treatments-xml/data/03/CB/02/03CB023BAC19FF974E58F88015D5FD98.xml

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<mods:title id="8A04C565F896C5C199B333AB3F11A1AB">Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:namePart id="639E7B6D42C684649AE253F5DDD15C8F">Maruyama, Munetoshi</mods:namePart>
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<taxonomicName id="4C62C8AEAC19FF914E58F880157FF85E" ID-CoL="8XW4H" authority="Liu &amp; Ono &amp; Maruyama, 2021" authorityName="Liu &amp; Ono &amp; Maruyama" authorityYear="2021" box="[272,544,1915,1939]" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="182" phylum="Arthropoda" rank="species" species="fulvipennis" status="sp. nov.">
<emphasis id="B9166F3FAC19FF914E58F880157FF85E" bold="true" box="[272,544,1915,1939]" italics="true" pageId="19" pageNumber="182">Bryothinusa fulvipennis</emphasis>
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<paragraph id="8BDDB32DAC19FF914E10F8611568F860" blockId="19.[151,759,1946,2066]" box="[344,567,1946,1965]" pageId="19" pageNumber="182">
(
<figureCitation id="1359AFA8AC19FF914E16F86116FFF860" box="[350,416,1946,1965]" captionStart="Fig" captionStartId="2.[151,182,1972,1991]" captionTargetBox="[189,1405,205,1944]" captionTargetId="figure-13@2.[175,1411,189,1947]" captionTargetPageId="2" captionText="Fig. 1. Habitus of Bryothinusa spp. A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn &amp; Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono &amp; Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn &amp; Jeon, 2004. Scale bars: 1.00 mm." figureDoi="http://doi.org/10.5281/zenodo.5037332" httpUri="https://zenodo.org/record/5037332/files/figure.png" pageId="19" pageNumber="182">Figs 1J</figureCitation>
; 
<figureCitation id="1359AFA8AC19FF914EE1F86116E0F860" box="[425,447,1946,1965]" captionStart="Fig" captionStartId="4.[151,182,1978,1997]" captionTargetBox="[196,1391,190,1955]" captionTargetId="figure-13@4.[193,1394,189,1958]" captionTargetPageId="4" captionText="Fig. 2. Compound eyes of Bryothinusa spp.A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn &amp; Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono &amp; Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn &amp; Jeon, 2004. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5037334" httpUri="https://zenodo.org/record/5037334/files/figure.png" pageId="19" pageNumber="182">2J</figureCitation>
; 
<figureCitation id="1359AFA8AC19FF914E81F86116BEF860" box="[457,481,1946,1965]" captionStart="Fig" captionStartId="20.[151,182,2029,2048]" captionTargetBox="[154,1414,208,1976]" captionTargetId="figure-13@20.[151,1436,206,1976]" captionTargetPageId="20" captionText="Fig. 12. Bryothinusa fulvipennis sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037356" httpUri="https://zenodo.org/record/5037356/files/figure.png" pageId="19" pageNumber="182">12</figureCitation>
; 
<figureCitation id="1359AFA8AC19FF914EA2F861154FF860" box="[490,528,1946,1965]" captionStart="Fig" captionStartId="34.[151,182,1955,1974]" captionTargetBox="[167,1408,193,1935]" captionTargetId="figure-13@34.[161,1426,189,1938]" captionTargetPageId="34" captionText="Fig. 21. Collection localities of Bryothinusa species in Japan: A – B. koreana Ahn &amp; Jeon, 2004 and B. japonica Liu, Ono &amp; Maruyama, 2020 in Shima-nogita, Fukuoka; B – B. aikoae sp. nov., B. fulvipennis sp. nov., B. japonica and B. nigra sp. nov. in Kuroto, Chiba; C – B. hokkaidensis sp. nov. in Shunkunitai, Hokkaidô; D – B. nigra sp. nov. and B. tsutsuii (Sawada, 1955) in Edateku-jima; E – B. nakanei (Sawada, 1955) in Nagura-anparu, Ishigaki-jima; F – B. gangjinensis Ahn &amp; Jeon, 2004 in Iki-no-matsubara, Fukuoka; G – B. algarum Sawada, 1971, B. japonica and B. minuta (Sawada, 1955) in Sunosaki, Fukuoka." figureDoi="http://doi.org/10.5281/zenodo.5037374" httpUri="https://zenodo.org/record/5037374/files/figure.png" pageId="19" pageNumber="182">21B</figureCitation>
; 
<figureCitation id="1359AFA8AC19FF914D52F861156EF860" box="[538,561,1946,1965]" captionStart="Fig" captionStartId="35.[150,181,1388,1407]" captionTargetBox="[150,1434,191,1370]" captionTargetId="figure-236@35.[150,1435,191,1371]" captionTargetPageId="35" captionText="Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle – B. algarum Sawada, 1971; black inverted triangle – B. minuta (Sawada, 1955); white star – B. tsutsuii (Sawada, 1955); black star – B. koreana Ahn &amp; Jeon, 2004; black square – B. yoshigoui sp. nov.; white triangle – B. japonica Liu, Ono &amp; Maruyama, 2020; black diamond – B. sakishimana Sawada, 1991; white diamond – B. constricta sp. nov.; white square – B. hokkaidensis sp. nov.; black triangle – B. fulvipennis sp. nov.; double circle – B. itsuroi sp. nov.; arrow – B. aikoae sp. nov.; black hart – B. okinawana sp. nov.; circled dot – B. nigra sp. nov.; white circle – B. nakanei (Sawada, 1955); white hart – B. moriguchii sp. nov.; black circle – B. gangjinensis Ahn &amp; Jeon, 2004." figureDoi="http://doi.org/10.5281/zenodo.5037376" httpUri="https://zenodo.org/record/5037376/files/figure.png" pageId="19" pageNumber="182">22</figureCitation>
)
</paragraph>
<paragraph id="8BDDB32DAC19FF914FB7F84115CFF800" blockId="19.[151,759,1946,2066]" box="[255,656,1978,1997]" pageId="19" pageNumber="182">(Japanese name: Usucha-nagisa-hanekakushi)</paragraph>
</subSubSection>
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<emphasis id="B9166F3FAC19FF914FDFF81F167FF83A" bold="true" box="[151,288,2020,2039]" pageId="19" pageNumber="182">
<typeStatus id="54D90D8FAC19FF914FDFF81F179BF83A" box="[151,196,2020,2039]" pageId="19" pageNumber="182">Type</typeStatus>
material.
</emphasis>
Hඈඅඈඍඒඉൾ: J, 
<emphasis id="B9166F3FAC19FF914EE4F81F1509F83A" bold="true" box="[428,598,2020,2039]" pageId="19" pageNumber="182">
<collectingCountry id="F375F3BDAC19FF914EE4F81F16AAF83A" box="[428,501,2020,2039]" name="Japan" pageId="19" pageNumber="182">JAPAN</collectingCountry>
: HඈඇඌΗǕ:
</emphasis>
Mouth of Obitsugawa, Kuzuma, Kisarazu-shi, Chiba-ken, 
<date id="FFDC95EDAC19FF914D41F8041500F7DF" box="[521,607,2047,2066]" pageId="19" pageNumber="182" value="2007-03-02">2.iii.2007</date>
, H. Ono (
<collectionCode id="ED732BE8AC19FF914DFFF80415B3F7DF" box="[695,748,2047,2066]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="19" pageNumber="182" type="Herbarium">KUM</collectionCode>
).
</paragraph>
<paragraph id="8BDDB32DAC19FF914C74FF4512D0FD2B" blockId="19.[828,1437,183,742]" pageId="19" pageNumber="182">
Pൺ*©ൺඍඒඉൾඌ: 
<emphasis id="B9166F3FAC19FF914CEBFF4514A6FF1C" bold="true" box="[931,1017,190,209]" pageId="19" pageNumber="182">HඈඇඌΗǕ:</emphasis>
<specimenCount id="9D6478A4AC19FF914CB7FF45137FFF1D" box="[1023,1056,183,209]" pageId="19" pageNumber="182" type="female">1 ♀</specimenCount>
, same data as 
<typeStatus id="54D90D8FAC19FF914BEBFF4513B0FF1C" box="[1187,1263,190,209]" pageId="19" pageNumber="182" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED732BE8AC19FF914BB4FF45126CFF1C" box="[1276,1331,190,209]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="19" pageNumber="182" type="Herbarium">KUM</collectionCode>
); 1 unsexed spec., same locality as 
<typeStatus id="54D90D8FAC19FF914B46FF221302FF21" box="[1038,1117,217,236]" pageId="19" pageNumber="182" type="holotype">holotype</typeStatus>
, 
<date id="FFDC95EDAC19FF914B2FFF22139DFF21" box="[1127,1218,217,236]" pageId="19" pageNumber="182" value="2007-02-19">19.ii.2007</date>
, H. Ono (
<collectionCode id="ED732BE8AC19FF914A56FF22120AFF21" box="[1310,1365,217,236]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="19" pageNumber="182" type="Herbarium">KUM</collectionCode>
); 1 J 2 unsexed spec., Mouth of Obitsu-gawa, Kuroto, Kisarazu-shi, Chiba-ken, 
<date id="FFDC95EDAC19FF914C74FEF514C9FEEC" box="[828,918,270,289]" pageId="19" pageNumber="182" value="2007-02-19">19.ii.2007</date>
, H. Ono (
<collectionCode id="ED732BE8AC19FF914CAFFEF51341FEEC" box="[999,1054,270,289]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="19" pageNumber="182" type="Herbarium">KUM</collectionCode>
); 1 unsexed spec., same locality, 
<date id="FFDC95EDAC19FF914A70FEF512C6FEEC" box="[1336,1433,270,289]" pageId="19" pageNumber="182" value="2009-03-12">12.iii.2009</date>
, H. Ono (
<collectionCode id="ED732BE8AC19FF914CC3FED2149DFEF1" box="[907,962,297,316]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="19" pageNumber="182" type="Herbarium">KUM</collectionCode>
); 
<specimenCount id="9D6478A4AC19FF914C9BFED214AAFEF6" box="[979,1013,290,316]" pageId="19" pageNumber="182" type="female">1 ♀</specimenCount>
, 3 unsexed spec., same locality, 
<date id="FFDC95EDAC19FF914A54FED21223FEF1" box="[1308,1404,297,316]" pageId="19" pageNumber="182" value="2009-03-16">16.iii.2009</date>
, H. Ono (
<collectionCode id="ED732BE8AC19FF914C25FEB814FBFE9B" box="[877,932,323,342]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="19" pageNumber="182" type="Herbarium">KUM</collectionCode>
); 1 J, same locality, 
<date id="FFDC95EDAC19FF914B16FEB813F7FE9B" box="[1118,1192,323,342]" pageId="19" pageNumber="182" value="2010-01-04">4.i.2010</date>
, H. Ono (
<collectionCode id="ED732BE8AC19FF914BB5FEB8126BFE9B" box="[1277,1332,323,342]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="19" pageNumber="182" type="Herbarium">KUM</collectionCode>
); 1 unsexed spec., same locality, 
<date id="FFDC95EDAC19FF914B48FEA51308FEBC" box="[1024,1111,350,369]" pageId="19" pageNumber="182" value="2011-01-12">12.i.2011</date>
, H. Ono (
<collectionCode id="ED732BE8AC19FF914BFFFEA513AFFEBC" box="[1207,1264,350,369]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="19" pageNumber="182" type="Herbarium">KUM</collectionCode>
); 1 unsexed spec., same locality, 
<date id="FFDC95EDAC19FF914CF5FE82134FFE41" box="[957,1040,377,396]" pageId="19" pageNumber="182" value="2011-01-14">14.i.2011</date>
, H. Ono (
<collectionCode id="ED732BE8AC19FF914B22FE8213FEFE41" box="[1130,1185,377,396]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="19" pageNumber="182" type="Herbarium">KUM</collectionCode>
); 1 unsexed spec., Kushidagawa, Matsuzaka-shi, Mie-ken, 
<date id="FFDC95EDAC19FF914B1EFE6813EBFE6B" box="[1110,1204,403,422]" pageId="19" pageNumber="182" value="2002-04-28">28.iv.2002</date>
, Y. Kawakami (
<collectionCode id="ED732BE8AC19FF914A77FE681229FE6B" box="[1343,1398,403,422]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="19" pageNumber="182" type="Herbarium">KUM</collectionCode>
); 10 unsexed spec., Himeshima, Yodo-gawa, Nishiyodo-gawa-ku, Ôsaka-fu, 
<date id="FFDC95EDAC19FF914C74FE3214C5FE11" box="[828,922,457,476]" pageId="19" pageNumber="182" value="2002-04-12">12.iv.2002</date>
, Y. Kawakami (
<collectionCode id="ED732BE8AC19FF914B61FE321300FE11" box="[1065,1119,457,476]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="19" pageNumber="182" type="Herbarium">KUM</collectionCode>
, 
<collectionCode id="ED732BE8AC19FF914B22FE3213F5FE11" box="[1130,1194,457,476]" pageId="19" pageNumber="182">MNPC</collectionCode>
, 
<collectionCode id="ED732BE8AC19FF914BFCFE3213A6FE11" box="[1204,1273,457,476]" pageId="19" pageNumber="182">BMNH</collectionCode>
); 3 unsexed spec., Hanakawa Yodo-gawa, Nishiyodo-gawa-ku, Ôsaka-fu, 
<date id="FFDC95EDAC19FF914A70FE1812C7FE3B" box="[1336,1432,483,502]" pageId="19" pageNumber="182" value="2002-04-25">25.iv.2002</date>
, Y. Kawakami (
<collectionCode id="ED732BE8AC19FF914C88FE0514A8FDDC" box="[960,1015,510,529]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="19" pageNumber="182" type="Herbarium">KUM</collectionCode>
); 4 unsexed spec., same locality, 
<date id="FFDC95EDAC19FF914A68FE0512DFFDDC" box="[1312,1408,510,529]" pageId="19" pageNumber="182" value="2002-03-28">28.iii.2002</date>
, Y. Kawakami (
<collectionCode id="ED732BE8AC19FF914CEFFDE21481FDE1" box="[935,990,537,556]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="19" pageNumber="182" type="Herbarium">KUM</collectionCode>
); 1 unsexed spec., Kinburo, Kitagi-shima, Kasaoka-shi, Okayama-ken, 
<date id="FFDC95EDAC19FF914B49FDC81305FD8B" box="[1025,1114,563,582]" pageId="19" pageNumber="182" value="2008-10-11">11.x.2008</date>
, O. Yamaji (
<collectionCode id="ED732BE8AC19FF914B8EFDC813A2FD8B" box="[1222,1277,563,582]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="19" pageNumber="182" type="Herbarium">KUM</collectionCode>
); 3 unsexed spec., Kurahashi-jima (Umikoshi), Kurahashi-chô, Kure-shi, Hiroshima-ken, 
<date id="FFDC95EDAC19FF914C74FD9214D4FDB1" box="[828,907,617,636]" pageId="19" pageNumber="182" value="2009-02-01">1.ii.2009</date>
, K. Nakano (
<collectionCode id="ED732BE8AC19FF914CB7FD921369FDB1" box="[1023,1078,617,636]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="19" pageNumber="182" type="Herbarium">KUM</collectionCode>
). 
<emphasis id="B9166F3FAC19FF914B0DFD9213FFFDB1" bold="true" box="[1093,1184,617,636]" pageId="19" pageNumber="182">SΗංκඈκඎ:</emphasis>
11 unsexed spec., Kaigan-ji, Nishishirakata, Tadotsu-chô, Kagawa-ken, 
<date id="FFDC95EDAC19FF914B88FD78127EFD5B" box="[1216,1313,643,662]" pageId="19" pageNumber="182" value="2008-03-31">31.iii.2008</date>
, H. Fujimoto (
<collectionCode id="ED732BE8AC19FF914C0AFD651426FD7C" box="[834,889,670,689]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="19" pageNumber="182" type="Herbarium">KUM</collectionCode>
); 1 unsexed spec., Ôyatomi-machi, Sakaide-shi, Kagawa-ken, 
<date id="FFDC95EDAC19FF914C74FD4214C3FD01" box="[828,924,697,716]" pageId="19" pageNumber="182" value="2008-03-29">29.iii.2008</date>
, H. Fujimoto (
<collectionCode id="ED732BE8AC19FF914B69FD421307FD01" box="[1057,1112,697,716]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="19" pageNumber="182" type="Herbarium">KUM</collectionCode>
); 1 unsexed spec., Kôtô-gawa, Gôtô -chô, Takamatsu-shi, Kagawa-ken, 
<date id="FFDC95EDAC19FF914B38FD281391FD2B" box="[1136,1230,723,742]" pageId="19" pageNumber="182" value="2008-04-12">12.iv.2008</date>
, H. Fujimoto (
<collectionCode id="ED732BE8AC19FF914A06FD2812DAFD2B" box="[1358,1413,723,742]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="19" pageNumber="182" type="Herbarium">KUM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C378E0A6AC19FF914C74FD04123AF81B" pageId="19" pageNumber="182" type="description">
<paragraph id="8BDDB32DAC19FF914C74FD041270FCBB" blockId="19.[828,1437,767,2070]" pageId="19" pageNumber="182">
<emphasis id="B9166F3FAC19FF914C74FD041496FCDB" bold="true" box="[828,969,767,790]" pageId="19" pageNumber="182">Description.</emphasis>
Body (
<figureCitation id="1359AFA8AC19FF914B51FD04133CFCDB" box="[1049,1123,767,790]" captionStart="Fig" captionStartId="2.[151,182,1972,1991]" captionTargetBox="[189,1405,205,1944]" captionTargetId="figure-13@2.[175,1411,189,1947]" captionTargetPageId="2" captionText="Fig. 1. Habitus of Bryothinusa spp. A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn &amp; Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono &amp; Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn &amp; Jeon, 2004. Scale bars: 1.00 mm." figureDoi="http://doi.org/10.5281/zenodo.5037332" httpUri="https://zenodo.org/record/5037332/files/figure.png" pageId="19" pageNumber="182">Fig. 1J</figureCitation>
) small, about 1.6–2.0 mm in length, covered with minute setae. Abdomen almost black and darker than head and pronotum; elytra lighter than pronotum; antenna and legs yellowish brown.
</paragraph>
<paragraph id="8BDDB32DAC19FF914C12FC841320FBBB" blockId="19.[828,1437,767,2070]" pageId="19" pageNumber="182">
Head evenly rounded, more or less produced anteriorly, wider than long, shorter than pronotum and elytra; eyes (
<figureCitation id="1359AFA8AC19FF914C0CFC4414D1FC1B" box="[836,910,959,982]" captionStart="Fig" captionStartId="4.[151,182,1978,1997]" captionTargetBox="[196,1391,190,1955]" captionTargetId="figure-13@4.[193,1394,189,1958]" captionTargetPageId="4" captionText="Fig. 2. Compound eyes of Bryothinusa spp.A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn &amp; Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono &amp; Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn &amp; Jeon, 2004. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5037334" httpUri="https://zenodo.org/record/5037334/files/figure.png" pageId="19" pageNumber="182">Fig. 2J</figureCitation>
) large, setae present between about 22–24 facets, slightly longer than tempora length; antenna yellowish brown, almost equals to total length of head and pronotum; segment I 1.7 times longer than II, almost 4 times longer than III, and longer than XI; antennal segments IV–X almost as long as wide.
</paragraph>
<paragraph id="8BDDB32DAC19FF914C12FB84135FFA3B" blockId="19.[828,1437,767,2070]" pageId="19" pageNumber="182">
Mouthparts (
<figureCitation id="1359AFA8AC19FF914CA0FB841330FB5B" box="[1000,1135,1151,1174]" captionStart="Fig" captionStartId="20.[151,182,2029,2048]" captionTargetBox="[154,1414,208,1976]" captionTargetId="figure-13@20.[151,1436,206,1976]" captionTargetPageId="20" captionText="Fig. 12. Bryothinusa fulvipennis sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037356" httpUri="https://zenodo.org/record/5037356/files/figure.png" pageId="19" pageNumber="182">Figs 12A–E</figureCitation>
) with distribution of labral setae as in 
<figureCitation id="1359AFA8AC19FF914CF1FB641364FB7B" box="[953,1083,1183,1206]" captionStart="Fig" captionStartId="20.[151,182,2029,2048]" captionTargetBox="[154,1414,208,1976]" captionTargetId="figure-13@20.[151,1436,206,1976]" captionTargetPageId="20" captionText="Fig. 12. Bryothinusa fulvipennis sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037356" httpUri="https://zenodo.org/record/5037356/files/figure.png" pageId="19" pageNumber="182">Figure 12A</figureCitation>
, with large and thick spiniform seta antero-laterally; mandibles (
<figureCitation id="1359AFA8AC19FF914BF7FB441277FB1B" box="[1215,1320,1215,1238]" captionStart="Fig" captionStartId="20.[151,182,2029,2048]" captionTargetBox="[154,1414,208,1976]" captionTargetId="figure-13@20.[151,1436,206,1976]" captionTargetPageId="20" captionText="Fig. 12. Bryothinusa fulvipennis sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037356" httpUri="https://zenodo.org/record/5037356/files/figure.png" pageId="19" pageNumber="182">Fig. 12B</figureCitation>
) elongate triangle, asymmetrical, right with median tooth; median tooth shorter than 1/4 of mandibles; left without tooth, blunt at apex; maxilla with apical margin of lacinia (
<figureCitation id="1359AFA8AC19FF914A38FAE41435FA9B" captionStart="Fig" captionStartId="20.[151,182,2029,2048]" captionTargetBox="[154,1414,208,1976]" captionTargetId="figure-13@20.[151,1436,206,1976]" captionTargetPageId="20" captionText="Fig. 12. Bryothinusa fulvipennis sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037356" httpUri="https://zenodo.org/record/5037356/files/figure.png" pageId="19" pageNumber="182">Fig. 12C</figureCitation>
) second spine very small, not obvious; 3 setae on inner margin of lacinia; labial palpus (
<figureCitation id="1359AFA8AC19FF914BBBFAA41205FABB" box="[1267,1370,1375,1398]" captionStart="Fig" captionStartId="20.[151,182,2029,2048]" captionTargetBox="[154,1414,208,1976]" captionTargetId="figure-13@20.[151,1436,206,1976]" captionTargetPageId="20" captionText="Fig. 12. Bryothinusa fulvipennis sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037356" httpUri="https://zenodo.org/record/5037356/files/figure.png" pageId="19" pageNumber="182">Fig. 12D</figureCitation>
) with most setae located basally; mentum (
<figureCitation id="1359AFA8AC19FF914BB8FA841207FA5B" box="[1264,1368,1407,1430]" captionStart="Fig" captionStartId="20.[151,182,2029,2048]" captionTargetBox="[154,1414,208,1976]" captionTargetId="figure-13@20.[151,1436,206,1976]" captionTargetPageId="20" captionText="Fig. 12. Bryothinusa fulvipennis sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037356" httpUri="https://zenodo.org/record/5037356/files/figure.png" pageId="19" pageNumber="182">Fig. 12E</figureCitation>
) with antero-lateral angles prolonged into spinose processes; anterior margin deeply truncate; small “v seta” far away from apical setae.
</paragraph>
<paragraph id="8BDDB32DAC19FF914C12FA0412DCF9DB" blockId="19.[828,1437,767,2070]" box="[858,1411,1535,1558]" pageId="19" pageNumber="182">Pronotum almost as long as elytra, wider than head.</paragraph>
<paragraph id="8BDDB32DAC19FF914C12F9E414FBF9BB" blockId="19.[828,1437,767,2070]" pageId="19" pageNumber="182">Elytra subquadrate but wider than long, almost as wide as pronotum; hind wings present but very small, apparently flightless.</paragraph>
<paragraph id="8BDDB32DAC19FF914C12F984148DF93B" blockId="19.[828,1437,767,2070]" pageId="19" pageNumber="182">
Male abdominal sternite VIII (
<figureCitation id="1359AFA8AC19FF914BD6F98413A5F95B" box="[1182,1274,1663,1686]" captionStart="Fig" captionStartId="20.[151,182,2029,2048]" captionTargetBox="[154,1414,208,1976]" captionTargetId="figure-13@20.[151,1436,206,1976]" captionTargetPageId="20" captionText="Fig. 12. Bryothinusa fulvipennis sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037356" httpUri="https://zenodo.org/record/5037356/files/figure.png" pageId="19" pageNumber="182">Fig. 12F</figureCitation>
) with posterior margin slightly serrate. Median lobe of aedeagus (
<figureCitation id="1359AFA8AC19FF914A39F9641434F91B" captionStart="Fig" captionStartId="20.[151,182,2029,2048]" captionTargetBox="[154,1414,208,1976]" captionTargetId="figure-13@20.[151,1436,206,1976]" captionTargetPageId="20" captionText="Fig. 12. Bryothinusa fulvipennis sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037356" httpUri="https://zenodo.org/record/5037356/files/figure.png" pageId="19" pageNumber="182">Fig. 12H</figureCitation>
) and dorsal view as in 
<figureCitation id="1359AFA8AC19FF914B3EF94413B2F91B" box="[1142,1261,1727,1750]" captionStart="Fig" captionStartId="20.[151,182,2029,2048]" captionTargetBox="[154,1414,208,1976]" captionTargetId="figure-13@20.[151,1436,206,1976]" captionTargetPageId="20" captionText="Fig. 12. Bryothinusa fulvipennis sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037356" httpUri="https://zenodo.org/record/5037356/files/figure.png" pageId="19" pageNumber="182">Figure 12I</figureCitation>
; paramere (
<figureCitation id="1359AFA8AC19FF914A39F944143EF93B" captionStart="Fig" captionStartId="20.[151,182,2029,2048]" captionTargetBox="[154,1414,208,1976]" captionTargetId="figure-13@20.[151,1436,206,1976]" captionTargetPageId="20" captionText="Fig. 12. Bryothinusa fulvipennis sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037356" httpUri="https://zenodo.org/record/5037356/files/figure.png" pageId="19" pageNumber="182">Fig. 12J</figureCitation>
) elongate.
</paragraph>
<paragraph id="8BDDB32DAC19FF914C12F904139CF89B" blockId="19.[828,1437,767,2070]" pageId="19" pageNumber="182">
Female abdominal sternite VIII (
<figureCitation id="1359AFA8AC19FF914B98F9041268F8DB" box="[1232,1335,1791,1814]" captionStart="Fig" captionStartId="20.[151,182,2029,2048]" captionTargetBox="[154,1414,208,1976]" captionTargetId="figure-13@20.[151,1436,206,1976]" captionTargetPageId="20" captionText="Fig. 12. Bryothinusa fulvipennis sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037356" httpUri="https://zenodo.org/record/5037356/files/figure.png" pageId="19" pageNumber="182">Fig. 12G</figureCitation>
) slightly serrate alike male on posterior margin. Spermatheca (
<figureCitation id="1359AFA8AC19FF914A39F8E41434F89B" captionStart="Fig" captionStartId="20.[151,182,2029,2048]" captionTargetBox="[154,1414,208,1976]" captionTargetId="figure-13@20.[151,1436,206,1976]" captionTargetPageId="20" captionText="Fig. 12. Bryothinusa fulvipennis sp. nov. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037356" httpUri="https://zenodo.org/record/5037356/files/figure.png" pageId="19" pageNumber="182">Fig. 12K</figureCitation>
) about 2 times coiled near base.
</paragraph>
<paragraph id="8BDDB32DAC19FF914C74F8A4123AF81B" blockId="19.[828,1437,767,2070]" pageId="19" pageNumber="182">
<emphasis id="B9166F3FAC19FF914C74F8A414B5F8BB" bold="true" box="[828,1002,1887,1910]" pageId="19" pageNumber="182">Measurements.</emphasis>
BL 1.60–2.00, FBL 0.90–1.00, HL 0.28– 0.30, HW 0.36–0.43, PL 0.36–0.40, PW 0.40–0.45, EL 0.35–0.41, EW 0.36–0.45, HTL 0.23–0.33, AL 0.61–0.76, AS-I 0.13–0.14, AS-II 0.07–0.08, AS-III 0.03–0.04.
</paragraph>
</subSubSection>
<subSubSection id="C378E0A6AC19FF974C74F824168CFEF8" lastPageId="21" lastPageNumber="184" pageId="19" pageNumber="182" type="diagnosis">
<paragraph id="8BDDB32DAC19FF974C74F824168CFEF8" blockId="19.[828,1437,767,2070]" lastBlockId="21.[151,761,189,597]" lastPageId="21" lastPageNumber="184" pageId="19" pageNumber="182">
<emphasis id="B9166F3FAC19FF914C74F824136CF83B" bold="true" box="[828,1075,2015,2038]" pageId="19" pageNumber="182">Differential diagnosis.</emphasis>
In general appearance the species is very similar to 
<taxonomicName id="4C62C8AEAC19FF914CA7F80413CBF7DB" authority="Liu &amp; Ono &amp; Maruyama, 2021" authorityName="Liu &amp; Ono &amp; Maruyama" authorityYear="2021" box="[1007,1172,2046,2070]" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="182" phylum="Arthropoda" rank="species" species="hokkaidensis" status="sp. nov.">
<emphasis id="B9166F3FAC19FF914CA7F80413CBF7DB" box="[1007,1172,2046,2070]" italics="true" pageId="19" pageNumber="182">B. hokkaidensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A225D244AC19FF914BD1F80413B6F7DB" box="[1177,1257,2047,2070]" pageId="19" pageNumber="182" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="4C62C8AEAC19FF914A51F8041226F7DB" authority="Liu &amp; Ono &amp; Maruyama, 2021" authorityName="Liu &amp; Ono &amp; Maruyama" authorityYear="2021" box="[1305,1401,2047,2070]" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="182" phylum="Arthropoda" rank="species" species="itsuroi" status="sp. nov.">
<emphasis id="B9166F3FAC19FF914A51F8041226F7DB" box="[1305,1401,2047,2070]" italics="true" pageId="19" pageNumber="182">B. itsuroi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A225D244AC19FF974A36F8041799FF18" lastPageId="21" lastPageNumber="184" pageId="19" pageNumber="182" rank="species">sp. nov.</taxonomicNameLabel>
, but easily distinguished from 
<taxonomicName id="4C62C8AEAC1FFF974D48FF4515FBFF18" authority="Liu &amp; Ono &amp; Maruyama, 2021" authorityName="Liu &amp; Ono &amp; Maruyama" authorityYear="2021" box="[512,676,189,213]" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="184" phylum="Arthropoda" rank="species" species="hokkaidensis" status="sp. nov.">
<emphasis id="B9166F3FAC1FFF974D48FF4515FBFF18" box="[512,676,189,213]" italics="true" pageId="21" pageNumber="184">B. hokkaidensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A225D244AC1FFF974DE0FF4515A6FF18" box="[680,761,190,213]" pageId="21" pageNumber="184" rank="species">sp. nov.</taxonomicNameLabel>
by the paler body colour and the shape of the spermatheca, and from 
<taxonomicName id="4C62C8AEAC1FFF974E49FF05163AFED8" authority="Liu &amp; Ono &amp; Maruyama, 2021" authorityName="Liu &amp; Ono &amp; Maruyama" authorityYear="2021" box="[257,357,254,277]" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="184" phylum="Arthropoda" rank="species" species="itsuroi" status="sp. nov.">
<emphasis id="B9166F3FAC1FFF974E49FF05163AFED8" box="[257,357,254,277]" italics="true" pageId="21" pageNumber="184">B. itsuroi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A225D244AC1FFF974E25FF05169EFED8" box="[365,449,254,277]" pageId="21" pageNumber="184" rank="species">sp. nov.</taxonomicNameLabel>
by the number of facets and the shape of the spermatheca.
</paragraph>
</subSubSection>
<caption id="DF1DE3A5AC1EFF964FDFF8161237F7D7" ID-DOI="http://doi.org/10.5281/zenodo.5037356" ID-Zenodo-Dep="5037356" httpUri="https://zenodo.org/record/5037356/files/figure.png" pageId="20" pageNumber="183" startId="20.[151,182,2029,2048]" targetBox="[154,1414,208,1976]" targetPageId="20">
<paragraph id="8BDDB32DAC1EFF964FDFF8161237F7D7" blockId="20.[151,1436,2028,2074]" pageId="20" pageNumber="183">
Fig. 12. 
<taxonomicName id="4C62C8AEAC1EFF964F97F81716F4F832" authority="Liu &amp; Ono &amp; Maruyama, 2021" authorityName="Liu &amp; Ono &amp; Maruyama" authorityYear="2021" box="[223,427,2028,2047]" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="183" phylum="Arthropoda" rank="species" species="fulvipennis" status="sp. nov.">
<emphasis id="B9166F3FAC1EFF964F97F81716F4F832" box="[223,427,2028,2047]" italics="true" pageId="20" pageNumber="183">Bryothinusa fulvipennis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A225D244AC1EFF964EF9F81616ABF7CD" box="[433,500,2029,2048]" pageId="20" pageNumber="183" rank="species">sp. nov.</taxonomicNameLabel>
A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm.
</paragraph>
</caption>
<subSubSection id="C378E0A6AC1FFF974FDFFEC516C2FEB8" pageId="21" pageNumber="184" type="etymology">
<paragraph id="8BDDB32DAC1FFF974FDFFEC516C2FEB8" blockId="21.[151,761,189,597]" pageId="21" pageNumber="184">
<emphasis id="B9166F3FAC1FFF974FDFFEC51647FE98" bold="true" box="[151,280,318,341]" pageId="21" pageNumber="184">Etymology.</emphasis>
The species epithet refers to the yellowish -brown elytra; adjective.
</paragraph>
</subSubSection>
<subSubSection id="C378E0A6AC1FFF974FDFFE851650FE18" pageId="21" pageNumber="184" type="distribution">
<paragraph id="8BDDB32DAC1FFF974FDFFE851650FE18" blockId="21.[151,761,189,597]" pageId="21" pageNumber="184">
<emphasis id="B9166F3FAC1FFF974FDFFE85166CFE58" bold="true" box="[151,307,382,405]" pageId="21" pageNumber="184">Distribution.</emphasis>
<collectingCountry id="F375F3BDAC1FFF974E77FE851620FE58" box="[319,383,382,405]" name="Japan" pageId="21" pageNumber="184">Japan</collectingCountry>
(
<emphasis id="B9166F3FAC1FFF974EDDFE8516A3FE58" bold="true" box="[405,508,382,405]" pageId="21" pageNumber="184">Honshû:</emphasis>
Chiba-ken, Mie-ken, Ôsaka-fu, Okayama-ken, Hiroshima-ken; 
<emphasis id="B9166F3FAC1FFF974D17FE651598FE78" bold="true" box="[607,711,414,437]" pageId="21" pageNumber="184">Shikoku:</emphasis>
Kagawa-ken).
</paragraph>
</subSubSection>
<subSubSection id="C378E0A6AC1FFF974FDFFE2515D5FD98" pageId="21" pageNumber="184" type="discussion">
<paragraph id="8BDDB32DAC1FFF974FDFFE2515D5FD98" blockId="21.[151,761,189,597]" pageId="21" pageNumber="184">
<emphasis id="B9166F3FAC1FFF974FDFFE251659FE38" bold="true" box="[151,262,478,501]" pageId="21" pageNumber="184">Remarks.</emphasis>
This species has been found mainly in coasts of bay and inland sea such as Tôkyô-wan, Ise-wan and Setonaikai. A distribution map of examined specimens of this species in 
<collectingCountry id="F375F3BDAC1FFF974E7BFDC5162FFD98" box="[307,368,574,597]" name="Japan" pageId="21" pageNumber="184">Japan</collectingCountry>
is illustrated in 
<figureCitation id="1359AFA8AC1FFF974D54FDC515D9FD98" box="[540,646,574,597]" captionStart="Fig" captionStartId="35.[150,181,1388,1407]" captionTargetBox="[150,1434,191,1370]" captionTargetId="figure-236@35.[150,1435,191,1371]" captionTargetPageId="35" captionText="Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle – B. algarum Sawada, 1971; black inverted triangle – B. minuta (Sawada, 1955); white star – B. tsutsuii (Sawada, 1955); black star – B. koreana Ahn &amp; Jeon, 2004; black square – B. yoshigoui sp. nov.; white triangle – B. japonica Liu, Ono &amp; Maruyama, 2020; black diamond – B. sakishimana Sawada, 1991; white diamond – B. constricta sp. nov.; white square – B. hokkaidensis sp. nov.; black triangle – B. fulvipennis sp. nov.; double circle – B. itsuroi sp. nov.; arrow – B. aikoae sp. nov.; black hart – B. okinawana sp. nov.; circled dot – B. nigra sp. nov.; white circle – B. nakanei (Sawada, 1955); white hart – B. moriguchii sp. nov.; black circle – B. gangjinensis Ahn &amp; Jeon, 2004." figureDoi="http://doi.org/10.5281/zenodo.5037376" httpUri="https://zenodo.org/record/5037376/files/figure.png" pageId="21" pageNumber="184">Figure 22</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>