362 lines
44 KiB
XML
362 lines
44 KiB
XML
<document id="AB73F1322BF51B7EF43EBA1E53AB70E2" ID-DOI="10.11646/zootaxa.3999.1.7" ID-GBIF-Dataset="12d6ad4d-eb31-4cc5-811e-c9dd1879ff99" ID-ISSN="1175-5326" ID-Zenodo-Dep="243357" ID-ZooBank="60B5AB6B-C586-41E5-9370-80BF56025D1F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461280129055" checkinUser="plazi" docAuthor="Zamotajlov, Alexandr S., Solodovnikov, Igor A. & Fominykh, Dmitriy D." docDate="2015" docId="03F28B45FFA2FFBCFF555FBD933CF889" docLanguage="en" docName="zt03999p124.pdf" docOrigin="Zootaxa 3999 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pterostichus (Aphaonus) svetlanae Zamotajlov, Solodovnikov & Fominykh, 2015, sp. nov." docType="treatment" docVersion="8" lastPageNumber="119" masterDocId="FFCBF33DFFA7FFB4FFC25E359157FF8C" masterDocTitle="Description of two new species related to Pterostichus (Aphaonus) arcanoides Lorenz, 1998 from the West Transcaucasia (Coleoptera: Carabidae: Pterostichini)" masterLastPageNumber="124" masterPageNumber="111" pageNumber="116" updateTime="1698613781523" updateUser="plazi">
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<mods:mods id="12FFDF8160189FBE5977DA2DD06EF201" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="62E28557FC437DA331C550DE80FDF597">Description of two new species related to Pterostichus (Aphaonus) arcanoides Lorenz, 1998 from the West Transcaucasia (Coleoptera: Carabidae: Pterostichini)</mods:title>
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<mods:roleTerm id="06D2FB7C16D276831D3720BD4FB3FC3D">Author</mods:roleTerm>
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<mods:namePart id="3BE9FBEC08D15A7DB0BB3E6BA2AD673D">Zamotajlov, Alexandr S.</mods:namePart>
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<mods:namePart id="15668D4DFC0FE8037CAC75DC0BD648E3">Solodovnikov, Igor A.</mods:namePart>
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<mods:roleTerm id="9AD2C48B1E8B44B1D586D8CA41EF6E8E">Author</mods:roleTerm>
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<mods:namePart id="67E2F8370A8A01274A4BBF422F6EBAC5">Fominykh, Dmitriy D.</mods:namePart>
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<mods:title id="C6509556E46B85C3E8A468FB4DA68331">Zootaxa</mods:title>
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<mods:part id="0E29CD12BB1C43E03BD6FDFCB9221787">
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<mods:date id="96D4CEFEC354C9E2CBA665EBF39D87AF">2015</mods:date>
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<mods:number id="8338662A17AB494BCEE522B57FBDF656">3999</mods:number>
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<mods:classification id="31BA56401BA30C6A8A56F023FC3AB4D7">journal article</mods:classification>
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<mods:identifier id="3C7104D4CDBEE869A0A3B4EB943EFEB5" type="DOI">10.11646/zootaxa.3999.1.7</mods:identifier>
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<treatment id="03F28B45FFA2FFBCFF555FBD933CF889" ID-DOI="http://doi.org/10.5281/zenodo.6114398" ID-GBIF-Taxon="119640448" ID-Zenodo-Dep="6114398" LSID="urn:lsid:plazi:treatment:03F28B45FFA2FFBCFF555FBD933CF889" httpUri="http://treatment.plazi.org/id/03F28B45FFA2FFBCFF555FBD933CF889" lastPageId="8" lastPageNumber="119" pageId="5" pageNumber="116">
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<subSubSection id="C34169D8FFA2FFB1FF555FBD901EFE48" pageId="5" pageNumber="116" type="nomenclature">
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<paragraph id="8BE43A53FFA2FFB1FF555FBD93C8FE2F" blockId="5.[151,671,392,453]" box="[151,671,392,419]" pageId="5" pageNumber="116">
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<heading id="D0AC8D3FFFA2FFB1FF555FBD93C8FE2F" bold="true" box="[151,671,392,419]" fontSize="11" level="1" pageId="5" pageNumber="116" reason="1">
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<emphasis id="B92FE641FFA2FFB1FF555FBD93C8FE2F" bold="true" box="[151,671,392,419]" pageId="5" pageNumber="116">
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<taxonomicName id="4C5B41D0FFA2FFB1FF555FBD936CFE2E" ID-CoL="8MGM2" box="[151,571,392,419]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="116" phylum="Arthropoda" rank="species" species="svetlanae" status="sp. nov." subGenus="Aphaonus">
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<emphasis id="B92FE641FFA2FFB1FF555FBD907AFE2E" bold="true" box="[151,301,392,418]" italics="true" pageId="5" pageNumber="116">Pterostichus</emphasis>
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(
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<emphasis id="B92FE641FFA2FFB1FEFF5FBD90EEFE2E" bold="true" box="[317,441,392,418]" italics="true" pageId="5" pageNumber="116">Aphaonus</emphasis>
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)
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<emphasis id="B92FE641FFA2FFB1FE095FBD936CFE2E" bold="true" box="[459,571,392,418]" italics="true" pageId="5" pageNumber="116">svetlanae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21C5B3AFFA2FFB1FD805FBC93C8FE2F" box="[578,671,393,419]" pageId="5" pageNumber="116" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BE43A53FFA2FFB1FF555F99901EFE48" blockId="5.[151,671,392,453]" box="[151,329,428,453]" pageId="5" pageNumber="116">
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(
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<figureCitation id="136026D6FFA2FFB1FF5D5F9991B9FE49" box="[159,238,428,453]" captionStart="FIGURE 6" captionStartId="5.[151,250,2000,2022]" captionTargetBox="[513,1436,572,1970]" captionTargetId="figure@5.[502,1085,631,1979]" captionTargetPageId="5" captionText="FIGURE 6. Pterostichus svetlanae sp. nov., dorsal habitus, holotype, female. Scale bar 1.0 mm." httpUri="https://zenodo.org/record/243362/files/figure.png" pageId="5" pageNumber="116">Figs. 6</figureCitation>
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,
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<figureCitation id="136026D6FFA2FFB1FF385F99904FFE48" box="[250,280,428,452]" captionStart="FIGURES 10 – 14" captionStartId="7.[151,264,1460,1483]" captionTargetBox="[169,1412,218,1409]" captionTargetId="figure@7.[151,1436,193,1440]" captionTargetPageId="7" captionText="FIGURES 10 – 14. Pterostichus spp., mentum, dorsal view: 10 — Pterostichus mariamae sp. nov., holotype, male; 11 — Pterostichus mariamae sp. nov., paratype, female; 12 — Pterostichus arcanoides Lorenz, male, Chedym Mt. Range; 13 — Pterostichus arcanoides Lorenz, female, Chedym Mt. Range; 14 — Pterostichus svetlanae sp. nov., holotype, female. Scale bar 1.0 mm." httpUri="https://zenodo.org/record/243364/files/figure.png" pageId="5" pageNumber="116">14</figureCitation>
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,
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<figureCitation id="136026D6FFA2FFB1FEE75F999015FE48" box="[293,322,428,452]" captionStart="FIGURE 23" captionStartId="11.[151,250,938,960]" captionTargetBox="[282,1307,202,913]" captionTargetId="figure@11.[275,1312,193,916]" captionTargetPageId="11" captionText="FIGURE 23. Female head comparative proportions of Pterostichus- species: left— Pt. svetlanae sp. nov.; middle— Pt. mariamae sp. nov.; right— Pt. arcanoides Lorenz." httpUri="https://zenodo.org/record/243367/files/figure.png" pageId="5" pageNumber="116">23</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<paragraph id="8BE43A53FFA2FFB1FF555FC190FCFDD8" pageId="5" pageNumber="116">
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<table id="F95BC8F3FFA2004BFF555FC194CBFDD9" box="[151,1436,500,597]" gridcols="3" gridrows="3" pageId="5" pageNumber="116">
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<tr id="356B3811FFA2004BFF555FC194CBFD81" box="[151,1436,500,525]" gridrow="0" pageId="5" pageNumber="116">
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<th id="76BA516DFFA2004BFF555FC193A0FD81" box="[151,759,500,525]" gridcol="0" gridrow="0" pageId="5" pageNumber="116">
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<emphasis id="B92FE641FFA2FFB1FF555FC19012FD81" bold="true" box="[151,325,500,525]" pageId="5" pageNumber="116">Type material.</emphasis>
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Holotype: female, labelled: Abkhazia,
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</th>
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<th id="76BA516DFFA2004BFD3C5FC19429FD81" box="[766,1406,500,525]" gridcol="1" gridrow="0" pageId="5" pageNumber="116">Gudauta Distr., vill. Mzara (= Chiryuta) vicinities, valley</th>
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<th id="76BA516DFFA2004BFA475FC194CBFD81" box="[1413,1436,500,525]" gridcol="2" gridrow="0" pageId="5" pageNumber="116">of</th>
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</tr>
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<tr id="356B3811FFA2004BFF555C2294CBFDBC" box="[151,1436,535,560]" gridrow="1" pageId="5" pageNumber="116">
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<th id="76BA516DFFA2004BFF555C2293A0FDBC" box="[151,759,535,560]" gridcol="0" gridrow="1" pageId="5" pageNumber="116">River Mzara, Zashirbara Mt. Range, N 43º10'18" /</th>
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<td id="76BA516DFFA2004BFD3C5C229429FDBC" box="[766,1406,535,560]" gridcol="1" gridrow="1" pageId="5" pageNumber="116">E 040º48'23", 705 m, 15.05.2014 (I. Solodovnikov &</td>
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<td id="76BA516DFFA2004BFA475C2294CBFDBC" box="[1413,1436,535,560]" gridcol="2" gridrow="1" pageId="5" pageNumber="116">S.</td>
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</tr>
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<tr id="356B3811FFA2004BFF555C0994CBFDD9" box="[151,1436,572,597]" gridrow="2" pageId="5" pageNumber="116" rowspan-1="1" rowspan-2="1">
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<th id="76BA516DFFA2004BFF555C0993A0FDD9" box="[151,759,572,597]" gridcol="0" gridrow="2" pageId="5" pageNumber="116">Solodovnikova) (CSIA).</th>
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</tr>
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</table>
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</paragraph>
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<caption id="DF246ADBFFA2FFB1FF5559E59538F86A" box="[151,1135,2000,2022]" httpUri="https://zenodo.org/record/243362/files/figure.png" pageId="5" pageNumber="116" targetBox="[513,1436,572,1970]" targetPageId="5">
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<paragraph id="8BE43A53FFA2FFB1FF5559E59538F86A" blockId="5.[151,1135,2000,2022]" box="[151,1135,2000,2022]" pageId="5" pageNumber="116">
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<emphasis id="B92FE641FFA2FFB1FF5559E59045F869" bold="true" box="[151,274,2000,2022]" pageId="5" pageNumber="116">FIGURE 6.</emphasis>
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<taxonomicName id="4C5B41D0FFA2FFB1FEDB59E590AAF86A" box="[281,509,2000,2022]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="116" phylum="Arthropoda" rank="species" species="svetlanae" status="sp. nov.">
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<emphasis id="B92FE641FFA2FFB1FEDB59E590AAF86A" box="[281,509,2000,2022]" italics="true" pageId="5" pageNumber="116">Pterostichus svetlanae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21C5B3AFFA2FFB1FDC159E49304F86A" box="[515,595,2001,2022]" pageId="5" pageNumber="116" rank="species">
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<emphasis id="B92FE641FFA2FFB1FDC159E49304F86A" bold="true" box="[515,595,2001,2022]" pageId="5" pageNumber="116">sp. nov.</emphasis>
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</taxonomicNameLabel>
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, dorsal habitus, holotype, female. Scale bar 1.0 mm.
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</paragraph>
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</caption>
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<caption id="DF246ADBFFA1FFB2FF55599595C6F85A" httpUri="https://zenodo.org/record/243363/files/figure.png" pageId="6" pageNumber="117" targetBox="[327,1261,213,1908]" targetPageId="6">
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<paragraph id="8BE43A53FFA1FFB2FF55599595C6F85A" blockId="6.[151,1436,1952,2006]" pageId="6" pageNumber="117">
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<emphasis id="B92FE641FFA1FFB2FF555995906DF83A" bold="true" box="[151,314,1952,1975]" pageId="6" pageNumber="117">FIGURES 7–9.</emphasis>
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<taxonomicName id="4C5B41D0FFA1FFB2FE83599493D1F83B" authority="Lorenz" authorityName="Lorenz" box="[321,646,1953,1975]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="117" phylum="Arthropoda" rank="species" species="arcanoides">
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<emphasis id="B92FE641FFA1FFB2FE8359949362F83B" box="[321,565,1953,1975]" italics="true" pageId="6" pageNumber="117">Pterostichus arcanoides</emphasis>
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Lorenz
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</taxonomicName>
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: 7–8—dorsal habitus: 7—male, Chedym Mt. Range; 8—female, Chedym Mt. Range; 9–right metafemur and metatrochanter apices, male, Chedym Mt. Range. Scale bars 1.0 mm.
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</paragraph>
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</caption>
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<caption id="DF246ADBFFA0FFB3FF555B8191BCF9AB" httpUri="https://zenodo.org/record/243364/files/figure.png" pageId="7" pageNumber="118" targetBox="[169,1412,218,1409]" targetPageId="7">
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<paragraph id="8BE43A53FFA0FFB3FF555B8191BCF9AB" blockId="7.[151,1436,1460,1575]" pageId="7" pageNumber="118">
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<emphasis id="B92FE641FFA0FFB3FF555B81900DFA46" bold="true" box="[151,346,1460,1483]" pageId="7" pageNumber="118">FIGURES 10–14.</emphasis>
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<taxonomicName id="4C5B41D0FFA0FFB3FEA45B8090B3FA47" box="[358,484,1461,1483]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="118" phylum="Arthropoda" rank="genus">
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<emphasis id="B92FE641FFA0FFB3FEA45B8090B3FA47" box="[358,484,1461,1483]" italics="true" pageId="7" pageNumber="118">Pterostichus</emphasis>
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</taxonomicName>
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spp., mentum, dorsal view: 10—
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<taxonomicName id="4C5B41D0FFA0FFB3FC915B809514FA47" box="[851,1091,1461,1483]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="118" phylum="Arthropoda" rank="species" species="mariamae" status="sp. nov.">
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<emphasis id="B92FE641FFA0FFB3FC915B809514FA47" box="[851,1091,1461,1483]" italics="true" pageId="7" pageNumber="118">Pterostichus mariamae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21C5B3AFFA0FFB3FB925B8395F0FA47" box="[1104,1191,1462,1483]" pageId="7" pageNumber="118" rank="species">
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<emphasis id="B92FE641FFA0FFB3FB925B8395F0FA47" bold="true" box="[1104,1191,1462,1483]" pageId="7" pageNumber="118">sp. nov.</emphasis>
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</taxonomicNameLabel>
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, holotype, male; 11—
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<taxonomicName id="4C5B41D0FFA0FFB3FF555BE190D2FA66" box="[151,389,1492,1514]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="118" phylum="Arthropoda" rank="species" species="mariamae" status="sp. nov.">
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<emphasis id="B92FE641FFA0FFB3FF555BE190D2FA66" box="[151,389,1492,1514]" italics="true" pageId="7" pageNumber="118">Pterostichus mariamae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21C5B3AFFA0FFB3FE535BE190B1FA65" box="[401,486,1492,1513]" pageId="7" pageNumber="118" rank="species">
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<emphasis id="B92FE641FFA0FFB3FE535BE190B1FA65" bold="true" box="[401,486,1492,1513]" pageId="7" pageNumber="118">sp. nov.</emphasis>
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</taxonomicNameLabel>
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, paratype, female; 12—
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<taxonomicName id="4C5B41D0FFA0FFB3FD2B5BE19561FA66" authority="Lorenz" authorityName="Lorenz" box="[745,1078,1492,1514]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="118" phylum="Arthropoda" rank="species" species="arcanoides">
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<emphasis id="B92FE641FFA0FFB3FD2B5BE192B6FA66" box="[745,993,1492,1514]" italics="true" pageId="7" pageNumber="118">Pterostichus arcanoides</emphasis>
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Lorenz
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</taxonomicName>
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, male, Chedym Mt. Range; 13—
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<taxonomicName id="4C5B41D0FFA0FFB3FF555BC7908CF984" authority="Lorenz" authorityName="Lorenz" box="[151,475,1522,1544]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="118" phylum="Arthropoda" rank="species" species="arcanoides">
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<emphasis id="B92FE641FFA0FFB3FF555BC790DCF984" box="[151,395,1522,1544]" italics="true" pageId="7" pageNumber="118">Pterostichus arcanoides</emphasis>
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Lorenz
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</taxonomicName>
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, female, Chedym Mt. Range; 14—
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<taxonomicName id="4C5B41D0FFA0FFB3FCFD5BC79574F984" box="[831,1059,1522,1544]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="118" phylum="Arthropoda" rank="species" species="svetlanae" status="sp. nov.">
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<emphasis id="B92FE641FFA0FFB3FCFD5BC79574F984" box="[831,1059,1522,1544]" italics="true" pageId="7" pageNumber="118">Pterostichus svetlanae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21C5B3AFFA0FFB3FBEB5BC6952DF984" box="[1065,1146,1523,1544]" pageId="7" pageNumber="118" rank="species">
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<emphasis id="B92FE641FFA0FFB3FBEB5BC6952DF984" bold="true" box="[1065,1146,1523,1544]" pageId="7" pageNumber="118">sp. nov.</emphasis>
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</taxonomicNameLabel>
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, holotype, female. Scale bar 1.0 mm.
