423 lines
51 KiB
XML
423 lines
51 KiB
XML
<document id="7A68849043CBCC0C69A9506933A6D8C0" ID-DOI="10.11646/zootaxa.5397.2.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="10468563" ID-ZooBank="C55178A5-DABB-4615-8443-59B388F99A36" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1704706602791" checkinUser="plazi" docAuthor="Zamani, Alireza & Marusik, Yuri M." docDate="2024" docId="03DC9030FFACFFA66EFAFA7EFAD9E1D8" docLanguage="en" docName="zootaxa.5397.2.2.pdf" docOrigin="Zootaxa 5397 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5397.2.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Harpactea abasgiana Zamani & Marusik 2024, sp. n." docType="treatment" docVersion="2" lastPageNumber="210" masterDocId="FFE5E848FFA7FFA96E6DFFB4FFFBE051" masterDocTitle="New data on Dysdera Latreille, 1804 and Harpactea Bristowe, 1939 (Araneae: Dysderidae) of the Caucasus, with new species and records" masterLastPageNumber="217" masterPageNumber="195" pageNumber="206" updateTime="1704707340498" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="8D8B995F5DE8F12C65535DCD5C3BED63">
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<mods:title id="B65403D7BD5A9A308E11968D62A78016">New data on Dysdera Latreille, 1804 and Harpactea Bristowe, 1939 (Araneae: Dysderidae) of the Caucasus, with new species and records</mods:title>
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</mods:titleInfo>
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<mods:name id="74A961E1B38A4207F7DBD1A0611E153B" type="personal">
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<mods:roleTerm id="17FAFE16EA5041409B35C1538DA31768">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3A321F26F357D11A99576B69ED015849">Zamani, Alireza</mods:namePart>
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<mods:nameIdentifier id="BF0C7566AA97CDF1E561EAB46591B343" type="ORCID">0000-0002-8084-9666</mods:nameIdentifier>
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<mods:affiliation id="3D5249DAD1F6D143E156A8670639453B">Zoological Museum, Biodiversity Unit, FI- 20014 University of Turku, Turku 20500, Finland.</mods:affiliation>
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<mods:nameIdentifier id="0FEE70A9A763A82B35A2E79D52780631" type="email">zamani.alireza5@gmail.com</mods:nameIdentifier>
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</mods:role>
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<mods:namePart id="B7B496FABBDF1475F3A51E531F651698">Marusik, Yuri M.</mods:namePart>
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<mods:nameIdentifier id="5F270F08AA220DDB5F9AB3F4991C5C4C" type="ORCID">0000-0002-4499-5148</mods:nameIdentifier>
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<mods:affiliation id="A75153C60DF10077E753D84665F587A9">Department of Zoology & Entomology, University of the Free State, Bloemfontein 9300, South Africa. & Altai State University, Lenina Pr., 61, Barnaul, RF- 656049, Russia. & Institute for Biological Problems of the North, Portovaya Str. 18, Magadan 685000, Russia.</mods:affiliation>
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<mods:nameIdentifier id="7A5ED7927BFDB7BCFE29308EA5CC9076" type="email">yurmar@mail.ru</mods:nameIdentifier>
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<mods:title id="ACEFB398EEC8997A64E0E8DA60C6EA40">Zootaxa</mods:title>
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<mods:part id="FCEC6486970D463B3CEE9F280A5D5743">
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<mods:date id="7E7EBCC47C6FAD7796321717C5442062">2024</mods:date>
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<mods:number id="7650464AD48700DB99E10CDF94E9E951">2024-01-04</mods:number>
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<mods:number id="7F6FE5184CC28B80B9A01106DAE171B5">5397</mods:number>
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<mods:url id="2FE0B51EB813B37417164A3A15106C25">http://dx.doi.org/10.11646/zootaxa.5397.2.2</mods:url>
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<treatment id="03DC9030FFACFFA66EFAFA7EFAD9E1D8" ID-DOI="http://doi.org/10.5281/zenodo.10468532" ID-Zenodo-Dep="10468532" LSID="urn:lsid:plazi:treatment:03DC9030FFACFFA66EFAFA7EFAD9E1D8" httpUri="http://treatment.plazi.org/id/03DC9030FFACFFA66EFAFA7EFAD9E1D8" lastPageId="15" lastPageNumber="210" pageId="11" pageNumber="206">
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<subSubSection id="C36F72ADFFACFFA26EFAFA7EFE18E5B5" box="[151,483,1482,1509]" pageId="11" pageNumber="206" type="nomenclature">
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<paragraph id="8BCA2126FFACFFA26EFAFA7EFE18E5B5" blockId="11.[151,483,1482,1545]" box="[151,483,1482,1509]" pageId="11" pageNumber="206">
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<heading id="D082964AFFACFFA26EFAFA7EFE18E5B5" bold="true" box="[151,483,1482,1509]" fontSize="11" level="1" pageId="11" pageNumber="206" reason="1">
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<emphasis id="B901FD34FFACFFA26EFAFA7EFE18E5B5" bold="true" box="[151,483,1482,1509]" pageId="11" pageNumber="206">
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<taxonomicName id="4C755AA5FFACFFA26EFAFA7EFE61E5B4" authority="Zamani & Marusik, 2024" authorityName="Zamani & Marusik" authorityYear="2024" box="[151,410,1482,1509]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="206" phylum="Arthropoda" rank="species" species="abasgiana" status="sp. nov.">
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<emphasis id="B901FD34FFACFFA26EFAFA7EFE61E5B4" bold="true" box="[151,410,1482,1509]" italics="true" pageId="11" pageNumber="206">Harpactea abasgiana</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A232404FFFACFFA26FCCFA7EFE18E5B5" box="[417,483,1482,1508]" pageId="11" pageNumber="206" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36F72ADFFACFFA26EFAFA5AFE53E659" box="[151,424,1518,1545]" pageId="11" pageNumber="206" type="description">
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<paragraph id="8BCA2126FFACFFA26EFAFA5AFE53E659" blockId="11.[151,483,1482,1545]" box="[151,424,1518,1545]" pageId="11" pageNumber="206">
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<figureCitation id="134E3DA3FFACFFA26EFAFA5AFED9E658" box="[151,290,1518,1545]" captionStart="FIGURE 14" captionStartId="13.[152,255,1876,1900]" captionTargetBox="[157,1422,181,1852]" captionTargetId="figure-21@13.[151,1436,181,1852]" captionTargetPageId="13" captionText="FIGURE 14. Habitus of Harpactea abasgiana sp. n. (A, B), H. dunini sp. n. (C) and H. caucasia (D, E). A–C, E dorsal; D lateral. A, C, D–R male B female." figureDoi="http://doi.org/10.5281/zenodo.10468593" httpUri="https://zenodo.org/record/10468593/files/figure.png" pageId="11" pageNumber="206">Figs 14A, B</figureCitation>
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,
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<figureCitation id="134E3DA3FFACFFA26F40FA5AFE7AE658" box="[301,385,1518,1545]" captionStart="FIGURE 15" captionStartId="14.[152,255,1880,1904]" captionTargetBox="[151,1426,164,1863]" captionTargetId="figure-32@14.[151,1436,143,1863]" captionTargetPageId="14" captionText="FIGURE 15. Males of Harpactea abasgiana sp. n. (A–C) and H. caucasia (D–J), left bulbs. A, D, H–J prolateral; B, E retrolateral, C, F, G ventral. D, H–I different aspects illustrating how the point of view affects the perception of the shape of the embolus and conductor. D–I lectotype, G from Kulczyński (1895), J from Dunin (1992b, most likely misidentified). Abbreviations: Co—‘conductor’, EH—embolic heel, T—tegulum." figureDoi="http://doi.org/10.5281/zenodo.10468597" httpUri="https://zenodo.org/record/10468597/files/figure.png" pageId="11" pageNumber="206">15A–C</figureCitation>
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,
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<figureCitation id="134E3DA3FFACFFA26FE1FA5AFE53E659" box="[396,424,1518,1544]" captionStart="FIGURE 16" captionStartId="15.[152,255,1570,1594]" captionTargetBox="[411,1176,417,1547]" captionTargetId="figure-144@15.[411,1176,417,1547]" captionTargetPageId="15" captionText="FIGURE 16. Female of Harpactea abasgiana sp. n., endogyne, dorsal. Scale bar = 0.2 mm. Abbreviation: AA—anterior arch, LMA—lateral margins of the anterior arch, Re—receptacle." figureDoi="http://doi.org/10.5281/zenodo.10468599" httpUri="https://zenodo.org/record/10468599/files/figure.png" pageId="11" pageNumber="206">16</figureCitation>
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</paragraph>
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<subSubSection id="C36F72ADFFACFFA26EFAF982FBE1E6B1" pageId="11" pageNumber="206" type="materials_examined">
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<paragraph id="8BCA2126FFACFFA26EFAF982FBA8E6C9" blockId="11.[151,1437,1590,2013]" pageId="11" pageNumber="206">
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<emphasis id="B901FD34FFACFFA26EFAF982FEBCE601" bold="true" box="[151,327,1590,1616]" pageId="11" pageNumber="206">Type material.</emphasis>
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<materialsCitation id="3B1D2B7BFFACFFA26F3CF982FDD4E625" collectingDate="2004-10-16" collectionCode="ZMMU" collectorName="Y. M. Marusik" country="Georgia" latitude="43.166668" location="Pitsunda" longLatPrecision="1145" longitude="40.366665" municipality="Republic" pageId="11" pageNumber="206" specimenCount="1" stateProvince="Abkhazia" typeStatus="holotype">
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<typeStatus id="54CE9F84FFACFFA26F3CF982FE42E601" box="[337,441,1590,1616]" pageId="11" pageNumber="206" type="holotype">Holotype</typeStatus>
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³ (
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<collectionCode id="ED64B9E3FFACFFA26F84F982FDBDE601" box="[489,582,1590,1616]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="11" pageNumber="206" type="Museum">ZMMU</collectionCode>
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),
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<collectingCountry id="F36261B6FFACFFA26C36F982FD25E601" box="[603,734,1590,1616]" name="Georgia" pageId="11" pageNumber="206">GEORGIA</collectingCountry>
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:
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<emphasis id="B901FD34FFACFFA26C86F982FC25E601" box="[747,990,1590,1616]" italics="true" pageId="11" pageNumber="206">
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<collectingMunicipality id="6BAEBB5CFFACFFA26C86F982FCB4E601" box="[747,847,1590,1616]" pageId="11" pageNumber="206">Republic</collectingMunicipality>
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of
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<collectingRegion id="49B1EFC4FFACFFA26D1BF982FC25E601" box="[886,990,1590,1616]" country="Georgia" name="Abkhazia" pageId="11" pageNumber="206">Abkhazia</collectingRegion>
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</emphasis>
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: env. of
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<location id="8EAA77FDFFACFFA26A25F982FB50E601" LSID="urn:lsid:plazi:treatment:03DC9030FFACFFA66EFAFA7EFAD9E1D8:8EAA77FDFFACFFA26A25F982FB50E601" box="[1096,1195,1590,1616]" country="Georgia" latitude="43.166668" longLatPrecision="1145" longitude="40.366665" municipality="Republic" name="Pitsunda" pageId="11" pageNumber="206" stateProvince="Abkhazia">Pitsunda</location>
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, Ldzaa Vil.,
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<geoCoordinate id="EE4147E1FFACFFA26B2DF982FA66E601" box="[1344,1437,1590,1616]" degrees="43" direction="north" minutes="10" orientation="latitude" pageId="11" pageNumber="206" precision="925" value="43.166668">43°10'N</geoCoordinate>
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<geoCoordinate id="EE4147E1FFACFFA26EFAF9EEFF08E625" box="[151,243,1626,1652]" degrees="40" direction="east" minutes="22" orientation="longitude" pageId="11" pageNumber="206" precision="925" value="40.366665">40°22'E</geoCoordinate>
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,
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<date id="FFCB07E6FFACFFA26E91F9EEFE81E625" box="[252,378,1626,1652]" pageId="11" pageNumber="206" value="2004-10-16">
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<collectingDate id="EF8FFE0EFFACFFA26E91F9EEFE81E625" box="[252,378,1626,1652]" pageId="11" pageNumber="206" value="2004-10-16">16.10.2004</collectingDate>
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</date>
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(
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<collectorName id="268044F0FFACFFA26FEAF9EEFDDCE625" box="[391,551,1626,1652]" pageId="11" pageNumber="206">Y.M. Marusik</collectorName>
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)
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</materialsCitation>
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.
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<materialsCitation id="3B1D2B7BFFACFFA26C5BF9EFFB30E625" box="[566,1227,1626,1655]" collectingDate="2004-10-16" collectionCode="ZMMU" collectorName="Y. M. Marusik" country="Georgia" latitude="43.166668" location="Pitsunda" longLatPrecision="1145" longitude="40.366665" municipality="Republic" pageId="11" pageNumber="206" specimenCount="9" specimenCount-female="9" stateProvince="Abkhazia" typeStatus="paratype">
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<typeStatus id="54CE9F84FFACFFA26C5BF9EFFD5CE624" box="[566,679,1627,1653]" pageId="11" pageNumber="206" type="paratype">Paratypes</typeStatus>
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: 5³
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<specimenCount id="9D73EAAFFFACFFA26CA6F9E8FD14E626" box="[715,751,1628,1655]" count="9" pageId="11" pageNumber="206" type="female">9♀</specimenCount>
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2j. (
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<collectionCode id="ED64B9E3FFACFFA26D73F9EEFC80E625" box="[798,891,1626,1652]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="11" pageNumber="206" type="Museum">ZMMU</collectionCode>
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), same data as for the holotype
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</materialsCitation>
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; 9³
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<materialsCitation id="3B1D2B7BFFACFFA26A9DF9E8FBB4E6C9" collectingDate="2004-10-10" collectionCode="ZMMU" collectorName="Y. M. Marusik" country="Georgia" latitude="42.973335" longLatPrecision="92" longitude="41.067333" pageId="11" pageNumber="206" specimenCount="10" specimenCount-female="10" stateProvince="Abkhazia" typeStatus="paratype">
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<specimenCount id="9D73EAAFFFACFFA26A9DF9E8FADCE626" box="[1264,1319,1628,1655]" count="10" pageId="11" pageNumber="206" type="female">10♀</specimenCount>
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(
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<collectionCode id="ED64B9E3FFACFFA26B5EF9EEFA6BE625" box="[1331,1424,1626,1652]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="11" pageNumber="206" type="Museum">ZMMU</collectionCode>
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||
), Sukhumi, University campus,
|
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<geoCoordinate id="EE4147E1FFACFFA26F86F9CAFDA7E6C8" box="[491,604,1662,1689]" degrees="42" direction="north" minutes="58.4" orientation="latitude" pageId="11" pageNumber="206" precision="92" value="42.973335">42°58.4'N</geoCoordinate>
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<geoCoordinate id="EE4147E1FFACFFA26C09F9CAFD1BE6C8" box="[612,736,1662,1689]" degrees="41" direction="east" minutes="04.04" orientation="longitude" pageId="11" pageNumber="206" precision="9" value="41.067333">41°04.04'E</geoCoordinate>
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, 09–
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<date id="FFCB07E6FFACFFA26D7AF9CAFC6EE6C9" box="[791,917,1662,1688]" pageId="11" pageNumber="206" value="2004-10-10">
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<collectingDate id="EF8FFE0EFFACFFA26D7AF9CAFC6EE6C9" box="[791,917,1662,1688]" pageId="11" pageNumber="206" value="2004-10-10">10.10.2004</collectingDate>
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</date>
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(
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||
<collectorName id="268044F0FFACFFA26DC8F9CAFBB3E6C9" box="[933,1096,1662,1688]" pageId="11" pageNumber="206">Y.M. Marusik</collectorName>
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)
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</materialsCitation>
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.
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||
</paragraph>
|
||
<paragraph id="8BCA2126FFACFFA26EAAF916FBE1E6B1" blockId="11.[151,1437,1590,2013]" pageId="11" pageNumber="206">
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||
<emphasis id="B901FD34FFACFFA26EAAF916FE20E6ED" bold="true" box="[199,475,1698,1724]" pageId="11" pageNumber="206">Comparative material.</emphasis>
|
||
<taxonomicName id="4C755AA5FFACFFA26F88F917FC4DE6EC" authority="(Kulczynski, 1895)" baseAuthorityName="Kulczynski" baseAuthorityYear="1895" box="[485,950,1698,1725]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="206" phylum="Arthropoda" rank="species" species="caucasia">
|
||
<emphasis id="B901FD34FFACFFA26F88F917FD30E6ED" box="[485,715,1699,1724]" italics="true" pageId="11" pageNumber="206">Harpactea caucasia</emphasis>
|
||
(
|
||
<materialsCitation id="3B1D2B7BFFACFFA26CB0F916FC4DE6EC" box="[733,950,1698,1725]" collectingDate="1893-05-28" collectionCode="MIZP" collectorName="G. Horvath" country="Georgia" location="Gelati" pageId="11" pageNumber="206" specimenCount="1" stateProvince="Imereti" typeStatus="holotype">
|
||
<bibRefCitation id="EFE45CD7FFACFFA26CB0F916FC56E6EC" author="Kulczynski, W." box="[733,941,1698,1725]" pageId="11" pageNumber="206" pagination="3 - 38" refId="ref10449" refString="Kulczynski, W. (1895) Araneae a Dre G. Horvath in Bessarabia, Chersoneso Taurico, Transcaucasia et Armenia Russica collectae. Termesztrajzi Fuzetek, 18, 3 - 38." type="journal article" year="1895">Kulczyński, 1895</bibRefCitation>
|
||
)
|
||
</materialsCitation>
|
||
</taxonomicName>
|
||
:
|
||
<typeStatus id="54CE9F84FFACFFA26DAEF916FBD0E6ED" box="[963,1067,1698,1724]" pageId="11" pageNumber="206" type="holotype">Holotype</typeStatus>
|
||
³ (
|
||
<collectionCode id="ED64B9E3FFACFFA26A30F916FB5EE6ED" box="[1117,1189,1698,1724]" pageId="11" pageNumber="206">MIZP</collectionCode>
|
||
),
|
||
<collectingCountry id="F36261B6FFACFFA26AD1F916FAC4E6ED" box="[1212,1343,1698,1724]" name="Georgia" pageId="11" pageNumber="206">GEORGIA</collectingCountry>
|
||
:
|
||
<emphasis id="B901FD34FFACFFA26B21F917FF1DE6B1" italics="true" pageId="11" pageNumber="206">
|
||
<collectingRegion id="49B1EFC4FFACFFA26B21F917FF1DE6B1" country="Georgia" name="Imereti" pageId="11" pageNumber="206">Imereti Region</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<location id="8EAA77FDFFACFFA26E98F972FEC7E6B1" LSID="urn:lsid:plazi:treatment:03DC9030FFACFFA66EFAFA7EFAD9E1D8:8EAA77FDFFACFFA26E98F972FEC7E6B1" box="[245,316,1734,1760]" country="Georgia" name="Gelati" pageId="11" pageNumber="206" stateProvince="Imereti">Gelati</location>
|
||
,
|
||
<date id="FFCB07E6FFACFFA26F25F972FE3DE6B0" box="[328,454,1734,1761]" pageId="11" pageNumber="206" value="1893-05-28">
|
||
<collectingDate id="EF8FFE0EFFACFFA26F25F972FE3DE6B0" box="[328,454,1734,1761]" pageId="11" pageNumber="206" value="1893-05-28">28.05.1893</collectingDate>
|
||
</date>
|
||
(
|
||
<collectorName id="268044F0FFACFFA26FBBF972FDA2E6B1" box="[470,601,1734,1760]" pageId="11" pageNumber="206">G. Horvath</collectorName>
|
||
). (
|
||
<figureCitation id="134E3DA3FFACFFA26C19F972FD02E6B0" box="[628,761,1734,1761]" captionStart="FIGURE 14" captionStartId="13.[152,255,1876,1900]" captionTargetBox="[157,1422,181,1852]" captionTargetId="figure-21@13.[151,1436,181,1852]" captionTargetPageId="13" captionText="FIGURE 14. Habitus of Harpactea abasgiana sp. n. (A, B), H. dunini sp. n. (C) and H. caucasia (D, E). A–C, E dorsal; D lateral. A, C, D–R male B female." figureDoi="http://doi.org/10.5281/zenodo.10468593" httpUri="https://zenodo.org/record/10468593/files/figure.png" pageId="11" pageNumber="206">Fig. 14D, E</figureCitation>
|
||
,
|
||
<figureCitation id="134E3DA3FFACFFA26D68F972FC70E6B1" box="[773,907,1734,1761]" captionStart="FIGURE 15" captionStartId="14.[152,255,1880,1904]" captionTargetBox="[151,1426,164,1863]" captionTargetId="figure-32@14.[151,1436,143,1863]" captionTargetPageId="14" captionText="FIGURE 15. Males of Harpactea abasgiana sp. n. (A–C) and H. caucasia (D–J), left bulbs. A, D, H–J prolateral; B, E retrolateral, C, F, G ventral. D, H–I different aspects illustrating how the point of view affects the perception of the shape of the embolus and conductor. D–I lectotype, G from Kulczyński (1895), J from Dunin (1992b, most likely misidentified). Abbreviations: Co—‘conductor’, EH—embolic heel, T—tegulum." figureDoi="http://doi.org/10.5281/zenodo.10468597" httpUri="https://zenodo.org/record/10468597/files/figure.png" pageId="11" pageNumber="206">15D–F, H, I</figureCitation>
|
||
) [examined]
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36F72ADFFACFFA26EAAF95EFB97E779" pageId="11" pageNumber="206" type="etymology">
|
||
<paragraph id="8BCA2126FFACFFA26EAAF95EFB97E779" blockId="11.[151,1437,1590,2013]" pageId="11" pageNumber="206">
|
||
<emphasis id="B901FD34FFACFFA26EAAF95EFEB5E755" bold="true" box="[199,334,1770,1796]" pageId="11" pageNumber="206">Etymology.</emphasis>
|
||
The specific epithet refers to the Kingdom of
|
||
<collectingRegion id="49B1EFC4FFACFFA26D1DF95EFC25E755" box="[880,990,1770,1796]" country="Georgia" name="Abkhazia" pageId="11" pageNumber="206">Abkhazia</collectingRegion>
|
||
, which was also known as Abasgia or Egrisi-Abkhazia. It was a medieval feudal state established in the Caucasus in the 780s.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36F72ADFFACFFA56EAAF886FCA5E14C" lastPageId="12" lastPageNumber="207" pageId="11" pageNumber="206" type="diagnosis">
|
||
<paragraph id="8BCA2126FFACFFA56EAAF886FCA5E14C" blockId="11.[151,1437,1590,2013]" lastBlockId="12.[151,1437,151,897]" lastPageId="12" lastPageNumber="207" pageId="11" pageNumber="206">
|
||
<emphasis id="B901FD34FFACFFA26EAAF886FEB9E71D" bold="true" box="[199,322,1842,1868]" pageId="11" pageNumber="206">Diagnosis.</emphasis>
|
||
The new species is most similar to the species that has been identified and illustrated as
|
||
<taxonomicName id="4C755AA5FFACFFA26B7AF887FA66E71D" baseAuthorityName="Kulczynski" baseAuthorityYear="1895" box="[1303,1437,1843,1869]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="206" phylum="Arthropoda" rank="species" species="caucasia">
|
||
<emphasis id="B901FD34FFACFFA26B7AF887FAD0E71D" box="[1303,1323,1843,1868]" italics="true" pageId="11" pageNumber="206">H</emphasis>
|
||
.
|
||
<emphasis id="B901FD34FFACFFA26B54F887FA66E71D" box="[1337,1437,1843,1868]" italics="true" pageId="11" pageNumber="206">caucasia</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFE45CD7FFACFFA26ED4F8E2FEA6E721" author="Dunin, P. M." box="[185,349,1878,1904]" pageId="11" pageNumber="206" pagination="35 - 76" refId="ref10097" refString="Dunin, P. M. (1992 b) The spider family Dysderidae of the Caucasian fauna (Arachnida Aranei Haplogynae). Arthropoda Selecta, 1 (3), 35 - 76. [in Russian]" type="journal article" year="1992">Dunin (1992b)</bibRefCitation>
|
||
in the shape of the copulatory organs. The male can be distinguished by thinner bulb (ca. 1.3 times longer than wide,
|
||
<emphasis id="B901FD34FFACFFA26F0FF8CFFE82E7C5" box="[354,377,1915,1940]" italics="true" pageId="11" pageNumber="206">vs</emphasis>
|
||
. about as long as wide), shape of embolus base (presence of a kind of a heel (
|
||
<emphasis id="B901FD34FFACFFA26A8FF8CFFAFCE7C5" box="[1250,1287,1915,1940]" italics="true" pageId="11" pageNumber="206">EH</emphasis>
|
||
),
|
||
<emphasis id="B901FD34FFACFFA26B72F8CFFACDE7C5" box="[1311,1334,1915,1940]" italics="true" pageId="11" pageNumber="206">vs</emphasis>
|
||
. absent), and shorter and heavily sclerotized ‘conductor’ (
|
||
<emphasis id="B901FD34FFACFFA26CA1F82BFD16E7E8" box="[716,749,1951,1977]" italics="true" pageId="11" pageNumber="206">Co</emphasis>
|
||
) directed posteriorly (
|
||
<emphasis id="B901FD34FFACFFA26D9FF82BFBF2E7E9" box="[1010,1033,1951,1976]" italics="true" pageId="11" pageNumber="206">vs</emphasis>
|
||
. membranous ‘conductor’ directed laterally) (cf.
|
||
<figureCitation id="134E3DA3FFACFFA26F42F876FE3BE78C" box="[303,448,1986,2013]" captionStart="FIGURE 15" captionStartId="14.[152,255,1880,1904]" captionTargetBox="[151,1426,164,1863]" captionTargetId="figure-32@14.[151,1436,143,1863]" captionTargetPageId="14" captionText="FIGURE 15. Males of Harpactea abasgiana sp. n. (A–C) and H. caucasia (D–J), left bulbs. A, D, H–J prolateral; B, E retrolateral, C, F, G ventral. D, H–I different aspects illustrating how the point of view affects the perception of the shape of the embolus and conductor. D–I lectotype, G from Kulczyński (1895), J from Dunin (1992b, most likely misidentified). Abbreviations: Co—‘conductor’, EH—embolic heel, T—tegulum." figureDoi="http://doi.org/10.5281/zenodo.10468597" httpUri="https://zenodo.org/record/10468597/files/figure.png" pageId="11" pageNumber="206">Fig. 15A 15J</figureCitation>
|
||
). The female differs by the widened and abrupt tip of receptacle (
|
||
<emphasis id="B901FD34FFACFFA26AFEF877FB51E78D" box="[1171,1194,1987,2012]" italics="true" pageId="11" pageNumber="206">vs</emphasis>
|
||
. rounded and slightly tapering), and by the claw-like lateral margins (
|
||
<emphasis id="B901FD34FFABFFA56CDCFF23FD12E0E0" box="[689,745,151,177]" italics="true" pageId="12" pageNumber="207">LMA</emphasis>
|
||
) of anterior arch (
|
||
<emphasis id="B901FD34FFABFFA56DD4FF23FC2BE0E1" box="[953,976,151,176]" italics="true" pageId="12" pageNumber="207">vs</emphasis>
|
||
. rounded) (cf.
|
||
<figureCitation id="134E3DA3FFABFFA56A14FF23FB32E0E0" box="[1145,1225,151,177]" captionStart="FIGURE 16" captionStartId="15.[152,255,1570,1594]" captionTargetBox="[411,1176,417,1547]" captionTargetId="figure-144@15.[411,1176,417,1547]" captionTargetPageId="15" captionText="FIGURE 16. Female of Harpactea abasgiana sp. n., endogyne, dorsal. Scale bar = 0.2 mm. Abbreviation: AA—anterior arch, LMA—lateral margins of the anterior arch, Re—receptacle." figureDoi="http://doi.org/10.5281/zenodo.10468599" httpUri="https://zenodo.org/record/10468599/files/figure.png" pageId="12" pageNumber="207">Fig. 16</figureCitation>
|
||
and
|
||
<bibRefCitation id="EFE45CD7FFABFFA56B6CFF23FA62E0E0" author="Dunin, P. M." box="[1281,1433,151,177]" pageId="12" pageNumber="207" pagination="35 - 76" refId="ref10097" refString="Dunin, P. M. (1992 b) The spider family Dysderidae of the Caucasian fauna (Arachnida Aranei Haplogynae). Arthropoda Selecta, 1 (3), 35 - 76. [in Russian]" type="journal article" year="1992">Dunin 1992b</bibRefCitation>
|
||
: fig.
|
||
<quantity id="4C8D8CC3FFABFFA56EABFF0FFF0AE084" box="[198,241,187,213]" metricMagnitude="-2" metricUnit="kg" metricValue="1.4" pageId="12" pageNumber="207" unit="g" value="14.0">14г</quantity>
|
||
). From the
|
||
<typeStatus id="54CE9F84FFABFFA56F15FF0FFE21E084" box="[376,474,187,213]" pageId="12" pageNumber="207" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C755AA5FFABFFA56C6EFF0FFD70E085" baseAuthorityName="Kulczynski" baseAuthorityYear="1895" box="[515,651,187,213]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="207" phylum="Arthropoda" rank="species" species="caucasia">
|
||
<emphasis id="B901FD34FFABFFA56C6EFF0FFDECE085" box="[515,535,187,212]" italics="true" pageId="12" pageNumber="207">H</emphasis>
|
||
.
|
||
<emphasis id="B901FD34FFABFFA56C4AFF0FFD70E085" box="[551,651,187,212]" italics="true" pageId="12" pageNumber="207">caucasia</emphasis>
|
||
</taxonomicName>
|
||
, the new species differs by having an embolic base heel (
|
||
<emphasis id="B901FD34FFABFFA56B49FF0FFAC0E085" box="[1316,1339,187,212]" italics="true" pageId="12" pageNumber="207">vs</emphasis>
|
||
. absent) and shorter proximal outgrowth of the conductor (cf.
|
||
<figureCitation id="134E3DA3FFABFFA56C87FF6BFBEBE0A8" box="[746,1040,222,249]" captionStart="FIGURE 15" captionStartId="14.[152,255,1880,1904]" captionTargetBox="[151,1426,164,1863]" captionTargetId="figure-32@14.[151,1436,143,1863]" captionTargetPageId="14" captionText="FIGURE 15. Males of Harpactea abasgiana sp. n. (A–C) and H. caucasia (D–J), left bulbs. A, D, H–J prolateral; B, E retrolateral, C, F, G ventral. D, H–I different aspects illustrating how the point of view affects the perception of the shape of the embolus and conductor. D–I lectotype, G from Kulczyński (1895), J from Dunin (1992b, most likely misidentified). Abbreviations: Co—‘conductor’, EH—embolic heel, T—tegulum." figureDoi="http://doi.org/10.5281/zenodo.10468597" httpUri="https://zenodo.org/record/10468597/files/figure.png" pageId="12" pageNumber="207">Fig. 15A, B and D, E, H, I</figureCitation>
|
||
). The two species also differ by the relative length of legs (leg I/carapace length ratio: 2.5
|
||
<emphasis id="B901FD34FFABFFA56C9AFEB7FCF5E14D" box="[759,782,259,284]" italics="true" pageId="12" pageNumber="207">vs</emphasis>
|
||
. 3.53).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36F72ADFFABFFA56EAAFE92FEB5E3D0" pageId="12" pageNumber="207" type="description">
|
||
<paragraph id="8BCA2126FFABFFA56EAAFE92FAE3E1A4" blockId="12.[151,1437,151,897]" pageId="12" pageNumber="207">
|
||
<emphasis id="B901FD34FFABFFA56EAAFE92FEA2E111" bold="true" box="[199,345,294,320]" pageId="12" pageNumber="207">Description.</emphasis>
|
||
Male (
|
||
<typeStatus id="54CE9F84FFABFFA56FC7FE93FDE2E110" box="[426,537,295,321]" pageId="12" pageNumber="207" type="holotype">Holotype</typeStatus>
|
||
). Habitus as in
|
||
<figureCitation id="134E3DA3FFABFFA56CAAFE93FCD4E110" box="[711,815,295,321]" captionStart="FIGURE 14" captionStartId="13.[152,255,1876,1900]" captionTargetBox="[157,1422,181,1852]" captionTargetId="figure-21@13.[151,1436,181,1852]" captionTargetPageId="13" captionText="FIGURE 14. Habitus of Harpactea abasgiana sp. n. (A, B), H. dunini sp. n. (C) and H. caucasia (D, E). A–C, E dorsal; D lateral. A, C, D–R male B female." figureDoi="http://doi.org/10.5281/zenodo.10468593" httpUri="https://zenodo.org/record/10468593/files/figure.png" pageId="12" pageNumber="207">Fig. 14A</figureCitation>
|
||
. Total length 5.20. Carapace 2.50 long, 2.00 wide. Eye sizes: AE 0.11, PME 0.10, PLE 0.11. Carapace, chelicerae, sternum, labium and maxillae light brown; carapace slightly lighter posteriorly. Legs yellowish brown. Abdomen greyish, without any pattern. Spinnerets uniformly light greyish. Measurements of legs: I: 6.36 (1.85, 1.04, 1.55, 1.45, 0.47), II: 5.68 (1.58, 0.97, 1.41, 1.24, 0.48), III: 4.49 (1.25, 0.66, 0.94, 1.25, 0.39), IV: 6.32 (1.78, 0.86, 1.47, 1.74, 0.47). Spination: I: Fe: 2pl. II: Fe: 2pl. III: Fe: 1d, 2pl; Pa: 1pl; Ti: 1d, 3pl, 2rl, 5v; Mt: 3pl, 2rl, 4v. IV: Fe: 4d, 1rl; Ti: 1d, 3pl, 3rl, 5v; Mt: 2d, 4pl, 3rl, 5v.
|
||
</paragraph>
|
||
<paragraph id="8BCA2126FFABFFA56EAAFE4BFC5EE230" blockId="12.[151,1437,151,897]" pageId="12" pageNumber="207">
|
||
Bulb as in
|
||
<figureCitation id="134E3DA3FFABFFA56F2FFE4BFE31E248" box="[322,458,510,537]" captionStart="FIGURE 15" captionStartId="14.[152,255,1880,1904]" captionTargetBox="[151,1426,164,1863]" captionTargetId="figure-32@14.[151,1436,143,1863]" captionTargetPageId="14" captionText="FIGURE 15. Males of Harpactea abasgiana sp. n. (A–C) and H. caucasia (D–J), left bulbs. A, D, H–J prolateral; B, E retrolateral, C, F, G ventral. D, H–I different aspects illustrating how the point of view affects the perception of the shape of the embolus and conductor. D–I lectotype, G from Kulczyński (1895), J from Dunin (1992b, most likely misidentified). Abbreviations: Co—‘conductor’, EH—embolic heel, T—tegulum." figureDoi="http://doi.org/10.5281/zenodo.10468597" httpUri="https://zenodo.org/record/10468597/files/figure.png" pageId="12" pageNumber="207">Fig. 15A–C</figureCitation>
|
||
; ca. 1.3 times longer than wide, ‘conductor’ (
|
||
<emphasis id="B901FD34FFABFFA56DA1FE4BFC16E248" box="[972,1005,511,537]" italics="true" pageId="12" pageNumber="207">Co</emphasis>
|
||
) with subtriangular membranous base and well sclerotized digitiform extension directed anteriorly; embolus bent at base at ca. right angle, terminal part bent on acute angle, proximal part of embolus with kind of heel (
|
||
<emphasis id="B901FD34FFABFFA56D1DFDF3FC6EE231" box="[880,917,583,608]" italics="true" pageId="12" pageNumber="207">EH</emphasis>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCA2126FFABFFA56EAAFDDFFEBAE344" blockId="12.[151,1437,151,897]" pageId="12" pageNumber="207">
|
||
Female. Habitus as in
|
||
<figureCitation id="134E3DA3FFABFFA56FA5FDDFFDD4E2D4" box="[456,559,619,645]" captionStart="FIGURE 14" captionStartId="13.[152,255,1876,1900]" captionTargetBox="[157,1422,181,1852]" captionTargetId="figure-21@13.[151,1436,181,1852]" captionTargetPageId="13" captionText="FIGURE 14. Habitus of Harpactea abasgiana sp. n. (A, B), H. dunini sp. n. (C) and H. caucasia (D, E). A–C, E dorsal; D lateral. A, C, D–R male B female." figureDoi="http://doi.org/10.5281/zenodo.10468593" httpUri="https://zenodo.org/record/10468593/files/figure.png" pageId="12" pageNumber="207">Fig. 14B</figureCitation>
|
||
. Total length 4.20. Carapace 1.83 long, 1.45 wide. Eye sizes: AE 0.10, PME 0.09, PLE 0.10. Coloration as in male. Measurements of legs: I: 4.94 (1.36, 0.94, 1.15, 1.06, 0.43), II: 4.54 (1.21, 0.80, 1.11, 1.00, 0.42), III: 3.85 (1.04, 0.57, 0.81, 1.00, 0.43), IV: 5.60 (1.57, 0.80, 1.30, 1.45, 0.48). Spination: I: Fe: 2pl. II: Fe: 2pl. III: Fe: 1d, 2pl; Pa: 1pl; Ti: 1d, 3pl, 2rl, 4v; Mt: 3pl, 2rl, 2v. IV: Fe: 3d; Ti: 1d, 3pl, 3rl, 5v; Mt: 2d, 3pl, 3rl, 3v.
|
||
</paragraph>
|
||
<paragraph id="8BCA2126FFABFFA56EAAFCABFEB5E3D0" blockId="12.[151,1437,151,897]" pageId="12" pageNumber="207">
|
||
Endogyne as in
|
||
<figureCitation id="134E3DA3FFABFFA56F10FCABFE2BE368" box="[381,464,799,825]" captionStart="FIGURE 16" captionStartId="15.[152,255,1570,1594]" captionTargetBox="[411,1176,417,1547]" captionTargetId="figure-144@15.[411,1176,417,1547]" captionTargetPageId="15" captionText="FIGURE 16. Female of Harpactea abasgiana sp. n., endogyne, dorsal. Scale bar = 0.2 mm. Abbreviation: AA—anterior arch, LMA—lateral margins of the anterior arch, Re—receptacle." figureDoi="http://doi.org/10.5281/zenodo.10468599" httpUri="https://zenodo.org/record/10468599/files/figure.png" pageId="12" pageNumber="207">Fig. 16</figureCitation>
|
||
; with long receptacle (
|
||
<emphasis id="B901FD34FFABFFA56CBCFCABFD14E369" box="[721,751,799,824]" italics="true" pageId="12" pageNumber="207">Re</emphasis>
|
||
) slightly longer than anterior arch (
|
||
<emphasis id="B901FD34FFABFFA56AEFFCABFB5FE368" box="[1154,1188,799,825]" italics="true" pageId="12" pageNumber="207">AA</emphasis>
|
||
), tip slightly widened and abrupt at the top edge, lacking bulbous widening; anterior arch with claw-like lateral margins (
|
||
<emphasis id="B901FD34FFABFFA56A94FCF7FACAE30C" box="[1273,1329,835,861]" italics="true" pageId="12" pageNumber="207">LMA</emphasis>
|
||
) directed meso-anteriorly.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF0A71AEFFABFFA56EF5F834FEA4E7B1" ID-DOI="http://doi.org/10.5281/zenodo.10468591" ID-Zenodo-Dep="10468591" httpUri="https://zenodo.org/record/10468591/files/figure.png" pageId="12" pageNumber="207" startId="12.[152,255,1920,1944]" targetBox="[151,1436,921,1897]" targetPageId="12" targetType="figure">
|
||
<paragraph id="8BCA2126FFABFFA56EF5F834FEA4E7B1" blockId="12.[151,1437,1920,2016]" pageId="12" pageNumber="207">
|
||
<emphasis id="B901FD34FFABFFA56EF5F834FEDEE7C9" bold="true" box="[152,293,1920,1944]" pageId="12" pageNumber="207">FIGURE 13.</emphasis>
|
||
Females of
|
||
<taxonomicName id="4C755AA5FFABFFA56FCEF834FDADE7C9" authority="Zamani & Marusik, 2024" authorityName="Zamani & Marusik" authorityYear="2024" box="[419,598,1920,1944]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="207" phylum="Arthropoda" rank="species" species="atabekia" status="sp. nov.">
|
||
<emphasis id="B901FD34FFABFFA56FCEF834FDADE7C9" box="[419,598,1920,1944]" italics="true" pageId="12" pageNumber="207">Dysdera atabekia</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B901FD34FFABFFA56C30F835FD6CE7C8" bold="true" box="[605,663,1921,1945]" pageId="12" pageNumber="207">
|
||
<taxonomicNameLabel id="A232404FFFABFFA56C30F835FD6CE7C8" box="[605,663,1921,1945]" pageId="12" pageNumber="207" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(A),
|
||
<taxonomicName id="4C755AA5FFABFFA56CA0F835FC6FE7C9" authorityName="Charitonov" authorityYear="1956" box="[717,916,1920,1944]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="207" phylum="Arthropoda" rank="species" species="azerbajdzhanica">
|
||
<emphasis id="B901FD34FFABFFA56CA0F835FD24E7C9" box="[717,735,1921,1944]" italics="true" pageId="12" pageNumber="207">D</emphasis>
|
||
.
|
||
<emphasis id="B901FD34FFABFFA56C80F834FC6FE7C9" box="[749,916,1920,1944]" italics="true" pageId="12" pageNumber="207">azerbajdzhanica</emphasis>
|
||
</taxonomicName>
|
||
(B),
|
||
<taxonomicName id="4C755AA5FFABFFA56DA7F835FBD1E7C9" authorityName="Deeleman-Reinhold" authorityYear="1988" box="[970,1066,1920,1944]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="207" phylum="Arthropoda" rank="species" species="dunini">
|
||
<emphasis id="B901FD34FFABFFA56DA7F835FC27E7C9" box="[970,988,1921,1944]" italics="true" pageId="12" pageNumber="207">D</emphasis>
|
||
.
|
||
<emphasis id="B901FD34FFABFFA56D84F834FBD1E7C9" box="[1001,1066,1920,1944]" italics="true" pageId="12" pageNumber="207">dunini</emphasis>
|
||
</taxonomicName>
|
||
(C),
|
||
<taxonomicName id="4C755AA5FFABFFA56A32F835FAFEE7C9" authorityName="Dunin" authorityYear="1991" box="[1119,1285,1920,1944]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="207" phylum="Arthropoda" rank="species" species="daghestanica">
|
||
<emphasis id="B901FD34FFABFFA56A32F835FB8AE7C9" box="[1119,1137,1921,1944]" italics="true" pageId="12" pageNumber="207">D</emphasis>
|
||
.
|
||
<emphasis id="B901FD34FFABFFA56A13F834FAFEE7C9" box="[1150,1285,1920,1944]" italics="true" pageId="12" pageNumber="207">daghestanica</emphasis>
|
||
</taxonomicName>
|
||
(D),
|
||
<taxonomicName id="4C755AA5FFABFFA56B51F835FA67E7C9" authorityName="Deeleman-Reinhold" authorityYear="1988" box="[1340,1436,1920,1944]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="207" phylum="Arthropoda" rank="species" species="dunini">
|
||
<emphasis id="B901FD34FFABFFA56B51F835FAB5E7C9" box="[1340,1358,1921,1944]" italics="true" pageId="12" pageNumber="207">D</emphasis>
|
||
.
|
||
<emphasis id="B901FD34FFABFFA56B36F834FA67E7C9" box="[1371,1436,1920,1944]" italics="true" pageId="12" pageNumber="207">dunini</emphasis>
|
||
</taxonomicName>
|
||
(E), and
|
||
<taxonomicName id="4C755AA5FFABFFA56E81F811FE5AE7ED" authorityName="Mcheidze" authorityYear="1979" box="[236,417,1956,1980]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="207" phylum="Arthropoda" rank="subSpecies" species="hungarica" subSpecies="atra">
|
||
<emphasis id="B901FD34FFABFFA56E81F811FF05E7ED" box="[236,254,1957,1980]" italics="true" pageId="12" pageNumber="207">D</emphasis>
|
||
.
|
||
<emphasis id="B901FD34FFABFFA56F67F810FE5AE7ED" box="[266,417,1956,1980]" italics="true" pageId="12" pageNumber="207">hungarica atra</emphasis>
|
||
</taxonomicName>
|
||
(F), dorsal view of endogynes. Scale bars = 0.2 mm.Abbreviations:
|
||
<emphasis id="B901FD34FFABFFA56A21F811FB7FE7ED" box="[1100,1156,1957,1980]" italics="true" pageId="12" pageNumber="207">DA—</emphasis>
|
||
dorsal arch,
|
||
<emphasis id="B901FD34FFABFFA56A93F811FAD4E7ED" box="[1278,1327,1957,1980]" italics="true" pageId="12" pageNumber="207">Re—</emphasis>
|
||
receptacle,
|
||
<emphasis id="B901FD34FFABFFA56EFAF87DFF30E7B1" box="[151,203,1993,2016]" italics="true" pageId="12" pageNumber="207">TB—</emphasis>
|
||
transverse bar.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF0A71AEFFAAFFA46EF5F8E0FE0FE7C1" ID-DOI="http://doi.org/10.5281/zenodo.10468593" ID-Zenodo-Dep="10468593" httpUri="https://zenodo.org/record/10468593/files/figure.png" pageId="13" pageNumber="208" startId="13.[152,255,1876,1900]" targetBox="[157,1422,181,1852]" targetPageId="13" targetType="figure">
|
||
<paragraph id="8BCA2126FFAAFFA46EF5F8E0FE0FE7C1" blockId="13.[151,1436,1876,1937]" pageId="13" pageNumber="208">
|
||
<emphasis id="B901FD34FFAAFFA46EF5F8E0FEDDE73D" bold="true" box="[152,294,1876,1900]" pageId="13" pageNumber="208">FIGURE 14.</emphasis>
|
||
Habitus of
|
||
<taxonomicName id="4C755AA5FFAAFFA46FCCF8E1FD84E73D" authority="Zamani & Marusik, 2024" authorityName="Zamani & Marusik" authorityYear="2024" box="[417,639,1876,1900]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="208" phylum="Arthropoda" rank="species" species="abasgiana" status="sp. nov.">
|
||
<emphasis id="B901FD34FFAAFFA46FCCF8E1FD84E73D" box="[417,639,1876,1900]" italics="true" pageId="13" pageNumber="208">Harpactea abasgiana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B901FD34FFAAFFA46CEAF8E1FD3AE73C" bold="true" box="[647,705,1877,1901]" pageId="13" pageNumber="208">
|
||
<taxonomicNameLabel id="A232404FFFAAFFA46CEAF8E1FD3AE73C" box="[647,705,1877,1901]" pageId="13" pageNumber="208" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(A, B),
|
||
<taxonomicName id="4C755AA5FFAAFFA46D74F8E1FC81E73D" authority="Zamani & Marusik, 2024" authorityName="Zamani & Marusik" authorityYear="2024" box="[793,890,1876,1900]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="208" phylum="Arthropoda" rank="species" species="dunini" status="sp. n.">
|
||
<emphasis id="B901FD34FFAAFFA46D74F8E1FC81E73D" box="[793,890,1876,1900]" italics="true" pageId="13" pageNumber="208">H. dunini</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B901FD34FFAAFFA46DECF8E1FC47E73C" bold="true" box="[897,956,1877,1901]" pageId="13" pageNumber="208">
|
||
<taxonomicNameLabel id="A232404FFFAAFFA46DECF8E1FC47E73C" box="[897,956,1877,1901]" pageId="13" pageNumber="208" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(C) and
|
||
<taxonomicName id="4C755AA5FFAAFFA46A74F8E1FB68E73D" baseAuthorityName="Kulczynski" baseAuthorityYear="1895" box="[1049,1171,1877,1900]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="208" phylum="Arthropoda" rank="species" species="caucasia">
|
||
<emphasis id="B901FD34FFAAFFA46A74F8E1FB68E73D" box="[1049,1171,1877,1900]" italics="true" pageId="13" pageNumber="208">H. caucasia</emphasis>
|
||
</taxonomicName>
|
||
(D, E). A–C, E dorsal; D lateral.
|
||
<emphasis id="B901FD34FFAAFFA46E8EF8CCFEA9E7C0" bold="true" box="[227,338,1912,1937]" pageId="13" pageNumber="208">A, C, D–R</emphasis>
|
||
male
|
||
<emphasis id="B901FD34FFAAFFA46FFDF8CDFE5AE7C0" bold="true" box="[400,417,1913,1937]" pageId="13" pageNumber="208">B</emphasis>
|
||
female.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF0A71AEFFA9FFA76EF5F8ECFCC1E78D" ID-DOI="http://doi.org/10.5281/zenodo.10468597" ID-Zenodo-Dep="10468597" httpUri="https://zenodo.org/record/10468597/files/figure.png" pageId="14" pageNumber="209" startId="14.[152,255,1880,1904]" targetBox="[151,1426,164,1863]" targetPageId="14" targetType="figure">
|
||
<paragraph id="8BCA2126FFA9FFA76EF5F8ECFCC1E78D" blockId="14.[151,1437,1880,2012]" pageId="14" pageNumber="209">
|
||
<emphasis id="B901FD34FFA9FFA76EF5F8ECFED2E721" bold="true" box="[152,297,1880,1904]" pageId="14" pageNumber="209">FIGURE 15.</emphasis>
|
||
Males of
|
||
<taxonomicName id="4C755AA5FFA9FFA76FF4F8EDFD82E721" authority="Zamani & Marusik, 2024" authorityName="Zamani & Marusik" authorityYear="2024" box="[409,633,1880,1904]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="209" phylum="Arthropoda" rank="species" species="abasgiana" status="sp. nov.">
|
||
<emphasis id="B901FD34FFA9FFA76FF4F8EDFD82E721" box="[409,633,1880,1904]" italics="true" pageId="14" pageNumber="209">Harpactea abasgiana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B901FD34FFA9FFA76CEFF8ECFD44E721" bold="true" box="[642,703,1880,1904]" pageId="14" pageNumber="209">
|
||
<taxonomicNameLabel id="A232404FFFA9FFA76CEFF8ECFD44E721" box="[642,703,1880,1904]" pageId="14" pageNumber="209" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(A–C) and
|
||
<taxonomicName id="4C755AA5FFA9FFA76D2CF8EDFC45E721" baseAuthorityName="Kulczynski" baseAuthorityYear="1895" box="[833,958,1881,1904]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="209" phylum="Arthropoda" rank="species" species="caucasia">
|
||
<emphasis id="B901FD34FFA9FFA76D2CF8EDFC45E721" box="[833,958,1881,1904]" italics="true" pageId="14" pageNumber="209">H. caucasia</emphasis>
|
||
</taxonomicName>
|
||
(D–J), left bulbs. A, D, H–J prolateral; B, E retrolateral, C, F, G ventral. D, H–I different aspects illustrating how the point of view affects the perception of the shape of the embolus and conductor. D–I lectotype, G from
|
||
<bibRefCitation id="EFE45CD7FFA9FFA76CA2F814FC6AE7E9" author="Kulczynski, W." box="[719,913,1952,1976]" pageId="14" pageNumber="209" pagination="3 - 38" refId="ref10449" refString="Kulczynski, W. (1895) Araneae a Dre G. Horvath in Bessarabia, Chersoneso Taurico, Transcaucasia et Armenia Russica collectae. Termesztrajzi Fuzetek, 18, 3 - 38." type="journal article" year="1895">Kulczyński (1895)</bibRefCitation>
|
||
, J from
|
||
<bibRefCitation id="EFE45CD7FFA9FFA76D86F814FB86E7E9" author="Dunin, P. M." box="[1003,1149,1952,1976]" pageId="14" pageNumber="209" pagination="35 - 76" refId="ref10097" refString="Dunin, P. M. (1992 b) The spider family Dysderidae of the Caucasian fauna (Arachnida Aranei Haplogynae). Arthropoda Selecta, 1 (3), 35 - 76. [in Russian]" type="journal article" year="1992">Dunin (1992b</bibRefCitation>
|
||
, most likely misidentified). Abbreviations:
|
||
<emphasis id="B901FD34FFA9FFA76F59F870FEA9E78A" box="[308,338,1988,2011]" italics="true" pageId="14" pageNumber="209">Co</emphasis>
|
||
—‘conductor’,
|
||
<emphasis id="B901FD34FFA9FFA76F82F871FDEAE78D" box="[495,529,1989,2012]" italics="true" pageId="14" pageNumber="209">EH</emphasis>
|
||
—embolic heel,
|
||
<emphasis id="B901FD34FFA9FFA76CD4F871FD3CE78D" box="[697,711,1989,2012]" italics="true" pageId="14" pageNumber="209">T</emphasis>
|
||
—tegulum.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C36F72ADFFA8FFA66EAAFF22FCE8E134" pageId="15" pageNumber="210" type="discussion">
|
||
<paragraph id="8BCA2126FFA8FFA66EAAFF22FCE8E134" blockId="15.[151,1437,150,393]" pageId="15" pageNumber="210">
|
||
<emphasis id="B901FD34FFA8FFA66EAAFF22FEB4E0E1" bold="true" box="[199,335,150,176]" pageId="15" pageNumber="210">Comments.</emphasis>
|
||
We had the opportunity to examine photographs of the
|
||
<typeStatus id="54CE9F84FFA8FFA66DD0FF23FBE4E0E0" box="[957,1055,151,177]" pageId="15" pageNumber="210" type="holotype">holotype</typeStatus>
|
||
male of
|
||
<taxonomicName id="4C755AA5FFA8FFA66AECFF23FAFDE0E1" baseAuthorityName="Kulczynski" baseAuthorityYear="1895" box="[1153,1286,151,177]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="210" phylum="Arthropoda" rank="species" species="caucasia">
|
||
<emphasis id="B901FD34FFA8FFA66AECFF23FB6EE0E1" box="[1153,1173,151,176]" italics="true" pageId="15" pageNumber="210">H</emphasis>
|
||
.
|
||
<emphasis id="B901FD34FFA8FFA66ACFFF23FAFDE0E1" box="[1186,1286,151,176]" italics="true" pageId="15" pageNumber="210">caucasia</emphasis>
|
||
</taxonomicName>
|
||
and compare them with our new species. Although
|
||
<bibRefCitation id="EFE45CD7FFA8FFA66C50FF0FFD17E084" author="Brignoli, P. M." box="[573,748,187,213]" pageId="15" pageNumber="210" pagination="75 - 82" refId="ref9778" refString="Brignoli, P. M. (1980) Some new or interesting eastern Mediterranean Dysderidae and Agelenidae (Araneae). Annales Zoologici, Warszawa, 35, 75 - 82." type="journal article" year="1980">Brignoli (1980)</bibRefCitation>
|
||
mentioned that he had studied this material, his figures of the male palp are similar to those in
|
||
<bibRefCitation id="EFE45CD7FFA8FFA66C64FF6BFD4BE0A8" author="Dunin, P. M." box="[521,688,223,249]" pageId="15" pageNumber="210" pagination="35 - 76" refId="ref10097" refString="Dunin, P. M. (1992 b) The spider family Dysderidae of the Caucasian fauna (Arachnida Aranei Haplogynae). Arthropoda Selecta, 1 (3), 35 - 76. [in Russian]" type="journal article" year="1992">Dunin (1992b)</bibRefCitation>
|
||
and yet different from the
|
||
<typeStatus id="54CE9F84FFA8FFA66D8DFF6BFBB9E0A8" box="[992,1090,223,249]" pageId="15" pageNumber="210" type="holotype">holotype</typeStatus>
|
||
(cf.
|
||
<figureCitation id="134E3DA3FFA8FFA66A15FF6BFAA0E0A8" box="[1144,1371,222,249]" captionStart="FIGURE 15" captionStartId="14.[152,255,1880,1904]" captionTargetBox="[151,1426,164,1863]" captionTargetId="figure-32@14.[151,1436,143,1863]" captionTargetPageId="14" captionText="FIGURE 15. Males of Harpactea abasgiana sp. n. (A–C) and H. caucasia (D–J), left bulbs. A, D, H–J prolateral; B, E retrolateral, C, F, G ventral. D, H–I different aspects illustrating how the point of view affects the perception of the shape of the embolus and conductor. D–I lectotype, G from Kulczyński (1895), J from Dunin (1992b, most likely misidentified). Abbreviations: Co—‘conductor’, EH—embolic heel, T—tegulum." figureDoi="http://doi.org/10.5281/zenodo.10468597" httpUri="https://zenodo.org/record/10468597/files/figure.png" pageId="15" pageNumber="210">Fig. 15J and D, H, I</figureCitation>
|
||
). It is not clear what specimens were erroneously illustrated as
|
||
<taxonomicName id="4C755AA5FFA8FFA66D70FEB7FC5EE14D" baseAuthorityName="Kulczynski" baseAuthorityYear="1895" box="[797,933,259,285]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="210" phylum="Arthropoda" rank="species" species="caucasia">
|
||
<emphasis id="B901FD34FFA8FFA66D70FEB7FCCAE14D" box="[797,817,259,284]" italics="true" pageId="15" pageNumber="210">H</emphasis>
|
||
.
|
||
<emphasis id="B901FD34FFA8FFA66D2CFEB7FC5EE14D" box="[833,933,259,284]" italics="true" pageId="15" pageNumber="210">caucasia</emphasis>
|
||
</taxonomicName>
|
||
by these authors, but they seem to represent an undescribed species. Most likely, all records of
|
||
<taxonomicName id="4C755AA5FFA8FFA66CA6FE93FCAAE111" baseAuthorityName="Kulczynski" baseAuthorityYear="1895" box="[715,849,295,321]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="210" phylum="Arthropoda" rank="species" species="caucasia">
|
||
<emphasis id="B901FD34FFA8FFA66CA6FE93FD24E111" box="[715,735,295,320]" italics="true" pageId="15" pageNumber="210">H</emphasis>
|
||
.
|
||
<emphasis id="B901FD34FFA8FFA66C80FE93FCAAE111" box="[749,849,295,320]" italics="true" pageId="15" pageNumber="210">caucasia</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingRegion id="49B1EFC4FFA8FFA66DFFFE93FC06E110" box="[914,1021,295,321]" country="Georgia" name="Abkhazia" pageId="15" pageNumber="210">Abkhazia</collectingRegion>
|
||
and probably Northern Caucasus are based on misidentified specimens of
|
||
<taxonomicName id="4C755AA5FFA8FFA66C5BFEFFFD36E134" authority="Zamani & Marusik, 2024" authorityName="Zamani & Marusik" authorityYear="2024" box="[566,717,331,357]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="210" phylum="Arthropoda" rank="species" species="abasgiana" status="sp. n.">
|
||
<emphasis id="B901FD34FFA8FFA66C5BFEFFFDB1E135" box="[566,586,331,356]" italics="true" pageId="15" pageNumber="210">H</emphasis>
|
||
.
|
||
<emphasis id="B901FD34FFA8FFA66C35FEFFFD36E134" box="[600,717,331,357]" italics="true" pageId="15" pageNumber="210">abasgiana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B901FD34FFA8FFA66CB9FEFFFCE8E134" bold="true" box="[724,787,331,357]" pageId="15" pageNumber="210">
|
||
<taxonomicNameLabel id="A232404FFFA8FFA66CB9FEFFFCE8E134" box="[724,787,331,357]" pageId="15" pageNumber="210" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36F72ADFFA8FFA66EAAFEDAFAD9E1D8" box="[199,1314,366,393]" pageId="15" pageNumber="210" type="distribution">
|
||
<paragraph id="8BCA2126FFA8FFA66EAAFEDAFAD9E1D8" blockId="15.[151,1437,150,393]" box="[199,1314,366,393]" pageId="15" pageNumber="210">
|
||
<emphasis id="B901FD34FFA8FFA66EAAFEDAFE9AE1D9" bold="true" box="[199,353,366,392]" pageId="15" pageNumber="210">Distribution.</emphasis>
|
||
Known only from the listed localities in the Republic of
|
||
<collectingRegion id="49B1EFC4FFA8FFA66DB2FEDBFBB6E1D8" box="[991,1101,367,393]" country="Georgia" name="Abkhazia" pageId="15" pageNumber="210">Abkhazia</collectingRegion>
|
||
,
|
||
<collectingCountry id="F36261B6FFA8FFA66A35FEDBFB49E1D8" box="[1112,1202,367,393]" name="Georgia" pageId="15" pageNumber="210">Georgia</collectingCountry>
|
||
(
|
||
<figureCitation id="134E3DA3FFA8FFA66AACFEDBFAEEE1D8" box="[1217,1301,367,393]" captionStart="FIGURE 19" captionStartId="20.[152,255,1371,1395]" captionTargetBox="[151,1436,705,1348]" captionTargetId="figure-288@20.[151,1436,705,1348]" captionTargetPageId="20" captionText="FIGURE 19. Distribution records of Dysdera atabekia sp. n. (circles), Harpactea abasgiana sp. n. (squares), and H. dunini sp. n. (triangle). Type localities are marked with red." figureDoi="http://doi.org/10.5281/zenodo.10468607" httpUri="https://zenodo.org/record/10468607/files/figure.png" pageId="15" pageNumber="210">Fig. 19</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |