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<document id="C9108C9B9771D686EEF21783C1F62404" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629697810943" checkinUser="felipe" docAuthor="Chani-Posse, Mariana" docDate="2010" docId="03EAF012FFB18C1C5DA7799CFC2D5493" docLanguage="en" docName="zt02595p070.pdf" docOrigin="Zootaxa 2595" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Philonthus varians" docType="treatment" docVersion="2" lastPageNumber="62" masterDocId="FFD3886AFF8C8C215D30790BFFA85341" masterDocTitle="Revision of the southern South American species of Philonthus Stephens (Coleoptera: Staphylinidae) 2595" masterLastPageNumber="70" masterPageNumber="1" pageNumber="62" updateTime="1699097306278" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="D19FF8A1F21025A8A3AC2C7A8306B991">Revision of the southern South American species of Philonthus Stephens (Coleoptera: Staphylinidae) 2595</mods:title>
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<mods:namePart id="23F8D0DC1BDE1EC56C3DDCFDF5435AC0">Chani-Posse, Mariana</mods:namePart>
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<mods:date id="BB195E02A9A6EAB768C378670730DF60">2010</mods:date>
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<mods:number id="30E6B2A09B0FE159DB862803A149FE75">2010-08-31</mods:number>
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<mods:number id="3F18A2E1C7A5F2F1E4FE4EF4CA482346">2595</mods:number>
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<mods:classification id="14A387072ECA280D51E32122B1CEFB3B">journal article</mods:classification>
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<treatment id="03EAF012FFB18C1C5DA7799CFC2D5493" LSID="urn:lsid:plazi:treatment:03EAF012FFB18C1C5DA7799CFC2D5493" httpUri="http://treatment.plazi.org/id/03EAF012FFB18C1C5DA7799CFC2D5493" lastPageNumber="62" pageId="61" pageNumber="62">
<subSubSection id="C359128FFFB18C1C5DA7799CFDE953F0" box="[151,577,151,177]" pageId="61" pageNumber="62" type="nomenclature">
<paragraph id="8BFC4104FFB18C1C5DA7799CFDE953F0" blockId="61.[151,577,151,212]" box="[151,577,151,177]" pageId="61" pageNumber="62">
<heading id="D0B4F668FFB18C1C5DA7799CFDE953F0" bold="true" box="[151,577,151,177]" fontSize="11" level="1" pageId="61" pageNumber="62" reason="1">
<taxonomicName id="4C433A87FFB18C1C5DA7799CFDE953F0" ID-CoL="76XX2" authority="(Paykull, 1789)" baseAuthorityName="Paykull" baseAuthorityYear="1789" box="[151,577,151,177]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="varians">
<emphasis id="B9379D16FFB18C1C5DA7799CFDE953F0" bold="true" box="[151,577,151,177]" pageId="61" pageNumber="62">
<emphasis id="B9379D16FFB18C1C5DA7799CFED653F0" bold="true" box="[151,382,151,177]" italics="true" pageId="61" pageNumber="62">Philonthus varians</emphasis>
(
<bibRefCitation id="EFD23CF5FFB18C1C5CBC799CFD9153F0" author="Paykull, G. de" box="[396,569,151,177]" pageId="61" pageNumber="62" refId="ref50949" refString="Paykull, G. de (1789) Monographia Staphylinorum Sueciae. Johann Edman, Upsaliae, 8 + 81 pp." type="book" year="1789">Paykull, 1789</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C359128FFFB18C1C5DA779B1FE255395" box="[151,397,186,212]" pageId="61" pageNumber="62" type="description">
<paragraph id="8BFC4104FFB18C1C5DA779B1FE255395" blockId="61.[151,577,151,212]" box="[151,397,186,212]" pageId="61" pageNumber="62">
(
<figureCitation id="13785D81FFB18C1C5DAF79B1FEA65395" box="[159,270,186,212]" captionStart="FIGURES 116–143" captionStartId="51.[151,270,1778,1802]" captionTargetBox="[382,1201,196,1755]" captionTargetId="figure-149@51.[367,1205,189,1767]" captionTargetPageId="51" captionText="FIGURES 116–143. P. rubromaculatus: sternum 8, male (116); sternum 9 (117); aedoeagus, dorsal view (118); tergum 10, female (119); gonocoxites of female genital segment (120). P. ruficauda: sternum 8, male (121); sternum 9 (122); tergum 10, male (123); aedoeagus, lateral view (124); aedoeagus, dorsal view (125); tergum 10, female (126); gonocoxites of female genital segment (127). P. sericans: sternum 9 (128); tergum 10, male (129); aedoeagus, dorsal view (130); paramere, ventral side (131). P. stenocephalus: sternum 9 (132); tergum 10, male (133); aedoeagus, dorsal view (134); paramere, ventral side (135); aedoeagus, lateral view (136); tergum 10, female (137); gonocoxites of female genital segment (138). P. varians: tergum 10, male (139); sternum 9 (140); aedoeagus, lateral view (141); paramere, ventral side (142); aedoeagus, dorsal view (143); gonocoxites of female genital segment (144)." pageId="61" pageNumber="62">Figs. 139</figureCitation>
–144, 149)
</paragraph>
</subSubSection>
<subSubSection id="C359128FFFB18C1C5DA7780EFB8B525D" box="[151,1059,261,284]" pageId="61" pageNumber="62" type="reference_group">
<paragraph id="8BFC4104FFB18C1C5DA7780EFB8B525D" blockId="61.[151,1059,261,284]" box="[151,1059,261,284]" pageId="61" pageNumber="62">
<treatmentCitationGroup id="AB53662AFFB18C1C5DA7780EFB8B525D" box="[151,1059,261,284]" pageId="61" pageNumber="62">
<treatmentCitation id="0AE26715FFB18C1C5DA7780EFD87525D" author="Paykull, G. de" box="[151,559,261,284]" page="45" pageId="61" pageNumber="62" year="1789">
<taxonomicName id="4C433A87FFB18C1C5DA7780EFD87525D" authority="Paykull, 1789: 45" authorityName="Paykull" authorityPageNumber="45" authorityYear="1789" box="[151,559,261,284]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="varians">
<emphasis id="B9379D16FFB18C1C5DA7780EFEC1525D" box="[151,361,261,284]" italics="true" pageId="61" pageNumber="62">Staphylinus varians</emphasis>
<bibRefCitation id="EFD23CF5FFB18C1C5C40780EFD87525D" author="Paykull, G. de" box="[368,559,261,284]" pageId="61" pageNumber="62" refId="ref50949" refString="Paykull, G. de (1789) Monographia Staphylinorum Sueciae. Johann Edman, Upsaliae, 8 + 81 pp." type="book" year="1789">Paykull, 1789: 45</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. For complete references see
<bibRefCitation id="EFD23CF5FFB18C1C5E5A780EFBB7525D" author="Herman, L. H." box="[874,1055,261,284]" pageId="61" pageNumber="62" pagination="2441 - 3020" refId="ref50050" refString="Herman, L. H. (2001 b) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. Part V. Bulletin of the American Museum of Natural History, 265, 2441 - 3020." type="journal article" year="2001">Herman (2001b)</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C359128FFFB18C1C5DA77841FE4052CB" pageId="61" pageNumber="62" type="diagnosis">
<paragraph id="8BFC4104FFB18C1C5DA77841FE4052CB" blockId="61.[151,1437,330,2002]" pageId="61" pageNumber="62">
<emphasis id="B9379D16FFB18C1C5DA77841FEBF5225" bold="true" box="[151,279,330,356]" pageId="61" pageNumber="62">Diagnosis.</emphasis>
<taxonomicName id="4C433A87FFB18C1C5C2D7841FE535222" baseAuthorityName="Paykull" baseAuthorityYear="1789" box="[285,507,330,355]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="varians">
<emphasis id="B9379D16FFB18C1C5C2D7841FE535222" box="[285,507,330,355]" italics="true" pageId="61" pageNumber="62">Philonthus varians</emphasis>
</taxonomicName>
may be identified, and distinguished from
<taxonomicName id="4C433A87FFB18C1C5EDF7841FBE55222" authorityName="Herman" authorityYear="2001" box="[1007,1101,330,355]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="argus">
<emphasis id="B9379D16FFB18C1C5EDF7841FBE55222" box="[1007,1101,330,355]" italics="true" pageId="61" pageNumber="62">P. argus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C433A87FFB18C1C59B47841FAEE5222" authorityName="Scheerpeltz" authorityYear="1972" box="[1156,1350,330,355]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="stenocephalus">
<emphasis id="B9379D16FFB18C1C59B47841FAEE5222" box="[1156,1350,330,355]" italics="true" pageId="61" pageNumber="62">P. stenocephalus</emphasis>
</taxonomicName>
, by the character states given below.
