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<document id="6E79566C62E3F55704CC9C7FD11DB219" ID-CLB-Dataset="5890" ID-DOI="10.11646/zootaxa.3773.1.1" ID-GBIF-Dataset="6da85b06-8386-41bc-9b34-b16eba941eab" ID-ISSN="1175-5326" ID-Zenodo-Dep="4909796" ID-ZooBank="19F28753-B2D0-4D1F-9D47-88886F7333FD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623114397150" checkinUser="felipe" docAuthor="Ng, Peter K. L. &amp; Rahayu, Dwi Listyo" docDate="2014" docId="03E287AE5467E2198A9D45AEFC8E0A3F" docLanguage="en" docName="zt03773p063.pdf" docOrigin="Zootaxa 3773 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Raoulia calva Ng &amp; Rahayu 2014, sp. nov." docType="treatment" docVersion="7" lastPageNumber="35" masterDocId="FFDBFFD65445E23B8A0A4416FF8A0F11" masterDocTitle="Revision of the family Acidopsidae Števčić, 2005, and the systematic position of Typhlocarcinodes Alcock, 1900, Caecopilumnus Borradaile, 1902, and Raoulia Ng, 1987, with descriptions of two new genera and five new species (Crustacea: Brachyura: Goneplacoidea)" masterLastPageNumber="63" masterPageNumber="1" pageNumber="35" updateTime="1699003055515" updateUser="ExternalLinkService">
<mods:mods id="C79EC3081A6027DEFDC0281536BBE28C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BD535C693BB32A8F5DB476036B9B3F63">
<mods:title id="D26B0C75BCFC9AB9AFD71BECEB5DA722">Revision of the family Acidopsidae Števčić, 2005, and the systematic position of Typhlocarcinodes Alcock, 1900, Caecopilumnus Borradaile, 1902, and Raoulia Ng, 1987, with descriptions of two new genera and five new species (Crustacea: Brachyura: Goneplacoidea)</mods:title>
</mods:titleInfo>
<mods:name id="9DF09259B383F55F6511FB43F30C4F10" type="personal">
<mods:role id="005C82370D4522D5829E590E7C6C878F">
<mods:roleTerm id="04B36C122F93FA5062E74D78EFE749A0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D0B0E0F7C527204E82A3B2954602DF60">Ng, Peter K. L.</mods:namePart>
<mods:nameIdentifier id="152A0EE2170C0123561310F00F3DEBE7" type="email">dbsngkl@nus.edu.sg</mods:nameIdentifier>
</mods:name>
<mods:name id="8F607A1B9CE7C13BF59D7ADCFEA07344" type="personal">
<mods:role id="F2D0B115AA8CDCE86C1CC18F20792780">
<mods:roleTerm id="02CDB60941EED9C79492F186DBF32BDC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B9F13FDC19BEDC15C0358DB23B90B3C0">Rahayu, Dwi Listyo</mods:namePart>
<mods:nameIdentifier id="1D1FE202DA6B2A86D0E89377BDB65F08" type="email">dbsngkl@nus.edu.sg</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="BEC2A1696C3EAB630FA6EB751BCCF9A5">text</mods:typeOfResource>
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<mods:titleInfo id="07B3E421BAE0E468A8B5495BFC2423D5">
<mods:title id="14296446F1F0C6FDD462D3E63379D471">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="37F2EBFB763DC0FC18F87B5A82315B92">
<mods:date id="8452C15C717166CCC3559C6BF41437CF">2014</mods:date>
<mods:detail id="5F1B6F02B79DAC606025C54E84AA7BBC" type="pubDate">
<mods:number id="0EF159551667878846194164E29DE043">2014-03-03</mods:number>
</mods:detail>
<mods:detail id="632A51F4D92FAB28AC4FA3CFBA86C2D8" type="volume">
<mods:number id="A553BF77C846905C119AC95CD6CB43CA">3773</mods:number>
</mods:detail>
<mods:detail id="F995B60CD29BD2E9E497B4659F9D4D13" type="issue">
<mods:number id="F296ABAC626E7E81AFA9977F2B382661">1</mods:number>
</mods:detail>
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<mods:start id="72F035D83D9CEB7A1CBD04D2FECB44F1">1</mods:start>
<mods:end id="69D8A7C7F09B8D29A0C31BCDBA7DCC35">63</mods:end>
</mods:extent>
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<mods:classification id="36B1E4DF2E28498898A9E33DE439D6C1">journal article</mods:classification>
<mods:identifier id="36EAE6579971E5B5DEE587ED7C9DFC45" type="CLB-Dataset">5890</mods:identifier>
<mods:identifier id="A5F7C24BD41C6F674F482B8AAC928E6D" type="DOI">10.11646/zootaxa.3773.1.1</mods:identifier>
<mods:identifier id="29748B778A10610E08358CED970B61EE" type="GBIF-Dataset">6da85b06-8386-41bc-9b34-b16eba941eab</mods:identifier>
<mods:identifier id="DA19E3975A1BCC53BE16573B39E9035B" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="AF34937EBF4DAAD85985591CA56E62AD" type="Zenodo-Dep">4909796</mods:identifier>
<mods:identifier id="61BF1A18001EDD44FBC3274C870EACC0" type="ZooBank">19F28753-B2D0-4D1F-9D47-88886F7333FD</mods:identifier>
</mods:mods>
