183 lines
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183 lines
15 KiB
XML
<document ID-DOI="10.1206/0003-0090.457.1.1" ID-ISSN="0003-0090" ID-Zenodo-Dep="6971356" approvalRequired="59" approvalRequired_for_taxonomicNames="59" checkinTime="1659882917898" checkinUser="felipe" docAuthor="Beck, Robin M. D., Voss, Robert S. & Jansa, Sharon A." docDate="2022" docId="03EFDD5DF768697ADB2AFF4F1927FA67" docLanguage="en" docName="BulAmeMusNatHist.2022.457.1-350.pdf" docOrigin="Bulletin of the American Museum of Natural History 2022 (457)" docSource="https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-457/issue-1/0003-0090.457.1.1/Craniodental-Morphology-and-Phylogeny-of-Marsupials/10.1206/0003-0090.457.1.1.full" docStyle="DocumentStyle:915933466F796C9C739DF4DB6B8DCFA6.8:BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleId="915933466F796C9C739DF4DB6B8DCFA6" docStyleName="BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleVersion="8" docTitle="Yalkaparidon undetermined" docType="treatment" docVersion="5" lastPageNumber="339" masterDocId="FFD6A525F63B6829DA0BFFAE1A62FFC7" masterDocTitle="Craniodental Morphology And Phylogeny Of Marsupials" masterLastPageNumber="353" masterPageNumber="1" pageNumber="339" updateTime="1659987007554" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo>
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<mods:title>Craniodental Morphology And Phylogeny Of Marsupials</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Beck, Robin M. D.</mods:namePart>
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<mods:affiliation>School of Science, Engineering and Environment University of Salford, U. K. & School of Biological, Earth & Environmental Sciences University of New South Wales, Australia & Division of Vertebrate Zoology (Mammalogy) American Museum of Natural History</mods:affiliation>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Voss, Robert S.</mods:namePart>
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<mods:affiliation>Division of Vertebrate Zoology (Mammalogy) American Museum of Natural History</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jansa, Sharon A.</mods:namePart>
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<mods:affiliation>Bell Museum and Department of Ecology, Evolution, and Behavior University of Minnesota</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Bulletin of the American Museum of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-06-28</mods:number>
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<mods:detail type="volume">
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<mods:number>2022</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>457</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>353</mods:end>
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<mods:url>https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-457/issue-1/0003-0090.457.1.1/Craniodental-Morphology-and-Phylogeny-of-Marsupials/10.1206/0003-0090.457.1.1.full</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1206/0003-0090.457.1.1</mods:identifier>
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<mods:identifier type="ISSN">0003-0090</mods:identifier>
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<mods:identifier type="Zenodo-Dep">6971356</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6974527" ID-Zenodo-Dep="6974527" LSID="urn:lsid:plazi:treatment:03EFDD5DF768697ADB2AFF4F1927FA67" httpUri="http://treatment.plazi.org/id/03EFDD5DF768697ADB2AFF4F1927FA67" lastPageNumber="339" pageId="339" pageNumber="339">
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<subSubSection box="[289,455,225,249]" pageId="339" pageNumber="339" type="nomenclature">
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<paragraph blockId="339.[289,455,225,249]" box="[289,455,225,249]" pageId="339" pageNumber="339">
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<heading box="[289,455,225,249]" centered="true" fontSize="10" level="2" pageId="339" pageNumber="339" reason="4">
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<emphasis bold="true" box="[289,455,225,249]" pageId="339" pageNumber="339">
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†
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<taxonomicName box="[303,455,225,249]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" kingdom="Animalia" pageId="339" pageNumber="339" phylum="Chordata" rank="species">
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<emphasis bold="true" box="[303,455,225,249]" italics="true" pageId="339" pageNumber="339">Yalkaparidon</emphasis>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="339" pageNumber="339" type="synonymic_list">
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<paragraph blockId="339.[108,638,293,1442]" pageId="339" pageNumber="339">
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SPECIES SCORED:
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<emphasis bold="true" box="[336,350,293,317]" pageId="339" pageNumber="339">†</emphasis>
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<taxonomicName box="[350,571,293,317]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="339" pageNumber="339" phylum="Chordata" rank="species">
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<emphasis box="[350,571,293,317]" italics="true" pageId="339" pageNumber="339">Yalkaparidon coheni</emphasis>
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</taxonomicName>
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(
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<typeStatus box="[587,636,293,317]" pageId="339" pageNumber="339">type</typeStatus>
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species), †
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<emphasis box="[221,371,326,350]" italics="true" pageId="339" pageNumber="339">
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<taxonomicName box="[221,368,326,350]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="339" pageNumber="339" phylum="Chordata" rank="genus">Yalkaparidon</taxonomicName>
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.
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</emphasis>
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sp.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="339" pageNumber="339" type="materials_examined">
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<paragraph blockId="339.[108,638,293,1442]" pageId="339" pageNumber="339">
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GEOLOGICAL PROVENANCE OF SCORED
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<collectionCode box="[108,168,393,417]" pageId="339" pageNumber="339">SPECIMENS</collectionCode>
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: Riversleigh Faunal Zone B, Riversleigh World Heritage Area,
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<collectingRegion box="[350,482,426,450]" country="Australia" name="Queensland" pageId="339" pageNumber="339">Queensland</collectingRegion>
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,
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<collectingCountry box="[494,596,426,450]" name="Australia" pageId="339" pageNumber="339">Australia</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="339" pageNumber="339" type="description">
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<paragraph blockId="339.[108,638,293,1442]" pageId="339" pageNumber="339">AGE OF SCORED SPECIMENS: Riversleigh Faunal Zone B is interpreted to be early Miocene, based on biostratigraphy (see above). In the absence of radiometric dates, we have assumed the entire span of the early Miocene (Aquitanian to Burdigalian; Cohen et al., 2013 [updated]) for this terminal.</paragraph>
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<paragraph blockId="339.[108,638,293,1442]" box="[140,621,690,715]" pageId="339" pageNumber="339">
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ASSIGNED AGE
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<collectionCode box="[311,388,690,714]" pageId="339" pageNumber="339">RANGE</collectionCode>
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:
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<geoCoordinate box="[402,475,690,714]" degrees="23.030" direction="north" orientation="latitude" pageId="339" pageNumber="339" precision="55" value="23.03">23.030</geoCoordinate>
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<geoCoordinate box="[474,559,690,714]" degrees="15.970" direction="west" orientation="longitude" pageId="339" pageNumber="339" precision="55" value="-15.97">–15.970</geoCoordinate>
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Mya.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="339" pageNumber="339" type="discussion">
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<paragraph blockId="339.[108,638,293,1442]" lastBlockId="339.[684,1214,226,1440]" pageId="339" pageNumber="339">
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REMARKS: †
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<taxonomicName box="[280,430,723,747]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="339" pageNumber="339" phylum="Chordata" rank="genus">
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<emphasis box="[280,430,723,747]" italics="true" pageId="339" pageNumber="339">Yalkaparidon</emphasis>
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</taxonomicName>
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is unique among Australian fossil metatherians in that it is known from extensive craniodental material, including a well-preserved partial skull (
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<collectionCode box="[116,163,856,880]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="339" pageNumber="339" type="Museum">QM</collectionCode>
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<accessionNumber box="[172,256,856,880]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/F13008" pageId="339" pageNumber="339">F13008</accessionNumber>
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, the
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<typeStatus box="[314,413,855,879]" pageId="339" pageNumber="339">holotype</typeStatus>
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of †
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<taxonomicName box="[467,568,855,879]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="339" pageNumber="339" phylum="Chordata" rank="species">
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<emphasis box="[467,568,855,879]" italics="true" pageId="339" pageNumber="339">Y. coheni</emphasis>
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</taxonomicName>
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), and yet cannot be confidently referred to one of the seven extant Australian marsupial orders (Archer et al., 1988; Beck et al., 2014). This is due to its unique combination of dental features, including hypselodont first upper and lower incisors; a procumbent, gliriform lower first incisor; zalambdodont molars; and a cranial morphology that appears plesiomorphic compared with other known Australian marsupials yet also exhibits some striking apomorphic features (such as a very reduced postglenoid process, a transverse canal foramen that is posterior to the carotid foramen, and very robust and posteriorly extensive entopterygoid crests; Archer et al., 1988; Beck, 2009; Beck et al., 2014). Marshall et al. (1990) and Szalay (1994) suggested that †
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<taxonomicName box="[518,726,226,1442]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="339" pageNumber="339" phylum="Chordata" rank="genus">
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<emphasis box="[518,726,226,1442]" italics="true" pageId="339" pageNumber="339">Yalkaparidon</emphasis>
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</taxonomicName>
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is a member of
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<taxonomicName authorityName="Owen" authorityYear="1866" box="[935,1103,226,250]" class="Mammalia" kingdom="Animalia" order="Diprotodontia" pageId="339" pageNumber="339" phylum="Chordata" rank="order">Diprotodontia</taxonomicName>
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, whereas Woodburne and Case (1996) placed it in
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<taxonomicName authorityName="Kirsch" authorityYear="1977" class="Mammalia" kingdom="Animalia" order="Notoryctemorphia" pageId="339" pageNumber="339" phylum="Chordata" rank="order">Notoryctemorphia</taxonomicName>
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. However, the most detailed studies of †
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<taxonomicName box="[821,972,325,349]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="339" pageNumber="339" phylum="Chordata" rank="genus">
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<emphasis box="[821,972,325,349]" italics="true" pageId="339" pageNumber="339">Yalkaparidon</emphasis>
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</taxonomicName>
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to date suggest that it be placed in its own order, †Yalkaparidontia (Archer et al., 1988; Beck et al., 2014). Isolated tarsals tentatively referred to †
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<taxonomicName box="[1060,1212,424,448]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="339" pageNumber="339" phylum="Chordata" rank="genus">
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<emphasis box="[1060,1212,424,448]" italics="true" pageId="339" pageNumber="339">Yalkaparidon</emphasis>
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</taxonomicName>
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preserve apomorphic features that appear to be characteristic of australidelphians (Beck et al., 2014), and some phylogenetic analyses have placed †
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<taxonomicName box="[784,936,556,580]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="339" pageNumber="339" phylum="Chordata" rank="genus">
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<emphasis box="[784,936,556,580]" italics="true" pageId="339" pageNumber="339">Yalkaparidon</emphasis>
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</taxonomicName>
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within Australidelphia (Beck et al., 2014; Beck et al., 2016); however, other analyses have placed †
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<taxonomicName box="[1011,1160,622,646]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="339" pageNumber="339" phylum="Chordata" rank="genus">
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<emphasis box="[1011,1160,622,646]" italics="true" pageId="339" pageNumber="339">Yalkaparidon</emphasis>
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</taxonomicName>
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in a clade with paucituberculatans (Beck, 2017b; Zimicz and Goin, 2020), with which it shares presence of a gliriform lower first incisor.
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</paragraph>
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<paragraph blockId="339.[684,1214,226,1440]" pageId="339" pageNumber="339">
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Archer et al. (1988) described two species, both from Riversleigh: †
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<taxonomicName box="[977,1212,788,812]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="339" pageNumber="339" phylum="Chordata" rank="species">
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<emphasis box="[977,1212,788,812]" italics="true" pageId="339" pageNumber="339">Yalkaparidon coheni</emphasis>
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</taxonomicName>
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(based on specimens from Faunal Zone B) and †
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<taxonomicName box="[749,841,855,878]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="339" pageNumber="339" phylum="Chordata" rank="species">
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<emphasis box="[749,841,855,878]" italics="true" pageId="339" pageNumber="339">Y. jonesi</emphasis>
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</taxonomicName>
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(based on specimens from Faunal Zone C). Beck et al. (2014) examined all known material of †
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<taxonomicName box="[921,1069,920,944]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="339" pageNumber="339" phylum="Chordata" rank="genus">
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<emphasis box="[921,1069,920,944]" italics="true" pageId="339" pageNumber="339">Yalkaparidon</emphasis>
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</taxonomicName>
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collected to date from Riversleigh, and identified additional specimens of †
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<taxonomicName box="[854,955,986,1010]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="339" pageNumber="339" phylum="Chordata" rank="species">
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<emphasis box="[854,955,986,1010]" italics="true" pageId="339" pageNumber="339">Y. coheni</emphasis>
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</taxonomicName>
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and other †
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<taxonomicName class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="339" pageNumber="339" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="339" pageNumber="339">Yalkaparidon</emphasis>
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</taxonomicName>
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material not identifiable to species level from one Riversleigh Faunal Zone A site (White Hunter) and multiple Faunal Zone B sites (see Beck et al., 2014: electronic supplementary material); only Faunal Zone B specimens have been used for scoring purposes here. We did not score character data from specimens of †
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<taxonomicName box="[698,794,1251,1274]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="339" pageNumber="339" phylum="Chordata" rank="species">
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<emphasis box="[698,794,1251,1274]" italics="true" pageId="339" pageNumber="339">Y. jonesi</emphasis>
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</taxonomicName>
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, which appears to be more derived than †
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<taxonomicName box="[760,864,1284,1308]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="339" pageNumber="339" phylum="Chordata" rank="species">
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<emphasis box="[760,864,1284,1308]" italics="true" pageId="339" pageNumber="339">Y. coheni</emphasis>
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</taxonomicName>
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in lacking any teeth between the enlarged procumbent anteriormost lower incisor and the tooth that Archer et al. (1988) referred to as p3 but which Beck et al. (2014) argued is m1.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |