA85FBB2" blockId="8.[151,1437,151,1041]" pageId="8" pageNumber="494">
<emphasis id="BAFDEAEEFFAC947364F8FC06FED9FC6A" bold="true" box="[199,315,943,969]" pageId="8" pageNumber="494">Remarks.</emphasis>
<taxonomicName id="4F894D7FFFAC9473657AFC06FDB6FC6B" authorityName="Cameron" authorityYear="1931" box="[325,596,943,969]" class="Insecta" family="Staphylinidae" genus="Phloeonomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="papuanus">
<emphasis id="BAFDEAEEFFAC9473657AFC06FDB6FC6B" box="[325,596,943,969]" italics="true" pageId="8" pageNumber="494">Phloeonomus papuanus</emphasis>
</taxonomicName>
was originally described based on the female from “Mount Lamington”, collected by C.T. McNamara.
<bibRefCitation id="EC184B0DFFAC947365D7FC7AFD90FC4E" author="Steel, W. O." box="[488,626,979,1005]" pageId="8" pageNumber="494" pagination="141 - 172" refId="ref9011" refString="Steel, W. O. (1960) Three new omaliine genera from Asia and Australasia previously confused with Phloeonomus Thomson (Coleoptera: Staphylinidae). The Transactions of the Royal Entomological Society of London, 112, 141 - 172. https: // doi. org / 10.1111 / j. 1365 - 2311.1960. tb 00497. x" type="journal article" year="1960">Steel (1960)</bibRefCitation>
redescribed it and transferred to the genus
<taxonomicName id="4F894D7FFFAC9473606BFC7AFAEDFC4E" authorityName="Steel" authorityYear="1960" box="[1108,1295,979,1005]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="494" phylum="Arthropoda" rank="genus">
<emphasis id="BAFDEAEEFFAC9473606BFC7AFAEDFC4E" box="[1108,1295,979,1005]" italics="true" pageId="8" pageNumber="494">Paraphloeostiba</emphasis>
</taxonomicName>
. I re-studied the female which was previously studied by Steel. The spermatheca of
<taxonomicName id="4F894D7FFFAC9473678CFC5EFBD2FBB3" baseAuthorityName="Cameron" baseAuthorityYear="1952" box="[947,1072,1015,1040]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="papuana">
<emphasis id="BAFDEAEEFFAC9473678CFC5EFBD2FBB3" box="[947,1072,1015,1040]" italics="true" pageId="8" pageNumber="494">P. papuana</emphasis>
</taxonomicName>
is figured for the first time.
</paragraph>
</subSubSection>
</treatment>
</document>