<treatment id="03E287AE5467E2198A9D45AEFC8E0A3F" ID-DOI="http://doi.org/10.5281/zenodo.4909778" ID-GBIF-Taxon="183228356" ID-Zenodo-Dep="4909778" LSID="urn:lsid:plazi:treatment:03E287AE5467E2198A9D45AEFC8E0A3F" httpUri="http://treatment.plazi.org/id/03E287AE5467E2198A9D45AEFC8E0A3F" lastPageNumber="35" pageId="34" pageNumber="35">
<subSubSection id="C35165335467E2198A9D45AEFE280EC3" box="[151,418,440,466]" pageId="34" pageNumber="35" type="nomenclature">
<paragraph id="8BF436B85467E2198A9D45AEFE280EC3" blockId="34.[151,418,440,499]" box="[151,418,440,466]" pageId="34" pageNumber="35">
<heading id="D0BC81D45467E2198A9D45AEFE280EC3" bold="true" box="[151,418,440,466]" fontSize="11" level="2" pageId="34" pageNumber="35" reason="4">
<emphasis id="B93FEAAA5467E2198A9D45AEFE280EC3" bold="true" box="[151,418,440,466]" pageId="34" pageNumber="35">
<taxonomicName id="4C4B4D3B5467E2198A9D45AEFEB40EC3" authority="Ng &amp; Rahayu, 2014" authorityName="Ng &amp; Rahayu" authorityYear="2014" box="[151,318,440,466]" class="Malacostraca" family="Chasmocarcinidae" genus="Raoulia" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="calva" status="sp. nov.">
<emphasis id="B93FEAAA5467E2198A9D45AEFEB40EC3" bold="true" box="[151,318,440,466]" italics="true" pageId="34" pageNumber="35">Raoulia calva</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20C57D15467E2198B4F45AEFE280EC3" box="[325,418,440,466]" pageId="34" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35165335467E2198A9D45CCFE1D0EE2" box="[151,407,474,499]" pageId="34" pageNumber="35" type="description">
<paragraph id="8BF436B85467E2198A9D45CCFE1D0EE2" blockId="34.[151,418,440,499]" box="[151,407,474,499]" pageId="34" pageNumber="35">
(
<figureCitation id="13702A3D5467E2198A9545CCFE8A0EE2" box="[159,256,474,499]" captionStart="FIGURE 5" captionStartId="11.[151,249,1549,1571]" captionTargetBox="[151,1436,193,1527]" captionTargetId="figure-230@11.[151,1436,193,1527]" captionTargetPageId="11" captionText="FIGURE 5. Right third maxillipeds. A, Acidops cessacii (A. Milne-Edwards, 1878), syntype female (14.1 × 10.0 mm) (NHM 99.3.23.66) Cape Verde (laterally inverted); B, Parapilumnus cristimanus (A. Milne-Edwards, 1873), male (16.3 × 12.5 mm) (ZRC 2003.90), Nansha Is. (Spratly Is.), South China Sea (laterally inverted); C, Raoulia limosa Ng, 1987, holotype male (8.3 × 6.3 mm) (ZMUC M58), off Marongas, Jolo, Philippines; D, Raoulia piroculata (Rathbun, 1911), lectotype female (7.3 × 5.5 mm) (USNM 41359a), Amirante Is., Indian Ocean; E, Raoulia galea sp. nov., holotype male (9.4 × 7.4 mm) (ZRC 2000.0980), Phuket, Thailand; F, Raoulia fortis sp. nov., holotype male (12.0 × 9.2 mm) (MNHN), Santo, Vanuatu (laterally inverted); G, Raoulia calva sp. nov., holotype male (6.6 × 5.2 mm) (ZRC 2013.1721), Society Is., French Polynesia; H, Caecopilumnus hirsutus (Borradaile, 1902), male (7.1 × 5.6 mm) (ZMA 261027), Flores, Indonesia (laterally inverted); I, Caecopilumnus crassipes (Tesch, 1918), male (11.8 × 10.1 mm) (ZRC 2013.1710) Lombok, Indonesia; J, Caecopilumnus loculatus sp. nov., holotype male (8.0 × 6.7 mm) (ZRC 2013.1723), Moorea, French Polynesia (laterally inverted); K, Thecaplax capillosa gen. et. sp. nov., holotype male (11.6 × 9.5 mm) (MZB), Lombok, Indonesia; L, Typhlocarcinodes integrifrons (Miers, 1881), lectotype male (8.5 × 6.9 mm) (NHM 81.24a), Goree I., Senegal." figureDoi="http://doi.org/10.5281/zenodo.4909806" httpUri="https://zenodo.org/record/4909806/files/figure.png" pageId="34" pageNumber="35">Figs. 5G</figureCitation>
,
<figureCitation id="13702A3D5467E2198B0145CCFEA30EE2" box="[267,297,474,499]" captionStart="FIGURE 22" captionStartId="35.[151,249,1941,1963]" captionTargetBox="[151,1436,483,1919]" captionTargetId="figure-188@35.[151,1436,483,1919]" captionTargetPageId="35" captionText="FIGURE 22. Raoulia calva sp. nov., holotype male (6.6 × 5.2 mm) (ZRC 2013.1721), Society Is., French Polynesia. A, dorsal view; B, anterior thoracic sternum and sterno-abdominal cavity; C, frontal view of carapace; D, posterior thoracic sternum; E, outer view of right chela; F, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.4909840" httpUri="https://zenodo.org/record/4909840/files/figure.png" pageId="34" pageNumber="35">22</figureCitation>
,
<figureCitation id="13702A3D5467E2198B3F45CCFEDE0EE2" box="[309,340,474,499]" captionStart="FIGURE 23" captionStartId="36.[151,249,1821,1843]" captionTargetBox="[176,1422,210,1789]" captionTargetId="figure-96@36.[151,1436,193,1800]" captionTargetPageId="36" captionText="FIGURE 23. A, B, E–H, Raoulia calva sp. nov., holotype male (6.6 × 5.2 mm) (ZRC 2013.1721), Society Is., French Polynesia; H, paratype female (6.5 × 5.1 mm) (ZRC 2013.1722), Society Is., French Polynesia. A, male abdomen; B, left third maxilliped; C, D, left G1; E, F, distal part of left G1; G, left G2; H, female abdomen. Scales: A, B, H = 1.0 mm; C, D, G = 0.5 mm; E, F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4909842" httpUri="https://zenodo.org/record/4909842/files/figure.png" pageId="34" pageNumber="35">23</figureCitation>
,
<figureCitation id="13702A3D5467E2198B5545CCFE050EE2" box="[351,399,474,499]" captionStart="FIGURE 41" captionStartId="59.[151,249,1634,1656]" captionTargetBox="[151,1436,193,1613]" captionTargetId="figure-208@59.[151,1436,193,1613]" captionTargetPageId="59" captionText="FIGURE 41. A, Raoulia fortis sp. nov., holotype male (12.0 × 9.2 mm) (MNHN), Santo, Vanuatu; B, Raoulia calva sp. nov., holotype male (6.6 × 5.2 mm) (ZRC 2013.1721), Society Is., French Polynesia [photograph A. Anker]; C, Caecopilumnus crassipes (Tesch, 1918), female specimen, Ryukyus (not examined) [photograph Y. Yamada, courtesy of T. Naruse]; D, Caecopilumnus loculatus sp. nov., holotype male (8.0 × 6.7 mm) (ZRC 2013.1723), Society Is., French Polynesia; E, Caecopilumnus loculatus sp. nov., paratype female (9.4 × 7.7 mm) (ZRC 2013.1724), Society Is., French Polynesia; F, Caecopilumnus loculatus sp. nov., paratype female (15.7 × 13.6 mm) (ZRC 2013.1725), Society Is., French Polynesia; G, Thecaplax capillosa gen. et sp. nov., one of males from paratype series, precise identity cannot be ascertained." figureDoi="http://doi.org/10.5281/zenodo.4909880" httpUri="https://zenodo.org/record/4909880/files/figure.png" pageId="34" pageNumber="35">41B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35165335467E2198A9D4634FED90DFE" pageId="34" pageNumber="35" type="materials_examined">
<paragraph id="8BF436B85467E2198A9D4634FED90DFE" blockId="34.[151,1436,546,1327]" pageId="34" pageNumber="35">
<emphasis id="B93FEAAA5467E2198A9D4634FEF00D2A" bold="true" box="[151,378,546,571]" pageId="34" pageNumber="35">Material examined</emphasis>
.
<materialsCitation id="3B233CE55467E2198B834634FCA40D92" ID-GBIF-Occurrence="3312327313" collectingDate="2009-11-09" collectionCode="ZRC" elevation="15" latitude="-17.4901" longLatPrecision="7" longitude="-149.8263" pageId="34" pageNumber="35" specimenCode="ZRC 2013.1721" specimenCount="1" typeStatus="holotype">
<typeStatus id="54F0881A5467E2198B834634FE7C0D2A" box="[393,502,546,571]" pageId="34" pageNumber="35" type="holotype">Holotype</typeStatus>
: male (6.6 ×
<quantity id="4CB39B5D5467E21988814634FD690D2A" box="[651,739,546,571]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="34" pageNumber="35" unit="mm" value="5.2">5.2 mm</quantity>
) (
<specimenCode id="DBED9EC35467E21988F04634FC3D0D2A" box="[762,951,546,571]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="34" pageNumber="35">ZRC 2013.1721</specimenCode>
), station BMOO-08601, BIZ-138,
<collectorName id="26BE536E5467E2198F344635FA120D2B" box="[1342,1432,547,570]" pageId="34" pageNumber="35">Moorea</collectorName>
,
<collectorName id="26BE536E5467E2198A9D4650FE840D4F" box="[151,270,582,607]" pageId="34" pageNumber="35">Society Is.</collectorName>
,
<collectorName id="26BE536E5467E2198B104650FE150D4E" box="[282,415,582,607]" pageId="34" pageNumber="35">Gump Reef</collectorName>
,
<collectorName id="26BE536E5467E2198BA34650FDE70D4E" box="[425,621,582,607]" pageId="34" pageNumber="35">French Polynesia</collectorName>
,
<geoCoordinate id="EE7F507F5467E21988724650FD670D4E" box="[632,749,582,607]" degrees="17.4901" direction="south" orientation="latitude" pageId="34" pageNumber="35" precision="5" value="-17.4901">17.4901°S</geoCoordinate>
<geoCoordinate id="EE7F507F5467E21988FE4650FC090D4E" box="[756,899,582,607]" degrees="149.8263" direction="west" orientation="longitude" pageId="34" pageNumber="35" precision="5" value="-149.8263">149.8263°W</geoCoordinate>
, silty fringing reef wall, under rubble and rocks,
<quantity id="4CB39B5D5467E2198A9D467CFF770D93" box="[151,253,618,643]" metricMagnitude="1" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="34" pageNumber="35" unit="m" value="15.0" valueMax="20.0" valueMin="10.0">
<elevation id="0066D18B5467E2198A9D467CFF770D93" box="[151,253,618,643]" metricMagnitude="1" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="34" pageNumber="35" unit="m" value="15.0" valueMax="20.0" valueMin="10.0">10–20 m</elevation>
</quantity>
, brushing, coll.
<collectorName id="26BE536E5467E2198BB8467CFDD80D92" box="[434,594,618,643]" pageId="34" pageNumber="35">S. McPherson</collectorName>
,
<date id="FFF510785467E2198854467CFCA40D92" box="[606,814,618,643]" pageId="34" pageNumber="35" value="2009-11-09">
<collectingDate id="EFB1E9905467E2198854467CFCA40D92" box="[606,814,618,643]" pageId="34" pageNumber="35" value="2009-11-09">9 November 2009</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B233CE55467E2198933467DFEDA0DFE" ID-GBIF-Occurrence="3312327322" collectingDate="2009-11-01" collectionCode="ZRC" country="French Polynesia" elevation="9" latitude="-17.4908" location="Gump Station" longLatPrecision="7" longitude="-149.8261" municipality="Moorea" pageId="34" pageNumber="35" specimenCode="ZRC 2013.1722" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="54F0881A5467E2198933467DFC150D92" box="[825,927,619,643]" pageId="34" pageNumber="35" type="paratype">Paratype</typeStatus>
:
<emphasis id="B93FEAAA5467E21989A0467CFBF20D92" bold="true" box="[938,1144,618,643]" pageId="34" pageNumber="35">
<collectingCountry id="F35C76285467E21989A0467CFBF20D92" box="[938,1144,618,643]" name="French Polynesia" pageId="34" pageNumber="35">French Polynesia</collectingCountry>
</emphasis>
:
<specimenCount id="9D4DFD315467E2198E82467CFB630D92" box="[1160,1257,618,643]" pageId="34" pageNumber="35" type="female">1 female</specimenCount>
(6.5 ×
<quantity id="4CB39B5D5467E2198F36467CFA1E0D92" box="[1340,1428,618,643]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="34" pageNumber="35" unit="mm" value="5.1">5.1 mm</quantity>
) (
<specimenCode id="DBED9EC35467E2198A944698FED40DB6" box="[158,350,654,679]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="34" pageNumber="35">ZRC 2013.1722</specimenCode>
), station BMOO-07884, BIZ-089, in front of
<location id="8E9460635467E21989734698FB970DB6" LSID="urn:lsid:plazi:treatment:03E287AE5467E2198A9D45AEFC8E0A3F:8E9460635467E21989734698FB970DB6" box="[889,1053,654,679]" country="French Polynesia" latitude="-17.4908" longLatPrecision="7" longitude="-149.8261" municipality="Moorea" name="Gump Station" pageId="34" pageNumber="35">Gump Station</location>
, near dock,
<collectingMunicipality id="6B90ACC25467E2198EBB4699FA810DB7" box="[1201,1291,655,678]" pageId="34" pageNumber="35">Moorea</collectingMunicipality>
,
<collectorName id="26BE536E5467E2198F104698FA1C0DB7" box="[1306,1430,654,679]" pageId="34" pageNumber="35">Society Is.</collectorName>
,
<geoCoordinate id="EE7F507F5467E2198A9D46A4FE870DDA" box="[151,269,690,715]" degrees="17.4908" direction="south" orientation="latitude" pageId="34" pageNumber="35" precision="5" value="-17.4908">17.4908°S</geoCoordinate>
<geoCoordinate id="EE7F507F5467E2198B1D46A4FE2C0DDA" box="[279,422,690,715]" degrees="149.8261" direction="west" orientation="longitude" pageId="34" pageNumber="35" precision="5" value="-149.8261">149.8261°W</geoCoordinate>
, silty fringing reef wall, under silty rocks,
<quantity id="4CB39B5D5467E219899546A4FC730DDB" box="[927,1017,690,715]" metricMagnitude="0" metricUnit="m" metricValue="9.0" metricValueMax="15.0" metricValueMin="3.0" pageId="34" pageNumber="35" unit="m" value="9.0" valueMax="15.0" valueMin="3.0">
<elevation id="0066D18B5467E219899546A4FC730DDB" box="[927,1017,690,715]" metricMagnitude="0" metricUnit="m" metricValue="9.0" metricValueMax="15.0" metricValueMin="3.0" pageId="34" pageNumber="35" unit="m" value="9.0" valueMax="15.0" valueMin="3.0">3–15 m</elevation>
</quantity>
, coll.
<collectorName id="26BE536E5467E2198E4E46A4FB6F0DDA" box="[1092,1253,690,715]" pageId="34" pageNumber="35">S. McPherson</collectorName>
&amp;
<collectorName id="26BE536E5467E2198F1A46A4FA0A0DDA" box="[1296,1408,690,715]" pageId="34" pageNumber="35">G. Paulay</collectorName>
,
<date id="FFF510785467E2198F8446A4FEDA0DFE" pageId="34" pageNumber="35" value="2009-11-01">
<collectingDate id="EFB1E9905467E2198F8446A4FEDA0DFE" pageId="34" pageNumber="35" value="2009-11-01">1 November 2009</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35165335467E2198ACD46ECFE370CFA" pageId="34" pageNumber="35" type="diagnosis">
<paragraph id="8BF436B85467E2198ACD46ECFE3C0CB2" blockId="34.[151,1436,546,1327]" pageId="34" pageNumber="35">
<emphasis id="B93FEAAA5467E2198ACD46ECFEB00C02" bold="true" box="[199,314,762,787]" pageId="34" pageNumber="35">Diagnosis</emphasis>
. Carapace width 1.3 times length (
<figureCitation id="13702A3D5467E21988B446ECFCAC0C02" box="[702,806,762,787]" captionStart="FIGURE 22" captionStartId="35.[151,249,1941,1963]" captionTargetBox="[151,1436,483,1919]" captionTargetId="figure-188@35.[151,1436,483,1919]" captionTargetPageId="35" captionText="FIGURE 22. Raoulia calva sp. nov., holotype male (6.6 × 5.2 mm) (ZRC 2013.1721), Society Is., French Polynesia. A, dorsal view; B, anterior thoracic sternum and sterno-abdominal cavity; C, frontal view of carapace; D, posterior thoracic sternum; E, outer view of right chela; F, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.4909840" httpUri="https://zenodo.org/record/4909840/files/figure.png" pageId="34" pageNumber="35">Fig. 22A</figureCitation>
). Junction between frontal, supraorbital margins appears almost right-angled in frontal view (
<figureCitation id="13702A3D5467E21988344708FD230C26" box="[574,681,798,823]" captionStart="FIGURE 22" captionStartId="35.[151,249,1941,1963]" captionTargetBox="[151,1436,483,1919]" captionTargetId="figure-188@35.[151,1436,483,1919]" captionTargetPageId="35" captionText="FIGURE 22. Raoulia calva sp. nov., holotype male (6.6 × 5.2 mm) (ZRC 2013.1721), Society Is., French Polynesia. A, dorsal view; B, anterior thoracic sternum and sterno-abdominal cavity; C, frontal view of carapace; D, posterior thoracic sternum; E, outer view of right chela; F, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.4909840" httpUri="https://zenodo.org/record/4909840/files/figure.png" pageId="34" pageNumber="35">Fig. 22C</figureCitation>
). Third maxilliped merus short, less than half length of ischium; ischium subquadrate, 1.3 as long as broad (
<figureCitation id="13702A3D5467E219888C4754FD620C4A" box="[646,744,834,859]" captionStart="FIGURE 5" captionStartId="11.[151,249,1549,1571]" captionTargetBox="[151,1436,193,1527]" captionTargetId="figure-230@11.[151,1436,193,1527]" captionTargetPageId="11" captionText="FIGURE 5. Right third maxillipeds. A, Acidops cessacii (A. Milne-Edwards, 1878), syntype female (14.1 × 10.0 mm) (NHM 99.3.23.66) Cape Verde (laterally inverted); B, Parapilumnus cristimanus (A. Milne-Edwards, 1873), male (16.3 × 12.5 mm) (ZRC 2003.90), Nansha Is. (Spratly Is.), South China Sea (laterally inverted); C, Raoulia limosa Ng, 1987, holotype male (8.3 × 6.3 mm) (ZMUC M58), off Marongas, Jolo, Philippines; D, Raoulia piroculata (Rathbun, 1911), lectotype female (7.3 × 5.5 mm) (USNM 41359a), Amirante Is., Indian Ocean; E, Raoulia galea sp. nov., holotype male (9.4 × 7.4 mm) (ZRC 2000.0980), Phuket, Thailand; F, Raoulia fortis sp. nov., holotype male (12.0 × 9.2 mm) (MNHN), Santo, Vanuatu (laterally inverted); G, Raoulia calva sp. nov., holotype male (6.6 × 5.2 mm) (ZRC 2013.1721), Society Is., French Polynesia; H, Caecopilumnus hirsutus (Borradaile, 1902), male (7.1 × 5.6 mm) (ZMA 261027), Flores, Indonesia (laterally inverted); I, Caecopilumnus crassipes (Tesch, 1918), male (11.8 × 10.1 mm) (ZRC 2013.1710) Lombok, Indonesia; J, Caecopilumnus loculatus sp. nov., holotype male (8.0 × 6.7 mm) (ZRC 2013.1723), Moorea, French Polynesia (laterally inverted); K, Thecaplax capillosa gen. et. sp. nov., holotype male (11.6 × 9.5 mm) (MZB), Lombok, Indonesia; L, Typhlocarcinodes integrifrons (Miers, 1881), lectotype male (8.5 × 6.9 mm) (NHM 81.24a), Goree I., Senegal." figureDoi="http://doi.org/10.5281/zenodo.4909806" httpUri="https://zenodo.org/record/4909806/files/figure.png" pageId="34" pageNumber="35">Figs. 5G</figureCitation>
,
<figureCitation id="13702A3D5467E21988FF4754FCA20C4A" box="[757,808,834,859]" captionStart="FIGURE 23" captionStartId="36.[151,249,1821,1843]" captionTargetBox="[176,1422,210,1789]" captionTargetId="figure-96@36.[151,1436,193,1800]" captionTargetPageId="36" captionText="FIGURE 23. A, B, E–H, Raoulia calva sp. nov., holotype male (6.6 × 5.2 mm) (ZRC 2013.1721), Society Is., French Polynesia; H, paratype female (6.5 × 5.1 mm) (ZRC 2013.1722), Society Is., French Polynesia. A, male abdomen; B, left third maxilliped; C, D, left G1; E, F, distal part of left G1; G, left G2; H, female abdomen. Scales: A, B, H = 1.0 mm; C, D, G = 0.5 mm; E, F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4909842" httpUri="https://zenodo.org/record/4909842/files/figure.png" pageId="34" pageNumber="35">23B</figureCitation>
). Ambulatory legs short; merus of last ambulatory legs about 2.8 times as long as broad (
<figureCitation id="13702A3D5467E21988194770FDF70C6E" box="[531,637,870,895]" captionStart="FIGURE 22" captionStartId="35.[151,249,1941,1963]" captionTargetBox="[151,1436,483,1919]" captionTargetId="figure-188@35.[151,1436,483,1919]" captionTargetPageId="35" captionText="FIGURE 22. Raoulia calva sp. nov., holotype male (6.6 × 5.2 mm) (ZRC 2013.1721), Society Is., French Polynesia. A, dorsal view; B, anterior thoracic sternum and sterno-abdominal cavity; C, frontal view of carapace; D, posterior thoracic sternum; E, outer view of right chela; F, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.4909840" httpUri="https://zenodo.org/record/4909840/files/figure.png" pageId="34" pageNumber="35">Fig. 22A</figureCitation>
). G1 with distal half distinctly curving laterally outwards in situ, distal part dilated (
<figureCitation id="13702A3D5467E2198B2F479CFE200CB2" box="[293,426,906,931]" captionStart="FIGURE 23" captionStartId="36.[151,249,1821,1843]" captionTargetBox="[176,1422,210,1789]" captionTargetId="figure-96@36.[151,1436,193,1800]" captionTargetPageId="36" captionText="FIGURE 23. A, B, E–H, Raoulia calva sp. nov., holotype male (6.6 × 5.2 mm) (ZRC 2013.1721), Society Is., French Polynesia; H, paratype female (6.5 × 5.1 mm) (ZRC 2013.1722), Society Is., French Polynesia. A, male abdomen; B, left third maxilliped; C, D, left G1; E, F, distal part of left G1; G, left G2; H, female abdomen. Scales: A, B, H = 1.0 mm; C, D, G = 0.5 mm; E, F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4909842" httpUri="https://zenodo.org/record/4909842/files/figure.png" pageId="34" pageNumber="35">Fig. 23C–F</figureCitation>
).
</paragraph>
<paragraph id="8BF436B85467E2198ACD47B8FE370CFA" blockId="34.[151,1436,546,1327]" pageId="34" pageNumber="35">
<emphasis id="B93FEAAA5467E2198ACD47B8FE900CD6" bold="true" box="[199,282,942,967]" pageId="34" pageNumber="35">Colour</emphasis>
. In life, the carapaces of the specimens are dirty white, with the pereopods light brown, and the setae reddish-brown (
<figureCitation id="13702A3D5467E2198B4347C4FE3A0CFA" box="[329,432,978,1003]" captionStart="FIGURE 41" captionStartId="59.[151,249,1634,1656]" captionTargetBox="[151,1436,193,1613]" captionTargetId="figure-208@59.[151,1436,193,1613]" captionTargetPageId="59" captionText="FIGURE 41. A, Raoulia fortis sp. nov., holotype male (12.0 × 9.2 mm) (MNHN), Santo, Vanuatu; B, Raoulia calva sp. nov., holotype male (6.6 × 5.2 mm) (ZRC 2013.1721), Society Is., French Polynesia [photograph A. Anker]; C, Caecopilumnus crassipes (Tesch, 1918), female specimen, Ryukyus (not examined) [photograph Y. Yamada, courtesy of T. Naruse]; D, Caecopilumnus loculatus sp. nov., holotype male (8.0 × 6.7 mm) (ZRC 2013.1723), Society Is., French Polynesia; E, Caecopilumnus loculatus sp. nov., paratype female (9.4 × 7.7 mm) (ZRC 2013.1724), Society Is., French Polynesia; F, Caecopilumnus loculatus sp. nov., paratype female (15.7 × 13.6 mm) (ZRC 2013.1725), Society Is., French Polynesia; G, Thecaplax capillosa gen. et sp. nov., one of males from paratype series, precise identity cannot be ascertained." figureDoi="http://doi.org/10.5281/zenodo.4909880" httpUri="https://zenodo.org/record/4909880/files/figure.png" pageId="34" pageNumber="35">Fig. 41B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35165335467E2198ACD47E0FE4D0B22" pageId="34" pageNumber="35" type="etymology">
<paragraph id="8BF436B85467E2198ACD47E0FE4D0B22" blockId="34.[151,1436,546,1327]" pageId="34" pageNumber="35">
<emphasis id="B93FEAAA5467E2198ACD47E0FECC0B1E" bold="true" box="[199,326,1014,1039]" pageId="34" pageNumber="35">Etymology</emphasis>
. From the Latin
<emphasis id="B93FEAAA5467E219881E47E0FDD10B1E" box="[532,603,1014,1039]" italics="true" pageId="34" pageNumber="35">calvus</emphasis>
for “smooth” and “bald”, alluding to the smooth carapace and chelae, without setae and tubercles
</paragraph>
</subSubSection>
<subSubSection id="C35165335467E2198ACD4028FEDD0A1A" pageId="34" pageNumber="35" type="discussion">
<paragraph id="8BF436B85467E2198ACD4028FEDD0A1A" blockId="34.[151,1436,546,1327]" pageId="34" pageNumber="35">
<emphasis id="B93FEAAA5467E2198ACD4028FEBE0B46" bold="true" box="[199,308,1086,1111]" pageId="34" pageNumber="35">Remarks</emphasis>
.
<taxonomicName id="4C4B4D3B5467E2198B424028FE610B46" authority="Ng &amp; Rahayu, 2014" authorityName="Ng &amp; Rahayu" authorityYear="2014" box="[328,491,1086,1111]" class="Malacostraca" family="Chasmocarcinidae" genus="Raoulia" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="calva" status="sp. nov.">
<emphasis id="B93FEAAA5467E2198B424028FE610B46" box="[328,491,1086,1111]" italics="true" pageId="34" pageNumber="35">Raoulia calva</emphasis>
</taxonomicName>
<emphasis id="B93FEAAA5467E2198BF34028FDD30B46" bold="true" box="[505,601,1086,1111]" pageId="34" pageNumber="35">
<taxonomicNameLabel id="A20C57D15467E2198BF34028FDD30B46" box="[505,601,1086,1111]" pageId="34" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
shares with
<taxonomicName id="4C4B4D3B5467E21988F34029FC190B46" baseAuthorityName="Rathbun" baseAuthorityYear="1911" box="[761,915,1086,1111]" class="Malacostraca" family="Chasmocarcinidae" genus="Raoulia" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="piroculata">
<emphasis id="B93FEAAA5467E21988F34029FC190B46" box="[761,915,1086,1111]" italics="true" pageId="34" pageNumber="35">R. piroculata</emphasis>
</taxonomicName>
a relatively elongated ischium of the third maxilliped (length to width ratio about 1.3), but their carapaces are different, with that of
<taxonomicName id="4C4B4D3B5467E2198EDA4075FABB0B6A" authority="Ng &amp; Rahayu, 2014" authorityName="Ng &amp; Rahayu" authorityYear="2014" box="[1232,1329,1122,1147]" class="Malacostraca" family="Chasmocarcinidae" genus="Raoulia" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="calva" status="sp. nov.">
<emphasis id="B93FEAAA5467E2198EDA4075FABB0B6A" box="[1232,1329,1122,1147]" italics="true" pageId="34" pageNumber="35">R. calva</emphasis>
</taxonomicName>
<emphasis id="B93FEAAA5467E2198F374074FA160B6A" bold="true" box="[1341,1436,1122,1147]" pageId="34" pageNumber="35">
<taxonomicNameLabel id="A20C57D15467E2198F374074FA160B6A" box="[1341,1436,1122,1147]" pageId="34" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
proportionately less broad with the front margin more or less straight (
<figureCitation id="13702A3D5467E21989B14090FBA80B8E" box="[955,1058,1158,1183]" captionStart="FIGURE 22" captionStartId="35.[151,249,1941,1963]" captionTargetBox="[151,1436,483,1919]" captionTargetId="figure-188@35.[151,1436,483,1919]" captionTargetPageId="35" captionText="FIGURE 22. Raoulia calva sp. nov., holotype male (6.6 × 5.2 mm) (ZRC 2013.1721), Society Is., French Polynesia. A, dorsal view; B, anterior thoracic sternum and sterno-abdominal cavity; C, frontal view of carapace; D, posterior thoracic sternum; E, outer view of right chela; F, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.4909840" httpUri="https://zenodo.org/record/4909840/files/figure.png" pageId="34" pageNumber="35">Fig. 22C</figureCitation>
) while in
<taxonomicName id="4C4B4D3B5467E2198E9D4091FAA10B8E" baseAuthorityName="Rathbun" baseAuthorityYear="1911" box="[1175,1323,1158,1183]" class="Malacostraca" family="Chasmocarcinidae" genus="Raoulia" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="piroculata">
<emphasis id="B93FEAAA5467E2198E9D4091FAA10B8E" box="[1175,1323,1158,1183]" italics="true" pageId="34" pageNumber="35">R. piroculata</emphasis>
</taxonomicName>
, the front margin is widely triangular (
<figureCitation id="13702A3D5467E2198BDF40BCFDB10BD2" box="[469,571,1194,1219]" captionStart="FIGURE 17" captionStartId="27.[151,249,1905,1927]" captionTargetBox="[151,1436,373,1884]" captionTargetId="figure-144@27.[151,1436,373,1884]" captionTargetPageId="27" captionText="FIGURE 17. Raoulia piroculata (Rathbun, 1911), lectotype female (7.3 × 5.5 mm) (USNM 41359a), Amirante Is., Indian Ocean. A, dorsal view; B, anterior thoracic sternum and sterno-abdominal cavity; C, frontal view of carapace; D, posterior thoracic sternum; E, outer view of right chela; F, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.4909830" httpUri="https://zenodo.org/record/4909830/files/figure.png" pageId="34" pageNumber="35">Fig. 17C</figureCitation>
). Most importantly, while the G1 of
<taxonomicName id="4C4B4D3B5467E21989D940BDFBA40BD2" authority="Ng &amp; Rahayu, 2014" authorityName="Ng &amp; Rahayu" authorityYear="2014" box="[979,1070,1194,1219]" class="Malacostraca" family="Chasmocarcinidae" genus="Raoulia" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="calva" status="sp. nov.">
<emphasis id="B93FEAAA5467E21989D940BDFBA40BD2" box="[979,1070,1194,1219]" italics="true" pageId="34" pageNumber="35">R. calva</emphasis>
</taxonomicName>
<emphasis id="B93FEAAA5467E2198E3F40BCFB050BD2" bold="true" box="[1077,1167,1194,1219]" pageId="34" pageNumber="35">
<taxonomicNameLabel id="A20C57D15467E2198E3F40BCFB050BD2" box="[1077,1167,1194,1219]" pageId="34" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is curved like
<taxonomicName id="4C4B4D3B5467E2198F3840BDFA160BD2" authorityName="Ng" authorityYear="1987" box="[1330,1436,1194,1219]" class="Malacostraca" family="Chasmocarcinidae" genus="Raoulia" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="limosa">
<emphasis id="B93FEAAA5467E2198F3840BDFA160BD2" box="[1330,1436,1194,1219]" italics="true" pageId="34" pageNumber="35">R. limosa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4B4D3B5467E2198ACD40D9FEAE0BF6" authority="Ng &amp; Rahayu, 2014" authorityName="Ng &amp; Rahayu" authorityYear="2014" box="[199,292,1230,1255]" class="Malacostraca" family="Chasmocarcinidae" genus="Raoulia" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="galea" status="sp. nov.">
<emphasis id="B93FEAAA5467E2198ACD40D9FEAE0BF6" box="[199,292,1230,1255]" italics="true" pageId="34" pageNumber="35">R. galea</emphasis>
</taxonomicName>
<emphasis id="B93FEAAA5467E2198B2640D8FE0C0BF6" bold="true" box="[300,390,1230,1255]" pageId="34" pageNumber="35">
<taxonomicNameLabel id="A20C57D15467E2198B2640D8FE0C0BF6" box="[300,390,1230,1255]" pageId="34" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, its distal part is distinctly dilated (
<figureCitation id="13702A3D5467E219891E40D8FC1C0BF6" box="[788,918,1230,1255]" captionStart="FIGURE 23" captionStartId="36.[151,249,1821,1843]" captionTargetBox="[176,1422,210,1789]" captionTargetId="figure-96@36.[151,1436,193,1800]" captionTargetPageId="36" captionText="FIGURE 23. A, B, E–H, Raoulia calva sp. nov., holotype male (6.6 × 5.2 mm) (ZRC 2013.1721), Society Is., French Polynesia; H, paratype female (6.5 × 5.1 mm) (ZRC 2013.1722), Society Is., French Polynesia. A, male abdomen; B, left third maxilliped; C, D, left G1; E, F, distal part of left G1; G, left G2; H, female abdomen. Scales: A, B, H = 1.0 mm; C, D, G = 0.5 mm; E, F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4909842" httpUri="https://zenodo.org/record/4909842/files/figure.png" pageId="34" pageNumber="35">Fig. 23C–F</figureCitation>
) whereas it is tapering in the other taxa (
<figureCitation id="13702A3D5467E2198F6E40D8FF630A1A" captionStart="FIGURE 16" captionStartId="26.[151,249,1634,1656]" captionTargetBox="[168,1435,209,1606]" captionTargetId="figure-197@26.[151,1436,193,1613]" captionTargetPageId="26" captionText="FIGURE 16. Raoulia limosa Ng, 1987. A–H, holotype male (8.3 × 6.3 mm) (ZMUC M58), Jolo, Philippines; I, female (9.5 × 7.3 mm) (ZRC 2013.1705), Lombok, Indonesia. A, male abdomen; B, left third maxilliped; C, frontal view of carapace showing antennules, antennae, epistome and orbit; D, E, left G1; F, G, distal part of left G1; H, left G2; I, female abdomen. Scales: A–C, I = 1.0 mm; D, E, H = 0.5 mm; F, G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4909828" httpUri="https://zenodo.org/record/4909828/files/figure.png" pageId="34" pageNumber="35">Figs. 16D–G</figureCitation>
,
<figureCitation id="13702A3D5467E2198AFE40E4FEC10A1A" box="[244,331,1266,1291]" captionStart="FIGURE 19" captionStartId="31.[151,249,1936,1958]" captionTargetBox="[204,1363,213,1901]" captionTargetId="figure-93@31.[179,1399,193,1917]" captionTargetPageId="31" captionText="FIGURE 19. A–E, Raoulia piroculata (Rathbun, 1911), male (11.0 × 8.5 mm), South Africa; F–L, Raoulia galea sp. nov., holotype male (9.4 × 7.4 mm) (ZRC 2000.0980), Phuket, Thailand. A, male abdomen; B, left third maxilliped; C, D, left G1; E, F, distal part of left G1; G, left G2. Scales: A, B = 1.0 mm; C, D, G = 0.5 mm; E, F = 0.1 mm. A–E after Barnard (1955: fig. 16)." figureDoi="http://doi.org/10.5281/zenodo.4909834" httpUri="https://zenodo.org/record/4909834/files/figure.png" pageId="34" pageNumber="35">19H–K</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35165335467E2198ACD4100FC8E0A3F" box="[199,772,1302,1327]" pageId="34" pageNumber="35" type="distribution">
<paragraph id="8BF436B85467E2198ACD4100FC8E0A3F" blockId="34.[151,1436,546,1327]" box="[199,772,1302,1327]" pageId="34" pageNumber="35">
<emphasis id="B93FEAAA5467E2198ACD4100FED30A3E" bold="true" box="[199,345,1302,1327]" pageId="34" pageNumber="35">Distribution</emphasis>
. Moorea (
<collectingCountry id="F35C76285467E2198BC64100FD1F0A3E" box="[460,661,1302,1327]" name="French Polynesia" pageId="34" pageNumber="35">French Polynesia</collectingCountry>
);
<quantity id="4CB39B5D5467E21988A24100FC8A0A3F" box="[680,768,1302,1327]" metricMagnitude="0" metricUnit="m" metricValue="9.0" metricValueMax="15.0" metricValueMin="3.0" pageId="34" pageNumber="35" unit="m" value="9.0" valueMax="15.0" valueMin="3.0">
3–
<specimenCount id="9D4DFD315467E21988CF4100FC8A0A3F" box="[709,768,1302,1327]" pageId="34" pageNumber="35" type="generic">15 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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