</paragraph>
</subSubSection>
<subSubSection id="C359128FFFB18C1C5DF5789EFC2A564D" pageId="61" pageNumber="62" type="description">
<paragraph id="8BFC4104FFB18C1C5DF5789EFD515105" blockId="61.[151,1437,330,2002]" pageId="61" pageNumber="62">
<emphasis id="B9379D16FFB18C1C5DF5789EFE2C52EE" bold="true" box="[197,388,405,431]" pageId="61" pageNumber="62">Redescription.</emphasis>
Length of the body
<quantity id="4CBBECE1FFB18C1C5FB8789EFC8952EE" box="[648,801,405,431]" metricMagnitude="-3" metricUnit="m" metricValue="7.15" metricValueMax="7.5" metricValueMin="6.8" pageId="61" pageNumber="62" unit="mm" value="7.15" valueMax="7.5" valueMin="6.8">6.8–7.5 mm</quantity>
. In most character states similar to
<taxonomicName id="4C433A87FFB18C1C59D2789EFAEF52EF" authorityName="Herman" authorityYear="2001" box="[1250,1351,405,430]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="argus">
<emphasis id="B9379D16FFB18C1C59D2789EFAEF52EF" box="[1250,1351,405,430]" italics="true" pageId="61" pageNumber="62">P. argus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C433A87FFB18C1C58B5789EFE965292" authorityName="Scheerpeltz" authorityYear="1972" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="stenocephalus">
<emphasis id="B9379D16FFB18C1C58B5789EFE965292" italics="true" pageId="61" pageNumber="62">P. stenocephalus</emphasis>
</taxonomicName>
, but different as follows: body color castaneous-piceous to piceous-black, basomedial area of reddish to reddish-brunneous. Head similar to that of
<taxonomicName id="4C433A87FFB18C1C5E2978EBFC7752B8" authorityName="Scheerpeltz" authorityYear="1972" box="[793,991,480,505]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="stenocephalus">
<emphasis id="B9379D16FFB18C1C5E2978EBFC7752B8" box="[793,991,480,505]" italics="true" pageId="61" pageNumber="62">P. stenocephalus</emphasis>
</taxonomicName>
, but submentum 1.5 times as long as mentum. Pronotum about as long as wide (PW/PL=1.0), and dorsal rows of punstures each with five punctures. Tergum 8 truncate apically in both sexes.
</paragraph>
<paragraph id="8BFC4104FFB18C1C5DF57B5BFD52500B" blockId="61.[151,1437,330,2002]" pageId="61" pageNumber="62">
Male genitalia. Sternum 8 similar to
<taxonomicName id="4C433A87FFB18C1C5FB47B5BFD4D5128" authorityName="Herman" authorityYear="2001" box="[644,741,592,617]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="argus">
<emphasis id="B9379D16FFB18C1C5FB47B5BFD4D5128" box="[644,741,592,617]" italics="true" pageId="61" pageNumber="62">P. argus</emphasis>
</taxonomicName>
, deeply emarginate apically.Tergum 10 subacute apically (
<figureCitation id="13785D81FFB18C1C5D907B7EFEA051CE" box="[160,264,629,655]" captionStart="FIGURES 116–143" captionStartId="51.[151,270,1778,1802]" captionTargetBox="[382,1201,196,1755]" captionTargetId="figure-149@51.[367,1205,189,1767]" captionTargetPageId="51" captionText="FIGURES 116–143. P. rubromaculatus: sternum 8, male (116); sternum 9 (117); aedoeagus, dorsal view (118); tergum 10, female (119); gonocoxites of female genital segment (120). P. ruficauda: sternum 8, male (121); sternum 9 (122); tergum 10, male (123); aedoeagus, lateral view (124); aedoeagus, dorsal view (125); tergum 10, female (126); gonocoxites of female genital segment (127). P. sericans: sternum 9 (128); tergum 10, male (129); aedoeagus, dorsal view (130); paramere, ventral side (131). P. stenocephalus: sternum 9 (132); tergum 10, male (133); aedoeagus, dorsal view (134); paramere, ventral side (135); aedoeagus, lateral view (136); tergum 10, female (137); gonocoxites of female genital segment (138). P. varians: tergum 10, male (139); sternum 9 (140); aedoeagus, lateral view (141); paramere, ventral side (142); aedoeagus, dorsal view (143); gonocoxites of female genital segment (144)." pageId="61" pageNumber="62">Fig. 139</figureCitation>
), with two long subapical macrosetae and two apical setae. Sternum 9 similar to
<taxonomicName id="4C433A87FFB18C1C59FF7B7EFA3C51CF" authorityName="Scheerpeltz" authorityYear="1972" box="[1231,1428,629,654]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="stenocephalus">
<emphasis id="B9379D16FFB18C1C59FF7B7EFA3C51CF" box="[1231,1428,629,654]" italics="true" pageId="61" pageNumber="62">P. stenocephalus</emphasis>
</taxonomicName>
, deeply emarginate apically, with two long subapical macrosetae, and two apical setae at each side (
<figureCitation id="13785D81FFB18C1C58177B91FA2751F5" box="[1319,1423,666,692]" captionStart="FIGURES 116–143" captionStartId="51.[151,270,1778,1802]" captionTargetBox="[382,1201,196,1755]" captionTargetId="figure-149@51.[367,1205,189,1767]" captionTargetPageId="51" captionText="FIGURES 116–143. P. rubromaculatus: sternum 8, male (116); sternum 9 (117); aedoeagus, dorsal view (118); tergum 10, female (119); gonocoxites of female genital segment (120). P. ruficauda: sternum 8, male (121); sternum 9 (122); tergum 10, male (123); aedoeagus, lateral view (124); aedoeagus, dorsal view (125); tergum 10, female (126); gonocoxites of female genital segment (127). P. sericans: sternum 9 (128); tergum 10, male (129); aedoeagus, dorsal view (130); paramere, ventral side (131). P. stenocephalus: sternum 9 (132); tergum 10, male (133); aedoeagus, dorsal view (134); paramere, ventral side (135); aedoeagus, lateral view (136); tergum 10, female (137); gonocoxites of female genital segment (138). P. varians: tergum 10, male (139); sternum 9 (140); aedoeagus, lateral view (141); paramere, ventral side (142); aedoeagus, dorsal view (143); gonocoxites of female genital segment (144)." pageId="61" pageNumber="62">Fig. 140</figureCitation>
). Aedeagus with median lobe narrowed from base of paramere, apex of median lobe subtruncate apically (in dorsal view) and simple in its apical fourth (in lateral view); paramere elongate and entire, leaf-like, reaching four fifths of median lobe length, with two lateral rows of sensory peg setae joined at apex (
<figureCitation id="13785D81FFB18C1C59E97A01FA2C5065" box="[1241,1412,778,804]" captionStart="FIGURES 116–143" captionStartId="51.[151,270,1778,1802]" captionTargetBox="[382,1201,196,1755]" captionTargetId="figure-149@51.[367,1205,189,1767]" captionTargetPageId="51" captionText="FIGURES 116–143. P. rubromaculatus: sternum 8, male (116); sternum 9 (117); aedoeagus, dorsal view (118); tergum 10, female (119); gonocoxites of female genital segment (120). P. ruficauda: sternum 8, male (121); sternum 9 (122); tergum 10, male (123); aedoeagus, lateral view (124); aedoeagus, dorsal view (125); tergum 10, female (126); gonocoxites of female genital segment (127). P. sericans: sternum 9 (128); tergum 10, male (129); aedoeagus, dorsal view (130); paramere, ventral side (131). P. stenocephalus: sternum 9 (132); tergum 10, male (133); aedoeagus, dorsal view (134); paramere, ventral side (135); aedoeagus, lateral view (136); tergum 10, female (137); gonocoxites of female genital segment (138). P. varians: tergum 10, male (139); sternum 9 (140); aedoeagus, lateral view (141); paramere, ventral side (142); aedoeagus, dorsal view (143); gonocoxites of female genital segment (144)." pageId="61" pageNumber="62">Figs. 141–143</figureCitation>
). Internal sac with no distinctly sclerotized structures.
</paragraph>
<paragraph id="8BFC4104FFB18C1C5DF57A5EFEBB50FB" blockId="61.[151,1437,330,2002]" pageId="61" pageNumber="62">Female genitalia. Tergum 10 subacute medio-apically, with four long subapical setae and without apical setae. Second gonocoxite with 2–5 strong setae along its outer margin, and stylus with two long apical setae (Fig. 144).</paragraph>
<paragraph id="8BFC4104FFB18C1C5DF57ACEFEF25736" blockId="61.[151,1437,330,2002]" pageId="61" pageNumber="62">
<emphasis id="B9379D16FFB18C1C5DF57ACEFDB9509E" bold="true" box="[197,529,965,991]" pageId="61" pageNumber="62">Geographical distribution.</emphasis>
<taxonomicName id="4C433A87FFB18C1C5F287ACEFD5E509F" baseAuthorityName="Paykull" baseAuthorityYear="1789" box="[536,758,965,990]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="varians">
<emphasis id="B9379D16FFB18C1C5F287ACEFD5E509F" box="[536,758,965,990]" italics="true" pageId="61" pageNumber="62">Philonthus varians</emphasis>
</taxonomicName>
is a Palaearctic species which was probably introduced in the New World during the second half of the 19
<superScript id="7C36EC4CFFB18C1C5FC27AE1FD5750B9" attach="left" box="[754,767,1002,1016]" fontSize="6" pageId="61" pageNumber="62">th</superScript>
century (
<bibRefCitation id="EFD23CF5FFB18C1C5E497AE6FB875746" author="Smetana, A." box="[889,1071,1005,1031]" pageId="61" pageNumber="62" pagination="1 - 946" refId="ref52003" refString="Smetana, A. (1995) Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae): Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International, 3, 1 - 946." type="journal article" year="1995">Smetana 1995</bibRefCitation>
;
<bibRefCitation id="EFD23CF5FFB18C1C590E7AE6FB555746" author="Herman, L. H." box="[1086,1277,1005,1031]" pageId="61" pageNumber="62" pagination="2441 - 3020" refId="ref50050" refString="Herman, L. H. (2001 b) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. Part V. Bulletin of the American Museum of Natural History, 265, 2441 - 3020." type="journal article" year="2001">Herman 2001b</bibRefCitation>
). It was first recorded in southern South America by
<bibRefCitation id="EFD23CF5FFB18C1C5FB67D19FCDB576D" author="Scheerpeltz, O." box="[646,883,1042,1068]" pageId="61" pageNumber="62" refId="ref51351" refString="Scheerpeltz, O. (1972) Wissenschaftliche Ergebnisse der Studienreise von Gy. Topal nach Sudwest-Argentinien (Coleoptera: Staphylinidae). Folia Entomologica Hungarica (N. S.) 25 (Suppl.), 268 pp., 5 pls." type="book" year="1972">Scheerpeltz (1972)</bibRefCitation>
. After examining material of southern South American species of
<taxonomicName id="4C433A87FFB18C1C5CA47D33FDBD5710" baseAuthorityName="Smetana" baseAuthorityYear="1995" box="[404,533,1080,1105]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="B9379D16FFB18C1C5CA47D33FDBD5710" box="[404,533,1080,1105]" italics="true" pageId="61" pageNumber="62">Philonthus</emphasis>
</taxonomicName>
from different collections, I have only found specimens of
<taxonomicName id="4C433A87FFB18C1C59D27D33FAF25710" baseAuthorityName="Paykull" baseAuthorityYear="1789" box="[1250,1370,1080,1105]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="varians">
<emphasis id="B9379D16FFB18C1C59D27D33FAF25710" box="[1250,1370,1080,1105]" italics="true" pageId="61" pageNumber="62">P. varians</emphasis>
</taxonomicName>
from
<collectingCountry id="F3540194FFB18C1C5DA77D56FF7F5736" box="[151,215,1117,1143]" name="Chile" pageId="61" pageNumber="62">Chile</collectingCountry>
(
<figureCitation id="13785D81FFB18C1C5DD67D56FEE55736" box="[230,333,1117,1143]" captionStart="FIGURE 149" captionStartId="58.[151,255,1833,1857]" captionTargetBox="[316,1281,182,1824]" captionTargetId="figure-85@58.[306,1281,182,1834]" captionTargetPageId="58" captionText="FIGURE 149. Distribution of P. figulus, P. jenseni, P. ruficauda and P. varians in southern South America." pageId="61" pageNumber="62">Fig. 149</figureCitation>
).
</paragraph>
<paragraph id="8BFC4104FFB18C1C5DF57D89FC2A564D" blockId="61.[151,1437,330,2002]" pageId="61" pageNumber="62">
<emphasis id="B9379D16FFB18C1C5DF57D89FEFB57DD" bold="true" box="[197,339,1154,1180]" pageId="61" pageNumber="62">Bionomics.</emphasis>
<taxonomicName id="4C433A87FFB18C1C5C6C7D88FDEB57DD" baseAuthorityName="Paykull" baseAuthorityYear="1789" box="[348,579,1155,1180]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="varians">
<emphasis id="B9379D16FFB18C1C5C6C7D88FDEB57DD" box="[348,579,1155,1180]" italics="true" pageId="61" pageNumber="62">Philonthus varians</emphasis>
</taxonomicName>
is found in decaying organic materials, such as carrion, dung, weeds, rotting tissues, compost piles, etc. Specimens have been collected in flight intercept traps, carrion and dung baits. Trapping data suggest that this species is attracted both to carrion and dung.
<bibRefCitation id="EFD23CF5FFB18C1C596C7DC6FB5E57A6" author="Frank, J. H." box="[1116,1270,1229,1255]" pageId="61" pageNumber="62" pagination="1 - 118" refId="ref49676" refString="Frank, J. H. (1982) The parasites of the Staphylinidae (Coleoptera). A contribution towards an encyclopedia of the Staphylinidae. University of Florida Agr. Exp. Stations, Technical Bulletin, 824, i - vii + 1 - 118 pp." type="book chapter" year="1982">Frank (1982)</bibRefCitation>
also cites this species as parasitized by Laboulbeniales (Fungi, Ascomycetes).
</paragraph>
</subSubSection>
<subSubSection id="C359128FFFB18C1C5DF57C13FC2D5493" pageId="61" pageNumber="62" type="materials_examined">
<paragraph id="8BFC4104FFB18C1C5DF57C13FCEF563D" blockId="61.[151,1437,330,2002]" pageId="61" pageNumber="62">
<emphasis id="B9379D16FFB18C1C5DF57C13FED35673" bold="true" box="[197,379,1304,1330]" pageId="61" pageNumber="62">
<typeStatus id="54F8FFA6FFB18C1C5DF57C13FEAD5673" box="[197,261,1304,1330]" pageId="61" pageNumber="62">Type</typeStatus>
material.
</emphasis>
<bibRefCitation id="EFD23CF5FFB18C1C5CB37C13FDF75673" author="Paykull, G. de" box="[387,607,1304,1330]" pageId="61" pageNumber="62" refId="ref50949" refString="Paykull, G. de (1789) Monographia Staphylinorum Sueciae. Johann Edman, Upsaliae, 8 + 81 pp." type="book" year="1789">Paykull (1789: 45)</bibRefCitation>
described the species from specimens from
<collectingCountry id="F3540194FFB18C1C595C7C13FB655673" box="[1132,1229,1304,1330]" name="Sweden" pageId="61" pageNumber="62">Sweden</collectingCountry>
. I was not able to see the original material but there is little doubt that this species, as interpreted by modern authors (e.g. Lohse 1964;
<bibRefCitation id="EFD23CF5FFB18C1C5DD17C69FE21563D" author="Smetana, A." box="[225,393,1378,1404]" pageId="61" pageNumber="62" pagination="1 - 946" refId="ref52003" refString="Smetana, A. (1995) Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae): Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International, 3, 1 - 946." type="journal article" year="1995">Smetana 1995</bibRefCitation>
) is the species that Paykull described.
</paragraph>
<paragraph id="8BFC4104FFB18C1C5DF57C83FCD256E3" blockId="61.[151,1437,330,2002]" box="[197,890,1415,1442]" pageId="61" pageNumber="62">
<emphasis id="B9379D16FFB18C1C5DF57C83FE6D56E3" bold="true" box="[197,453,1416,1442]" pageId="61" pageNumber="62">Additional material:</emphasis>
<specimenCount id="9D458A8DFFB18C1C5CFC7C83FDC356E3" box="[460,619,1416,1442]" pageId="61" pageNumber="62" type="generic">21 specimens</specimenCount>
(
<specimenCount id="9D458A8DFFB18C1C5F4C7C83FD1C56DA" box="[636,692,1415,1442]" pageId="61" pageNumber="62" type="male">11 ♂</specimenCount>
,
<specimenCount id="9D458A8DFFB18C1C5FF27C83FD4456DA" box="[706,748,1415,1442]" pageId="61" pageNumber="62" type="female">8 ♀</specimenCount>
, 2 unsexed)
</paragraph>
<paragraph id="8BFC4104FFB18C1C5DF57CA6FC2D5493" blockId="61.[151,1437,330,2002]" pageId="61" pageNumber="62">
<materialsCitation id="3B2B4B59FFB18C1C5DF57CA6FDCA551D" collectingDate="1985-12-15" collectingDateMax="1997-02-05" collectingDateMin="1985-12-15" collectionCode="FMNH" collectorName="A. Newton &amp; M. Thayer &amp; P. N. Conguillio" country="Chile" elevation="310" location="North Shore Lago Villarica" municipality="Bellavista" pageId="61" pageNumber="62" specimenCount="7" specimenCount-female="3" specimenCount-male="4" stateProvince="Cautin">
<emphasis id="B9379D16FFB18C1C5DF57CA6FE345686" bold="true" box="[197,412,1453,1479]" pageId="61" pageNumber="62">
<collectingCountry id="F3540194FFB18C1C5DF57CA6FE835686" box="[197,299,1453,1479]" name="Chile" pageId="61" pageNumber="62">CHILE</collectingCountry>
:
<collectingRegion id="49878FE6FFB18C1C5C097CA6FE3F5686" box="[313,407,1453,1479]" country="Chile" name="Cautin" pageId="61" pageNumber="62">Cautín</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6B98DB7EFFB18C1C5C957CA6FD8B5686" box="[421,547,1453,1479]" pageId="61" pageNumber="62">Bellavista</collectingMunicipality>
,
<location id="8E9C17DFFFB18C1C5F017CA6FC2C5686" LSID="urn:lsid:plazi:treatment:03EAF012FFB18C1C5DA7799CFC2D5493:8E9C17DFFFB18C1C5F017CA6FC2C5686" box="[561,900,1453,1479]" country="Chile" municipality="Bellavista" name="North Shore Lago Villarica" pageId="61" pageNumber="62" stateProvince="Cautin">North Shore Lago Villarica</location>
,
<date id="FFFD67C4FFB18C1C5EA27CA6FBF05686" box="[914,1112,1453,1479]" pageId="61" pageNumber="62" value="1985-12-15" valueMax="1985-12-29" valueMin="1985-12-15">
<collectingDate id="EFB99E2CFFB18C1C5EA27CA6FBF05686" box="[914,1112,1453,1479]" pageId="61" pageNumber="62" value="1985-12-15" valueMax="1985-12-29" valueMin="1985-12-15">15–29.XII.1985</collectingDate>
</date>
,
<collectorName id="26B624D2FFB18C1C59567CA6FB435686" box="[1126,1259,1453,1479]" pageId="61" pageNumber="62">A. Newton</collectorName>
&amp;
<collectorName id="26B624D2FFB18C1C58247CA6FA3F5686" box="[1300,1431,1453,1479]" pageId="61" pageNumber="62">M. Thayer</collectorName>
,
<quantity id="4CBBECE1FFB18C1C5DA77CD9FF7456AD" box="[151,220,1490,1516]" metricMagnitude="2" metricUnit="m" metricValue="3.1" pageId="61" pageNumber="62" unit="m" value="310.0">
<elevation id="006EA637FFB18C1C5DA77CD9FF7456AD" box="[151,220,1490,1516]" metricMagnitude="2" metricUnit="m" metricValue="3.1" pageId="61" pageNumber="62" unit="m" value="310.0">310m</elevation>
</quantity>
,
<collectingMethod id="52023913FFB18C1C5DD97CD9FED956AD" box="[233,369,1490,1516]" pageId="61" pageNumber="62">carrion trap</collectingMethod>
(squid),
<specimenCount id="9D458A8DFFB18C1C5CEB7CD9FDAE56A7" box="[475,518,1490,1516]" pageId="61" pageNumber="62" type="male">1 ♂</specimenCount>
(
<collectionCode id="ED52D9C1FFB18C1C5F257CD9FDD856AD" box="[533,624,1490,1516]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="61" pageNumber="62" type="Museum">FMNH</collectionCode>
);
<collectorName id="26B624D2FFB18C1C5FB57CD9FC9556AD" box="[645,829,1490,1516]" pageId="61" pageNumber="62">P.N. Conguillio</collectorName>
,
<locationDeviation id="9A4E5826FFB18C1C5E7A7CD9FBF356AD" box="[842,1115,1490,1516]" location="Laguna Captren" pageId="61" pageNumber="62">
<quantity id="4CBBECE1FFB18C1C5E7A7CD9FC3C56AD" box="[842,916,1490,1516]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="61" pageNumber="62" unit="km" value="1.5">1.5km</quantity>
Laguna Captren
</locationDeviation>
,
<date id="FFFD67C4FFB18C1C59577CD9FB5256AD" box="[1127,1274,1490,1516]" pageId="61" pageNumber="62" value="1996-12-23">
<collectingDate id="EFB99E2CFFB18C1C59577CD9FB5256AD" box="[1127,1274,1490,1516]" pageId="61" pageNumber="62" value="1996-12-23">23.XII.1996</collectingDate>
</date>
–
<date id="FFFD67C4FFB18C1C583A7CD9FADB56AD" box="[1290,1395,1490,1516]" pageId="61" pageNumber="62" value="1997-02-05">
<collectingDate id="EFB99E2CFFB18C1C583A7CD9FADB56AD" box="[1290,1395,1490,1516]" pageId="61" pageNumber="62" value="1997-02-05">5.II.1997</collectingDate>
</date>
,
<collectorName id="26B624D2FFB18C1C584F7CD9FF5E5553" pageId="61" pageNumber="62">A. Newton</collectorName>
&amp;
<collectorName id="26B624D2FFB18C1C5C2E7CF3FE375553" box="[286,415,1528,1554]" pageId="61" pageNumber="62">M. Thayer</collectorName>
,
<quantity id="4CBBECE1FFB18C1C5C9C7CF3FDAA5553" box="[428,514,1528,1554]" metricMagnitude="3" metricUnit="m" metricValue="1.365" pageId="61" pageNumber="62" unit="m" value="1365.0">
<elevation id="006EA637FFB18C1C5C9C7CF3FDAA5553" box="[428,514,1528,1554]" metricMagnitude="3" metricUnit="m" metricValue="1.365" pageId="61" pageNumber="62" unit="m" value="1365.0">1365m</elevation>
</quantity>
,
<collectingMethod id="52023913FFB18C1C5F3F7CF3FD545553" box="[527,764,1528,1554]" pageId="61" pageNumber="62">flight intercept trap</collectingMethod>
,
<taxonomicName id="4C433A87FFB18C1C5E3A7CF3FACB5553" authority="Payk Det. M. K. Thayer 2004" authorityName="Payk Det. M. K. Thayer" authorityYear="2004" box="[778,1379,1528,1554]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="varians">
<emphasis id="B9379D16FFB18C1C5E3A7CF3FC465550" box="[778,1006,1528,1553]" italics="true" pageId="61" pageNumber="62">Philonthus varians</emphasis>
Payk Det. M. K. Thayer 2004
</taxonomicName>
,
<specimenCount id="9D458A8DFFB18C1C58417CF3FA35554A" box="[1393,1437,1527,1554]" pageId="61" pageNumber="62" type="male">1 ♂</specimenCount>
(
<collectionCode id="ED52D9C1FFB18C1C5DAF7F16FF565576" box="[159,254,1565,1591]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="61" pageNumber="62" type="Museum">FMNH</collectionCode>
); Volcan Villarica, road to ski center,
<date id="FFFD67C4FFB18C1C5FE67F16FCD85576" box="[726,880,1565,1591]" pageId="61" pageNumber="62" value="1996-12-23">
<collectingDate id="EFB99E2CFFB18C1C5FE67F16FCD85576" box="[726,880,1565,1591]" pageId="61" pageNumber="62" value="1996-12-23">23.XII.1996</collectingDate>
</date>
–
<date id="FFFD67C4FFB18C1C5EB27F16FC475576" box="[898,1007,1565,1591]" pageId="61" pageNumber="62" value="1997-02-05">
<collectingDate id="EFB99E2CFFB18C1C5EB27F16FC475576" box="[898,1007,1565,1591]" pageId="61" pageNumber="62" value="1997-02-05">5.II.1997</collectingDate>
</date>
,
<collectorName id="26B624D2FFB18C1C5ECE7F16FB2D5576" box="[1022,1157,1565,1591]" pageId="61" pageNumber="62">A. Newton</collectorName>
&amp;
<collectorName id="26B624D2FFB18C1C599E7F16FA9B5576" box="[1198,1331,1565,1591]" pageId="61" pageNumber="62">M. Thayer</collectorName>
,
<quantity id="4CBBECE1FFB18C1C58717F16FA3F5576" box="[1345,1431,1565,1591]" metricMagnitude="3" metricUnit="m" metricValue="1.18" pageId="61" pageNumber="62" unit="m" value="1180.0">
<elevation id="006EA637FFB18C1C58717F16FA3F5576" box="[1345,1431,1565,1591]" metricMagnitude="3" metricUnit="m" metricValue="1.18" pageId="61" pageNumber="62" unit="m" value="1180.0">1180m</elevation>
</quantity>
,
<collectingMethod id="52023913FFB18C1C5DA77F49FE88551D" box="[151,288,1602,1628]" pageId="61" pageNumber="62">carrion trap</collectingMethod>
(squid),
<specimenCount id="9D458A8DFFB18C1C5CBA7F49FE1E5517" box="[394,438,1602,1628]" pageId="61" pageNumber="62" type="male">2 ♂</specimenCount>
,
<specimenCount id="9D458A8DFFB18C1C5CF47F49FE585517" box="[452,496,1602,1628]" pageId="61" pageNumber="62" type="female">3 ♀</specimenCount>
(
<collectionCode id="ED52D9C1FFB18C1C5CCF7F49FDF3551D" box="[511,603,1602,1628]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="61" pageNumber="62" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B2B4B59FFB18C1C5F5E7F49FA30551D" box="[622,1432,1602,1628]" collectingDate="1995-12-01" collectionCode="MNHN" collectorName="P. Comadres &amp; S. Roitman" country="Chile" location="Chillan" pageId="61" pageNumber="62" specimenCount="1" stateProvince="Chillan">
<emphasis id="B9379D16FFB18C1C5F5E7F49FD7D551D" bold="true" box="[622,725,1602,1628]" pageId="61" pageNumber="62">
<collectingRegion id="49878FE6FFB18C1C5F5E7F49FD78551D" box="[622,720,1602,1628]" country="Chile" name="Chillan" pageId="61" pageNumber="62">Chillán</collectingRegion>
:
</emphasis>
<collectorName id="26B624D2FFB18C1C5FED7F49FCDC551D" box="[733,884,1602,1628]" pageId="61" pageNumber="62">P. Comadres</collectorName>
,
<date id="FFFD67C4FFB18C1C5EB07F49FC50551D" box="[896,1016,1602,1628]" pageId="61" pageNumber="62" value="1995-12-01">
<collectingDate id="EFB99E2CFFB18C1C5EB07F49FC50551D" box="[896,1016,1602,1628]" pageId="61" pageNumber="62" value="1995-12-01">12.1.1995</collectingDate>
</date>
,
<collectorName id="26B624D2FFB18C1C59357F49FB25551D" box="[1029,1165,1602,1628]" pageId="61" pageNumber="62">S. Roitman</collectorName>
,
<specimenCount id="9D458A8DFFB18C1C59AA7F49FAB7551D" box="[1178,1311,1602,1628]" pageId="61" pageNumber="62" type="generic">1 specimen</specimenCount>
(
<collectionCode id="ED52D9C1FFB18C1C581F7F49FA38551D" box="[1327,1424,1602,1628]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="61" pageNumber="62">MNHN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B2B4B59FFB18C1C5DA77F63FC50558D" collectingDate="2002-12" collectionCode="FMNH" collectorName="Castro, S. &amp; Solodovnikov &amp; Newton &amp; Chani &amp; Thayer" country="Chile" elevation="600" location="Coihaique" longitude="-71.6895" municipality="Cabanas Trayen" pageId="61" pageNumber="62" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Chiloe">
<emphasis id="B9379D16FFB18C1C5DA77F63FF5A55C3" bold="true" box="[151,242,1640,1666]" pageId="61" pageNumber="62">
<collectingRegion id="49878FE6FFB18C1C5DA77F63FF4555C3" box="[151,237,1640,1666]" country="Chile" name="Chiloe" pageId="61" pageNumber="62">Chiloe</collectingRegion>
:
</emphasis>
<collectorName id="26B624D2FFB18C1C5DC97F63FEC455C3" box="[249,364,1640,1666]" pageId="61" pageNumber="62">Castro, S.</collectorName>
of
<collectingMunicipality id="6B98DB7EFFB18C1C5CA37F63FDFB55C3" box="[403,595,1640,1666]" pageId="61" pageNumber="62">Cabañas Trayen</collectingMunicipality>
,
<date id="FFFD67C4FFB18C1C5F6F7F63FD6555C3" box="[607,717,1640,1666]" pageId="61" pageNumber="62" value="2002-12">
<collectingDate id="EFB99E2CFFB18C1C5F6F7F63FD6555C3" box="[607,717,1640,1666]" pageId="61" pageNumber="62" value="2002-12">XII.2002</collectingDate>
</date>
,
<collectorName id="26B624D2FFB18C1C5FE87F63FCD555C3" box="[728,893,1640,1666]" pageId="61" pageNumber="62">Solodovnikov</collectorName>
,
<collectorName id="26B624D2FFB18C1C5EB97F63FC4155C3" box="[905,1001,1640,1666]" pageId="61" pageNumber="62">Newton</collectorName>
,
<collectorName id="26B624D2FFB18C1C5EC57F63FB9655C3" box="[1013,1086,1640,1666]" pageId="61" pageNumber="62">Chani</collectorName>
,
<collectorName id="26B624D2FFB18C1C597A7F63FB3755C3" box="[1098,1183,1640,1666]" pageId="61" pageNumber="62">Thayer</collectorName>
, 38° 67' S,
<geoCoordinate id="EE7727C3FFB18C1C58147F63FF1D55E6" degrees="71" direction="west" minutes="41.37" orientation="longitude" pageId="61" pageNumber="62" precision="9" value="-71.6895">71° 41.37' W</geoCoordinate>
,
<specimenCount id="9D458A8DFFB18C1C5D8F7F86FF4155E0" box="[191,233,1677,1703]" pageId="61" pageNumber="62" type="male">2 ♂</specimenCount>
,
<specimenCount id="9D458A8DFFB18C1C5DC77F86FE8A55E0" box="[247,290,1677,1703]" pageId="61" pageNumber="62" type="female">2 ♀</specimenCount>
(
<collectionCode id="ED52D9C1FFB18C1C5C017F86FE2555E6" box="[305,397,1677,1703]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="61" pageNumber="62" type="Museum">FMNH</collectionCode>
).
<location id="8E9C17DFFFB18C1C5C907F86FDB555E6" LSID="urn:lsid:plazi:treatment:03EAF012FFB18C1C5DA7799CFC2D5493:8E9C17DFFFB18C1C5C907F86FDB555E6" box="[416,541,1677,1703]" country="Chile" longitude="-71.6895" municipality="Cabanas Trayen" name="Coihaique" pageId="61" pageNumber="62" stateProvince="Chiloe">Coihaique</location>
:
<locationDeviation id="9A4E5826FFB18C1C5F187F86FCE555E6" box="[552,845,1677,1703]" location="Coihaique" pageId="61" pageNumber="62">
<quantity id="4CBBECE1FFB18C1C5F187F86FDC255E6" box="[552,618,1677,1703]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="61" pageNumber="62" unit="km" value="20.0">20km</quantity>
ENE of
<locationDeviation id="9A4E5826FFB18C1C5FE27F86FCE555E6" box="[722,845,1677,1703]" pageId="61" pageNumber="62">Coihaique</locationDeviation>
</locationDeviation>
,
<locationDeviation id="9A4E5826FFB18C1C5E6A7F86FC5F55E6" box="[858,1015,1677,1703]" pageId="61" pageNumber="62">Dos Lagunas</locationDeviation>
,
<quantity id="4CBBECE1FFB18C1C59337F86FBE055E6" box="[1027,1096,1677,1703]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="61" pageNumber="62" unit="m" value="600.0">
<elevation id="006EA637FFB18C1C59337F86FBE055E6" box="[1027,1096,1677,1703]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="61" pageNumber="62" unit="m" value="600.0">600m</elevation>
</quantity>
,
<collectingMethod id="52023913FFB18C1C59647F86FB7455E6" box="[1108,1244,1677,1703]" pageId="61" pageNumber="62">carrion trap</collectingMethod>
beech groves in steppe,
<taxonomicName id="4C433A87FFB18C1C5DC97FB8FCE9558D" authority="Pk Det. Chani-Posse 2006" authorityName="Pk Det. Chani-Posse" authorityYear="2006" box="[249,833,1714,1740]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="varians">
<emphasis id="B9379D16FFB18C1C5DC97FB8FE41558D" box="[249,489,1715,1740]" italics="true" pageId="61" pageNumber="62">Philonthus varians</emphasis>
Pk Det. Chani-Posse 2006
</taxonomicName>
,
<specimenCount id="9D458A8DFFB18C1C5E607FB9FCD75587" box="[848,895,1714,1740]" pageId="61" pageNumber="62" type="male">1 ♂</specimenCount>
(
<collectionCode id="ED52D9C1FFB18C1C5EBF7FB9FC47558D" box="[911,1007,1714,1740]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="61" pageNumber="62" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B2B4B59FFB18C1C59367FB9FC1755B3" collectingDate="1997-12-22" collectingDateMax="1997-12-31" collectingDateMin="1997-12-22" collectorName="E. Curico &amp; Col. J. E. Barriga" country="Chile" location="Zapallar" municipality="Hueca-Huecan" pageId="61" pageNumber="62" specimenCount="1" specimenCount-female="1" stateProvince="Curico">
<emphasis id="B9379D16FFB18C1C59367FB9FBC4558D" bold="true" box="[1030,1132,1714,1740]" pageId="61" pageNumber="62">
<collectingRegion id="49878FE6FFB18C1C59367FB9FBCE558D" box="[1030,1126,1714,1740]" country="Chile" name="Curico" pageId="61" pageNumber="62">Curicó</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6B98DB7EFFB18C1C59467FB9FA9A558D" box="[1142,1330,1714,1740]" pageId="61" pageNumber="62">Hueca-Huecan</collectingMunicipality>
, camino
<location id="8E9C17DFFFB18C1C5DA77FD3FF5455B3" LSID="urn:lsid:plazi:treatment:03EAF012FFB18C1C5DA7799CFC2D5493:8E9C17DFFFB18C1C5DA77FD3FF5455B3" box="[151,252,1752,1778]" country="Chile" municipality="Hueca-Huecan" name="Zapallar" pageId="61" pageNumber="62" stateProvince="Curico">Zapallar</location>
,
<quantity id="4CBBECE1FFB18C1C5C397FD3FEE455B3" box="[265,332,1752,1778]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="61" pageNumber="62" unit="km" value="10.0">10km</quantity>
<collectorName id="26B624D2FFB18C1C5C657FD3FE6355B3" box="[341,459,1752,1778]" pageId="61" pageNumber="62">E. Curicó</collectorName>
,
<date id="FFFD67C4FFB18C1C5CE87FD3FD3D55B3" box="[472,661,1752,1778]" pageId="61" pageNumber="62" value="1997-12-22" valueMax="1997-12-31" valueMin="1997-12-22">
<collectingDate id="EFB99E2CFFB18C1C5CE87FD3FD3D55B3" box="[472,661,1752,1778]" pageId="61" pageNumber="62" value="1997-12-22" valueMax="1997-12-31" valueMin="1997-12-22">22–31 dic 1997</collectingDate>
</date>
,
<specimenCount id="9D458A8DFFB18C1C5F937FD3FD6755AA" box="[675,719,1751,1778]" pageId="61" pageNumber="62" type="female">1 ♀</specimenCount>
(
<collectorName id="26B624D2FFB18C1C5FEF7FD3FC1F55B3" box="[735,951,1752,1778]" pageId="61" pageNumber="62">Col. J. E. Barriga</collectorName>
)
</materialsCitation>
.
<emphasis id="B9379D16FFB18C1C5EFC7FD3FC4655B3" bold="true" box="[972,1006,1752,1778]" pageId="61" pageNumber="62">La</emphasis>
<materialsCitation id="3B2B4B59FFB18C1C5EC77FD3FB165456" collectingDate="2004-02-15" collectingDateMax="2004-02-16" collectingDateMin="2004-02-15" collectionCode="FMNH" collectorName="A. Smith &amp; M. Paulsen &amp; J. Mondaca" country="Chile" elevation="1120" location="Pehuenco" municipality="Nahuelbuta" pageId="61" pageNumber="62" specimenCount="1" specimenCount-female="1" stateProvince="Araucania">
<emphasis id="B9379D16FFB18C1C5EC77FD3FB2E55B3" bold="true" box="[1015,1158,1752,1778]" pageId="61" pageNumber="62">
<collectingRegion id="49878FE6FFB18C1C5EC77FD3FB2955B3" box="[1015,1153,1752,1778]" country="Chile" name="Araucania" pageId="61" pageNumber="62">Araucania</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6B98DB7EFFB18C1C59BF7FD3FAB355B3" box="[1167,1307,1752,1778]" pageId="61" pageNumber="62">Nahuelbuta</collectingMunicipality>
,
<location id="8E9C17DFFFB18C1C58187FD3FA3355B3" LSID="urn:lsid:plazi:treatment:03EAF012FFB18C1C5DA7799CFC2D5493:8E9C17DFFFB18C1C58187FD3FA3355B3" box="[1320,1435,1752,1778]" country="Chile" municipality="Nahuelbuta" name="Pehuenco" pageId="61" pageNumber="62" stateProvince="Araucania">Pehuenco</location>
near campground,
<date id="FFFD67C4FFB18C1C5C417FF6FDBF5456" box="[369,535,1789,1815]" pageId="61" pageNumber="62" value="2004-02-15" valueMax="2004-02-16" valueMin="2004-02-15">
<collectingDate id="EFB99E2CFFB18C1C5C417FF6FDBF5456" box="[369,535,1789,1815]" pageId="61" pageNumber="62" value="2004-02-15" valueMax="2004-02-16" valueMin="2004-02-15">15–16.ii.2004</collectingDate>
</date>
,
<collectorName id="26B624D2FFB18C1C5F137FF6FD275456" box="[547,655,1789,1815]" pageId="61" pageNumber="62">A. Smith</collectorName>
,
<collectorName id="26B624D2FFB18C1C5FAB7FF6FC8A5456" box="[667,802,1789,1815]" pageId="61" pageNumber="62">M. Paulsen</collectorName>
,
<collectorName id="26B624D2FFB18C1C5E1D7FF6FC1F5456" box="[813,951,1789,1815]" pageId="61" pageNumber="62">J. Mondaca</collectorName>
,
<quantity id="4CBBECE1FFB18C1C5EF37FF6FBBE5456" box="[963,1046,1789,1815]" metricMagnitude="3" metricUnit="m" metricValue="1.1199999999999999" pageId="61" pageNumber="62" unit="m" value="1120.0">
<elevation id="006EA637FFB18C1C5EF37FF6FBBE5456" box="[963,1046,1789,1815]" metricMagnitude="3" metricUnit="m" metricValue="1.1199999999999999" pageId="61" pageNumber="62" unit="m" value="1120.0">1120m</elevation>
</quantity>
,
<specimenCount id="9D458A8DFFB18C1C59117FF6FBE45450" box="[1057,1100,1789,1815]" pageId="61" pageNumber="62" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED52D9C1FFB18C1C596B7FF6FB1E5456" box="[1115,1206,1789,1815]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="61" pageNumber="62" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B2B4B59FFB18C1C59F97FF6FC745423" collectingDate="1997-01-02" collectingDateMax="1997-01-28" collectingDateMin="1997-01-02" collectionCode="FMNH" collectorName="P. N. Vicente Perez Rosales &amp; Newton &amp; M. Thayer" country="Chile" elevation="125" latitude="-41.17" location="Petrohue" longLatPrecision="11" longitude="-72.45167" pageId="61" pageNumber="62" specimenCount="1" specimenCount-female="1" stateProvince="Llanquihue">
<emphasis id="B9379D16FFB18C1C59F97FF6FACB5456" bold="true" box="[1225,1379,1789,1815]" pageId="61" pageNumber="62">
<collectingRegion id="49878FE6FFB18C1C59F97FF6FAF65456" box="[1225,1374,1789,1815]" country="Chile" name="Llanquihue" pageId="61" pageNumber="62">Llanquihue</collectingRegion>
:
</emphasis>
<collectorName id="26B624D2FFB18C1C585B7FF6FE03547D" pageId="61" pageNumber="62">P.N. Vicente Perez Rosales</collectorName>
,
<locationDeviation id="9A4E5826FFB18C1C5C897E29FD1C547D" box="[441,692,1826,1852]" location="Ensenada" pageId="61" pageNumber="62">
<quantity id="4CBBECE1FFB18C1C5C897E29FDAD547D" box="[441,517,1826,1852]" metricMagnitude="3" metricUnit="m" metricValue="9.2" pageId="61" pageNumber="62" unit="km" value="9.2">9.2km</quantity>
NE Ensenada
</locationDeviation>
on road to
<location id="8E9C17DFFFB18C1C5E747E29FC1B547D" LSID="urn:lsid:plazi:treatment:03EAF012FFB18C1C5DA7799CFC2D5493:8E9C17DFFFB18C1C5E747E29FC1B547D" box="[836,947,1826,1852]" country="Chile" latitude="-41.17" longLatPrecision="11" longitude="-72.45167" name="Petrohue" pageId="61" pageNumber="62" stateProvince="Llanquihue">Petrohue</location>
,
<date id="FFFD67C4FFB18C1C5EF17E29FBFE547D" box="[961,1110,1826,1852]" pageId="61" pageNumber="62" value="1997-01-02" valueMax="1997-01-28" valueMin="1997-01-02">
<collectingDate id="EFB99E2CFFB18C1C5EF17E29FBFE547D" box="[961,1110,1826,1852]" pageId="61" pageNumber="62" value="1997-01-02" valueMax="1997-01-28" valueMin="1997-01-02">2–28.i.1997</collectingDate>
</date>
, A.
<collectorName id="26B624D2FFB18C1C59BB7E29FB42547D" box="[1163,1258,1826,1852]" pageId="61" pageNumber="62">Newton</collectorName>
&amp;
<collectorName id="26B624D2FFB18C1C58247E29FA30547D" box="[1300,1432,1826,1852]" pageId="61" pageNumber="62">M. Thayer</collectorName>
,
<quantity id="4CBBECE1FFB18C1C5DA77E43FF765423" box="[151,222,1864,1890]" metricMagnitude="2" metricUnit="m" metricValue="1.25" pageId="61" pageNumber="62" unit="m" value="125.0">
<elevation id="006EA637FFB18C1C5DA77E43FF765423" box="[151,222,1864,1890]" metricMagnitude="2" metricUnit="m" metricValue="1.25" pageId="61" pageNumber="62" unit="m" value="125.0">125m</elevation>
</quantity>
,
<geoCoordinate id="EE7727C3FFB18C1C5DDB7E43FE2A5423" box="[235,386,1864,1890]" degrees="41" direction="south" minutes="10.20" orientation="latitude" pageId="61" pageNumber="62" precision="9" value="-41.17">41° 10.20' S</geoCoordinate>
,
<geoCoordinate id="EE7727C3FFB18C1C5CA17E43FD9A5423" box="[401,562,1864,1890]" degrees="72" direction="west" minutes="27.10" orientation="longitude" pageId="61" pageNumber="62" precision="9" value="-72.45167">72° 27.10' W</geoCoordinate>
,
<collectingMethod id="52023913FFB18C1C5F0F7E43FD645423" box="[575,716,1864,1890]" pageId="61" pageNumber="62">carrion trap</collectingMethod>
(squid),
<specimenCount id="9D458A8DFFB18C1C5E0A7E43FCCF541A" box="[826,871,1863,1890]" pageId="61" pageNumber="62" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED52D9C1FFB18C1C5E477E43FC7D5423" box="[887,981,1864,1890]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="61" pageNumber="62" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B2B4B59FFB18C1C5ED97E43FB2E54ED" collectingDate="1986-12-18" collectingDateMax="2002-12-18" collectingDateMin="1986-12-18" collectionCode="FMNH" collectorName="A. Newton &amp; M. Thayer &amp; Solodovnikov" country="Chile" county="Parque Nac. Puyehue" elevation="965" latitude="-40.75" location="Bertin" longLatPrecision="1161" longitude="-72.3" municipality="Aguas Calientes" pageId="61" pageNumber="62" specimenCount="3" specimenCount-male="3" stateProvince="Osorno">
<emphasis id="B9379D16FFB18C1C5ED97E43FBF95423" bold="true" box="[1001,1105,1864,1890]" pageId="61" pageNumber="62">
<collectingRegion id="49878FE6FFB18C1C5ED97E43FBE45423" box="[1001,1100,1864,1890]" country="Chile" name="Osorno" pageId="61" pageNumber="62">Osorno</collectingRegion>
:
</emphasis>
<collectingCounty id="629D3988FFB18C1C596B7E43FACA5423" box="[1115,1378,1864,1890]" pageId="61" pageNumber="62">Parque Nac. Puyehue</collectingCounty>
,
<date id="FFFD67C4FFB18C1C585F7E43FE8154C6" pageId="61" pageNumber="62" value="1986-12-18" valueMax="1986-12-25" valueMin="1986-12-18">
<collectingDate id="EFB99E2CFFB18C1C585F7E43FE8154C6" pageId="61" pageNumber="62" value="1986-12-18" valueMax="1986-12-25" valueMin="1986-12-18">18– 25.XII.1986</collectingDate>
</date>
,
<collectorName id="26B624D2FFB18C1C5C047E66FE1D54C6" box="[308,437,1901,1927]" pageId="61" pageNumber="62">A. Newton</collectorName>
&amp;
<collectorName id="26B624D2FFB18C1C5CE97E66FDFF54C6" box="[473,599,1901,1927]" pageId="61" pageNumber="62">M. Thayer</collectorName>
,
<quantity id="4CBBECE1FFB18C1C5F527E66FD0F54C6" box="[610,679,1901,1927]" metricMagnitude="2" metricUnit="m" metricValue="9.65" pageId="61" pageNumber="62" unit="m" value="965.0">
<elevation id="006EA637FFB18C1C5F527E66FD0F54C6" box="[610,679,1901,1927]" metricMagnitude="2" metricUnit="m" metricValue="9.65" pageId="61" pageNumber="62" unit="m" value="965.0">965m</elevation>
</quantity>
,
<collectingMethod id="52023913FFB18C1C5F837E66FC9154C6" box="[691,825,1901,1927]" pageId="61" pageNumber="62">carrion trap</collectingMethod>
(squid),
<specimenCount id="9D458A8DFFB18C1C5E927E66FC6354C0" box="[930,971,1901,1927]" pageId="61" pageNumber="62" type="male">2 ♂</specimenCount>
(
<collectionCode id="ED52D9C1FFB18C1C5EEA7E66FB9D54C6" box="[986,1077,1901,1927]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="61" pageNumber="62" type="Museum">FMNH</collectionCode>
); vic.
<collectingMunicipality id="6B98DB7EFFB18C1C594B7E66FA9254C6" box="[1147,1338,1901,1927]" pageId="61" pageNumber="62">Aguas Calientes</collectingMunicipality>
sendero
<location id="8E9C17DFFFB18C1C5DA77E99FF4254ED" LSID="urn:lsid:plazi:treatment:03EAF012FFB18C1C5DA7799CFC2D5493:8E9C17DFFFB18C1C5DA77E99FF4254ED" box="[151,234,1938,1964]" country="Chile" county="Parque Nac. Puyehue" latitude="-40.75" longLatPrecision="1161" longitude="-72.3" municipality="Aguas Calientes" name="Bertin" pageId="61" pageNumber="62" stateProvince="Osorno">Bertin</location>
,
<date id="FFFD67C4FFB18C1C5DC97E99FE3F54ED" box="[249,407,1938,1964]" pageId="61" pageNumber="62" value="2002-12-18">
<collectingDate id="EFB99E2CFFB18C1C5DC97E99FE3F54ED" box="[249,407,1938,1964]" pageId="61" pageNumber="62" value="2002-12-18">18.XII.2002</collectingDate>
</date>
,
<collectorName id="26B624D2FFB18C1C5C977E99FDF254ED" box="[423,602,1938,1964]" pageId="61" pageNumber="62">Solodovnikov</collectorName>
,
<quantity id="4CBBECE1FFB18C1C5F597E99FD1B54ED" box="[617,691,1938,1964]" metricMagnitude="2" metricUnit="m" metricValue="9.65" pageId="61" pageNumber="62" unit="m" value="965.0">
<elevation id="006EA637FFB18C1C5F597E99FD1B54ED" box="[617,691,1938,1964]" metricMagnitude="2" metricUnit="m" metricValue="9.65" pageId="61" pageNumber="62" unit="m" value="965.0">965m</elevation>
</quantity>
,
<geoCoordinate id="EE7727C3FFB18C1C5FF27E99FC9254ED" box="[706,826,1938,1964]" degrees="40" direction="south" minutes="45" orientation="latitude" pageId="61" pageNumber="62" precision="925" value="-40.75">40° 45' S</geoCoordinate>
,
<geoCoordinate id="EE7727C3FFB18C1C5E7B7E99FC6554ED" box="[843,973,1938,1964]" degrees="72" direction="west" minutes="18" orientation="longitude" pageId="61" pageNumber="62" precision="925" value="-72.3">72° 18' W</geoCoordinate>
,
<specimenCount id="9D458A8DFFB18C1C5EEB7E99FBA254E7" box="[987,1034,1938,1964]" pageId="61" pageNumber="62" type="male">1 ♂</specimenCount>
(
<collectionCode id="ED52D9C1FFB18C1C592B7E99FBD454ED" box="[1051,1148,1938,1964]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="61" pageNumber="62" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B2B4B59FFB18C1C59A47E99FC295493" collectingDate="1953-10-29" collectionCode="FMNH, MNHN" collectorName="L. E. Pena" country="Chile" location="Pudahuel" pageId="61" pageNumber="62" specimenCount="2" specimenCount-male="1" stateProvince="Region Metropolitana">
<emphasis id="B9379D16FFB18C1C59A47E99FABC54ED" bold="true" box="[1172,1300,1938,1964]" pageId="61" pageNumber="62">
<collectingRegion id="49878FE6FFB18C1C59A47E99FAA654ED" box="[1172,1294,1938,1964]" country="Chile" name="Region Metropolitana" pageId="61" pageNumber="62">Santiago</collectingRegion>
:
</emphasis>
<location id="8E9C17DFFFB18C1C582E7E99FA3F54ED" LSID="urn:lsid:plazi:treatment:03EAF012FFB18C1C5DA7799CFC2D5493:8E9C17DFFFB18C1C582E7E99FA3F54ED" box="[1310,1431,1938,1964]" country="Chile" name="Pudahuel" pageId="61" pageNumber="62" stateProvince="Region Metropolitana">Pudahuel</location>
,
<date id="FFFD67C4FFB18C1C5DA77EB3FEBD5493" box="[151,277,1976,2002]" pageId="61" pageNumber="62" value="1953-10-29">
<collectingDate id="EFB99E2CFFB18C1C5DA77EB3FEBD5493" box="[151,277,1976,2002]" pageId="61" pageNumber="62" value="1953-10-29">29.X.1953</collectingDate>
</date>
,
<collectorName id="26B624D2FFB18C1C5C117EB3FE335493" box="[289,411,1976,2002]" pageId="61" pageNumber="62">L. E. Peña</collectorName>
leg.,
<specimenCount id="9D458A8DFFB18C1C5CEC7EB3FDAE548A" box="[476,518,1975,2002]" pageId="61" pageNumber="62" type="male">1 ♂</specimenCount>
(
<collectionCode id="ED52D9C1FFB18C1C5F257EB3FDD85493" box="[533,624,1976,2002]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="61" pageNumber="62" type="Museum">FMNH</collectionCode>
);
<specimenCount id="9D458A8DFFB18C1C5FB57EB3FCA15493" box="[645,777,1976,2002]" pageId="61" pageNumber="62" type="generic">1 specimen</specimenCount>
(
<collectionCode id="ED52D9C1FFB18C1C5E297EB3FCD25493" box="[793,890,1976,2002]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="61" pageNumber="62">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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