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<document id="D5C8F50788EEB0143019EE2E636145B3" ID-DOI="10.11646/zootaxa.5397.3.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="10468872" ID-ZooBank="E14C585C-BF45-4BD5-8B12-AB9BB13BF66E" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1704708245043" checkinUser="plazi" docAuthor="Anichtchenko, Alexander &amp; Wiesner, Jürgen" docDate="2024" docId="03C78105101B7579DDAA358BFE88FB3B" docLanguage="en" docName="zootaxa.5397.3.3.pdf" docOrigin="Zootaxa 5397 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5397.3.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tricondyla (Stenotricondyla) conicicollis Chaudoir 1844" docType="treatment" docVersion="3" lastPageNumber="363" masterDocId="FFFEF97D10107575DD3D332DFFB8FF94" masterDocTitle="Tiger beetles of the genus Tricondyla Latreille, 1822 (Coleoptera: Cicindelidae) of the Philippines, with some considerations of the taxonomy of the genus" masterLastPageNumber="376" masterPageNumber="351" pageNumber="362" updateTime="1704709213912" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="91CD3EE783F7167415EC5195E6FC2AF1">Tiger beetles of the genus Tricondyla Latreille, 1822 (Coleoptera: Cicindelidae) of the Philippines, with some considerations of the taxonomy of the genus</mods:title>
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<mods:namePart id="E73444C8954CFC0A328399FF326C0CD3">Anichtchenko, Alexander</mods:namePart>
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<treatment id="03C78105101B7579DDAA358BFE88FB3B" ID-DOI="http://doi.org/10.5281/zenodo.10468790" ID-Zenodo-Dep="10468790" LSID="urn:lsid:plazi:treatment:03C78105101B7579DDAA358BFE88FB3B" httpUri="http://treatment.plazi.org/id/03C78105101B7579DDAA358BFE88FB3B" lastPageId="12" lastPageNumber="363" pageId="11" pageNumber="362">
<subSubSection id="C3746398101B757EDDAA358BFDFDF897" pageId="11" pageNumber="362" type="nomenclature">
<paragraph id="8BD13013101B757EDDAA358BFC30F955" blockId="11.[151,904,1702,1729]" box="[151,904,1702,1729]" pageId="11" pageNumber="362">
<heading id="D099877F101B757EDDAA358BFC30F955" bold="true" box="[151,904,1702,1729]" fontSize="11" level="1" pageId="11" pageNumber="362" reason="1">
<taxonomicName id="4C6E4B90101B757EDDAA358BFC30F955" ID-CoL="58MWQ" authority="Chaudoir, 1844: 458" authorityName="Chaudoir" authorityPageNumber="458" authorityYear="1844" box="[151,904,1702,1729]" class="Insecta" family="Carabidae" genus="Tricondyla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="362" phylum="Arthropoda" rank="species" species="conicicollis" subGenus="Stenotricondyla">
<emphasis id="B91AEC01101B757EDDAA358BFC30F955" bold="true" box="[151,904,1702,1729]" pageId="11" pageNumber="362">
<emphasis id="B91AEC01101B757EDDAA358BFD3CF955" bold="true" box="[151,644,1702,1729]" italics="true" pageId="11" pageNumber="362">Tricondyla (Stenotricondyla) conicicollis</emphasis>
Chaudoir, 1844: 458
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BD13013101B757EDDAA35C1FDFDF897" blockId="11.[151,721,1771,1857]" box="[151,581,1771,1796]" pageId="11" pageNumber="362">
<materialsCitation id="3B063A4E101B757EDDAA35C1FDFDF897" box="[151,581,1771,1796]" location="Manille" pageId="11" pageNumber="362" specimenCount="1" stateProvince="Manila" typeStatus="holotype">
Loc. typ.: “
<collectingRegion id="49AAFEF1101B757EDC3435C6FEE7F897" box="[265,351,1771,1795]" country="Philippines" name="Manila" pageId="11" pageNumber="362">Manille</collectingRegion>
”.
<typeStatus id="54D58EB1101B757EDC4C35C6FE77F897" box="[369,463,1771,1795]" pageId="11" pageNumber="362" type="holotype">Holotype</typeStatus>
in MNHN.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3746398101B757EDDAA3427FD3BF8D5" pageId="11" pageNumber="362" type="reference_group">
<paragraph id="8BD13013101B757EDDAA3427FD69F8B6" blockId="11.[151,721,1771,1857]" box="[151,721,1802,1826]" pageId="11" pageNumber="362">
<taxonomicName id="4C6E4B90101B757EDDAA3427FDD1F8B6" ID-CoL="58MWZ" authority="Naviaux, 2009" authorityName="Naviaux" authorityYear="2009" box="[151,617,1802,1826]" class="Insecta" family="Carabidae" genus="Tricondyla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="362" phylum="Arthropoda" rank="species" species="huloti" status="syn. nov." subGenus="Tricondyla">
<emphasis id="B91AEC01101B757EDDAA3427FE73F8B6" box="[151,459,1802,1826]" italics="true" pageId="11" pageNumber="362">Tricondyla (Tricondyla) huloti</emphasis>
<bibRefCitation id="EFFF4DE2101B757EDCEF3427FDD1F8B6" author="Naviaux, R." box="[466,617,1802,1826]" pageId="11" pageNumber="362" pagination="107 - 109" refId="ref14679" refString="Naviaux, R. (2009) Tricondyla (s. str.) huloti, nouvelle espece de Philippines (Coleoptera: Cicindelidae). Revue Francaise d'Entomologie, 31 (3 - 4), 107 - 109." type="journal article" year="2009">Naviaux, 2009</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91AEC01101B757EDF493427FD69F8B6" bold="true" box="[628,721,1802,1826]" pageId="11" pageNumber="362">
<taxonomicNameLabel id="A229517A101B757EDF493427FD69F8B6" box="[628,721,1802,1826]" pageId="11" pageNumber="362">syn. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BD13013101B757EDDAA3405FD3BF8D5" blockId="11.[151,721,1771,1857]" box="[151,643,1832,1857]" pageId="11" pageNumber="362">
<taxonomicName id="4C6E4B90101B757EDDAA3405FD3BF8D5" authority="Mandl, 1964: 78" authorityName="Mandl" authorityPageNumber="78" authorityYear="1964" box="[151,643,1832,1857]" class="Insecta" family="Carabidae" genus="Tricondyla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="362" phylum="Arthropoda" rank="subSpecies" species="cyanipes" subSpecies="fuscilabris">
<emphasis id="B91AEC01101B757EDDAA3405FE6DF8D4" box="[151,469,1832,1856]" italics="true" pageId="11" pageNumber="362">Tricondyla cyanipes fuscilabris</emphasis>
Mandl, 1964: 78
</taxonomicName>
</paragraph>
</subSubSection>
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<paragraph id="8BD13013101B757EDDAA3442FD0DF861" blockId="11.[151,1437,1902,2037]" pageId="11" pageNumber="362">
<emphasis id="B91AEC01101B757EDDAA3442FEF1F81C" bold="true" box="[151,329,1902,1929]" pageId="11" pageNumber="362">Type material.</emphasis>
<materialsCitation id="3B063A4E101B757EDC6E3442FD09F861" collectionCode="NHMB" collectorName="F. G. Werner &amp; Coll. G. Frey" determinerName="Dr. Ing. K. Mandl &amp; R. Naviaux" elevation="692" location="Mantalingajan Range" municipality="Mt. Balabag" pageId="11" pageNumber="362" specimenCount="2" specimenCount-male="2" stateProvince="Palawan" typeStatus="paratype">
<typeStatus id="54D58EB1101B757EDC6E3442FE7BF81D" box="[339,451,1903,1929]" pageId="11" pageNumber="362" type="paratype">Paratypes</typeStatus>
,
<specimenCount id="9D68FB9A101B757EDCEF3442FD89F81D" box="[466,561,1903,1929]" count="2" pageId="11" pageNumber="362" type="male">2 males</specimenCount>
: “S. slope
<collectingMunicipality id="6BB5AA69101B757EDF8E3442FCFCF81D" box="[691,836,1903,1929]" pageId="11" pageNumber="362">Mt. Balabag</collectingMunicipality>
, /
<location id="8EB166C8101B757EDE583442FBE4F81D" LSID="urn:lsid:plazi:treatment:03C78105101B7579DDAA358BFE88FB3B:8EB166C8101B757EDE583442FBE4F81D" box="[869,1116,1903,1929]" municipality="Mt. Balabag" name="Mantalingajan Range" pageId="11" pageNumber="362" stateProvince="Palawan">Mantalingajan Range</location>
, /
<collectingRegion id="49AAFEF1101B757ED9433442FB65F81D" box="[1150,1245,1903,1929]" country="Philippines" name="Palawan" pageId="11" pageNumber="362">Palawan</collectingRegion>
Is., /
<quantity id="4C969DF6101B757ED81A3442FA20F81E" box="[1319,1432,1903,1930]" metricMagnitude="2" metricUnit="m" metricValue="6.922008" metricValueMax="13.716" metricValueMin="0.128016" pageId="11" pageNumber="362" unit="ft" value="2271.0" valueMax="4500.0" valueMin="42.0">
<elevation id="0043D720101B757ED81A3442FA20F81E" box="[1319,1432,1903,1930]" metricMagnitude="2" metricUnit="m" metricValue="6.922008" metricValueMax="13.716" metricValueMin="0.128016" pageId="11" pageNumber="362" unit="ft" value="2271.0" valueMax="4500.0" valueMin="42.0">42-4500ft</elevation>
</quantity>
, V.10-16.47 // CNHM-Philippine / Zool. Exped. (1946-47) /
<collectorName id="269B55C5101B757EDE7134BEFC6FF839" box="[844,983,1939,1965]" pageId="11" pageNumber="362">F.G. Werner</collectorName>
leg. //
<typeStatus id="54D58EB1101B757ED91034BEFB23F839" box="[1069,1179,1939,1965]" pageId="11" pageNumber="362" type="paratype">Paratypus</typeStatus>
/
<taxonomicName id="4C6E4B90101B757ED98B34BEFA24F839" authorityName="Eschscholtz" authorityYear="1829" box="[1206,1436,1939,1965]" class="Insecta" family="Carabidae" genus="Tricondyla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="362" phylum="Arthropoda" rank="species" species="cyanipes">Tricondyla cyanipes</taxonomicName>
/ fuscilabris m. / det.
<determinerName id="7E0DDCD7101B757EDCB6349AFDE0F845" box="[395,600,1975,2001]" pageId="11" pageNumber="362">Dr. Ing. K. Mandl</determinerName>
1959 [red label] //
<collectorName id="269B55C5101B757EDE0A349AFC75F845" box="[823,973,1975,2001]" pageId="11" pageNumber="362">Coll. G. Frey</collectorName>
/ NMB //
<taxonomicName id="4C6E4B90101B757ED978349AFB07F845" authorityName="Latreille" authorityYear="1822" box="[1093,1215,1975,2001]" class="Insecta" family="Carabidae" genus="Tricondyla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="362" phylum="Arthropoda" rank="genus">Tricondyla</taxonomicName>
(s.str.) / b.
<taxonomicName id="4C6E4B90101B757ED879349AFE90F861" authority="Gestro, 1874" authorityName="Gestro" authorityYear="1874" class="Insecta" family="Carabidae" genus="Tricondyla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="362" phylum="Arthropoda" rank="species" species="beccarii">beccarii Gestro, 1874</taxonomicName>
/ det.
<determinerName id="7E0DDCD7101B757EDC5334F6FE49F861" box="[366,497,2011,2037]" pageId="11" pageNumber="362">R. Naviaux</determinerName>
, 2001” (
<collectionCode id="ED7FA8D6101B757EDF7234F6FD11F861" box="[591,681,2011,2037]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="11" pageNumber="362">NHMB</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BD13013101C7579DDFA33BAFB90FF41" blockId="12.[151,1437,151,1199]" pageId="12" pageNumber="363">
<materialsCitation id="3B063A4E101C7579DDFA33BAFB90FF41" collectionCode="MNHN" country="Iceland" determinerName="G. Babault" location="Island" pageId="12" pageNumber="363" specimenCount="2" typeStatus="holotype">
<typeStatus id="54D58EB1101C7579DDFA33BAFE97FF25" box="[199,303,151,177]" pageId="12" pageNumber="363" type="holotype">Holotype</typeStatus>
male: “
<collectingCountry id="F3797083101C7579DCB233BAFE6EFF25" box="[399,470,151,177]" name="Iceland" pageId="12" pageNumber="363">Island</collectingCountry>
of / Basilan / Baker // Guy Babault //
<taxonomicName id="4C6E4B90101C7579DE9233BAFBFCFF25" authorityName="Schaum" authorityYear="1862" box="[943,1092,151,177]" class="Insecta" family="Carabidae" genus="Tricondyla" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="363" phylum="Arthropoda" rank="species" species="planiceps">Tr. planiceps</taxonomicName>
/ Schau. /
<determinerName id="7E0DDCD7101C7579D9F333BAFAF4FF25" box="[1230,1356,151,177]" pageId="12" pageNumber="363">G. Babault</determinerName>
det. //
<typeStatus id="54D58EB1101C7579DDAA3396FE93FF41" box="[151,299,187,213]" pageId="12" pageNumber="363" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="4C6E4B90101C7579DC7D3396FE02FF41" authorityName="Latreille" authorityYear="1822" box="[320,442,187,213]" class="Insecta" family="Carabidae" genus="Tricondyla" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="363" phylum="Arthropoda" rank="genus">Tricondyla</taxonomicName>
(s.str.) /
<taxonomicName id="4C6E4B90101C7579DF203396FC92FF41" authority="NAVIAUX, 2010" authorityName="NAVIAUX" authorityYear="2010" box="[541,810,187,213]" class="Insecta" family="Carabidae" genus="Tricondyla" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="363" phylum="Arthropoda" rank="species" species="huloti">huloti NAVIAUX, 2010</taxonomicName>
[red label]” (
<collectionCode id="ED7FA8D6101C7579DEFD3396FBA4FF41" box="[960,1052,187,213]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="363">MNHN</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BD13013101C7579DDFA33F2FC4DFE24" blockId="12.[151,1437,151,1199]" pageId="12" pageNumber="363">
<emphasis id="B91AEC01101C7579DDFA33F2FE05FF6D" bold="true" box="[199,445,223,249]" pageId="12" pageNumber="363">Additional material.</emphasis>
<materialsCitation id="3B063A4E101C7579DCFA33CDFF43FE8A" collectionCode="DUBC" country="Philippines" elevation="1" location="Mindanao" pageId="12" pageNumber="363" specimenCount="1" stateProvince="Agusan del Sur">
MINDANAO:
<quantity id="4C969DF6101C7579DF4933CDFD21FF6E" box="[628,665,224,250]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="363" unit="m" value="1.0">
<elevation id="0043D720101C7579DF4933CDFD21FF6E" box="[628,665,224,250]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="363" unit="m" value="1.0">1m</elevation>
</quantity>
, 2f: “
<collectingCountry id="F3797083101C7579DFE133CDFCE7FF6E" box="[732,863,224,250]" name="Philippines" pageId="12" pageNumber="363">Philippines</collectingCountry>
,
<location id="8EB166C8101C7579DE5033CDFC59FF6E" LSID="urn:lsid:plazi:treatment:03C78105101B7579DDAA358BFE88FB3B:8EB166C8101C7579DE5033CDFC59FF6E" box="[877,993,224,250]" country="Philippines" name="Mindanao" pageId="12" pageNumber="363" stateProvince="Agusan del Sur">Mindanao</location>
,
<collectingRegion id="49AAFEF1101C7579DED333CDFB1BFF6E" box="[1006,1187,224,250]" country="Philippines" name="Agusan del Sur" pageId="12" pageNumber="363">Agusan del Sur</collectingRegion>
, Esperanza, 03.2014” (
<collectionCode id="ED7FA8D6101C7579DDA23229FF4BFE8A" box="[159,243,260,286]" pageId="12" pageNumber="363">DUBC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B063A4E101C7579DC383229FBEAFE8A" box="[261,1106,260,287]" collectingDate="2014-08" collectionCode="DUBC" country="Philippines" location="Masara" municipality="Mindanao" pageId="12" pageNumber="363" specimenCount="1" stateProvince="Compostela">
1f: “
<collectingCountry id="F3797083101C7579DC0E3229FE0EFE8A" box="[307,438,260,286]" name="Philippines" pageId="12" pageNumber="363">Philippines</collectingCountry>
,
<collectingMunicipality id="6BB5AA69101C7579DCFD3229FD8CFE8A" box="[448,564,260,286]" pageId="12" pageNumber="363">Mindanao</collectingMunicipality>
,
<collectingRegion id="49AAFEF1101C7579DF033229FCB7FE8A" box="[574,783,260,286]" country="Philippines" name="Compostela" pageId="12" pageNumber="363">Compostela valley</collectingRegion>
,
<location id="8EB166C8101C7579DE243228FCD6FE8B" LSID="urn:lsid:plazi:treatment:03C78105101B7579DDAA358BFE88FB3B:8EB166C8101C7579DE243228FCD6FE8B" box="[793,878,261,287]" country="Philippines" municipality="Mindanao" name="Masara" pageId="12" pageNumber="363" stateProvince="Compostela">Masara</location>
,
<date id="FFD016D3101C7579DE453229FC66FE8A" box="[888,990,260,286]" pageId="12" pageNumber="363" value="2014-08">
<collectingDate id="EF94EF3B101C7579DE453229FC66FE8A" box="[888,990,260,286]" pageId="12" pageNumber="363" value="2014-08">viii.2014</collectingDate>
</date>
” (
<collectionCode id="ED7FA8D6101C7579DECB3229FBF2FE8A" box="[1014,1098,260,286]" pageId="12" pageNumber="363">DUBC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B063A4E101C7579D9613229FC16FED7" collectingDate="2019-10-10" collectionCode="AAc" collectorName="Anichtchenko A." country="Philippines" county="Mindanao" elevation="1" location="Nabunturan" municipality="Cabalinan Creek" pageId="12" pageNumber="363" specimenCount="1">
<quantity id="4C969DF6101C7579D9613229FB3BFE8A" box="[1116,1155,260,286]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="363" unit="m" value="1.0">
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: “
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,
<collectingMunicipality id="6BB5AA69101C7579DDAA3204FEEDFED7" box="[151,341,297,323]" pageId="12" pageNumber="363">Cabalinan Creek</collectingMunicipality>
,
<location id="8EB166C8101C7579DC623204FE5EFED7" LSID="urn:lsid:plazi:treatment:03C78105101B7579DDAA358BFE88FB3B:8EB166C8101C7579DC623204FE5EFED7" box="[351,486,297,323]" country="Philippines" county="Mindanao" municipality="Cabalinan Creek" name="Nabunturan" pageId="12" pageNumber="363">Nabunturan</location>
,
<date id="FFD016D3101C7579DCCD3204FDDAFED7" box="[496,610,297,323]" pageId="12" pageNumber="363" value="2019-10-10">
<collectingDate id="EF94EF3B101C7579DCCD3204FDDAFED7" box="[496,610,297,323]" pageId="12" pageNumber="363" value="2019-10-10">10.x.2019</collectingDate>
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,
<collectorName id="269B55C5101C7579DF513204FC91FED7" box="[620,809,297,323]" pageId="12" pageNumber="363">Anichtchenko A.</collectorName>
leg” (
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)
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;
<materialsCitation id="3B063A4E101C7579DE843204FE4CFEF3" collectingDate="2019-01" collectionCode="WJc" country="Philippines" elevation="1" location="Philippines" pageId="12" pageNumber="363" specimenCount="1">
<quantity id="4C969DF6101C7579DE843204FC66FED7" box="[953,990,297,323]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="363" unit="m" value="1.0">
<elevation id="0043D720101C7579DE843204FC66FED7" box="[953,990,297,323]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="363" unit="m" value="1.0">1m</elevation>
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, 2f: “
<date id="FFD016D3101C7579D9243204FB0CFED7" box="[1049,1204,297,323]" pageId="12" pageNumber="363" value="2019-01">
<collectingDate id="EF94EF3B101C7579D9243204FB0CFED7" box="[1049,1204,297,323]" pageId="12" pageNumber="363" value="2019-01">January 2019</collectingDate>
</date>
, Marihatag, Surigao, Mindanao,
<collectingCountry id="F3797083101C7579DC243260FE22FEF3" box="[281,410,333,359]" name="Philippines" pageId="12" pageNumber="363">Philippines</collectingCountry>
” (
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.
<materialsCitation id="3B063A4E101C7579DF3C3263FB0FFEF3" box="[513,1207,333,360]" collectingDate="2017-11" collectionCode="DUBC" country="Philippines" elevation="1" location="Panay" municipality="Mataas" pageId="12" pageNumber="363" specimenCount="1" stateProvince="Antique">
PANAY:
<quantity id="4C969DF6101C7579DF573260FD2AFEF3" box="[618,658,333,359]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="363" unit="m" value="1.0">
<elevation id="0043D720101C7579DF573260FD2AFEF3" box="[618,658,333,359]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="363" unit="m" value="1.0">1m</elevation>
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: “
<collectingMunicipality id="6BB5AA69101C7579DF943260FD47FEF3" box="[681,767,333,359]" pageId="12" pageNumber="363">Mataas</collectingMunicipality>
,
<collectingRegion id="49AAFEF1101C7579DE363260FCD0FEF3" box="[779,872,333,359]" country="Philippines" name="Antique" pageId="12" pageNumber="363">Antique</collectingRegion>
,
<location id="8EB166C8101C7579DE483263FC03FEFC" LSID="urn:lsid:plazi:treatment:03C78105101B7579DDAA358BFE88FB3B:8EB166C8101C7579DE483263FC03FEFC" box="[885,955,334,360]" country="Philippines" municipality="Mataas" name="Panay" pageId="12" pageNumber="363" stateProvince="Antique">Panay</location>
,
<date id="FFD016D3101C7579DEFA3263FBF8FEF3" box="[967,1088,333,360]" pageId="12" pageNumber="363" value="2017-11">
<collectingDate id="EF94EF3B101C7579DEFA3263FBF8FEF3" box="[967,1088,333,360]" pageId="12" pageNumber="363" value="2017-11">Nov. 2017</collectingDate>
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<materialsCitation id="3B063A4E101C7579D9F93260FD52FE18" collectingDate="2018-10" collectionCode="DUBC" country="Philippines" elevation="1" location="Bordeus" municipality="Polillo" pageId="12" pageNumber="363" specimenCount="1">
POLILLO:
<quantity id="4C969DF6101C7579D8743260FAD7FEF3" box="[1353,1391,333,359]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="363" unit="m" value="1.0">
<elevation id="0043D720101C7579D8743260FAD7FEF3" box="[1353,1391,333,359]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="363" unit="m" value="1.0">1m</elevation>
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, 1f: “
<collectingCountry id="F3797083101C7579DD9C325FFE9CFE18" box="[161,292,370,396]" name="Philippines" pageId="12" pageNumber="363">Philippines</collectingCountry>
,
<collectingMunicipality id="6BB5AA69101C7579DC0E325FFE39FE18" box="[307,385,370,396]" pageId="12" pageNumber="363">Polillo</collectingMunicipality>
,
<location id="8EB166C8101C7579DCB2325FFE57FE18" LSID="urn:lsid:plazi:treatment:03C78105101B7579DDAA358BFE88FB3B:8EB166C8101C7579DCB2325FFE57FE18" box="[399,495,370,396]" country="Philippines" municipality="Polillo" name="Bordeus" pageId="12" pageNumber="363">Bordeus</location>
,
<date id="FFD016D3101C7579DCC3325FFDC9FE18" box="[510,625,370,396]" pageId="12" pageNumber="363" value="2018-10">
<collectingDate id="EF94EF3B101C7579DCC3325FFDC9FE18" box="[510,625,370,396]" pageId="12" pageNumber="363" value="2018-10">Oct. 2018</collectingDate>
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.
<materialsCitation id="3B063A4E101C7579DFC5325FFEEFFE24" collectingDate="2015-01" collectionCode="DUBC" country="Philippines" county="Visayas" location="Hinabangan" municipality="Samar" pageId="12" pageNumber="363" specimenCount="1">
SAMAR: 1f: “
<collectingCountry id="F3797083101C7579DE9C325FFB9CFE18" box="[929,1060,370,396]" name="Philippines" pageId="12" pageNumber="363">Philippines</collectingCountry>
, E
<collectingCounty id="62B0489F101C7579D973325FFB10FE18" box="[1102,1192,370,396]" pageId="12" pageNumber="363">Visayas</collectingCounty>
,
<collectingMunicipality id="6BB5AA69101C7579D98B325FFB47FE18" box="[1206,1279,370,396]" pageId="12" pageNumber="363">Samar</collectingMunicipality>
,
<location id="8EB166C8101C7579D833325FFA20FE18" LSID="urn:lsid:plazi:treatment:03C78105101B7579DDAA358BFE88FB3B:8EB166C8101C7579D833325FFA20FE18" box="[1294,1432,370,396]" country="Philippines" county="Visayas" municipality="Samar" name="Hinabangan" pageId="12" pageNumber="363">Hinabangan</location>
,
<date id="FFD016D3101C7579DDAA32BBFF59FE25" box="[151,225,406,433]" pageId="12" pageNumber="363" value="2015-01">
<collectingDate id="EF94EF3B101C7579DDAA32BBFF59FE25" box="[151,225,406,433]" pageId="12" pageNumber="363" value="2015-01">I.2015</collectingDate>
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” (
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;
<materialsCitation id="3B063A4E101C7579DC5F32BBFC49FE24" box="[354,1009,406,433]" collectingDate="2016-07" collectionCode="DUBC" country="Philippines" county="Samar" location="de Vega" municipality="Lope de Vega" pageId="12" pageNumber="363" specimenCount="1">
1f: “
<collectingCountry id="F3797083101C7579DCAF32BBFDADFE24" box="[402,533,406,432]" name="Philippines" pageId="12" pageNumber="363">Philippines</collectingCountry>
,
<collectingCounty id="62B0489F101C7579DF1C32BBFDD2FE24" box="[545,618,406,432]" pageId="12" pageNumber="363">Samar</collectingCounty>
,
<collectingMunicipality id="6BB5AA69101C7579DF4832BAFCAAFE25" box="[629,786,406,433]" pageId="12" pageNumber="363">
Lope
<location id="8EB166C8101C7579DF8B32BBFCAAFE25" LSID="urn:lsid:plazi:treatment:03C78105101B7579DDAA358BFE88FB3B:8EB166C8101C7579DF8B32BBFCAAFE25" box="[694,786,406,433]" country="Philippines" county="Samar" municipality="Lope de Vega" name="de Vega" pageId="12" pageNumber="363">de Vega</location>
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,
<date id="FFD016D3101C7579DE2032BBFCC3FE24" box="[797,891,406,432]" pageId="12" pageNumber="363" value="2016-07">
<collectingDate id="EF94EF3B101C7579DE2032BBFCC3FE24" box="[797,891,406,432]" pageId="12" pageNumber="363" value="2016-07">vii.2016</collectingDate>
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)
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.
</paragraph>
</subSubSection>
<subSubSection id="C3746398101C7579DDFA3296FB5AFCD6" pageId="12" pageNumber="363" type="diagnosis">
<paragraph id="8BD13013101C7579DDFA3296FB5AFCD6" blockId="12.[151,1437,151,1199]" pageId="12" pageNumber="363">
<emphasis id="B91AEC01101C7579DDFA3296FE6AFE41" bold="true" box="[199,466,443,469]" pageId="12" pageNumber="363">Differential diagnosis.</emphasis>
Habitus elongated, body size
<quantity id="4C969DF6101C7579DE063296FC03FE41" box="[827,955,443,469]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" metricValueMax="2.0" metricValueMin="1.6" pageId="12" pageNumber="363" unit="mm" value="18.0" valueMax="20.0" valueMin="16.0">1620 mm</quantity>
, dorsally olivaceous or dark blue, shiny; slightly variable in shape but consistent in size; head broad with prominent eyes and long temples; neck narrow; interocular cavity deep with the bottom flat; pronotum long, smooth; medial lobe conical, rather broad near the base, then regularly shaped and narrowed anteriorly, feebly inflated above; collum narrowly long and rounded; apical lobe long; basal lobe narrower than the medial one; elytra oval-shaped without obvious lateral constriction; apex slightly truncated and rounded; gibbosity moderately high, mainly in the males; sculpture shallow, punctate-rugate near the base, then punctate, the posterior half smooth. Aedeagus (
<figureCitation id="13552C96101C7579DE1731BBFC73FD24" box="[810,971,662,688]" captionStart="FIGURES 112129" captionStartId="23.[151,264,1649,1673]" captionTargetBox="[155,1432,185,1622]" captionTargetId="figure-17@23.[151,1435,181,1625]" captionTargetPageId="23" captionText="FIGURES 112129. Aedeagus of Tricondyla species. 112—T. aptera (Samar); 113—Holotype of T. johnsoni sp. nov.; 114—T. ovicollis (Ditumabo); 115—Holotype of T. aptera lumawigi syn. nov. = T. ovicollis; 116—T. ventricosa (Sta Ana); 117—T. elongata (Kalatungan); 118—T. elongata (Misamis Or.); 119—T. gracilis syn. nov. = T. elongata (Lake Holon, Mindanao); 120 - T. gracilis syn. nov. = T. elongata (Bataugan, Mindanao); 121—T. conicicollis (Mindanao); 122—Paratype of T. cyanipes fuscilabris Mandl, 1964 syn.; 123—Holotype of T. huloti syn. nov.; 124—Holotype of T. cavifrons syn. nov. = T. cyanipes; 125—T. cavifrons (Negros); 126—Holotype of T. genieri; 127—T. planiceps; 128—Holotype of T. oblita; 129—Paratype of T. beccarii cassolai." figureDoi="http://doi.org/10.5281/zenodo.10468916" httpUri="https://zenodo.org/record/10468916/files/figure.png" pageId="12" pageNumber="363">Figs 121123</figureCitation>
). Endophallus (
<figureCitation id="13552C96101C7579D94131BBFAB8FD25" box="[1148,1280,662,689]" captionStart="FIGURES 4356" captionStartId="11.[151,264,1488,1512]" captionTargetBox="[154,1433,184,1463]" captionTargetId="figure-17@11.[151,1435,181,1465]" captionTargetPageId="11" captionText="FIGURES 4356. Variability of Tricondyla (Stenotricondyla) conicicollis (4349) Habitus: 43—Cabalinan, Mindanao (male); 44—Esperanza, Mindanao (female); 45—Polillo (female); 4647—Paratype of Tricondyla (Stenotricondyla) cyanipes fuscilabris Mandl, 1964 syn. nov.; 4849 Holotype of Tricondyla (Tricondyla) huloti Naviaux, 2009 syn. nov. (50-56) Aedeagus and endophallus: 5052—Mataas, Panay; 53-56—Nabunturan, Mindanao." figureDoi="http://doi.org/10.5281/zenodo.10468898" httpUri="https://zenodo.org/record/10468898/files/figure.png" pageId="12" pageNumber="363">Figs 5056</figureCitation>
), endophallus extremely similar to that of
<taxonomicName id="4C6E4B90101C7579DCE43196FDE1FD41" authorityName="Horn" authorityYear="1906" box="[473,601,699,725]" class="Insecta" family="Carabidae" genus="Tricondyla" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="363" phylum="Arthropoda" rank="species" species="elongata">
<emphasis id="B91AEC01101C7579DCE43196FDE1FD41" box="[473,601,699,725]" italics="true" pageId="12" pageNumber="363">T. elongata</emphasis>
</taxonomicName>
. Protuberances: VB relatively small; pair of laterally protruding BL; DB relatively large and weakly biforked at apex; median part of endophallus short in comparison to other species of subgenus; V small, directed towards the apex of endophallus; VA relatively large, directed towards the base of endophallus, and overlap with V; A transverse, with 23 apices; DA long; flagellum long and thin.
</paragraph>
</subSubSection>
<subSubSection id="C3746398101C7579DDFA3061FBF4FBF2" pageId="12" pageNumber="363" type="discussion">
<paragraph id="8BD13013101C7579DDFA3061FBF4FBF2" blockId="12.[151,1437,151,1199]" pageId="12" pageNumber="363">
<emphasis id="B91AEC01101C7579DDFA3061FEF7FCF2" bold="true" box="[199,335,844,870]" pageId="12" pageNumber="363">Comments.</emphasis>
Study of type specimens
<taxonomicName id="4C6E4B90101C7579DF4E3063FCD7FCF3" authorityName="Mandl" authorityYear="1964" box="[627,879,845,871]" class="Insecta" family="Carabidae" genus="Tricondyla" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="363" phylum="Arthropoda" rank="subSpecies" species="cyanipes" subSpecies="fuscilabris">
<emphasis id="B91AEC01101C7579DF4E3063FCD7FCF3" box="[627,879,845,871]" italics="true" pageId="12" pageNumber="363">T. cyanipes fuscilabris</emphasis>
</taxonomicName>
showed that they were erroneously synonymized by
<bibRefCitation id="EFFF4DE2101C7579DD81305CFE24FC1F" author="Naviaux, R." box="[188,412,881,907]" pageId="12" pageNumber="363" pagination="1 - 106" refId="ref14600" refString="Naviaux, R. (2002) Tricondylina (Coleoptera: Cicindelidae): Revision des genres Tricondyla Latreille et Derocrania Chaudoir et descriptions de nouveaux taxons. Memoirs de la Sef, 5, 1 - 106." type="journal article" year="2002">Naviaux (2002: 72)</bibRefCitation>
with
<emphasis id="B91AEC01101C7579DCDD305FFD7EFC1F" box="[480,710,881,907]" italics="true" pageId="12" pageNumber="363">
<taxonomicName id="4C6E4B90101C7579DCDD305FFD7AFC1F" authorityName="Gestro" authorityYear="1874" box="[480,706,881,907]" class="Insecta" family="Carabidae" genus="Tricondyla" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="363" phylum="Arthropoda" rank="subSpecies" species="beccarii" subSpecies="beccarii">T. beccarii beccarii</taxonomicName>
.
</emphasis>
<typeStatus id="54D58EB1101C7579DFF2305FFC89FC18" box="[719,817,882,908]" pageId="12" pageNumber="363" type="paratype">Paratype</typeStatus>
specimens, studied by Naviaux in 2001, look slightly depigmented, brownish, and by coloration similar to
<taxonomicName id="4C6E4B90101C7579DFD530BAFCE7FC24" authorityName="Gestro" authorityYear="1874" box="[744,863,918,944]" class="Insecta" family="Carabidae" genus="Tricondyla" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="363" phylum="Arthropoda" rank="species" species="beccarii">
<emphasis id="B91AEC01101C7579DFD530BAFCE7FC24" box="[744,863,918,944]" italics="true" pageId="12" pageNumber="363">T. beccarii</emphasis>
</taxonomicName>
, but the wide and flat interocular cavity and elytral structure show them to be identical to
<taxonomicName id="4C6E4B90101C7579DF603096FD45FC40" authorityName="Chaudoir" authorityYear="1844" box="[605,765,954,980]" class="Insecta" family="Carabidae" genus="Tricondyla" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="363" phylum="Arthropoda" rank="species" species="conicicollis">
<emphasis id="B91AEC01101C7579DF603096FD45FC40" box="[605,765,954,980]" italics="true" pageId="12" pageNumber="363">T. conicicollis</emphasis>
</taxonomicName>
. Unfortunately, both aedeagi are damaged and a study of endophalli is therefore impossible. A new combination of synonymy is proposed here.
<taxonomicName id="4C6E4B90101C7579D94F30CDFB68FC6D" authorityName="NAVIAUX" authorityYear="2010" box="[1138,1232,991,1017]" class="Insecta" family="Carabidae" genus="Tricondyla" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="363" phylum="Arthropoda" rank="species" species="huloti">
<emphasis id="B91AEC01101C7579D94F30CDFB68FC6D" box="[1138,1232,991,1017]" italics="true" pageId="12" pageNumber="363">T. huloti</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="EFFF4DE2101C7579DDAA372EFE04FB8A" author="Naviaux, R." box="[151,444,1027,1054]" pageId="12" pageNumber="363" pagination="107 - 109" refId="ref14679" refString="Naviaux, R. (2009) Tricondyla (s. str.) huloti, nouvelle espece de Philippines (Coleoptera: Cicindelidae). Revue Francaise d'Entomologie, 31 (3 - 4), 107 - 109." type="journal article" year="2009">Naviaux (2009: 108109)</bibRefCitation>
based on a male from
<collectingRegion id="49AAFEF1101C7579DFFA372EFCA6FB89" box="[711,798,1027,1053]" country="Philippines" name="Basilan" pageId="12" pageNumber="363">Basilan</collectingRegion>
, as its pronotum and elytra should be different in shape from
<taxonomicName id="4C6E4B90101C7579DDEA3704FECDFBD6" authorityName="Chaudoir" authorityYear="1844" box="[215,373,1064,1090]" class="Insecta" family="Carabidae" genus="Tricondyla" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="363" phylum="Arthropoda" rank="species" species="conicicollis">
<emphasis id="B91AEC01101C7579DDEA3704FECDFBD6" box="[215,373,1064,1090]" italics="true" pageId="12" pageNumber="363">T. conicicollis</emphasis>
</taxonomicName>
. However, the differences observed in the
<typeStatus id="54D58EB1101C7579DE5E3705FC7DFBD6" box="[867,965,1064,1090]" pageId="12" pageNumber="363" type="holotype">holotype</typeStatus>
from the MNHN are within the range of variation of
<taxonomicName id="4C6E4B90101C7579DC1D3760FE7AFBF2" authorityName="Chaudoir" authorityYear="1844" box="[288,450,1100,1127]" class="Insecta" family="Carabidae" genus="Tricondyla" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="363" phylum="Arthropoda" rank="species" species="conicicollis">
<emphasis id="B91AEC01101C7579DC1D3760FE7AFBF2" box="[288,450,1100,1127]" italics="true" pageId="12" pageNumber="363">C. conicicollis</emphasis>
</taxonomicName>
, and thus
<taxonomicName id="4C6E4B90101C7579DF083760FD29FBF2" authorityName="NAVIAUX" authorityYear="2010" box="[565,657,1100,1126]" class="Insecta" family="Carabidae" genus="Tricondyla" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="363" phylum="Arthropoda" rank="species" species="huloti">
<emphasis id="B91AEC01101C7579DF083760FD29FBF2" box="[565,657,1100,1126]" italics="true" pageId="12" pageNumber="363">T. huloti</emphasis>
</taxonomicName>
is put into synonymy with this species.
</paragraph>
</subSubSection>
<subSubSection id="C3746398101C7579DDFA375CFE88FB3B" pageId="12" pageNumber="363" type="distribution">
<paragraph id="8BD13013101C7579DDFA375CFE88FB3B" blockId="12.[151,1437,151,1199]" pageId="12" pageNumber="363">
<emphasis id="B91AEC01101C7579DDFA375CFED9FB1F" bold="true" box="[199,353,1137,1163]" pageId="12" pageNumber="363">Distribution.</emphasis>
<collectingCountry id="F3797083101C7579DC4F375CFE48FB1F" box="[370,496,1137,1163]" name="Philippines" pageId="12" pageNumber="363">Philippines</collectingCountry>
(Mindanao,
<collectingRegion id="49AAFEF1101C7579DFA9375CFD4EFB1F" box="[660,758,1137,1163]" country="Philippines" name="Palawan" pageId="12" pageNumber="363">Palawan</collectingRegion>
, Panay, Polillo, Samar). Previous site for Luzon need confirmation.
</paragraph>
</subSubSection>
</treatment>
</document>