189 lines
21 KiB
XML
189 lines
21 KiB
XML
<document id="DD630FBAD92082AE32CD63C69C311E8F" ID-CLB-Dataset="21809" ID-DOI="10.5852/ejt.2020.652" ID-GBIF-Dataset="64fd24bb-ee1d-44d7-ac6a-25e8d5f96777" ID-ISSN="2118-9773" ID-Zenodo-Dep="3860236" ID-ZooBank="DC65C142-53F1-4416-A916-8F78C27DCF93" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1590592466678" checkinUser="valdenar" docAuthor="Garbin, Rafaella C., Bandyopadhyay, Saswati & Joyce, Walter G." docDate="2020" docId="03C7CF0BC02E1D6FFDF6AB43FE3696F9" docLanguage="en" docName="ejt-652_garbin_bandyopadhyay_joyce.pdf.imf" docOrigin="European Journal of Taxonomy 652" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Pangshura tatrotia Joyce & Lyson 2010" docType="treatment" docVersion="8" lastPageNumber="28" masterDocId="FFFEB773C0341D74FFD1A908FFA79736" masterDocTitle="A taxonomic revision of geoemydid turtles from Siwalik-age of India and Pakistan" masterLastPageNumber="67" masterPageNumber="1" pageNumber="27" updateTime="1698841626966" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="3550FA4E0B248C2A046082DF4B6A878E">
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<mods:title id="17E24D33CDEBB40AB53562523E219ED6">A taxonomic revision of geoemydid turtles from Siwalik-age of India and Pakistan</mods:title>
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<mods:name id="624277AABCE769AA7594FDB1E0232A3D" type="personal">
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<mods:roleTerm id="D458518F2575581FF34B85CE2730808E">Author</mods:roleTerm>
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<mods:namePart id="BD1A5732FD9CBC74F3F57FFACEB0A8CC">Garbin, Rafaella C.</mods:namePart>
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<mods:affiliation id="293EA087CF799D146EF05A0E3D4D3DE4">Department of Geosciences, University of Fribourg, Fribourg; Chemin du Musée 6, 1700 Fribourg, Switzerland. & urn: lsid: zoobank. org: author: A 8 FCEF 2 B- 7524 - 46 C 3 - 808 C-D 9 CBFBC 08017 & Corresponding author: garbin. rafaella @ gmail. com</mods:affiliation>
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<mods:nameIdentifier id="2D9C60B28291C2C855DA248367813065" type="email">garbin.rafaella@gmail.com</mods:nameIdentifier>
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</mods:role>
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<mods:namePart id="586F6B0D7A18C77975F47523C6A162D9">Bandyopadhyay, Saswati</mods:namePart>
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<mods:affiliation id="53F985CC0BBA27B29C34B1E04C23EC76">Geological Studies Unit, Indian Statistical Institute, Kolkata; Barrackpore Trunk Road 233, Kolkata, India. & Email: saswati @ isical. ac. in & urn: lsid: zoobank. org: author: 02005 D 35 - 6 DA 9 - 4 A 7 E-B 838 - 16 F 4 FEF 108 DA</mods:affiliation>
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<mods:nameIdentifier id="BD9CC82BA0CB2261C6366A57C9591396" type="email">saswati@isical.ac.in</mods:nameIdentifier>
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<mods:namePart id="2847F9C5C1DD3E714186F7CD0B5A5D88">Joyce, Walter G.</mods:namePart>
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<mods:affiliation id="65E5E750321531E05068CC9AC7857E09">Department of Geosciences, University of Fribourg, Fribourg; Chemin du Musée 6, 1700 Fribourg, Switzerland. & Email: walter. joyce @ unifr. ch & urn: lsid: zoobank. org: author: C 30 D 317 A- 0029 - 439 C- 8446 - 31281771 ADA 5</mods:affiliation>
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<mods:nameIdentifier id="7E9B2FD273A1425523B2C4351D10B8CF" type="email">walter.joyce@unifr.ch</mods:nameIdentifier>
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<mods:title id="143B58AA86BE54E2D4B9390518586341">European Journal of Taxonomy</mods:title>
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<mods:part id="89C2023197E1FE005C382DB98224F441">
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<mods:date id="2BD0AA8739897E8B951FEE5DBB7D43B4">2020</mods:date>
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<mods:number id="D6C69958BC564FE2EA206481C60448E6">2020-05-26</mods:number>
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<mods:number id="0BCF3AD34A9D7CC7D513A3C482D4C911">652</mods:number>
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<mods:identifier id="8EF94BDDEF6321D57B1C743DA35CDFCA" type="CLB-Dataset">21809</mods:identifier>
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<mods:identifier id="A1EE0E12DD1BFB65AEA4907AA249D45B" type="DOI">10.5852/ejt.2020.652</mods:identifier>
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<mods:identifier id="02CAC09A5EC7738C14347A89D01CDEE4" type="ISSN">2118-9773</mods:identifier>
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<treatment id="03C7CF0BC02E1D6FFDF6AB43FE3696F9" ID-DOI="http://doi.org/10.5281/zenodo.3861099" ID-GBIF-Taxon="164362575" ID-Zenodo-Dep="3861099" LSID="urn:lsid:plazi:treatment:03C7CF0BC02E1D6FFDF6AB43FE3696F9" httpUri="http://treatment.plazi.org/id/03C7CF0BC02E1D6FFDF6AB43FE3696F9" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
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<subSubSection id="C3742D96C02E1D6EFDF6AB43FBAA9553" box="[551,1037,587,614]" pageId="26" pageNumber="27" type="nomenclature">
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<paragraph id="8BD17E1DC02E1D6EFDF6AB43FBAA9553" blockId="26.[551,1037,587,614]" box="[551,1037,587,614]" pageId="26" pageNumber="27">
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<heading id="D099C971C02E1D6EFDF6AB43FBAA9553" box="[551,1037,587,614]" centered="true" fontSize="11" level="2" pageId="26" pageNumber="27" reason="2">
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<taxonomicName id="4C6E059EC02E1D6EFDF6AB43FBAA9553" authority="Joyce & Lyson, 2010" authorityName="Joyce & Lyson" authorityYear="2010" box="[551,1037,587,614]" class="Reptilia" family="Geoemydidae" genus="Pangshura" kingdom="Animalia" order="Testudines" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="tatrotia">
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<emphasis id="B91AA20FC02E1D6EFDF6AB43FCAB9553" bold="true" box="[551,780,587,614]" italics="true" pageId="26" pageNumber="27">Pangshura tatrotia</emphasis>
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<bibRefCitation id="EFFF03ECC02E1D6EFCC2AB44FBAA9553" author="Joyce W. G. & Lyson T. R." box="[787,1037,587,614]" pageId="26" pageNumber="27" pagination="449 - 458" refId="ref24174" refString="Joyce W. G. & Lyson T. R. 2010. Pangshura tatrotia, a new species of pond turtle (Testudinoidea) from the Pliocene Siwaliks of Pakistan. Journal of Systematic Palaeontology 8 (3): 449 - 458. https: // doi. org / 10.1080 / 14772019.2010.500879" type="journal article" year="2010">Joyce & Lyson, 2010</bibRefCitation>
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</taxonomicName>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3742D96C02E1D6EFF6CAB87FB02939F" pageId="26" pageNumber="27" type="materials_examined">
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<paragraph id="8BD17E1DC02E1D6EFF6CAB87FF5D959F" blockId="26.[189,250,655,681]" box="[189,250,655,681]" pageId="26" pageNumber="27">
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<heading id="D099C971C02E1D6EFF6CAB87FF5D959F" bold="true" box="[189,250,655,681]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
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<emphasis id="B91AA20FC02E1D6EFF6CAB87FF5D959F" bold="true" box="[189,250,655,681]" pageId="26" pageNumber="27">Type</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD17E1DC02E1D6EFF6CABC8FD3395EC" blockId="26.[189,660,704,730]" box="[189,660,704,730]" pageId="26" pageNumber="27">
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<collectionCode id="ED7FE6D8C02E1D6EFF6CABC8FF5A95EC" box="[189,253,704,730]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34880" lsid="urn:lsid:biocol.org:col:34880" name="Peabody Museum of Natural History" pageId="26" pageNumber="27" type="Museum">YPM</collectionCode>
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4127, a near complete fossil shell.
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</paragraph>
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<paragraph id="8BD17E1DC02E1D6EFF6CAA0BFE0D942B" blockId="26.[189,426,771,797]" box="[189,426,771,797]" pageId="26" pageNumber="27">
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<heading id="D099C971C02E1D6EFF6CAA0BFE0D942B" bold="true" box="[189,426,771,797]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
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<emphasis id="B91AA20FC02E1D6EFF6CAA0BFE0D942B" bold="true" box="[189,426,771,797]" pageId="26" pageNumber="27">Material examined</emphasis>
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</heading>
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</paragraph>
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<materialsCitation id="3B067440C02E1D6EFF3BAA3DFEE194AC" ID-GBIF-Occurrence="2626092310" collectionCode="YPM" country="Pakistan" county="Potwar Plateau" location="Potwar Plateau" municipality="Padhri" pageId="26" pageNumber="27" specimenCode="YPM 4127" specimenCount="1" stateProvince="Punjab" typeStatus="holotype">
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<paragraph id="8BD17E1DC02E1D6EFF3BAA3DFEFB9479" blockId="26.[189,1398,821,922]" box="[234,348,821,847]" pageId="26" pageNumber="27">
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<emphasis id="B91AA20FC02E1D6EFF3BAA3DFEFB9479" bold="true" box="[234,348,821,847]" pageId="26" pageNumber="27">
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<typeStatus id="54D5C0BFC02E1D6EFF3BAA3DFEFB9479" box="[234,348,821,847]" pageId="26" pageNumber="27" type="holotype">Holotype</typeStatus>
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</emphasis>
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||
</paragraph>
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||
<paragraph id="8BD17E1DC02E1D6EFF6CAA52FEE194AC" blockId="26.[189,1398,821,922]" pageId="26" pageNumber="27">
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||
<collectingCountry id="F3793E8DC02E1D6EFF6CAA52FEEE9442" box="[189,329,858,884]" name="Pakistan" pageId="26" pageNumber="27">PAKISTAN</collectingCountry>
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•
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<collectingRegion id="49AAB0FFC02E1D6EFEB6AA52FE1B9442" box="[359,444,858,884]" country="Pakistan" name="Punjab" pageId="26" pageNumber="27">Punjab</collectingRegion>
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,
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<location id="8EB128C6C02E1D6EFE1BAA53FD279442" LSID="urn:lsid:plazi:treatment:03C7CF0BC02E1D6FFDF6AB43FE3696F9:8EB128C6C02E1D6EFE1BAA53FD279442" box="[458,640,858,885]" country="Pakistan" county="Potwar Plateau" municipality="Padhri" name="Potwar Plateau" pageId="26" pageNumber="27" stateProvince="Punjab">Potwar Plateau</location>
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,
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<locationDeviation id="9A63673FC02E1D6EFD5FAA52FC459442" box="[654,994,858,884]" pageId="26" pageNumber="27">
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||
<quantity id="4C96D3F8C02E1D6EFD5FAA52FD409442" box="[654,743,858,884]" metricMagnitude="3" metricUnit="m" metricValue="3.218688" pageId="26" pageNumber="27" unit="mi" value="2.0">2 miles</quantity>
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north-east of Padhri
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</locationDeviation>
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; Tatrot Formation; Late Pliocene;
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<specimenCode id="DBC8D666C02E1D6EFF6CAA88FEE594AC" box="[189,322,896,922]" collectionCode="YPM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34880" lsid="urn:lsid:biocol.org:col:34880" name="Peabody Museum of Natural History" pageId="26" pageNumber="27" type="Museum">YPM 4127</specimenCode>
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.
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</paragraph>
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</materialsCitation>
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||
<paragraph id="8BD17E1DC02E1D6EFF6CAACCFEFA94EB" blockId="26.[189,349,963,990]" box="[189,349,963,990]" pageId="26" pageNumber="27">
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<heading id="D099C971C02E1D6EFF6CAACCFEFA94EB" bold="true" box="[189,349,963,990]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
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<emphasis id="B91AA20FC02E1D6EFF6CAACCFEFA94EB" bold="true" box="[189,349,963,990]" pageId="26" pageNumber="27">Type locality</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD17E1DC02E1D6EFF6CAAFCFE8F9302" blockId="26.[189,1398,1012,1076]" pageId="26" pageNumber="27">
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Yale North
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<collectingCountry id="F3793E8DC02E1D6EFE97AAFCFE259338" box="[326,386,1012,1038]" name="India" pageId="26" pageNumber="27">India</collectingCountry>
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Expedition locality 99, about two miles north-east of Padhri, Potwar Plateau,
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<collectingRegion id="49AAB0FFC02E1D6EFACFAAFCFAD49338" box="[1310,1395,1012,1038]" country="Pakistan" name="Punjab" pageId="26" pageNumber="27">Punjab</collectingRegion>
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,
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<collectingCountry id="F3793E8DC02E1D6EFF6CAD12FE839302" box="[189,292,1050,1076]" name="Pakistan" pageId="26" pageNumber="27">Pakistan</collectingCountry>
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.
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</paragraph>
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<paragraph id="8BD17E1DC02E1D6EFF6CAD55FEE89341" blockId="26.[189,335,1117,1143]" box="[189,335,1117,1143]" pageId="26" pageNumber="27">
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<heading id="D099C971C02E1D6EFF6CAD55FEE89341" bold="true" box="[189,335,1117,1143]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
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||
<emphasis id="B91AA20FC02E1D6EFF6CAD55FEE89341" bold="true" box="[189,335,1117,1143]" pageId="26" pageNumber="27">Occurrence</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="8BD17E1DC02E1D6EFF6CAD87FB02939F" blockId="26.[189,1189,1166,1193]" box="[189,1189,1166,1193]" pageId="26" pageNumber="27">Tatrot Formation, Late Pliocene (3.6 to 2.6 Ma, possibly Plio–Pleistocene boundary).</paragraph>
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</subSubSection>
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||
<subSubSection id="C3742D96C02E1D6EFF6CADDAFE92925E" pageId="26" pageNumber="27" type="diagnosis">
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<paragraph id="8BD17E1DC02E1D6EFF6CADDAFCD093DA" blockId="26.[189,887,1234,1260]" box="[189,887,1234,1260]" pageId="26" pageNumber="27">
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<heading id="D099C971C02E1D6EFF6CADDAFCD093DA" bold="true" box="[189,887,1234,1260]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
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||
<emphasis id="B91AA20FC02E1D6EFF6CADDAFCD093DA" bold="true" box="[189,887,1234,1260]" pageId="26" pageNumber="27">Differential osteological diagnosis using shell characters</emphasis>
|
||
</heading>
|
||
</paragraph>
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||
<paragraph id="8BD17E1DC02E1D6EFF6CAC0BFE92925E" blockId="26.[189,1399,1283,1384]" pageId="26" pageNumber="27">
|
||
<taxonomicName id="4C6E059EC02E1D6EFF6CAC0BFE39922B" authorityName="Joyce & Lyson" authorityYear="2010" box="[189,414,1283,1309]" class="Reptilia" family="Geoemydidae" genus="Pangshura" kingdom="Animalia" order="Testudines" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="tatrotia">
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<emphasis id="B91AA20FC02E1D6EFF6CAC0BFE39922B" box="[189,414,1283,1309]" italics="true" pageId="26" pageNumber="27">Pangshura tatrotia</emphasis>
|
||
</taxonomicName>
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can be differentiated from other species of
|
||
<taxonomicName id="4C6E059EC02E1D6EFC78AC0BFB8D922B" authorityName="Gray" authorityYear="1856" box="[937,1066,1283,1309]" class="Reptilia" family="Geoemydidae" genus="Pangshura" kingdom="Animalia" order="Testudines" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">
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||
<emphasis id="B91AA20FC02E1D6EFC78AC0BFB8D922B" box="[937,1066,1283,1309]" italics="true" pageId="26" pageNumber="27">Pangshura</emphasis>
|
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</taxonomicName>
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by the presence of a strong median keel projection on both the second and third vertebral, and a first vertebral scute that is constricted anteriorly.
|
||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3742D96C02E1D6EFF6CAC9AFD7190DD" pageId="26" pageNumber="27" type="description">
|
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<paragraph id="8BD17E1DC02E1D6EFF6CAC9AFE7D929A" blockId="26.[189,474,1426,1452]" box="[189,474,1426,1452]" pageId="26" pageNumber="27">
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<heading id="D099C971C02E1D6EFF6CAC9AFE7D929A" bold="true" box="[189,474,1426,1452]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
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<emphasis id="B91AA20FC02E1D6EFF6CAC9AFE7D929A" bold="true" box="[189,474,1426,1452]" pageId="26" pageNumber="27">
|
||
Description of the
|
||
<typeStatus id="54D5C0BFC02E1D6EFE75AC9AFE7D929A" box="[420,474,1426,1452]" pageId="26" pageNumber="27">type</typeStatus>
|
||
</emphasis>
|
||
</heading>
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||
</paragraph>
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<paragraph id="8BD17E1DC02E1D6EFF6CACCBFD7190DD" blockId="26.[189,1399,1475,2027]" pageId="26" pageNumber="27">
|
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YPM 4127,
|
||
<typeStatus id="54D5C0BFC02E1D6EFE85ACCBFE1C92EB" box="[340,443,1475,1501]" pageId="26" pageNumber="27" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C6E059EC02E1D6EFE39ACCBFD6C92EB" authorityName="Joyce & Lyson" authorityYear="2010" box="[488,715,1475,1501]" class="Reptilia" family="Geoemydidae" genus="Pangshura" kingdom="Animalia" order="Testudines" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="tatrotia">
|
||
<emphasis id="B91AA20FC02E1D6EFE39ACCBFD6C92EB" box="[488,715,1475,1501]" italics="true" pageId="26" pageNumber="27">Pangshura tatrotia</emphasis>
|
||
</taxonomicName>
|
||
– This is an almost complete, well-preserved specimen, with a tectiform carapace from the Early Pliocene Tatrot Formation of the Potwar Plateau of
|
||
<collectingRegion id="49AAB0FFC02E1D6EFACFACE0FAD49134" box="[1310,1395,1512,1538]" country="Pakistan" name="Punjab" pageId="26" pageNumber="27">Punjab</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3793E8DC02E1D6EFF6CAF06FE83911E" box="[189,292,1550,1576]" name="Pakistan" pageId="26" pageNumber="27">Pakistan</collectingCountry>
|
||
. The posterior peripheral bones and left peripheral bones IV–VI are missing. A median, wellpronounced carapacial keel is present from neurals II to VIII. Neural bones II, III, VI–VIII are hexagonal with anterior short sides. Neural IV is octagonal, with both anterior and posterior sides short. Neural V is quadrangular, without short sides. The fourth vertebral scute runs from neural IV to VIII and has a strong anterior bottle-neck-shaped constriction. The posterior margins of the first and second pleural scutes run over costals II and IV, respectively, and have a strong anterior projection that crosses to the anterior costal bone. The pygal bone is completely divided by the twelfth intermarginal sulcus. Parts of the anterior and posterior plastral margins are missing. The entoplastron is intersected anteriorly by the gularohumeral sulcus, but not by the humeropectoral sulcus. The hyo-hypoplastral suture contacts peripheral V and does not overlap with the pectoroabdominal sulcus. The fifth and sixth marginal scutes form the bridge and overlap onto the hyoplastra. Only the sixth marginal scute overlaps with the hypoplastron. A large inguinal scute is present, likely contacting the femoral scute. For a more comprehensive description of this specimen, refer to
|
||
<bibRefCitation id="EFFF03ECC02E1D6EFE18AED9FD7490DD" author="Joyce W. G. & Lyson T. R." box="[457,723,2001,2027]" pageId="26" pageNumber="27" pagination="449 - 458" refId="ref24174" refString="Joyce W. G. & Lyson T. R. 2010. Pangshura tatrotia, a new species of pond turtle (Testudinoidea) from the Pliocene Siwaliks of Pakistan. Journal of Systematic Palaeontology 8 (3): 449 - 458. https: // doi. org / 10.1080 / 14772019.2010.500879" type="journal article" year="2010">Joyce & Lyson (2010)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3742D96C02F1D6FFF6CA803FE3696F9" pageId="27" pageNumber="28" type="discussion">
|
||
<paragraph id="8BD17E1DC02F1D6FFF6CA803FEE39613" blockId="27.[189,324,267,293]" box="[189,324,267,293]" pageId="27" pageNumber="28">
|
||
<heading id="D099C971C02F1D6FFF6CA803FEE39613" bold="true" box="[189,324,267,293]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
|
||
<emphasis id="B91AA20FC02F1D6FFF6CA803FEE39613" bold="true" box="[189,324,267,293]" pageId="27" pageNumber="28">Comments</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BD17E1DC02F1D6FFF6CA836FE3696F9" blockId="27.[189,1399,318,463]" pageId="27" pageNumber="28">
|
||
<taxonomicName id="4C6E059EC02F1D6FFF6CA836FE06966E" authorityName="Joyce & Lyson" authorityYear="2010" box="[189,417,318,344]" class="Reptilia" family="Geoemydidae" genus="Pangshura" kingdom="Animalia" order="Testudines" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="tatrotia">
|
||
<emphasis id="B91AA20FC02F1D6FFF6CA836FE06966E" box="[189,417,318,344]" italics="true" pageId="27" pageNumber="28">Pangshura tatrotia</emphasis>
|
||
</taxonomicName>
|
||
was only recently named based on a well-preserved shell that documents with confidence a morphotype different from all extant representatives of
|
||
<taxonomicName id="4C6E059EC02F1D6FFBC9A86EFB3E96B6" authorityName="Gray" authorityYear="1856" box="[1048,1177,358,384]" class="Reptilia" family="Geoemydidae" genus="Pangshura" kingdom="Animalia" order="Testudines" pageId="27" pageNumber="28" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91AA20FC02F1D6FFBC9A86EFB3E96B6" box="[1048,1177,358,384]" italics="true" pageId="27" pageNumber="28">Pangshura</emphasis>
|
||
</taxonomicName>
|
||
. The specimen is furthermore associated with quality locality data. We therefore here find the validity of this species to be unproblematic.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |