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<mods:title id="386C58A5CFC31CAAD3EA1F922E8147CC">Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:namePart id="3D6C1A08DB29489D935404493A407BAF">Maruyama, Munetoshi</mods:namePart>
<mods:affiliation id="C6C8792C289315BA188EC7B2B00F6226">The Kyushu University Museum, Hakozaki</mods:affiliation>
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<heading id="D0950441AC2AFFA24F99FE6615E1FE78" box="[209,702,413,437]" centered="true" fontSize="10" level="2" pageId="32" pageNumber="195" reason="3">
<taxonomicName id="4C62C8AEAC2AFFA24F99FE6615E1FE78" ID-CoL="NJWH" authority="Ahn &amp; Jeon, 2004" authorityName="Ahn &amp; Jeon" authorityYear="2004" box="[209,702,413,437]" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="195" phylum="Arthropoda" rank="species" species="gangjinensis">
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<emphasis id="B9166F3FAC2AFFA24F99FE6616B4FE78" bold="true" box="[209,491,413,437]" italics="true" pageId="32" pageNumber="195">Bryothinusa gangjinensis</emphasis>
<bibRefCitation id="EFF3CEDCAC2AFFA24EBAFE6515E1FE78" author="AHN K. J. &amp; JEON M. J." box="[498,702,414,437]" pageId="32" pageNumber="195" pagination="29 - 35" refId="ref25135" refString="AHN K. J. &amp; JEON M. J. 2004: The Korean species of the genus Bryothinusa Casey with descriptions of two new species (Coleoptera: Staphylinidae: Aleocharinae). Coleopterists Bulletin 58: 29 - 35." type="journal article" year="2004">Ahn &amp; Jeon, 2004</bibRefCitation>
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</paragraph>
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<paragraph id="8BDDB32DAC2AFFA24E1DFE471565FE02" blockId="32.[151,761,444,684]" box="[341,570,444,463]" pageId="32" pageNumber="195">
(
<figureCitation id="1359AFA8AC2AFFA24E13FE471697FE02" box="[347,456,444,463]" captionStart="Fig" captionStartId="2.[151,182,1972,1991]" captionTargetBox="[189,1405,205,1944]" captionTargetId="figure-13@2.[175,1411,189,1947]" captionTargetPageId="2" captionText="Fig. 1. Habitus of Bryothinusa spp. A B. algarum Sawada, 1971; B B. minuta (Sawada, 1955); C B. tsutsuii (Sawada, 1955); D B. koreana Ahn &amp; Jeon, 2004; E B. yoshigoui sp. nov.; F B. japonica Liu, Ono &amp; Maruyama, 2020; G B. sakishimana Sawada, 1991; H B. constricta sp. nov.; I B. hokkaidensis sp. nov.; J B. fulvipennis sp. nov.; K B. itsuroi sp. nov.; L B. aikoae sp. nov.; M B. okinawana sp. nov.; N B. nigra sp. nov.; O B. nakanei (Sawada, 1955); P B. moriguchii sp. nov.; Q B. gangjinensis Ahn &amp; Jeon, 2004. Scale bars: 1.00 mm." figureDoi="http://doi.org/10.5281/zenodo.5037332" httpUri="https://zenodo.org/record/5037332/files/figure.png" pageId="32" pageNumber="195">Figs 1Q; 1Q</figureCitation>
;
<figureCitation id="1359AFA8AC2AFFA24E99FE4716B7FE02" box="[465,488,444,463]" captionStart="Fig" captionStartId="31.[150,181,2034,2053]" captionTargetBox="[150,1436,204,1992]" captionTargetId="figure-14@31.[150,1436,204,1992]" captionTargetPageId="31" captionText="Fig. 19. Bryothinusa gangjinensis Ahn &amp; Jeon, 2004.A labrum; B mandible, dorsal view; C lacinia, dorsal view; D labium; E mentum; F male sternite VIII; G female sternite VIII; H median lobe, ventral view; I median lobe, lateral view; J paramere; K spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037370" httpUri="https://zenodo.org/record/5037370/files/figure.png" pageId="32" pageNumber="195">19</figureCitation>
;
<figureCitation id="1359AFA8AC2AFFA24EB9FE47154AFE02" box="[497,533,444,463]" captionStart="Fig" captionStartId="34.[151,182,1955,1974]" captionTargetBox="[167,1408,193,1935]" captionTargetId="figure-13@34.[161,1426,189,1938]" captionTargetPageId="34" captionText="Fig. 21. Collection localities of Bryothinusa species in Japan: A B. koreana Ahn &amp; Jeon, 2004 and B. japonica Liu, Ono &amp; Maruyama, 2020 in Shima-nogita, Fukuoka; B B. aikoae sp. nov., B. fulvipennis sp. nov., B. japonica and B. nigra sp. nov. in Kuroto, Chiba; C B. hokkaidensis sp. nov. in Shunkunitai, Hokkaidô; D B. nigra sp. nov. and B. tsutsuii (Sawada, 1955) in Edateku-jima; E B. nakanei (Sawada, 1955) in Nagura-anparu, Ishigaki-jima; F B. gangjinensis Ahn &amp; Jeon, 2004 in Iki-no-matsubara, Fukuoka; G B. algarum Sawada, 1971, B. japonica and B. minuta (Sawada, 1955) in Sunosaki, Fukuoka." figureDoi="http://doi.org/10.5281/zenodo.5037374" httpUri="https://zenodo.org/record/5037374/files/figure.png" pageId="32" pageNumber="195">21F</figureCitation>
;
<figureCitation id="1359AFA8AC2AFFA24D56FE47156BFE02" box="[542,564,444,463]" captionStart="Fig" captionStartId="35.[150,181,1388,1407]" captionTargetBox="[150,1434,191,1370]" captionTargetId="figure-236@35.[150,1435,191,1371]" captionTargetPageId="35" captionText="Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle B. algarum Sawada, 1971; black inverted triangle B. minuta (Sawada, 1955); white star B. tsutsuii (Sawada, 1955); black star B. koreana Ahn &amp; Jeon, 2004; black square B. yoshigoui sp. nov.; white triangle B. japonica Liu, Ono &amp; Maruyama, 2020; black diamond B. sakishimana Sawada, 1991; white diamond B. constricta sp. nov.; white square B. hokkaidensis sp. nov.; black triangle B. fulvipennis sp. nov.; double circle B. itsuroi sp. nov.; arrow B. aikoae sp. nov.; black hart B. okinawana sp. nov.; circled dot B. nigra sp. nov.; white circle B. nakanei (Sawada, 1955); white hart B. moriguchii sp. nov.; black circle B. gangjinensis Ahn &amp; Jeon, 2004." figureDoi="http://doi.org/10.5281/zenodo.5037376" httpUri="https://zenodo.org/record/5037376/files/figure.png" pageId="32" pageNumber="195">22</figureCitation>
)
</paragraph>
<paragraph id="8BDDB32DAC2AFFA24E4EFE2D15D6FE24" blockId="32.[151,761,444,684]" box="[262,649,470,489]" pageId="32" pageNumber="195">(Japanese name: Kanjin-nagisa-hanekakushi)</paragraph>
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<treatmentCitationGroup id="AB729403AC2AFFA24FDFFE0315A6FDC1" box="[151,761,504,524]" pageId="32" pageNumber="195">
<treatmentCitation id="0AC3953CAC2AFFA24FDFFE031564FDC1" author="AHN K. J. &amp; JEON M. J." box="[151,571,504,524]" httpUri="http://treatment.plazi.org/id/03E387AAAA71FFA50AA0FA15F0EFFC77" page="29" pageId="32" pageNumber="195" year="2004">
<taxonomicName id="4C62C8AEAC2AFFA24FDFFE031564FDC1" ID-CoL="NJWH" authority="Ahn &amp; Jeon, 2004: 29" authorityName="Ahn &amp; Jeon" authorityPageNumber="29" authorityYear="2004" box="[151,571,504,524]" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="195" phylum="Arthropoda" rank="species" species="gangjinensis">
<emphasis id="B9166F3FAC2AFFA24FDFFE031628FDC6" box="[151,375,504,523]" italics="true" pageId="32" pageNumber="195">Bryothinusa gangjinensis</emphasis>
<bibRefCitation id="EFF3CEDCAC2AFFA24E31FE021564FDC1" author="AHN K. J. &amp; JEON M. J." box="[377,571,505,524]" pageId="32" pageNumber="195" pagination="29 - 35" refId="ref25135" refString="AHN K. J. &amp; JEON M. J. 2004: The Korean species of the genus Bryothinusa Casey with descriptions of two new species (Coleoptera: Staphylinidae: Aleocharinae). Coleopterists Bulletin 58: 29 - 35." type="journal article" year="2004">Ahn &amp; Jeon, 2004: 29</bibRefCitation>
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(original description;
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</paragraph>
<paragraph id="8BDDB32DAC2AFFA24FFDFDE815A7FDEB" blockId="32.[151,761,444,684]" box="[181,760,531,550]" pageId="32" pageNumber="195">
<typeStatus id="54D90D8FAC2AFFA24FFDFDE81785FDEB" box="[181,218,531,550]" pageId="32" pageNumber="195">type</typeStatus>
locality:
<collectingCountry id="F375F3BDAC2AFFA24E64FDE8163CFDEB" box="[300,355,531,550]" name="South Korea" pageId="32" pageNumber="195">Korea</collectingCountry>
:
<collectingRegion id="49A67DCFAC2AFFA24E24FDE816E3FDEB" box="[364,444,531,550]" country="South Korea" name="Jeonnam" pageId="32" pageNumber="195">Jeonnam</collectingRegion>
); Aඌ*üൾ (2004): 591 (in annotatedca-
</paragraph>
</subSubSection>
<subSubSection id="C378E0A6AC2AFFA24FFDFDD515A6FD61" pageId="32" pageNumber="195" type="description">
<paragraph id="8BDDB32DAC2AFFA24FFDFDD515A7FD8C" blockId="32.[151,761,444,684]" box="[181,760,558,577]" pageId="32" pageNumber="195">talogue of world species);Aඌൺඇඈ &amp; Kඈ*°ංආൺ (2010): 297 (new record</paragraph>
<paragraph id="8BDDB32DAC2AFFA24FFDFDB215A7FD91" blockId="32.[151,761,444,684]" box="[181,760,585,604]" pageId="32" pageNumber="195">
from
<collectingCountry id="F375F3BDAC2AFFA24FABFDB21648FD91" box="[227,279,585,604]" name="Japan" pageId="32" pageNumber="195">Japan</collectingCountry>
: Tanega-shima); F*©ൺඇκ &amp; A*üඇ (2011): 26 (in checklist);
</paragraph>
<paragraph id="8BDDB32DAC2AFFA24FFDFD9815A6FDBB" blockId="32.[151,761,444,684]" box="[181,761,611,630]" pageId="32" pageNumber="195">Sർ*ü*“අκൾ &amp; Sආൾඍൺඇൺ (2015): 676 (in catalogue of Palearctic species);</paragraph>
<paragraph id="8BDDB32DAC2AFFA24FFDFD8515A6FD5C" blockId="32.[151,761,444,684]" box="[181,761,638,657]" pageId="32" pageNumber="195">
A*üඇ et al. (2017): 307 (in checklist of the
<taxonomicName id="4C62C8AEAC2AFFA24D66FD8515FCFD5C" box="[558,675,638,657]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="195" phylum="Arthropoda" rank="family">Staphylinidae</taxonomicName>
in
<collectingCountry id="F375F3BDAC2AFFA24DF7FD8515B0FD5C" box="[703,751,638,657]" name="South Korea" pageId="32" pageNumber="195">Korea</collectingCountry>
);
</paragraph>
<paragraph id="8BDDB32DAC2AFFA24FFDFD6215A6FD61" blockId="32.[151,761,444,684]" box="[181,761,665,684]" pageId="32" pageNumber="195">Lඎ et al. (2020): 589 (in key of Japanese species), 592 (in checklist).</paragraph>
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<subSubSection id="C378E0A6AC2AFFA24FDFFD39165BF9DD" pageId="32" pageNumber="195" type="materials_examined">
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<emphasis id="B9166F3FAC2AFFA24FDFFD391643FD18" bold="true" box="[151,284,706,725]" pageId="32" pageNumber="195">
<typeStatus id="54D90D8FAC2AFFA24FDFFD39179BFD18" box="[151,196,706,725]" pageId="32" pageNumber="195">Type</typeStatus>
material.
</emphasis>
Not examined.
</paragraph>
<paragraph id="8BDDB32DAC2AFFA24FDFFD26165BF9DD" blockId="32.[151,761,706,1552]" pageId="32" pageNumber="195">
<emphasis id="B9166F3FAC2AFFA24FDFFD261551FD3D" bold="true" box="[151,526,733,752]" pageId="32" pageNumber="195">
Material examined.
<collectingCountry id="F375F3BDAC2AFFA24E17FD2616F6FD3D" box="[351,425,733,752]" name="Japan" pageId="32" pageNumber="195">JAPAN</collectingCountry>
: HඈඇඌΗǕ:
</emphasis>
2 JJ 17 unsexed spec., Mouth of Minato-gawa, Minato, Futtsu-shi, Chiba-ken,
<date id="FFDC95EDAC2AFFA24DC9FD031585FCC6" box="[641,730,760,779]" pageId="32" pageNumber="195" value="2013-05-27">27.v.2013</date>
, H. Ono; 4 JJ
<specimenCount id="9D6478A4AC2AFFA24FB5FCE91642FCE8" box="[253,285,780,805]" pageId="32" pageNumber="195" type="female">1 ♀</specimenCount>
, 24 unsexed spec., same locality,
<date id="FFDC95EDAC2AFFA24D0BFCE915F2FCE8" box="[579,685,786,805]" pageId="32" pageNumber="195" value="2009-08-24">24.viii.2009</date>
H. Ono; 2 JJ
<specimenCount id="9D6478A4AC2AFFA24F9AFCD6165AFC8D" box="[210,261,807,832]" pageId="32" pageNumber="195" type="female">2 ♀♀</specimenCount>
, 7 unsexed spec., Koajiro-wan, Koajiro, Misaki-machi, Miura-shi, Kanagawa-ken,
<date id="FFDC95EDAC2AFFA24ECEFCB31681FC96" box="[390,478,840,859]" pageId="32" pageNumber="195" value="2013-05-23">23.v.2013</date>
, H. Ono; 1 J
<specimenCount id="9D6478A4AC2AFFA24D11FCB315D6FC97" box="[601,649,833,859]" pageId="32" pageNumber="195" type="female">2 ♀♀</specimenCount>
, 67 unsexed spec., Miya-gawa, Iso-chô, Ise-shi, Mie-ken,
<date id="FFDC95EDAC2AFFA24D5FFC991535FCB8" box="[535,618,866,885]" pageId="32" pageNumber="195" value="2008-06-08">8.vi.2008</date>
, K.
<collectingRegion id="49A67DCFAC2AFFA24DC2FC9915E2FCB8" box="[650,701,866,885]" country="Japan" name="Akita" pageId="32" pageNumber="195">Akita</collectingRegion>
;
<specimenCount id="9D6478A4AC2AFFA24D8CFC9915ACFCB8" box="[708,755,860,885]" pageId="32" pageNumber="195" type="female">2 ♀♀</specimenCount>
, 6 unsexed spec., Hannan 2-ku, Kishiwada-shi, Ôsaka-fu,
<date id="FFDC95EDAC2AFFA24DC8FC861581FC5D" box="[640,734,893,912]" pageId="32" pageNumber="195" value="2010-06-11">11.vi.2010</date>
, Y. Kawakami;
<specimenCount id="9D6478A4AC2AFFA24FB6FC631643FC67" box="[254,284,913,939]" pageId="32" pageNumber="195" type="female">1 ♀</specimenCount>
, 5 unsexed spec., same locality,
<date id="FFDC95EDAC2AFFA24D64FC631527FC66" box="[556,632,920,939]" pageId="32" pageNumber="195" value="2010-10-07">7.x.2010</date>
, Y.Kawakami;
<specimenCount id="9D6478A4AC2AFFA24FDFFC4917E9FC08" box="[151,182,940,965]" pageId="32" pageNumber="195" type="female">1 ♀</specimenCount>
, Arida-gawa, Miyazaki-chô, Arida-shi, Wakayama-ken,
<date id="FFDC95EDAC2AFFA24DD5FC4915ABFC08" box="[669,756,946,965]" pageId="32" pageNumber="195" value="2002-05-28">28.v.2002</date>
, Y. Kawakami.
<emphasis id="B9166F3FAC2AFFA24E5FFC36162DFC2D" bold="true" box="[279,370,973,992]" pageId="32" pageNumber="195">SΗκඈκඎ:</emphasis>
<specimenCount id="9D6478A4AC2AFFA24E3DFC3616CBFC2D" box="[373,404,967,992]" pageId="32" pageNumber="195" type="female">1 ♀</specimenCount>
, 1 unsexed spec., Kabe-gawa, Kamoshô, Sanuki-shi, Kagawa-ken,
<date id="FFDC95EDAC2AFFA24ECAFC1316BEFC36" box="[386,481,1000,1019]" pageId="32" pageNumber="195" value="2008-04-13">13.iv.2008</date>
, H. Fujimoto;
<specimenCount id="9D6478A4AC2AFFA24D23FC1315D1FC37" box="[619,654,993,1019]" pageId="32" pageNumber="195" type="female">1 ♀</specimenCount>
, 2 unsexed spec., Kôtô-gawa, Gôtô-chô, Takamatsu-shi, Kagawa-ken,
<date id="FFDC95EDAC2AFFA24DDEFBF915ABFBD8" box="[662,756,1026,1045]" pageId="32" pageNumber="195" value="2008-04-12">12.iv.2008</date>
, H. Fujimoto; 1 J 3 unsexed spec., Kamobe-gawa, Kamoshô, Sanuki -shi, Kagawa-ken,
<date id="FFDC95EDAC2AFFA24E74FBC316C6FB86" box="[316,409,1080,1099]" pageId="32" pageNumber="195" value="2008-04-12">12.iv.2008</date>
, H. Fujimoto; 1 J, Aya-gawa, Ejiri-chô, Sakaide-shi, Kagawa-ken,
<date id="FFDC95EDAC2AFFA24ECCFBA916BDFBA8" box="[388,482,1106,1125]" pageId="32" pageNumber="195" value="2008-04-12">12.iv.2008</date>
, H. Fujimoto; 2 unsexed spec., Nishihabu-machi, Matsuyama-shi, Ehime-ken,
<date id="FFDC95EDAC2AFFA24D64FB9615D6FB4D" box="[556,649,1133,1152]" pageId="32" pageNumber="195" value="2009-04-19">19.iv.2009</date>
, H. Suenaga.
<emphasis id="B9166F3FAC2AFFA24FDFFB7317B4FB56" bold="true" box="[151,235,1160,1179]" pageId="32" pageNumber="195">KඒǕඌΗǕ:</emphasis>
1J, Iki-no-matsubara, Nishi-ku, Fukuoka-shi, Fukuoka-ken,
<date id="FFDC95EDAC2AFFA24FDFFB5917ADFB78" box="[151,242,1186,1205]" pageId="32" pageNumber="195" value="2018-07-01">1.vii.2018</date>
, Y. Hisasue; 1 J
<specimenCount id="9D6478A4AC2AFFA24ED8FB5916ECFB78" box="[400,435,1180,1205]" pageId="32" pageNumber="195" type="female">1 ♀</specimenCount>
, 13 unsexed spec., Chichibu-ga-ura, Shimabara, Nagasaki-ken,
<date id="FFDC95EDAC2AFFA24EC2FB4616BDFB1D" box="[394,482,1213,1232]" pageId="32" pageNumber="195" value="2001-05-10">10.v.2001</date>
,
<collectionCode id="ED732BE8AC2AFFA24EA6FB4616A3FB1D" box="[494,508,1213,1232]" name="Tavera, Department of Geology and Geophysics" pageId="32" pageNumber="195">T</collectionCode>
. Watanabe;
<specimenCount id="9D6478A4AC2AFFA24D24FB4615D0FB1D" box="[620,655,1207,1232]" pageId="32" pageNumber="195" type="female">1 ♀</specimenCount>
, 3 unsexed spec., Kawashima, Nobeoka-shi, Miyazaki-ken,
<date id="FFDC95EDAC2AFFA24D08FB2315FAFB26" box="[576,677,1240,1259]" pageId="32" pageNumber="195" value="2006-07-24">24.vii.2006</date>
, J. Nakajima;
<specimenCount id="9D6478A4AC2AFFA24F86FB0917B1FAC8" box="[206,238,1260,1285]" pageId="32" pageNumber="195" type="female">1 ♀</specimenCount>
, 4 unsexed spec., Nagayoshi-kawa, Fukiage-chô, Hioki-shi, Kagoshima-ken,
<date id="FFDC95EDAC2AFFA24E64FAF616D8FAED" box="[300,391,1293,1312]" pageId="32" pageNumber="195" value="2007-12-09">9.xii.2007</date>
, K. Nakamura;
<specimenCount id="9D6478A4AC2AFFA24D5AFAF6156CFAED" box="[530,563,1287,1312]" pageId="32" pageNumber="195" type="female">1 ♀</specimenCount>
, 6 unsexed spec., Manose-gawa, Masukawa, Kaseda, Minamisatsuma-shi, Kagoshima-ken,
<date id="FFDC95EDAC2AFFA24FDFFAB917AAFA98" box="[151,245,1346,1365]" pageId="32" pageNumber="195" value="2008-04-20">20.iv.2008</date>
, M. Maruyama; 1 J
<specimenCount id="9D6478A4AC2AFFA24EEEFAB9168CFA98" box="[422,467,1340,1365]" pageId="32" pageNumber="195" type="female">2♀♀</specimenCount>
, 11 unsexed spec., Aiboshi-gawa, Aiboshi, Minamisatsuma-shi, Kagoshima-ken,
<date id="FFDC95EDAC2AFFA24D64FAA615D6FABD" box="[556,649,1373,1392]" pageId="32" pageNumber="195" value="2008-04-20">20.iv.2008</date>
, M. Maruyama.
<emphasis id="B9166F3FAC2AFFA24FF6FA831614FA46" bold="true" box="[190,331,1400,1419]" pageId="32" pageNumber="195">Nൺඇඌൾ-ඌΗඈඍත:</emphasis>
3 JJ 6 unsexed spec., Ou-jima, Nago-shi, Okinawa-ken,
<date id="FFDC95EDAC2AFFA24FBFFA69160AFA68" box="[247,341,1426,1445]" pageId="32" pageNumber="195" value="2006-04-30">30.iv.2006</date>
, M. Moriguchi; 2 unsexed spec., Awase-Higata, Okinawa-shi, Okinawa-ken,
<date id="FFDC95EDAC2AFFA24ED2FA5616A6FA0D" box="[410,505,1453,1472]" pageId="32" pageNumber="195" value="2006-04-30">30.iv.2006</date>
, M. Moriguchi; 11 unsexed spec., Makiya, Nago-shi, Okinawa-jima, Okinawa-ken,
<date id="FFDC95EDAC2AFFA24D34FA331588FA16" box="[636,727,1480,1499]" pageId="32" pageNumber="195" value="2010-07-04">4.vii.2010</date>
, M. Moriguchi. Deposited at
<collectionCode id="ED732BE8AC2AFFA24E3AFA1916FBFA38" box="[370,420,1506,1525]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="32" pageNumber="195" type="Herbarium">KUM</collectionCode>
but some will be distributed to
<collectionCode id="ED732BE8AC2AFFA24DF2FA1915A8FA38" box="[698,759,1506,1525]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="32" pageNumber="195">NMPC</collectionCode>
and
<collectionCode id="ED732BE8AC2AFFA24FF4FA06165FF9DD" box="[188,256,1533,1552]" pageId="32" pageNumber="195">BMNH</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C378E0A6AC2AFFA24FDFF9D2123CFCB8" pageId="32" pageNumber="195" type="description">
<paragraph id="8BDDB32DAC2AFFA24FDFF9D21515F90D" blockId="32.[151,760,1577,2080]" pageId="32" pageNumber="195">
<emphasis id="B9166F3FAC2AFFA24FDFF9D21662F98D" bold="true" box="[151,317,1577,1600]" pageId="32" pageNumber="195">Redescription.</emphasis>
Body (
<figureCitation id="1359AFA8AC2AFFA24EC4F9D21681F98D" box="[396,478,1577,1600]" captionStart="Fig" captionStartId="2.[151,182,1972,1991]" captionTargetBox="[189,1405,205,1944]" captionTargetId="figure-13@2.[175,1411,189,1947]" captionTargetPageId="2" captionText="Fig. 1. Habitus of Bryothinusa spp. A B. algarum Sawada, 1971; B B. minuta (Sawada, 1955); C B. tsutsuii (Sawada, 1955); D B. koreana Ahn &amp; Jeon, 2004; E B. yoshigoui sp. nov.; F B. japonica Liu, Ono &amp; Maruyama, 2020; G B. sakishimana Sawada, 1991; H B. constricta sp. nov.; I B. hokkaidensis sp. nov.; J B. fulvipennis sp. nov.; K B. itsuroi sp. nov.; L B. aikoae sp. nov.; M B. okinawana sp. nov.; N B. nigra sp. nov.; O B. nakanei (Sawada, 1955); P B. moriguchii sp. nov.; Q B. gangjinensis Ahn &amp; Jeon, 2004. Scale bars: 1.00 mm." figureDoi="http://doi.org/10.5281/zenodo.5037332" httpUri="https://zenodo.org/record/5037332/files/figure.png" pageId="32" pageNumber="195">Fig. 1Q</figureCitation>
) small, about
<quantity id="4C9A1EC8AC2AFFA24D3FF9D215A8F98D" box="[631,759,1577,1600]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" metricValueMax="2.4" metricValueMin="1.9" pageId="32" pageNumber="195" unit="mm" value="2.15" valueMax="2.4" valueMin="1.9">1.92.4 mm</quantity>
in length, covered with minute setae. Abdomen almost black, darker than head and pronotum; elytra darker than pronotum, bicoloured but not clearly, base darker than apex; antenna and legs yellowish brown.
</paragraph>
<paragraph id="8BDDB32DAC2AFFA24FFDF932156BF80D" blockId="32.[151,760,1577,2080]" pageId="32" pageNumber="195">
Head evenly rounded, more or less produced anteriorly, wider than long, shorter than elytra; eyes (
<figureCitation id="1359AFA8AC2AFFA24D12F91215F1F8CD" box="[602,686,1769,1792]" captionStart="Fig" captionStartId="4.[151,182,1978,1997]" captionTargetBox="[196,1391,190,1955]" captionTargetId="figure-13@4.[193,1394,189,1958]" captionTargetPageId="4" captionText="Fig. 2. Compound eyes of Bryothinusa spp.A B. algarum Sawada, 1971; B B. minuta (Sawada, 1955); C B. tsutsuii (Sawada, 1955); D B. koreana Ahn &amp; Jeon, 2004; E B. yoshigoui sp. nov.; F B. japonica Liu, Ono &amp; Maruyama, 2020; G B. sakishimana Sawada, 1991; H B. constricta sp. nov.; I B. hokkaidensis sp. nov.; J B. fulvipennis sp. nov.; K B. itsuroi sp. nov.; L B. aikoae sp. nov.; M B. okinawana sp. nov.; N B. nigra sp. nov.; O B. nakanei (Sawada, 1955); P B. moriguchii sp. nov.; Q B. gangjinensis Ahn &amp; Jeon, 2004. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5037334" httpUri="https://zenodo.org/record/5037334/files/figure.png" pageId="32" pageNumber="195">Fig. 2Q</figureCitation>
) large, setae present between about 3234 facets, slightly longer than 2/3 of head length; antennal length slightly longer than distance from anterior margin of head to posterior margin of pronotum; segment I 1.5 times longer than II, almost 2.8 times longer than III, and longer than XI; antennal segments VX almost as long as wide.
</paragraph>
<paragraph id="8BDDB32DAC2AFFA24FFDF83213A6FE58" blockId="32.[151,760,1577,2080]" lastBlockId="32.[828,1437,190,1365]" pageId="32" pageNumber="195">
Mouthparts (
<figureCitation id="1359AFA8AC2AFFA24E0AF8321696F82D" box="[322,457,1993,2016]" captionStart="Fig" captionStartId="31.[150,181,2034,2053]" captionTargetBox="[150,1436,204,1992]" captionTargetId="figure-14@31.[150,1436,204,1992]" captionTargetPageId="31" captionText="Fig. 19. Bryothinusa gangjinensis Ahn &amp; Jeon, 2004.A labrum; B mandible, dorsal view; C lacinia, dorsal view; D labium; E mentum; F male sternite VIII; G female sternite VIII; H median lobe, ventral view; I median lobe, lateral view; J paramere; K spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037370" httpUri="https://zenodo.org/record/5037370/files/figure.png" pageId="32" pageNumber="195">Figs 19AE</figureCitation>
) with distribution of labral setae as in
<figureCitation id="1359AFA8AC2AFFA24E5EF81216C7F7CD" box="[278,408,2025,2048]" captionStart="Fig" captionStartId="31.[150,181,2034,2053]" captionTargetBox="[150,1436,204,1992]" captionTargetId="figure-14@31.[150,1436,204,1992]" captionTargetPageId="31" captionText="Fig. 19. Bryothinusa gangjinensis Ahn &amp; Jeon, 2004.A labrum; B mandible, dorsal view; C lacinia, dorsal view; D labium; E mentum; F male sternite VIII; G female sternite VIII; H median lobe, ventral view; I median lobe, lateral view; J paramere; K spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037370" httpUri="https://zenodo.org/record/5037370/files/figure.png" pageId="32" pageNumber="195">Figure 19A</figureCitation>
; mandibles (
<figureCitation id="1359AFA8AC2AFFA24D6FF81215D5F7CD" box="[551,650,2025,2048]" captionStart="Fig" captionStartId="31.[150,181,2034,2053]" captionTargetBox="[150,1436,204,1992]" captionTargetId="figure-14@31.[150,1436,204,1992]" captionTargetPageId="31" captionText="Fig. 19. Bryothinusa gangjinensis Ahn &amp; Jeon, 2004.A labrum; B mandible, dorsal view; C lacinia, dorsal view; D labium; E mentum; F male sternite VIII; G female sternite VIII; H median lobe, ventral view; I median lobe, lateral view; J paramere; K spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037370" httpUri="https://zenodo.org/record/5037370/files/figure.png" pageId="32" pageNumber="195">Fig. 19B</figureCitation>
) elongate triangle, asymmetrical, right with densely median tooth, left mandible without tooth, blunt at apex; maxilla with apical margin of lacinia (
<figureCitation id="1359AFA8AC2AFFA24B72FF2513C8FF38" box="[1082,1175,222,245]" captionStart="Fig" captionStartId="31.[150,181,2034,2053]" captionTargetBox="[150,1436,204,1992]" captionTargetId="figure-14@31.[150,1436,204,1992]" captionTargetPageId="31" captionText="Fig. 19. Bryothinusa gangjinensis Ahn &amp; Jeon, 2004.A labrum; B mandible, dorsal view; C lacinia, dorsal view; D labium; E mentum; F male sternite VIII; G female sternite VIII; H median lobe, ventral view; I median lobe, lateral view; J paramere; K spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037370" httpUri="https://zenodo.org/record/5037370/files/figure.png" pageId="32" pageNumber="195">Fig. 19C</figureCitation>
) at right angle; at tip with two spines not clearly; 7 setae on inner margin of lacinia; labial palpus (
<figureCitation id="1359AFA8AC2AFFA24C9CFEE51369FEF8" box="[980,1078,286,309]" captionStart="Fig" captionStartId="31.[150,181,2034,2053]" captionTargetBox="[150,1436,204,1992]" captionTargetId="figure-14@31.[150,1436,204,1992]" captionTargetPageId="31" captionText="Fig. 19. Bryothinusa gangjinensis Ahn &amp; Jeon, 2004.A labrum; B mandible, dorsal view; C lacinia, dorsal view; D labium; E mentum; F male sternite VIII; G female sternite VIII; H median lobe, ventral view; I median lobe, lateral view; J paramere; K spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037370" httpUri="https://zenodo.org/record/5037370/files/figure.png" pageId="32" pageNumber="195">Fig. 19D</figureCitation>
) with most setae located basally; mentum (
<figureCitation id="1359AFA8AC2AFFA24CECFEC5135BFE98" box="[932,1028,318,341]" captionStart="Fig" captionStartId="31.[150,181,2034,2053]" captionTargetBox="[150,1436,204,1992]" captionTargetId="figure-14@31.[150,1436,204,1992]" captionTargetPageId="31" captionText="Fig. 19. Bryothinusa gangjinensis Ahn &amp; Jeon, 2004.A labrum; B mandible, dorsal view; C lacinia, dorsal view; D labium; E mentum; F male sternite VIII; G female sternite VIII; H median lobe, ventral view; I median lobe, lateral view; J paramere; K spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037370" httpUri="https://zenodo.org/record/5037370/files/figure.png" pageId="32" pageNumber="195">Fig. 19E</figureCitation>
) with antero-lateral angles prolonged into spinose processes; anterior margin deeply truncate; small “v seta” far away from apical setae.
</paragraph>
<paragraph id="8BDDB32DAC2AFFA24C12FE651214FE78" blockId="32.[828,1437,190,1365]" box="[858,1355,414,437]" pageId="32" pageNumber="195">Pronotum shorter than elytra, wider than head.</paragraph>
<paragraph id="8BDDB32DAC2AFFA24C12FE4514B5FE38" blockId="32.[828,1437,190,1365]" pageId="32" pageNumber="195">Elytra subquadrate, wider than pronotum; hind wings fully developed.</paragraph>
<paragraph id="8BDDB32DAC2AFFA24C12FE05148DFDB8" blockId="32.[828,1437,190,1365]" pageId="32" pageNumber="195">
Male abdominal sternite VIII (
<figureCitation id="1359AFA8AC2AFFA24BD6FE0513A6FDD8" box="[1182,1273,510,533]" captionStart="Fig" captionStartId="31.[150,181,2034,2053]" captionTargetBox="[150,1436,204,1992]" captionTargetId="figure-14@31.[150,1436,204,1992]" captionTargetPageId="31" captionText="Fig. 19. Bryothinusa gangjinensis Ahn &amp; Jeon, 2004.A labrum; B mandible, dorsal view; C lacinia, dorsal view; D labium; E mentum; F male sternite VIII; G female sternite VIII; H median lobe, ventral view; I median lobe, lateral view; J paramere; K spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037370" httpUri="https://zenodo.org/record/5037370/files/figure.png" pageId="32" pageNumber="195">Fig. 19F</figureCitation>
) with posterior margin slightly serrate. Median lobe of aedeagus and dorsal view as in
<figureCitation id="1359AFA8AC2AFFA24CE4FDC51318FD98" box="[940,1095,574,597]" captionStart="Fig" captionStartId="31.[150,181,2034,2053]" captionTargetBox="[150,1436,204,1992]" captionTargetId="figure-14@31.[150,1436,204,1992]" captionTargetPageId="31" captionText="Fig. 19. Bryothinusa gangjinensis Ahn &amp; Jeon, 2004.A labrum; B mandible, dorsal view; C lacinia, dorsal view; D labium; E mentum; F male sternite VIII; G female sternite VIII; H median lobe, ventral view; I median lobe, lateral view; J paramere; K spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037370" httpUri="https://zenodo.org/record/5037370/files/figure.png" pageId="32" pageNumber="195">Figures 19H, I</figureCitation>
; paramere (
<figureCitation id="1359AFA8AC2AFFA24B8BFDC51246FD98" box="[1219,1305,574,597]" captionStart="Fig" captionStartId="31.[150,181,2034,2053]" captionTargetBox="[150,1436,204,1992]" captionTargetId="figure-14@31.[150,1436,204,1992]" captionTargetPageId="31" captionText="Fig. 19. Bryothinusa gangjinensis Ahn &amp; Jeon, 2004.A labrum; B mandible, dorsal view; C lacinia, dorsal view; D labium; E mentum; F male sternite VIII; G female sternite VIII; H median lobe, ventral view; I median lobe, lateral view; J paramere; K spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037370" httpUri="https://zenodo.org/record/5037370/files/figure.png" pageId="32" pageNumber="195">Fig. 19J</figureCitation>
) with apical lobe elongate.
</paragraph>
<paragraph id="8BDDB32DAC2AFFA24C12FD8513A5FD38" blockId="32.[828,1437,190,1365]" pageId="32" pageNumber="195">
Female abdominal sternite VIII (
<figureCitation id="1359AFA8AC2AFFA24B98FD851268FD58" box="[1232,1335,638,661]" captionStart="Fig" captionStartId="31.[150,181,2034,2053]" captionTargetBox="[150,1436,204,1992]" captionTargetId="figure-14@31.[150,1436,204,1992]" captionTargetPageId="31" captionText="Fig. 19. Bryothinusa gangjinensis Ahn &amp; Jeon, 2004.A labrum; B mandible, dorsal view; C lacinia, dorsal view; D labium; E mentum; F male sternite VIII; G female sternite VIII; H median lobe, ventral view; I median lobe, lateral view; J paramere; K spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037370" httpUri="https://zenodo.org/record/5037370/files/figure.png" pageId="32" pageNumber="195">Fig. 19G</figureCitation>
) slightly sunken at median posterior margin but wider and smoother than in male. Spermatheca (
<figureCitation id="1359AFA8AC2AFFA24B10FD4513EAFD18" box="[1112,1205,702,725]" captionStart="Fig" captionStartId="31.[150,181,2034,2053]" captionTargetBox="[150,1436,204,1992]" captionTargetId="figure-14@31.[150,1436,204,1992]" captionTargetPageId="31" captionText="Fig. 19. Bryothinusa gangjinensis Ahn &amp; Jeon, 2004.A labrum; B mandible, dorsal view; C lacinia, dorsal view; D labium; E mentum; F male sternite VIII; G female sternite VIII; H median lobe, ventral view; I median lobe, lateral view; J paramere; K spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037370" httpUri="https://zenodo.org/record/5037370/files/figure.png" pageId="32" pageNumber="195">Fig. 19K</figureCitation>
) large, about 1.5 times coiled near base and larger than
<quantity id="4C9A1EC8AC2AFFA24BDCFD2513A8FD38" box="[1172,1271,734,757]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="32" pageNumber="195" unit="mm" value="0.25">0.25 mm</quantity>
.
</paragraph>
<paragraph id="8BDDB32DAC2AFFA24C74FD05123CFCB8" blockId="32.[828,1437,190,1365]" pageId="32" pageNumber="195">
<emphasis id="B9166F3FAC2AFFA24C74FD0514B5FCD8" bold="true" box="[828,1002,766,789]" pageId="32" pageNumber="195">Measurements.</emphasis>
BL 1.902.40, FBL 0.901.20, HL 0.29 0.30, HW 0.360.45, PL 0.310.45, PW 0.370.49, EL 0.360.55, EW 0.430.56, HTL 0.280.43, AL 0.600.75, AS-I 0.110.14, AS-II 0.060.11, AS-III 0.040.05.
</paragraph>
</subSubSection>
<subSubSection id="C378E0A6AC2AFFA24C74FC8514C0FBF8" pageId="32" pageNumber="195" type="diagnosis">
<paragraph id="8BDDB32DAC2AFFA24C74FC8514C0FBF8" blockId="32.[828,1437,190,1365]" pageId="32" pageNumber="195">
<emphasis id="B9166F3FAC2AFFA24C74FC85136DFC58" bold="true" box="[828,1074,894,917]" pageId="32" pageNumber="195">Differential diagnosis.</emphasis>
In general appearance, the species closely resembles
<taxonomicName id="4C62C8AEAC2AFFA24B48FC651306FC78" authority="Liu &amp; Ono &amp; Maruyama, 2021" authorityName="Liu &amp; Ono &amp; Maruyama" authorityYear="2021" box="[1024,1113,926,949]" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="195" phylum="Arthropoda" rank="species" species="nigra" status="sp. nov.">
<emphasis id="B9166F3FAC2AFFA24B48FC651306FC78" box="[1024,1113,926,949]" italics="true" pageId="32" pageNumber="195">B. nigra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A225D244AC2AFFA24B28FC6513ECFC78" box="[1120,1203,926,949]" pageId="32" pageNumber="195" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C62C8AEAC2AFFA24BAFFC651206FC78" baseAuthorityName="Sawada" baseAuthorityYear="1955" box="[1255,1369,925,949]" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="195" phylum="Arthropoda" rank="species" species="nakanei">
<emphasis id="B9166F3FAC2AFFA24BAFFC651206FC78" box="[1255,1369,925,949]" italics="true" pageId="32" pageNumber="195">B. nakanei</emphasis>
</taxonomicName>
but is easily distinguished from
<taxonomicName id="4C62C8AEAC2AFFA24B00FC4513C2FC18" authority="Liu &amp; Ono &amp; Maruyama, 2021" authorityName="Liu &amp; Ono &amp; Maruyama" authorityYear="2021" box="[1096,1181,958,981]" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="195" phylum="Arthropoda" rank="species" species="nigra" status="sp. nov.">
<emphasis id="B9166F3FAC2AFFA24B00FC4513C2FC18" box="[1096,1181,958,981]" italics="true" pageId="32" pageNumber="195">B. nigra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A225D244AC2AFFA24BEAFC4513ADFC18" box="[1186,1266,958,981]" pageId="32" pageNumber="195" rank="species">sp. nov.</taxonomicNameLabel>
in the shape and size of the spermatheca and is different from
<taxonomicName id="4C62C8AEAC2AFFA24A61FC2512C3FC38" baseAuthorityName="Sawada" baseAuthorityYear="1955" box="[1321,1436,989,1013]" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="195" phylum="Arthropoda" rank="species" species="nakanei">
<emphasis id="B9166F3FAC2AFFA24A61FC2512C3FC38" box="[1321,1436,989,1013]" italics="true" pageId="32" pageNumber="195">B. nakanei</emphasis>
</taxonomicName>
in having the posterior margin of the tergite X rounded medially.
</paragraph>
</subSubSection>
<subSubSection id="C378E0A6AC2AFFA24C74FBC514EBFB18" pageId="32" pageNumber="195" type="distribution">
<paragraph id="8BDDB32DAC2AFFA24C74FBC514EBFB18" blockId="32.[828,1437,190,1365]" pageId="32" pageNumber="195">
<emphasis id="B9166F3FAC2AFFA24C74FBC51490FB98" bold="true" box="[828,975,1086,1109]" pageId="32" pageNumber="195">Distribution.</emphasis>
<collectingCountry id="F375F3BDAC2AFFA24C91FBC51349FB98" box="[985,1046,1086,1109]" name="Japan" pageId="32" pageNumber="195">Japan</collectingCountry>
(
<emphasis id="B9166F3FAC2AFFA24B62FBC513D3FB98" bold="true" box="[1066,1164,1086,1109]" pageId="32" pageNumber="195">Honshû:</emphasis>
<collectingRegion id="49A67DCFAC2AFFA24BDFFBC51385FB98" box="[1175,1242,1086,1109]" country="Japan" name="Chiba" pageId="32" pageNumber="195">Chiba</collectingRegion>
,
<collectingRegion id="49A67DCFAC2AFFA24BA0FBC51203FB98" box="[1256,1372,1086,1109]" country="Japan" name="Kanagawa" pageId="32" pageNumber="195">Kanagawa</collectingRegion>
,
<collectingRegion id="49A67DCFAC2AFFA24A22FBC512C7FB98" box="[1386,1432,1086,1109]" country="Japan" name="Mie" pageId="32" pageNumber="195">Mie</collectingRegion>
,
<collectingRegion id="49A67DCFAC2AFFA24C74FBA514DEFBB8" box="[828,897,1118,1141]" country="Japan" name="Osaka" pageId="32" pageNumber="195">Ôsaka</collectingRegion>
,
<collectingRegion id="49A67DCFAC2AFFA24CC4FBA5135AFBB8" box="[908,1029,1118,1141]" country="Japan" name="Wakayama" pageId="32" pageNumber="195">Wakayama</collectingRegion>
;
<emphasis id="B9166F3FAC2AFFA24B59FBA51326FBB8" bold="true" box="[1041,1145,1118,1141]" pageId="32" pageNumber="195">Shikoku:</emphasis>
<collectingRegion id="49A67DCFAC2AFFA24BC8FBA51385FBB8" box="[1152,1242,1118,1141]" country="Japan" name="Kagawa" pageId="32" pageNumber="195">Kagawa</collectingRegion>
,
<collectingRegion id="49A67DCFAC2AFFA24BAEFBA51270FBB8" box="[1254,1327,1118,1141]" country="Japan" name="Ehime" pageId="32" pageNumber="195">Ehime</collectingRegion>
;
<emphasis id="B9166F3FAC2AFFA24A72FBA512C3FBB8" bold="true" box="[1338,1436,1118,1141]" pageId="32" pageNumber="195">Kyûshû:</emphasis>
<collectingRegion id="49A67DCFAC2AFFA24C74FB8514C4FB58" box="[828,923,1150,1173]" country="Japan" name="Fukuoka" pageId="32" pageNumber="195">Fukuoka</collectingRegion>
,
<collectingRegion id="49A67DCFAC2AFFA24CEBFB851357FB58" box="[931,1032,1150,1173]" country="Japan" name="Nagasaki" pageId="32" pageNumber="195">Nagasaki</collectingRegion>
,
<collectingRegion id="49A67DCFAC2AFFA24B58FB851329FB58" box="[1040,1142,1150,1173]" country="Japan" name="Miyazaki" pageId="32" pageNumber="195">Miyazaki</collectingRegion>
,
<collectingRegion id="49A67DCFAC2AFFA24B37FB8513A5FB58" box="[1151,1274,1150,1173]" country="Japan" name="Kagoshima" pageId="32" pageNumber="195">Kagoshima</collectingRegion>
;
<emphasis id="B9166F3FAC2AFFA24A4BFB8512C2FB58" bold="true" box="[1283,1437,1150,1173]" pageId="32" pageNumber="195">Nansei-shotô:</emphasis>
Tanega-shima,
<collectingRegion id="49A67DCFAC2AFFA24C94FB65131FFB78" box="[988,1088,1182,1205]" country="Japan" name="Okinawa" pageId="32" pageNumber="195">Okinawa</collectingRegion>
: Ou-jima, Okinawa-jima);
<collectingCountry id="F375F3BDAC2AFFA24A13FB6512C3FB78" box="[1371,1436,1182,1205]" name="South Korea" pageId="32" pageNumber="195">Korea</collectingCountry>
(
<collectingRegion id="49A67DCFAC2AFFA24C0BFB4514F7FB18" box="[835,936,1214,1237]" country="South Korea" name="Jeonnam" pageId="32" pageNumber="195">Jeonnam</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C378E0A6AC2AFFA24C74FB2514F5FA98" pageId="32" pageNumber="195" type="discussion">
<paragraph id="8BDDB32DAC2AFFA24C74FB2514F5FA98" blockId="32.[828,1437,190,1365]" pageId="32" pageNumber="195">
<emphasis id="B9166F3FAC2AFFA24C74FB2514F4FB38" bold="true" box="[828,939,1246,1269]" pageId="32" pageNumber="195">Remarks.</emphasis>
This species is mainly found west of Kantô District and
<collectingRegion id="49A67DCFAC2AFFA24C8BFB05137CFAD8" box="[963,1059,1278,1301]" country="Japan" name="Okinawa" pageId="32" pageNumber="195">Okinawa</collectingRegion>
Island. A distribution map of examined specimens of this species in
<collectingCountry id="F375F3BDAC2AFFA24BF0FAE513AAFAF8" box="[1208,1269,1310,1333]" name="Japan" pageId="32" pageNumber="195">Japan</collectingCountry>
is illustrated in
<figureCitation id="1359AFA8AC2AFFA24C74FAC514F9FA98" box="[828,934,1342,1365]" captionStart="Fig" captionStartId="35.[150,181,1388,1407]" captionTargetBox="[150,1434,191,1370]" captionTargetId="figure-236@35.[150,1435,191,1371]" captionTargetPageId="35" captionText="Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle B. algarum Sawada, 1971; black inverted triangle B. minuta (Sawada, 1955); white star B. tsutsuii (Sawada, 1955); black star B. koreana Ahn &amp; Jeon, 2004; black square B. yoshigoui sp. nov.; white triangle B. japonica Liu, Ono &amp; Maruyama, 2020; black diamond B. sakishimana Sawada, 1991; white diamond B. constricta sp. nov.; white square B. hokkaidensis sp. nov.; black triangle B. fulvipennis sp. nov.; double circle B. itsuroi sp. nov.; arrow B. aikoae sp. nov.; black hart B. okinawana sp. nov.; circled dot B. nigra sp. nov.; white circle B. nakanei (Sawada, 1955); white hart B. moriguchii sp. nov.; black circle B. gangjinensis Ahn &amp; Jeon, 2004." figureDoi="http://doi.org/10.5281/zenodo.5037376" httpUri="https://zenodo.org/record/5037376/files/figure.png" pageId="32" pageNumber="195">Figure 22</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>