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</paragraph>
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</caption>
|
||
<subSubSection id="C34169D8FFA0FFBCFF05586192F3FAF4" lastPageId="8" lastPageNumber="119" pageId="7" pageNumber="118" type="description">
|
||
<paragraph id="8BE43A53FFA0FFB3FF05586195CBF91D" blockId="7.[151,1436,1620,2005]" pageId="7" pageNumber="118">
|
||
<emphasis id="B92FE641FFA0FFB3FF055861900EF9E1" bold="true" box="[199,345,1620,1645]" pageId="7" pageNumber="118">Description.</emphasis>
|
||
Habitus (
|
||
<figureCitation id="136026D6FFA0FFB3FE0B58619358F9E1" box="[457,527,1620,1645]" captionStart="FIGURE 6" captionStartId="5.[151,250,2000,2022]" captionTargetBox="[513,1436,572,1970]" captionTargetId="figure@5.[502,1085,631,1979]" captionTargetPageId="5" captionText="FIGURE 6. Pterostichus svetlanae sp. nov., dorsal habitus, holotype, female. Scale bar 1.0 mm." httpUri="https://zenodo.org/record/243362/files/figure.png" pageId="7" pageNumber="118">Fig. 6</figureCitation>
|
||
). Large and robust species, body length
|
||
<quantity id="4CA397B6FFA0FFB3FC235861951EF9E0" box="[993,1097,1620,1644]" metricMagnitude="-2" metricUnit="m" metricValue="2.34" pageId="7" pageNumber="118" unit="mm" value="23.4">23.4 mm</quantity>
|
||
, width 5.0 mm. Dorsum dark brown to black, shiny, underside dark brown to black, legs, antennae, and palpi dark brown.
|
||
</paragraph>
|
||
<paragraph id="8BE43A53FFA0FFBCFF0558A99437FF74" blockId="7.[151,1436,1620,2005]" lastBlockId="8.[151,1437,151,1797]" lastPageId="8" lastPageNumber="119" pageId="7" pageNumber="118">
|
||
Head broad, inflated, 1.03 times as long as wide and 0.97 times as wide as pronotum, nearly square (viewed dorsally), index 2 = 1.00: 0.90: 1.09, genae prominent, long, much stronger extended anteriorly than both in previous species and
|
||
<taxonomicName id="4C5B41D0FFA0FFB3FE4D58D0937EF970" box="[399,553,1764,1788]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="118" phylum="Arthropoda" rank="species" species="arcanoides">
|
||
<emphasis id="B92FE641FFA0FFB3FE4D58D0937EF970" box="[399,553,1764,1788]" italics="true" pageId="7" pageNumber="118">P. arcanoides</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="136026D6FFA0FFB3FDFE58D193C4F970" box="[572,659,1764,1789]" captionStart="FIGURE 23" captionStartId="11.[151,250,938,960]" captionTargetBox="[282,1307,202,913]" captionTargetId="figure@11.[275,1312,193,916]" captionTargetPageId="11" captionText="FIGURE 23. Female head comparative proportions of Pterostichus- species: left— Pt. svetlanae sp. nov.; middle— Pt. mariamae sp. nov.; right— Pt. arcanoides Lorenz." httpUri="https://zenodo.org/record/243367/files/figure.png" pageId="7" pageNumber="118">Fig. 23</figureCitation>
|
||
), compound eyes small, rudimentary, completely depressed, hardly developed, occupying somewhat dorso-lateral position, placed nearly equidistant between anterior angle of geanae and temples; temples long, tumid, more or less evenly rounded, resembling
|
||
<taxonomicName id="4C5B41D0FFA0FFB3FBC6591895C3F8C8" box="[1028,1172,1837,1860]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="118" phylum="Arthropoda" rank="species" species="mariamae" status="sp. nov.">
|
||
<emphasis id="B92FE641FFA0FFB3FBC6591895C3F8C8" box="[1028,1172,1837,1860]" italics="true" pageId="7" pageNumber="118">P. mariamae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21C5B3AFFA0FFB3FB5F591995AEF8C9" box="[1181,1273,1836,1861]" pageId="7" pageNumber="118" rank="species">
|
||
<emphasis id="B92FE641FFA0FFB3FB5F591995AEF8C9" bold="true" box="[1181,1273,1836,1861]" pageId="7" pageNumber="118">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, posttemporal tubercles completely missing, neck broad; frontal furrows prominent, faintly divergent posteriorly and nearly reaching eye level, extending forwards up to clypeal setae, resembling in shape
|
||
<taxonomicName id="4C5B41D0FFA0FFB3FBDF594095E1F800" box="[1053,1206,1908,1932]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="118" phylum="Arthropoda" rank="species" species="arcanoides">
|
||
<emphasis id="B92FE641FFA0FFB3FBDF594095E1F800" box="[1053,1206,1908,1932]" italics="true" pageId="7" pageNumber="118">P. arcanoides</emphasis>
|
||
</taxonomicName>
|
||
, frons with coarsely rugose transversal impression, arcuately connecting frontal furrows; surface smooth, 2 setiferous pores present on each side, anterior pore situated in supraorbital groove at eye level, and posterior—at hind extremity of supraorbital groove, strongly removed posteriorly, index 1 = 0.40: 1.00: 1.07; tooth of mentum (
|
||
<figureCitation id="136026D6FFAFFFBCFBA35EA295E8FF3C" box="[1121,1215,151,176]" captionStart="FIGURES 10 – 14" captionStartId="7.[151,264,1460,1483]" captionTargetBox="[169,1412,218,1409]" captionTargetId="figure@7.[151,1436,193,1440]" captionTargetPageId="7" captionText="FIGURES 10 – 14. Pterostichus spp., mentum, dorsal view: 10 — Pterostichus mariamae sp. nov., holotype, male; 11 — Pterostichus mariamae sp. nov., paratype, female; 12 — Pterostichus arcanoides Lorenz, male, Chedym Mt. Range; 13 — Pterostichus arcanoides Lorenz, female, Chedym Mt. Range; 14 — Pterostichus svetlanae sp. nov., holotype, female. Scale bar 1.0 mm." httpUri="https://zenodo.org/record/243364/files/figure.png" pageId="8" pageNumber="119">Figs. 14</figureCitation>
|
||
) rather short, bifid, with rather shallow apical hollow, resembling previous species; mandibles rather short with moderately bent apices; antennomere 2 rather short, index 3 = 1.89 (2.21): 1.00 (1.54): 1.72 (2,63): 1.73 (2.90): 1.46 (2.69).
|
||
</paragraph>
|
||
<paragraph id="8BE43A53FFAFFFBCFF055F30932FFE5C" blockId="8.[151,1437,151,1797]" pageId="8" pageNumber="119">Pronotum somewhat back-trapeziform, 0.97 times as long as wide, faintly convex, widest at apical quarter; apex fairly emarginate in the middle, lateral sides narrowly reflexed, inconspicuously rounded anteriorly, prominently sinuated before hind angles, basal margin emarginate in the middle, hind angles prominent, nearly rectangular, pointed; disk rather finely rugose, anterior transverse impression rather shallow, median line deep, reaching both extremities, basal foveae longitudinal, shallow; lateral margins with 5–7 marginal setae in anterior half and 1 seta in hind angles at both sides.</paragraph>
|
||
<paragraph id="8BE43A53FFAFFFBCFF055FE995CFFD09" blockId="8.[151,1437,151,1797]" pageId="8" pageNumber="119">
|
||
Elytra elongate, rather narrow, 1.76 times as long as wide and 1.17 times as wide as pronotum, sides faintly rounded, shoulders rounded, completely not angulated; striae fine but prominent, scutellary striole distinct, intervals nearly flat; interval 3 with 4 pores adjoining stria 3, umbilicate series distinctly subinterrupted in the middle, forming vague humeral (6–7 pores), median (1–2 pores), and apical (8–9 pores) groups; microsculpture composed of fine transverse wrinkles and nearly isodiametric meshes.
|
||
<collectingCountry id="F34C7AC3FFAFFFBCFC6D5C5992B0FD09" box="[943,999,620,645]" name="India" pageId="8" pageNumber="119">Hind</collectingCountry>
|
||
wings reduced.
|
||
</paragraph>
|
||
<paragraph id="8BE43A53FFAFFFBCFF055CA59316FC99" blockId="8.[151,1437,151,1797]" pageId="8" pageNumber="119">
|
||
Mesosternum coarsely punctate, proepisterna rather sparsely and smoothly punctate, mesepisterna, metepisterna, and lateral sides of metasternum smooth, lateral areas of sternites rather coarsely rugose. Metatrochantera nearly of the same shape as in
|
||
<taxonomicName id="4C5B41D0FFAFFFBCFD0B5CEC920DFD63" box="[713,858,728,752]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="119" phylum="Arthropoda" rank="species" species="mariamae" status="sp. nov.">
|
||
<emphasis id="B92FE641FFAFFFBCFD0B5CEC920DFD63" box="[713,858,728,752]" italics="true" pageId="8" pageNumber="119">P. mariamae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21C5B3AFFAFFFBCFCA65CE29296FD7C" box="[868,961,727,752]" pageId="8" pageNumber="119" rank="species">
|
||
<emphasis id="B92FE641FFAFFFBCFCA65CE29296FD7C" bold="true" box="[868,961,727,752]" pageId="8" pageNumber="119">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
female. Ventral side of tarsal segment 5 pubescent both dorsally and ventrally.
|
||
</paragraph>
|
||
<paragraph id="8BE43A53FFAFFFBCFF055D2A9020FCB4" blockId="8.[151,1437,151,1797]" box="[199,375,799,824]" pageId="8" pageNumber="119">Male unknown.</paragraph>
|
||
<paragraph id="8BE43A53FFAFFFBCFF055D7192F3FAF4" blockId="8.[151,1437,151,1797]" pageId="8" pageNumber="119">
|
||
<emphasis id="B92FE641FFAFFFBCFF055D719082FCD1" bold="true" box="[199,469,836,861]" pageId="8" pageNumber="119">Differential diagnosis.</emphasis>
|
||
The new species resembles in habitus
|
||
<taxonomicName id="4C5B41D0FFAFFFBCFC7E5D70950DFCD0" box="[956,1114,836,860]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="119" phylum="Arthropoda" rank="species" species="arcanoides">
|
||
<emphasis id="B92FE641FFAFFFBCFC7E5D70950DFCD0" box="[956,1114,836,860]" italics="true" pageId="8" pageNumber="119">P. arcanoides</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5B41D0FFAFFFBCFB5D5D709464FCD0" box="[1183,1331,837,860]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="119" phylum="Arthropoda" rank="species" species="mariamae" status="sp. nov.">
|
||
<emphasis id="B92FE641FFAFFFBCFB5D5D709464FCD0" box="[1183,1331,837,860]" italics="true" pageId="8" pageNumber="119">P. mariamae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21C5B3AFFAFFFBCFA805D7194C3FCD1" box="[1346,1428,836,861]" pageId="8" pageNumber="119" rank="species">
|
||
<emphasis id="B92FE641FFAFFFBCFA805D7194C3FCD1" bold="true" box="[1346,1428,836,861]" pageId="8" pageNumber="119">sp.nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, combining their individual characters, but also possesses some unique ones. In general it differs in the following main features (
|
||
<figureCitation id="136026D6FFAFFFBCFE825DB990FAFC28" box="[320,429,908,933]" captionStart="FIGURES 3 – 5" captionStartId="4.[151,264,1952,1975]" captionTargetBox="[332,1257,211,1906]" captionTargetId="figure@4.[311,1276,193,1932]" captionTargetPageId="4" captionText="FIGURES 3 – 5. Pterostichus mariamae sp. nov.: 3 – 4 — dorsal habitus: 3 — holotype, male; 4 — paratype, female; 5 — right metafemur and metatrochanter apices, holotype, male. Scale bars 1.0 mm." httpUri="https://zenodo.org/record/243361/files/figure.png" pageId="8" pageNumber="119">Figs. 3, 4</figureCitation>
|
||
,
|
||
<figureCitation id="136026D6FFAFFFBCFE785DB9909DFC29" box="[442,458,908,933]" captionStart="FIGURE 6" captionStartId="5.[151,250,2000,2022]" captionTargetBox="[513,1436,572,1970]" captionTargetId="figure@5.[502,1085,631,1979]" captionTargetPageId="5" captionText="FIGURE 6. Pterostichus svetlanae sp. nov., dorsal habitus, holotype, female. Scale bar 1.0 mm." httpUri="https://zenodo.org/record/243362/files/figure.png" pageId="8" pageNumber="119">6</figureCitation>
|
||
,
|
||
<figureCitation id="136026D6FFAFFFBCFE155DB89353FC28" box="[471,516,908,932]" captionStart="FIGURES 7 – 9" captionStartId="6.[151,264,1952,1975]" captionTargetBox="[327,1261,213,1908]" captionTargetId="figure@6.[307,1280,193,1932]" captionTargetPageId="6" captionText="FIGURES 7 – 9. Pterostichus arcanoides Lorenz: 7 – 8 — dorsal habitus: 7 — male, Chedym Mt. Range; 8 — female, Chedym Mt. Range; 9 – right metafemur and metatrochanter apices, male, Chedym Mt. Range. Scale bars 1.0 mm." httpUri="https://zenodo.org/record/243363/files/figure.png" pageId="8" pageNumber="119">7, 8</figureCitation>
|
||
): body larger and less slender; head of more parallel-side form (
|
||
<figureCitation id="136026D6FFAFFFBCFB1C5DB99465FC28" box="[1246,1330,908,933]" captionStart="FIGURE 23" captionStartId="11.[151,250,938,960]" captionTargetBox="[282,1307,202,913]" captionTargetId="figure@11.[275,1312,193,916]" captionTargetPageId="11" captionText="FIGURE 23. Female head comparative proportions of Pterostichus- species: left— Pt. svetlanae sp. nov.; middle— Pt. mariamae sp. nov.; right— Pt. arcanoides Lorenz." httpUri="https://zenodo.org/record/243367/files/figure.png" pageId="8" pageNumber="119">Fig. 23</figureCitation>
|
||
), index 2 being 1.00: 0.90: 1.09 (
|
||
<taxonomicName id="4C5B41D0FFAFFFBCFE655D849369FC44" box="[423,574,944,968]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="119" phylum="Arthropoda" rank="species" species="arcanoides">
|
||
<emphasis id="B92FE641FFAFFFBCFE655D849369FC44" box="[423,574,944,968]" italics="true" pageId="8" pageNumber="119">P. arcanoides</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5B41D0FFAFFFBCFDB15D8493A8FC4B" box="[627,767,944,968]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="119" phylum="Arthropoda" rank="species" species="mariamae" status="sp. nov.">
|
||
<emphasis id="B92FE641FFAFFFBCFDB15D8493A8FC4B" box="[627,767,944,968]" italics="true" pageId="8" pageNumber="119">P. mariamae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21C5B3AFFAFFFBCFCC45D9A9208FC44" box="[774,863,943,968]" pageId="8" pageNumber="119" rank="species">
|
||
<emphasis id="B92FE641FFAFFFBCFCC45D9A9208FC44" bold="true" box="[774,863,943,968]" pageId="8" pageNumber="119">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
possess more trapeziform head, their index 2 being 1.00: 0.92–0.94: 1.15–1.16); genae less developed, compound eyes occupy somewhat lateral position and seem to be removed posteriorly from the anterior margin of genae to posterior margin of temples, index 1 being 0.40: 1.00: 1.07 (in
|
||
<taxonomicName id="4C5B41D0FFAFFFBCFEC05A2890CEFBB8" box="[258,409,1052,1076]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="119" phylum="Arthropoda" rank="species" species="arcanoides">
|
||
<emphasis id="B92FE641FFAFFFBCFEC05A2890CEFBB8" box="[258,409,1052,1076]" italics="true" pageId="8" pageNumber="119">P. arcanoides</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5B41D0FFAFFFBCFE0F5A28930DFBB8" box="[461,602,1053,1076]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="119" phylum="Arthropoda" rank="species" species="mariamae" status="sp. nov.">
|
||
<emphasis id="B92FE641FFAFFFBCFE0F5A28930DFBB8" box="[461,602,1053,1076]" italics="true" pageId="8" pageNumber="119">P. mariamae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21C5B3AFFAFFFBCFDA25A2993E4FBB9" box="[608,691,1052,1077]" pageId="8" pageNumber="119" rank="species">
|
||
<emphasis id="B92FE641FFAFFFBCFDA25A2993E4FBB9" bold="true" box="[608,691,1052,1077]" pageId="8" pageNumber="119">sp.nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
it is 0.58–0.75: 1.00: 0.90–1.00); antennomere 2 is the shortest of all the three species, mandibles moderately bent apically to compare with
|
||
<taxonomicName id="4C5B41D0FFAFFFBCFC1E5A749524FBD4" box="[988,1139,1088,1112]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="119" phylum="Arthropoda" rank="species" species="arcanoides">
|
||
<emphasis id="B92FE641FFAFFFBCFC1E5A749524FBD4" box="[988,1139,1088,1112]" italics="true" pageId="8" pageNumber="119">P. arcanoides</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5B41D0FFAFFFBCFB6E5A74946DFBDB" box="[1196,1338,1088,1112]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="119" phylum="Arthropoda" rank="species" species="mariamae" status="sp. nov.">
|
||
<emphasis id="B92FE641FFAFFFBCFB6E5A74946DFBDB" box="[1196,1338,1088,1112]" italics="true" pageId="8" pageNumber="119">P. mariamae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21C5B3AFFAFFFBCFA805A0A94CBFBD4" box="[1346,1436,1087,1112]" pageId="8" pageNumber="119" rank="species">
|
||
<emphasis id="B92FE641FFAFFFBCFA805A0A94CBFBD4" bold="true" box="[1346,1436,1087,1112]" pageId="8" pageNumber="119">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
From
|
||
<taxonomicName id="4C5B41D0FFAFFFBCFF1E5A509021FBF0" box="[220,374,1124,1148]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="119" phylum="Arthropoda" rank="species" species="arcanoides">
|
||
<emphasis id="B92FE641FFAFFFBCFF1E5A509021FBF0" box="[220,374,1124,1148]" italics="true" pageId="8" pageNumber="119">P. arcanoides</emphasis>
|
||
</taxonomicName>
|
||
differs also in the shape of the tooth of mentum (
|
||
<figureCitation id="136026D6FFAFFFBCFC755A519514FBF0" box="[951,1091,1124,1149]" captionStart="FIGURES 10 – 14" captionStartId="7.[151,264,1460,1483]" captionTargetBox="[169,1412,218,1409]" captionTargetId="figure@7.[151,1436,193,1440]" captionTargetPageId="7" captionText="FIGURES 10 – 14. Pterostichus spp., mentum, dorsal view: 10 — Pterostichus mariamae sp. nov., holotype, male; 11 — Pterostichus mariamae sp. nov., paratype, female; 12 — Pterostichus arcanoides Lorenz, male, Chedym Mt. Range; 13 — Pterostichus arcanoides Lorenz, female, Chedym Mt. Range; 14 — Pterostichus svetlanae sp. nov., holotype, female. Scale bar 1.0 mm." httpUri="https://zenodo.org/record/243364/files/figure.png" pageId="8" pageNumber="119">Figs. 10–14</figureCitation>
|
||
); proportions of pronotum are nearly intermediate between
|
||
<taxonomicName id="4C5B41D0FFAFFFBCFE3A5ABC93CEFB2C" box="[504,665,1160,1184]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="119" phylum="Arthropoda" rank="species" species="arcanoides">
|
||
<emphasis id="B92FE641FFAFFFBCFE3A5ABC93CEFB2C" box="[504,665,1160,1184]" italics="true" pageId="8" pageNumber="119">P. arcanoides</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5B41D0FFAFFFBCFD205ABC922EFB13" box="[738,889,1160,1184]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="119" phylum="Arthropoda" rank="species" species="mariamae" status="sp. nov.">
|
||
<emphasis id="B92FE641FFAFFFBCFD205ABC922EFB13" box="[738,889,1160,1184]" italics="true" pageId="8" pageNumber="119">P. mariamae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21C5B3AFFAFFFBCFC4B5AB292BBFB2C" box="[905,1004,1159,1184]" pageId="8" pageNumber="119" rank="species">
|
||
<emphasis id="B92FE641FFAFFFBCFC4B5AB292BBFB2C" bold="true" box="[905,1004,1159,1184]" pageId="8" pageNumber="119">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, in general form and character of microreticulation it is closer however to
|
||
<taxonomicName id="4C5B41D0FFAFFFBCFD995A9893B3FB48" box="[603,740,1197,1220]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="119" phylum="Arthropoda" rank="subSpecies" species="mari" status="sp. nov." subSpecies="amae">
|
||
<emphasis id="B92FE641FFAFFFBCFD995A9893B3FB48" box="[603,740,1197,1220]" italics="true" pageId="8" pageNumber="119">P. mari amae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21C5B3AFFAFFFBCFD2C5A999210FB49" box="[750,839,1196,1221]" pageId="8" pageNumber="119" rank="species">
|
||
<emphasis id="B92FE641FFAFFFBCFD2C5A999210FB49" bold="true" box="[750,839,1196,1221]" pageId="8" pageNumber="119">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, but differs in a very shallow, faint basal foveae; disk of pronotum without prominent and locally coarse microsculpture, characteristic for
|
||
<emphasis id="B92FE641FFAFFFBCFB855AE495B2FB64" box="[1095,1253,1232,1256]" italics="true" pageId="8" pageNumber="119">
|
||
<taxonomicName id="4C5B41D0FFAFFFBCFB855AE495B6FB64" box="[1095,1249,1232,1256]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="119" phylum="Arthropoda" rank="species" species="arcanoides">P. arcanoides</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Elytra elongate, narrow (see description), shoulders widely rounded, without prominent angles, being 1.76 times as long as wide and 1.17 times as wide as pronotum (in
|
||
<taxonomicName id="4C5B41D0FFAFFFBCFDA25B2C93AFFABC" box="[608,760,1304,1328]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="119" phylum="Arthropoda" rank="species" species="arcanoides">
|
||
<emphasis id="B92FE641FFAFFFBCFDA25B2C93AFFABC" box="[608,760,1304,1328]" italics="true" pageId="8" pageNumber="119">P. arcanoides</emphasis>
|
||
</taxonomicName>
|
||
elytra are 1.78–1.82 times as long as wide, with prominent humeral angle, in
|
||
<taxonomicName id="4C5B41D0FFAFFFBCFEA75B0890A2FAD8" box="[357,501,1341,1364]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="119" phylum="Arthropoda" rank="species" species="mariamae" status="sp. nov.">
|
||
<emphasis id="B92FE641FFAFFFBCFEA75B0890A2FAD8" box="[357,501,1341,1364]" italics="true" pageId="8" pageNumber="119">P. mariamae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21C5B3AFFAFFFBCFE3E5B09930FFAD9" box="[508,600,1340,1365]" pageId="8" pageNumber="119" rank="species">
|
||
<emphasis id="B92FE641FFAFFFBCFE3E5B09930FFAD9" bold="true" box="[508,600,1340,1365]" pageId="8" pageNumber="119">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
elytra are 1.85–2.08 times as long as wide, with well-developed humeral angle); elytral striae fine, less prominent than in
|
||
<taxonomicName id="4C5B41D0FFAFFFBCFD745B549213FAFB" box="[694,836,1376,1400]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="119" phylum="Arthropoda" rank="species" species="mariamae" status="sp. nov.">
|
||
<emphasis id="B92FE641FFAFFFBCFD745B549213FAFB" box="[694,836,1376,1400]" italics="true" pageId="8" pageNumber="119">P. mariamae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21C5B3AFFAFFFBCFC885B6A92F3FAF4" box="[842,932,1375,1400]" pageId="8" pageNumber="119" rank="species">
|
||
<emphasis id="B92FE641FFAFFFBCFC885B6A92F3FAF4" bold="true" box="[842,932,1375,1400]" pageId="8" pageNumber="119">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34169D8FFAFFFBCFF055BB1933CF889" pageId="8" pageNumber="119" type="distribution">
|
||
<paragraph id="8BE43A53FFAFFFBCFF055BB194DFFA11" blockId="8.[151,1437,151,1797]" box="[199,1416,1412,1437]" pageId="8" pageNumber="119">
|
||
<emphasis id="B92FE641FFAFFFBCFF055BB19037FA11" bold="true" box="[199,352,1412,1437]" pageId="8" pageNumber="119">Distribution.</emphasis>
|
||
Transcaucasia (=South Caucasus), Abkhazia, valley of the river Mzara, Zashirbara Mt. Range.
|
||
</paragraph>
|
||
<paragraph id="8BE43A53FFAFFFBCFF055B92935DFA69" blockId="8.[151,1437,151,1797]" pageId="8" pageNumber="119">
|
||
<emphasis id="B92FE641FFAFFFBCFF055B92901DFA4C" bold="true" box="[199,330,1447,1472]" pageId="8" pageNumber="119">Bionomics.</emphasis>
|
||
<typeStatus id="54E084F1FFAFFFBCFE915B9C90DCFA4D" box="[339,395,1449,1473]" pageId="8" pageNumber="119">Type</typeStatus>
|
||
specimen was collected in a maple-beech forest on highly pronounced karstic landforms at altitudes of
|
||
<quantity id="4CA397B6FFAFFFBCFED95BF990CDFA68" box="[283,410,1484,1509]" metricMagnitude="2" metricUnit="m" metricValue="6.925" metricValueMax="7.05" metricValueMin="6.8" pageId="8" pageNumber="119" unit="m" value="692.5" valueMax="705.0" valueMin="680.0">680–705 m</quantity>
|
||
(
|
||
<figureCitation id="136026D6FFAFFFBCFE6B5BF990A9FA69" box="[425,510,1484,1509]" captionStart="FIGURE 25" captionStartId="12.[151,250,1178,1200]" captionTargetBox="[151,1435,193,1156]" captionTargetId="figure@12.[151,1436,193,1156]" captionTargetPageId="12" captionText="FIGURE 25. Habitat of Pterostichus svetlanae sp. nov., Zashirbara Mt. Range, 700 m, maple-beech forest." httpUri="https://zenodo.org/record/243369/files/figure.png" pageId="8" pageNumber="119">Fig. 25</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE43A53FFAFFFBCFF055BDA95FCF9A0" blockId="8.[151,1437,151,1797]" pageId="8" pageNumber="119">
|
||
<emphasis id="B92FE641FFAFFFBCFF055BDA90D1F984" bold="true" box="[199,390,1519,1544]" pageId="8" pageNumber="119">Specific epithet.</emphasis>
|
||
We are pleased to dedicate this new species to the wife of the second author, Svetlana, as a sign of gratitude for her infinite patience and support for his research travels in the Caucasus.
|
||
</paragraph>
|
||
<paragraph id="8BE43A53FFAFFFBCFF055802933CF889" blockId="8.[151,1437,151,1797]" pageId="8" pageNumber="119">
|
||
Several characters utilized earlier by Kurnakov for discrimination of his subgenus
|
||
<taxonomicName id="4C5B41D0FFAFFFBCFB70580D9432F9DC" box="[1202,1381,1592,1616]" class="Insecta" family="Carabidae" genus="Scaritomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="119" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92FE641FFAFFFBCFB70580D9432F9DC" box="[1202,1381,1592,1616]" italics="true" pageId="8" pageNumber="119">Scaritomorphus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5B41D0FFAFFFBCFF5558699050F9F8" box="[151,263,1628,1652]" class="Insecta" family="Carabidae" genus="Aphaonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="119" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92FE641FFAFFFBCFF5558699050F9F8" box="[151,263,1628,1652]" italics="true" pageId="8" pageNumber="119">Aphaonus</emphasis>
|
||
</taxonomicName>
|
||
(inflated head, microphthalmia, absence of the posttemporal tubercles) already appeared unreliable. Comparison of
|
||
<taxonomicName id="4C5B41D0FFAFFFBCFE8A58B490B7F914" box="[328,480,1664,1688]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="119" phylum="Arthropoda" rank="species" species="arcanoides">
|
||
<emphasis id="B92FE641FFAFFFBCFE8A58B490B7F914" box="[328,480,1664,1688]" italics="true" pageId="8" pageNumber="119">P. arcanoides</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5B41D0FFAFFFBCFE3258B49328F91B" box="[496,639,1664,1688]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="119" phylum="Arthropoda" rank="species" species="mariamae" status="sp. nov.">
|
||
<emphasis id="B92FE641FFAFFFBCFE3258B49328F91B" box="[496,639,1664,1688]" italics="true" pageId="8" pageNumber="119">P. mariamae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21C5B3AFFAFFFBCFD45584A93B5F914" box="[647,738,1663,1688]" pageId="8" pageNumber="119" rank="species">
|
||
<emphasis id="B92FE641FFAFFFBCFD45584A93B5F914" bold="true" box="[647,738,1663,1688]" pageId="8" pageNumber="119">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, and
|
||
<taxonomicName id="4C5B41D0FFAFFFBCFCE158B492FDF914" box="[803,938,1664,1688]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="119" phylum="Arthropoda" rank="species" species="svetlanae" status="sp. nov.">
|
||
<emphasis id="B92FE641FFAFFFBCFCE158B492FDF914" box="[803,938,1664,1688]" italics="true" pageId="8" pageNumber="119">P. svetlanae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21C5B3AFFAFFFBCFC71584A9559F914" box="[947,1038,1663,1688]" pageId="8" pageNumber="119" rank="species">
|
||
<emphasis id="B92FE641FFAFFFBCFC71584A9559F914" bold="true" box="[947,1038,1663,1688]" pageId="8" pageNumber="119">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
confirms, however, the stability in the pronotal and elytral chaetotaxy of this species-group (plurisetose pronotal lateral margin, umbilicate series hardly interrupted at middle, etc.) and in the presence of the frontal arcuate impression, easily distinguishing it from the other known
|
||
<taxonomicName id="4C5B41D0FFAFFFBCFE5358D99356F888" box="[401,513,1772,1796]" class="Insecta" family="Carabidae" genus="Aphaonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="119" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92FE641FFAFFFBCFE5358D99356F888" box="[401,513,1772,1796]" italics="true" pageId="8" pageNumber="119">Aphaonus</emphasis>
|
||
</taxonomicName>
|
||
lineages.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |